treatments-xml/data/03/A8/87/03A887A1A9641962AFE9FD8E453C63DC.xml
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<document id="C8234C5D6DEC6DB65B60DB643830CE9E" ID-CLB-Dataset="8896" ID-DOI="10.11646/zootaxa.4868.1.4" ID-GBIF-Dataset="cfd5c06e-06f0-4f1f-ae4b-40b7b46f44a3" ID-ISSN="1175-5326" ID-Zenodo-Dep="4417281" ID-ZooBank="DC4B8EC5-CBDA-49D2-B370-381C96AF3A10" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609805123592" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2020" docId="03A887A1A9641962AFE9FD8E453C63DC" docLanguage="en" docName="zootaxa.4868.1.4.pdf" docOrigin="Zootaxa 4868 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrochus astrictus Perkins 2020, new species" docType="treatment" docVersion="8" lastPageNumber="67" masterDocId="FF91FFD9A9611964AF7EFFC8470B6435" masterDocTitle="Taxonomy of Peruvian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)" masterLastPageNumber="89" masterPageNumber="61" pageNumber="66" updateTime="1698852338306" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EEF16D51E88D9C5DBE23494FD444A518">Taxonomy of Peruvian water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="D2492432E18CD97C1E4DF8B087FDBFD6">Perkins, Philip D.</mods:namePart>
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<mods:title id="403B75EF6C84451C06F4179B56853B75">Zootaxa</mods:title>
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<mods:part id="010EE371335C962EBE4A6714BCB82608">
<mods:date id="FCA5C21470DB75ED09EC610980357D92">2020</mods:date>
<mods:detail id="6DDA53BE4B6C94F70507B31FDD328ECE" type="pubDate">
<mods:number id="F55222A584F3FD9747E98F2BB5241DF6">2020-10-23</mods:number>
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<mods:classification id="958352F895DE986C953F4F2593EFED0B">journal article</mods:classification>
<mods:identifier id="A6BEFAB0ED1F71E90EC507FBDE6B2F14" type="CLB-Dataset">8896</mods:identifier>
<mods:identifier id="DFC36478205B8547DA82410DAA1C5302" type="DOI">10.11646/zootaxa.4868.1.4</mods:identifier>
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<treatment id="03A887A1A9641962AFE9FD8E453C63DC" ID-DOI="http://doi.org/10.5281/zenodo.4417460" ID-GBIF-Taxon="176581840" ID-Zenodo-Dep="4417460" LSID="urn:lsid:plazi:treatment:03A887A1A9641962AFE9FD8E453C63DC" httpUri="http://treatment.plazi.org/id/03A887A1A9641962AFE9FD8E453C63DC" lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66">
<subSubSection id="C31B653CA9641961AFE9FD8E45226655" box="[151,553,582,609]" pageId="5" pageNumber="66" type="nomenclature">
<paragraph id="8BBE36B7A9641961AFE9FD8E45226655" blockId="5.[151,553,582,645]" box="[151,553,582,609]" pageId="5" pageNumber="66">
<heading id="D0F681DBA9641961AFE9FD8E45226655" bold="true" box="[151,553,582,609]" fontSize="11" level="1" pageId="5" pageNumber="66" reason="1">
<emphasis id="B975EAA5A9641961AFE9FD8E45226655" bold="true" box="[151,553,582,609]" pageId="5" pageNumber="66">
<taxonomicName id="4C014D34A9641961AFE9FD8E46806654" ID-CoL="8Y4P8" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[151,395,582,609]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="astrictus" status="sp. nov.">
<emphasis id="B975EAA5A9641961AFE9FD8E46806654" bold="true" box="[151,395,582,609]" italics="true" pageId="5" pageNumber="66">Hydrochus astrictus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24657DEA9641961AEE4FD8F45226655" box="[410,553,582,609]" pageId="5" pageNumber="66" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA9641961AFE9FDA347F466B0" box="[151,255,619,645]" pageId="5" pageNumber="66" type="description">
<paragraph id="8BBE36B7A9641961AFE9FDA347F466B0" blockId="5.[151,553,582,645]" box="[151,255,619,645]" pageId="5" pageNumber="66">
<figureCitation id="133A2A32A9641961AFE9FDA347E866B0" box="[151,227,619,645]" captionStart="FIGURE 1" captionStartId="2.[151,250,1971,1995]" captionTargetBox="[157,1430,309,1948]" captionTargetId="figure-37@2.[157,1430,309,1948]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Hydrochus species." figureDoi="http://doi.org/10.5281/zenodo.4417283" httpUri="https://zenodo.org/record/4417283/files/figure.png" pageId="5" pageNumber="66">Figs. 1</figureCitation>
,
<figureCitation id="133A2A32A9641961AF8FFDA347F466B0" box="[241,255,619,645]" captionStart="FIGURE 4" captionStartId="6.[151,250,1820,1844]" captionTargetBox="[172,1412,198,1769]" captionTargetId="figure-26@6.[160,1427,181,1796]" captionTargetPageId="6" captionText="FIGURE 4. Hydrochus astrictus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417291" httpUri="https://zenodo.org/record/4417291/files/figure.png" pageId="5" pageNumber="66">4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA9641961AFE9FD7B422A6790" pageId="5" pageNumber="66" type="materials_examined">
<paragraph id="8BBE36B7A9641961AFE9FD7B422A6790" blockId="5.[151,1437,690,1977]" pageId="5" pageNumber="66">
<emphasis id="B975EAA5A9641961AFE9FD7B464266F9" bold="true" box="[151,329,690,717]" pageId="5" pageNumber="66">Type Material.</emphasis>
<materialsCitation id="3B693CEAA9641961AE2CFD7B464B6720" ID-GBIF-Occurrence="3015416367" collectingDate="1989-06-09" collectionCode="SEMC" collectorName="J. S. Ashe &amp; R. A. Leschen" country="Peru" county="Tambopata Prov." elevation="200" latitude="-12.55" location="Reserva" longLatPrecision="1293" longitude="-69.05" municipality="Dpto" pageId="5" pageNumber="66" specimenCount="1" stateProvince="Madre de Dios" typeStatus="holotype">
<emphasis id="B975EAA5A9641961AE2CFD7B451766F9" bold="true" box="[338,540,690,717]" pageId="5" pageNumber="66">
<typeStatus id="54BA8815A9641961AE2CFD7B46B466F8" box="[338,447,691,717]" pageId="5" pageNumber="66" type="holotype">Holotype</typeStatus>
(male):
</emphasis>
<collectingCountry id="F3167627A9641961AD51FD7B457666F8" box="[559,637,691,717]" name="Peru" pageId="5" pageNumber="66">PERU</collectingCountry>
:
<collectingCounty id="62DF4E3BA9641961ADF6FD7B444166F8" box="[648,842,691,717]" pageId="5" pageNumber="66">Tambopata Prov.</collectingCounty>
,
<collectingRegion id="49C5F855A9641961AC28FD7B44F466F8" box="[854,1023,691,717]" country="Peru" name="Madre de Dios" pageId="5" pageNumber="66">Madre de Dios</collectingRegion>
<collectingMunicipality id="6BDAACCDA9641961AB79FD7B434966F8" box="[1031,1090,691,717]" pageId="5" pageNumber="66">Dpto</collectingMunicipality>
,
<locationDeviation id="9A0C2F95A9641961AB31FD7A429366F8" box="[1103,1432,690,717]" location="Puerto Maldonado" pageId="5" pageNumber="66">
<quantity id="4CF99B52A9641961AB31FD7A438466F8" box="[1103,1167,690,717]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="66" unit="km" value="15.0">15km</quantity>
NE Puerto Maldonado
</locationDeviation>
,
<location id="8EDE606CA9641961AFE9FD1F47FA66C4" LSID="urn:lsid:plazi:treatment:03A887A1A9641962AFE9FD8E453C63DC:8EDE606CA9641961AFE9FD1F47FA66C4" box="[151,241,727,753]" country="Peru" county="Tambopata Prov." latitude="-12.55" longLatPrecision="1293" longitude="-69.05" municipality="Dpto" name="Reserva" pageId="5" pageNumber="66" stateProvince="Madre de Dios">Reserva</location>
<collectingRegion id="49C5F855A9641961AF84FD1F464966C4" box="[250,322,727,753]" country="Peru" name="Cusco" pageId="5" pageNumber="66">Cuzco</collectingRegion>
<location id="8EDE606CA9641961AE36FD1F46C766C4" LSID="urn:lsid:plazi:treatment:03A887A1A9641962AFE9FD8E453C63DC:8EDE606CA9641961AE36FD1F46C766C4" box="[328,460,727,753]" country="Peru" county="Tambopata Prov." latitude="-12.55" longLatPrecision="1293" longitude="-69.05" municipality="Dpto" name="Amazonico" pageId="5" pageNumber="66" stateProvince="Madre de Dios">Amazonico</location>
,
<geoCoordinate id="EE355070A9641961AEA7FD1F453C66C4" box="[473,567,727,753]" degrees="12" direction="south" minutes="33" orientation="latitude" pageId="5" pageNumber="66" precision="925" value="-12.55">12°33S</geoCoordinate>
,
<geoCoordinate id="EE355070A9641961AD3AFD1F459666C4" box="[580,669,727,753]" degrees="69" direction="west" minutes="3" orientation="longitude" pageId="5" pageNumber="66" precision="925" value="-69.05">69°3W</geoCoordinate>
,
<quantity id="4CF99B52A9641961ADD4FD1F45E766C4" box="[682,748,727,753]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="66" unit="m" value="200.0">
<elevation id="002CD184A9641961ADD4FD1F45E766C4" box="[682,748,727,753]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="66" unit="m" value="200.0">200m</elevation>
</quantity>
, camp,
<date id="FFBF1077A9641961AC3BFD1F44D966C4" box="[837,978,727,753]" pageId="5" pageNumber="66" value="1989-06-09">
<collectingDate id="EFFBE99FA9641961AC3BFD1F44D966C4" box="[837,978,727,753]" pageId="5" pageNumber="66" value="1989-06-09">9 June 1989</collectingDate>
</date>
,
<collectorName id="26F45361A9641961ACA0FD1F435966C4" box="[990,1106,727,753]" pageId="5" pageNumber="66">J. S. Ashe</collectorName>
,
<collectorName id="26F45361A9641961AB21FD1F420866C4" box="[1119,1283,727,753]" pageId="5" pageNumber="66">R. A. Leschen</collectorName>
, #009, ex., at light” (
<collectionCode id="ED10AE72A9641961AF98FD3346336720" box="[230,312,763,789]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="5" pageNumber="66" type="Museum">SEMC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B693CEAA9641961AE34FD3342426720" ID-GBIF-Occurrence="3015416319" box="[330,1353,762,789]" collectingDate="1969-04-10" collectingDateMax="1969-04-12" collectingDateMin="1969-04-10" collectionCode="NMNH" collectorName="P. &amp; P. Spangler" country="Peru" location="Pucallpa" pageId="5" pageNumber="66" specimenCount="1" stateProvince="Loreto" typeStatus="paratype">
<emphasis id="B975EAA5A9641961AE34FD33450A6721" bold="true" box="[330,513,762,789]" pageId="5" pageNumber="66">
<typeStatus id="54BA8815A9641961AE34FD3346C96720" box="[330,450,763,789]" pageId="5" pageNumber="66" type="paratype">Paratypes</typeStatus>
(35):
</emphasis>
<collectingCountry id="F3167627A9641961AD79FD3345586720" box="[519,595,763,789]" name="Peru" pageId="5" pageNumber="66">PERU</collectingCountry>
:
<collectingRegion id="49C5F855A9641961AD23FD3345A06720" box="[605,683,763,789]" country="Peru" name="Loreto" pageId="5" pageNumber="66">Loreto</collectingRegion>
,
<location id="8EDE606CA9641961ADCBFD3344136720" LSID="urn:lsid:plazi:treatment:03A887A1A9641962AFE9FD8E453C63DC:8EDE606CA9641961ADCBFD3344136720" box="[693,792,763,789]" country="Peru" name="Pucallpa" pageId="5" pageNumber="66" stateProvince="Loreto">Pucallpa</location>
,
<date id="FFBF1077A9641961AC5DFD3344CD6720" box="[803,966,763,789]" pageId="5" pageNumber="66" value="1969-04-10" valueMax="1969-04-12" valueMin="1969-04-10">
<collectingDate id="EFFBE99FA9641961AC5DFD3344CD6720" box="[803,966,763,789]" pageId="5" pageNumber="66" value="1969-04-10" valueMax="1969-04-12" valueMin="1969-04-10">1012.iv.1969</collectingDate>
</date>
, leg.
<collectorName id="26F45361A9641961AB7EFD3343BE6720" box="[1024,1205,763,789]" pageId="5" pageNumber="66">P. &amp; P. Spangler</collectorName>
(11
<collectionCode id="ED10AE72A9641961AB98FD33424A6720" box="[1254,1345,763,789]" country="USA" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="66" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B693CEAA9641961AA2DFD3346A96768" ID-GBIF-Occurrence="3015416337" collectingDate="1979-11-02" collectingDateMax="1979-11-05" collectingDateMin="1979-11-02" collectionCode="NMNH" collectorName="SW Pto. Maldonado &amp; J. B. Heppner" country="Peru" elevation="290" latitude="-13.0" location="Rio Tambopata Res" longLatPrecision="1291" longitude="-69.55" pageId="5" pageNumber="66" specimenCount="1" stateProvince="Madre de Dios" typeStatus="paratype">
<collectingRegion id="49C5F855A9641961AA2DFD3347F9670C" country="Peru" name="Madre de Dios" pageId="5" pageNumber="66">Madre de Dios</collectingRegion>
,
<location id="8EDE606CA9641961AF83FCD746D5670C" LSID="urn:lsid:plazi:treatment:03A887A1A9641962AFE9FD8E453C63DC:8EDE606CA9641961AF83FCD746D5670C" box="[253,478,799,825]" country="Peru" latitude="-13.0" longLatPrecision="1291" longitude="-69.55" name="Rio Tambopata Res" pageId="5" pageNumber="66" stateProvince="Madre de Dios">Rio Tambopata Res</location>
; 30 air km
<collectorName id="26F45361A9641961AD1EFCD7444C670C" box="[608,839,799,825]" pageId="5" pageNumber="66">SW Pto. Maldonado</collectorName>
, elev.
<quantity id="4CF99B52A9641961ACEEFCD744DC670C" box="[912,983,799,825]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="66" unit="m" value="290.0">
<elevation id="002CD184A9641961ACEEFCD744DC670C" box="[912,983,799,825]" metricMagnitude="2" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="66" unit="m" value="290.0">290 m</elevation>
</quantity>
(
<geoCoordinate id="EE355070A9641961AC97FCD74332670C" box="[1001,1081,799,825]" degrees="13" direction="south" minutes="0" orientation="latitude" pageId="5" pageNumber="66" precision="925" value="-13.0">13°0S</geoCoordinate>
,
<geoCoordinate id="EE355070A9641961AB3BFCD743A4670F" box="[1093,1199,799,826]" degrees="69" direction="west" minutes="33" orientation="longitude" pageId="5" pageNumber="66" precision="925" value="-69.55">69°33W</geoCoordinate>
),
<date id="FFBF1077A9641961ABBBFCD64246670C" box="[1221,1357,798,825]" pageId="5" pageNumber="66" value="1979-11-02" valueMax="1979-11-05" valueMin="1979-11-02">
<collectingDate id="EFFBE99FA9641961ABBBFCD64246670C" box="[1221,1357,798,825]" pageId="5" pageNumber="66" value="1979-11-02" valueMax="1979-11-05" valueMin="1979-11-02">25.xi.1979</collectingDate>
</date>
, leg.
<collectorName id="26F45361A9641961AAF4FCD746136768" pageId="5" pageNumber="66">J. B. Heppner</collectorName>
(4
<collectionCode id="ED10AE72A9641961AE41FC8B46916768" box="[319,410,835,861]" country="USA" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="66" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B693CEAA9641961AED0FC8B434C6768" ID-GBIF-Occurrence="3015416360" box="[430,1095,835,861]" collectingDate="1979-11-06" collectingDateMax="1979-11-10" collectingDateMin="1979-11-06" collectionCode="NMNH" collectorName="J. B. Heppner" country="Peru" elevation="290" latitude="-13.0" location="Rio Tambopata Res" longLatPrecision="1291" longitude="-69.55" pageId="5" pageNumber="66" specimenCount="1" stateProvince="Madre de Dios" typeStatus="paratype">
same locality,
<date id="FFBF1077A9641961AD31FC8B45EE6768" box="[591,741,835,861]" pageId="5" pageNumber="66" value="1979-11-06" valueMax="1979-11-10" valueMin="1979-11-06">
<collectingDate id="EFFBE99FA9641961AD31FC8B45EE6768" box="[591,741,835,861]" pageId="5" pageNumber="66" value="1979-11-06" valueMax="1979-11-10" valueMin="1979-11-06">610.xi.1979</collectingDate>
</date>
, leg.
<collectorName id="26F45361A9641961AC5DFC8B44B66768" box="[803,957,835,861]" pageId="5" pageNumber="66">J. B. Heppner</collectorName>
(2
<collectionCode id="ED10AE72A9641961AC9AFC8B43346768" box="[996,1087,835,861]" country="USA" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="66" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B693CEAA9641961AB2DFC8B46FC67B4" ID-GBIF-Occurrence="3015416375" collectingDate="1979-11-11" collectingDateMax="1979-11-15" collectingDateMin="1979-11-11" collectionCode="NMNH" collectorName="J. B. Heppner" country="Peru" elevation="290" latitude="-13.0" location="Rio Tambopata Res" longLatPrecision="1291" longitude="-69.55" pageId="5" pageNumber="66" specimenCount="1" stateProvince="Madre de Dios" typeStatus="paratype">
same locality,
<date id="FFBF1077A9641961AB8AFC8A429C6768" box="[1268,1431,834,861]" pageId="5" pageNumber="66" value="1979-11-11" valueMax="1979-11-15" valueMin="1979-11-11">
<collectingDate id="EFFBE99FA9641961AB8AFC8A429C6768" box="[1268,1431,834,861]" pageId="5" pageNumber="66" value="1979-11-11" valueMax="1979-11-15" valueMin="1979-11-11">1115.xi.1979</collectingDate>
</date>
, leg.
<collectorName id="26F45361A9641961AFB4FCAF466267B4" box="[202,361,871,897]" pageId="5" pageNumber="66">J. B. Heppner</collectorName>
(4
<collectionCode id="ED10AE72A9641961AEEBFCAF46FB67B4" box="[405,496,871,897]" country="USA" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="66" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B693CEAA9641961AD7BFCAF43B567B4" ID-GBIF-Occurrence="3015416343" box="[517,1214,871,897]" collectingDate="1979-11-16" collectingDateMax="1979-11-20" collectingDateMin="1979-11-16" collectionCode="NMNH" collectorName="J. B. Heppner" country="Peru" elevation="290" latitude="-13.0" location="Rio Tambopata Res" longLatPrecision="1291" longitude="-69.55" pageId="5" pageNumber="66" specimenCount="1" stateProvince="Madre de Dios" typeStatus="paratype">
same locality,
<date id="FFBF1077A9641961ADD5FCAF445B67B4" box="[683,848,871,897]" pageId="5" pageNumber="66" value="1979-11-16" valueMax="1979-11-20" valueMin="1979-11-16">
<collectingDate id="EFFBE99FA9641961ADD5FCAF445B67B4" box="[683,848,871,897]" pageId="5" pageNumber="66" value="1979-11-16" valueMax="1979-11-20" valueMin="1979-11-16">1620.xi.1979</collectingDate>
</date>
, leg.
<collectorName id="26F45361A9641961ACEFFCAF433B67B4" box="[913,1072,871,897]" pageId="5" pageNumber="66">J. B. Heppner</collectorName>
(8
<collectionCode id="ED10AE72A9641961AB22FCAF43BC67B4" box="[1116,1207,871,897]" country="USA" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="66" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B693CEAA9641961ABB2FCAF45646790" ID-GBIF-Occurrence="3015416332" collectingDate="1979-11-21" collectingDateMax="1979-11-25" collectingDateMin="1979-11-21" collectionCode="NMNH" collectorName="J. B. Heppner" country="Peru" elevation="290" latitude="-13.0" location="Rio Tambopata Res" longLatPrecision="1291" longitude="-69.55" pageId="5" pageNumber="66" specimenCount="1" stateProvince="Madre de Dios" typeStatus="paratype">
same locality,
<date id="FFBF1077A9641961AA0CFCAF461A6790" pageId="5" pageNumber="66" value="1979-11-21" valueMax="1979-11-25" valueMin="1979-11-21">
<collectingDate id="EFFBE99FA9641961AA0CFCAF461A6790" pageId="5" pageNumber="66" value="1979-11-21" valueMax="1979-11-25" valueMin="1979-11-21">2125.xi.1979</collectingDate>
</date>
, leg.
<collectorName id="26F45361A9641961AE33FC4346EC6790" box="[333,487,907,933]" pageId="5" pageNumber="66">J. B. Heppner</collectorName>
(3
<collectionCode id="ED10AE72A9641961AD72FC43456C6790" box="[524,615,907,933]" country="USA" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="66" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B693CEAA9641961AD04FC4342166790" ID-GBIF-Occurrence="3015416370" box="[634,1309,907,933]" collectingDate="1979-11-26" collectingDateMax="1979-11-30" collectingDateMin="1979-11-26" collectionCode="NMNH" collectorName="J. B. Heppner" country="Peru" elevation="290" latitude="-13.0" location="Rio Tambopata Res" longLatPrecision="1291" longitude="-69.55" pageId="5" pageNumber="66" specimenCount="1" stateProvince="Madre de Dios" typeStatus="paratype">
same locality,
<date id="FFBF1077A9641961AC65FC4344B46790" box="[795,959,907,933]" pageId="5" pageNumber="66" value="1979-11-26" valueMax="1979-11-30" valueMin="1979-11-26">
<collectingDate id="EFFBE99FA9641961AC65FC4344B46790" box="[795,959,907,933]" pageId="5" pageNumber="66" value="1979-11-26" valueMax="1979-11-30" valueMin="1979-11-26">2630.xi.1979</collectingDate>
</date>
, leg.
<collectorName id="26F45361A9641961AC85FC43439F6790" box="[1019,1172,907,933]" pageId="5" pageNumber="66">J. B. Heppner</collectorName>
(3
<collectionCode id="ED10AE72A9641961ABC4FC43421E6790" box="[1210,1301,907,933]" country="USA" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="66" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA9641961AFB9FC66452B606F" pageId="5" pageNumber="66" type="diagnosis">
<paragraph id="8BBE36B7A9641961AFB9FC66452B606F" blockId="5.[151,1437,690,1977]" pageId="5" pageNumber="66">
<emphasis id="B975EAA5A9641961AFB9FC6646DF67FD" bold="true" box="[199,468,942,968]" pageId="5" pageNumber="66">Differential Diagnosis.</emphasis>
Among known Peruvian species, differentiated from all, except
<taxonomicName id="4C014D34A9641961ABCEFC67424D67FC" authorityName="Perkins" authorityYear="2020" box="[1200,1350,943,969]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="bisinuatus">
<emphasis id="B975EAA5A9641961ABCEFC67424D67FC" box="[1200,1350,943,969]" italics="true" pageId="5" pageNumber="66">H. bisinuatus</emphasis>
</taxonomicName>
, by the deep cervical groove behind the eyes. This species is a member of the
<taxonomicName id="4C014D34A9641961ACD6FC1B432B67D8" authorityName="Perkins" authorityYear="2019" box="[936,1056,979,1005]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="collaris">
<emphasis id="B975EAA5A9641961ACD6FC1B432B67D8" box="[936,1056,979,1005]" italics="true" pageId="5" pageNumber="66">H. collaris</emphasis>
</taxonomicName>
group (
<emphasis id="B975EAA5A9641961AB06FC1B43BD67D9" box="[1144,1206,979,1004]" italics="true" pageId="5" pageNumber="66">sensu</emphasis>
<bibRefCitation id="EF904B46A9641961ABC3FC1B426B67D8" author="Perkins, P. D." box="[1213,1376,979,1005]" pageId="5" pageNumber="66" pagination="1 - 59" refId="ref12912" refString="Perkins, P. D. (2019 c) Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae). Zootaxa, 4708 (1), 1 - 59. https: // doi. org / 10.11646 / zootaxa. 4708.1.1" type="journal article" year="2019">Perkins 2019c</bibRefCitation>
); differentiated from
<taxonomicName id="4C014D34A9641961AE2BFC3F46E06024" authorityName="Perkins" authorityYear="2020" box="[341,491,1015,1041]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="bisinuatus">
<emphasis id="B975EAA5A9641961AE2BFC3F46E06024" box="[341,491,1015,1041]" italics="true" pageId="5" pageNumber="66">H. bisinuatus</emphasis>
</taxonomicName>
by the larger body size (ca. 3.003.38 vs.
<quantity id="4CF99B52A9641961ACB1FC3F43336024" box="[975,1080,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="2.74" pageId="5" pageNumber="66" unit="mm" value="2.74">2.74 mm</quantity>
), and the more strongly arcuate anterior margin of the pronotum. However, reliable determinations will require careful examination of the male genitalia (
<figureCitation id="133A2A32A9641961AE79FBF74647606C" box="[263,332,1087,1113]" captionStart="FIGURE 4" captionStartId="6.[151,250,1820,1844]" captionTargetBox="[172,1412,198,1769]" captionTargetId="figure-26@6.[160,1427,181,1796]" captionTargetPageId="6" captionText="FIGURE 4. Hydrochus astrictus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417291" httpUri="https://zenodo.org/record/4417291/files/figure.png" pageId="5" pageNumber="66">Fig. 4</figureCitation>
; described below).
</paragraph>
</subSubSection>
<subSubSection id="C31B653CA9641962AFB9FBAA43F86396" lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66" type="description">
<paragraph id="8BBE36B7A9641961AFB9FBAA44606104" blockId="5.[151,1437,690,1977]" pageId="5" pageNumber="66">
<emphasis id="B975EAA5A9641961AFB9FBAA465A6049" bold="true" box="[199,337,1122,1148]" pageId="5" pageNumber="66">Description</emphasis>
. Habitus as illustrated (
<figureCitation id="133A2A32A9641961AD23FBAB45A86048" box="[605,675,1123,1149]" captionStart="FIGURE 4" captionStartId="6.[151,250,1820,1844]" captionTargetBox="[172,1412,198,1769]" captionTargetId="figure-26@6.[160,1427,181,1796]" captionTargetPageId="6" captionText="FIGURE 4. Hydrochus astrictus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417291" httpUri="https://zenodo.org/record/4417291/files/figure.png" pageId="5" pageNumber="66">Fig. 4</figureCitation>
). Size:
<typeStatus id="54BA8815A9641961AD89FBAB44526048" box="[759,857,1123,1149]" pageId="5" pageNumber="66" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 3.16/1.10; head width 0.80; pronotum l/w 0.76/0.71; PA 0.71; PB 0.61; elytra 2.08/1.10. Dorsum grey to silver, with moderately strong iridescence, cervical groove black, elytra with black spots. Legs testaceous to brown, with tibiofemoral joints darker. Punctation on elytra ca. 2x that of pronotum. Elytra interstriae ca. 12x strial puncture diameter. Interstria 9
<superScript id="7C749BFFA9641961AEFFFB3A46856135" attach="left" box="[385,398,1266,1280]" fontSize="6" pageId="5" pageNumber="66">th</superScript>
more convex than others, overhanging 10
<superScript id="7C749BFFA9641961AC14FB3A447C6135" attach="left" box="[874,887,1266,1280]" fontSize="6" pageId="5" pageNumber="66">th</superScript>
interstria. Usual area of callus on 5
<superScript id="7C749BFFA9641961AA74FB3A421C6135" attach="left" box="[1290,1303,1266,1280]" fontSize="6" pageId="5" pageNumber="66">th</superScript>
moderately raised, moderately elongate, ca. 6 punctures in row on each side.
</paragraph>
<paragraph id="8BBE36B7A9641961AFB9FAF34342614C" blockId="5.[151,1437,690,1977]" pageId="5" pageNumber="66">Head with deep curving cervical groove from side to side behind eyes, groove punctate, cervical area behind groove impunctate, shining, with very fine, low, closely set longitudinal raised lines.</paragraph>
<paragraph id="8BBE36B7A9641961AFB9FA4B4629623C" blockId="5.[151,1437,690,1977]" pageId="5" pageNumber="66">Pronotum longer than wide (as ca. 76/71), widest at anterior margin, narrowed at base, sides straight or extremely slightly arcuate, smooth; anterior margin sinuate, median ½ markedly arcuate, fitting into cervical groove; anterior angles very slightly produced; depressions moderately deep, more densely punctate than reliefs between depressions.</paragraph>
<paragraph id="8BBE36B7A9641961AFB9F9DB42276264" blockId="5.[151,1437,690,1977]" pageId="5" pageNumber="66">Elytra with spaces between strial punctures ca. 0.51x their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with small angulation, impunctate area of apices rather large.</paragraph>
<paragraph id="8BBE36B7A9641961AFB9F993454F62AC" blockId="5.[151,1437,690,1977]" pageId="5" pageNumber="66">Ventral characters: Mentum coarsely densely punctate, with large, deep median fovea. Submentum with two deep foveae, posterior margin arcuate.</paragraph>
<paragraph id="8BBE36B7A9641961AFB9F96B43CA6344" blockId="5.[151,1437,690,1977]" pageId="5" pageNumber="66">
Male genitalia (
<figureCitation id="133A2A32A9641961AE04F96B46B46288" box="[378,447,1699,1725]" captionStart="FIGURE 4" captionStartId="6.[151,250,1820,1844]" captionTargetBox="[172,1412,198,1769]" captionTargetId="figure-26@6.[160,1427,181,1796]" captionTargetPageId="6" captionText="FIGURE 4. Hydrochus astrictus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.4417291" httpUri="https://zenodo.org/record/4417291/files/figure.png" pageId="5" pageNumber="66">Fig. 4</figureCitation>
) general characters: shaft of genitalia long and moderately narrow, basal piece slightly shorter than parameres; paramere tips slightly narrower than aedeagus shaft, medial margins strongly sclerotized, remainder of tips very weakly sclerotized; aedeagus widest at adbl, shaft moderately narrow, margins sinuate, shaft widest slightly before midlength (asgw), then narrowed at about proximal 1/3 of shaft (aslw), then becoming wider before tapering at tip; gonopore on both dorsal and ventral surfaces, the latter much larger than the former; basal piece almost cylindrical, with rim of orifice strongly sclerotized and distinctively hook-shaped.
</paragraph>
<paragraph id="8BBE36B7A9641961AFB9F8B34497638C" blockId="5.[151,1437,690,1977]" pageId="5" pageNumber="66">Dorsal surface: pdmm closely approximating alm, except at adbl; adtl at about midlength of shaft; triangular weakly sclerotized area of paramere very prominent; adbl very large.</paragraph>
<caption id="DF7E663FA9671962AFE9F8D444B76301" ID-DOI="http://doi.org/10.5281/zenodo.4417291" ID-Zenodo-Dep="4417291" box="[151,956,1820,1844]" httpUri="https://zenodo.org/record/4417291/files/figure.png" pageId="6" pageNumber="67" startId="6.[151,250,1820,1844]" targetBox="[172,1412,198,1769]" targetPageId="6">
<paragraph id="8BBE36B7A9671962AFE9F8D444B76301" blockId="6.[151,956,1820,1844]" box="[151,956,1820,1844]" pageId="6" pageNumber="67">
<emphasis id="B975EAA5A9671962AFE9F8D446186301" bold="true" box="[151,275,1820,1844]" pageId="6" pageNumber="67">FIGURE 4.</emphasis>
<taxonomicName id="4C014D34A9671962AE67F8D446EE6301" authority="Perkins, 2020" authorityName="Perkins" authorityYear="2020" box="[281,485,1820,1844]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="astrictus" status="sp. nov.">
<emphasis id="B975EAA5A9671962AE67F8D446EE6301" box="[281,485,1820,1844]" italics="true" pageId="6" pageNumber="67">Hydrochus astrictus</emphasis>
</taxonomicName>
<emphasis id="B975EAA5A9671962AE92F8D4452E6301" bold="true" box="[492,549,1820,1844]" pageId="6" pageNumber="67">
<taxonomicNameLabel id="A24657DEA9671962AE92F8D4452E6301" box="[492,549,1820,1844]" pageId="6" pageNumber="67" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, habitus and male genitalia of holotype.
</paragraph>
</caption>
<paragraph id="8BBE36B7A9671962AFB9F8AA43F86396" blockId="6.[151,1436,1890,2025]" pageId="6" pageNumber="67">Ventral surface: pvmm approximating alm except becoming very narrowly separated from one another and overlapping shaft in proxima1 1/3; gonopore long, distinct, with sclerotized margins in proximal ½.</paragraph>
</subSubSection>
<subSubSection id="C31B653CA9671962AFB9F862453C63DC" pageId="6" pageNumber="67" type="etymology">
<paragraph id="8BBE36B7A9671962AFB9F862453C63DC" blockId="6.[151,1436,1890,2025]" pageId="6" pageNumber="67">
<emphasis id="B975EAA5A9671962AFB9F862464563F1" bold="true" box="[199,334,1962,1988]" pageId="6" pageNumber="67">Etymology.</emphasis>
Named in reference to the tightly fitting components of the head and prothorax: cervical groove and anterior margin of the pronotum.
</paragraph>
</subSubSection>
</treatment>
</document>