462 lines
53 KiB
XML
462 lines
53 KiB
XML
<document id="EC3D741118F55D6F31FE0D808B728745" ID-DOI="10.1649/0010-065X-72.mo4.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5102840" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626277672172" checkinUser="carolina" docAuthor="Mullen, Logan J., Campbell, J. M. & Sikes, Derek S." docDate="2018" docId="03971078FFF9BC4BFD0CF9BEFC15CF3A" docLanguage="en" docName="TheColeopBull.mo.72.16.1-54.pdf" docOrigin="The Coleopterists Bulletin 72" docSource="http://dx.doi.org/10.1649/0010-065x-72.mo4.1" docStyle="DocumentStyle:A431C8F32F3D86E670C18C13DA9B5F2B.4:TheColeopBull.2000-2009.monograph" docStyleId="A431C8F32F3D86E670C18C13DA9B5F2B" docStyleName="TheColeopBull.2000-2009.monograph" docStyleVersion="4" docTitle="Phlaeopterus castaneus " docType="treatment" docVersion="3" lastPageNumber="16" masterDocId="FFAE6800FFF0BC44FFDAFFA6FFBDCE36" masterDocTitle="Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="54" masterPageNumber="1" pageNumber="10" updateTime="1699007718751" updateUser="plazi">
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<mods:title id="C40D8C0E7ED05DF0FB3251877CB43379">Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<mods:namePart id="B0D8D07FF488C216C666275F33010C0C">Mullen, Logan J.</mods:namePart>
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<mods:namePart id="53F834963AFB36612D4FFB23D91DFCC0">Campbell, J. M.</mods:namePart>
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<mods:namePart id="2AD44B0EAB2930D80CF51D6141ACB24E">Sikes, Derek S.</mods:namePart>
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<mods:typeOfResource id="89D4020616AC4E5B048D0D081FE8E477">text</mods:typeOfResource>
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<mods:title id="0B33F98117BB3FCD7CC558638CD1BB05">The Coleopterists Bulletin</mods:title>
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<mods:part id="70CA3D05953416CE04D30C5CB258D059">
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<mods:date id="CAF9E6DF26E7DEF2D8B41665FAADBC30">2018</mods:date>
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<mods:number id="2C05B2644BE7E8B359F788567AB240C4">2018-12-28</mods:number>
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<mods:number id="8F56C440341AF1D19F9EC77774E06657">72</mods:number>
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<paragraph id="8B81A16EFFF9BC4DFD0CF9BEFBD6C81B" blockId="9.[726,1131,1560,1608]" box="[726,1131,1560,1581]" pageId="9" pageNumber="10">
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<heading id="D0C91602FFF9BC4DFD0CF9BEFBD6C81B" box="[726,1131,1560,1581]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="10" reason="2">
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<emphasis id="B94A7D7CFFF9BC4DFD0CF9BEFBD6C81B" bold="true" box="[726,1131,1560,1581]" pageId="9" pageNumber="10">
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2.
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<taxonomicName id="4C3EDAEDFFF9BC4DFD2AF9BEFBD6C81B" ID-CoL="76ZZS" authority="(Casey, 1893)" authorityName="" baseAuthorityName="Casey" baseAuthorityYear="1893" box="[752,1131,1560,1581]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="castaneus">
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<emphasis id="B94A7D7CFFF9BC4DFD2AF9BEFC66C81B" bold="true" box="[752,987,1560,1581]" italics="true" pageId="9" pageNumber="10">Phlaeopterus castaneus</emphasis>
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(
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<bibRefCitation id="EFAFDC9FFFF9BC4DFC33F9BEFBD9C81B" author="Casey" box="[1001,1124,1560,1581]" pageId="9" pageNumber="10" pagination="281 - 624" refId="ref28683" refString="Casey, T. L. 1893. Coleopterological Notices V. Annals of the New York Academy of Sciences 7 (1): 281 - 624." type="journal article" year="1893">Casey, 1893</bibRefCitation>
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)
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C324F2E5FFF9BC4DFCD6F995FB88C87E" box="[780,1077,1587,1608]" pageId="9" pageNumber="10" type="description">
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<paragraph id="8B81A16EFFF9BC4DFCD6F995FB88C87E" blockId="9.[726,1131,1560,1608]" box="[780,1077,1587,1608]" pageId="9" pageNumber="10">
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(
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<figureCitation id="1305BDEBFFF9BC4DFCC9F995FCD5C87E" box="[787,872,1587,1608]" captionStart="Fig" captionStartId="10.[197,228,1731,1750]" captionTargetBox="[177,1188,196,1713]" captionTargetId="figure-0@10.[175,1190,194,1715]" captionTargetPageId="10" captionText="Fig. 1. Phlaeopterus species, habitus. A) P. bakerensis, female (UAMObs:Ento:233271), B) P. castaneus, female" figureDoi="http://doi.org/10.5281/zenodo.5102842" httpUri="https://zenodo.org/record/5102842/files/figure.png" pageId="9" pageNumber="10">Figs. 1B</figureCitation>
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,
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<figureCitation id="1305BDEBFFF9BC4DFCAEF995FC2FC87E" box="[884,914,1587,1608]" captionStart="Fig" captionStartId="15.[709,740,1495,1514]" captionTargetBox="[690,1168,666,1478]" captionTargetId="figure-473@15.[688,1170,664,1480]" captionTargetPageId="15" captionText="Fig. 6. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. bakerensis, B) P. castaneus. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected. In B, red triangles represent localities of P. castaneus castaneus, red squares represent localities of P. castaneus cascadiensis, and green stars represent areas of overlap of the two subspecies." figureDoi="http://doi.org/10.5281/zenodo.5102855" httpUri="https://zenodo.org/record/5102855/files/figure.png" pageId="9" pageNumber="10">6B</figureCitation>
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, 16B, 21F, 22A)
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</paragraph>
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</subSubSection>
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<subSubSection id="C324F2E5FFF9BC4EFD74F9FBFB1DC8D8" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="reference_group">
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<paragraph id="8B81A16EFFF9BC4DFD74F9FBFB28C844" blockId="9.[685,1173,1629,1730]" box="[686,1173,1629,1650]" pageId="9" pageNumber="10">
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<treatmentCitationGroup id="AB2E8640FFF9BC4DFD74F9FBFB28C844" box="[686,1173,1629,1650]" pageId="9" pageNumber="10">
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<treatmentCitation id="0A9F877FFFF9BC4DFD74F9FBFC70C844" author="Casey" box="[686,973,1629,1650]" page="403" pageId="9" pageNumber="10" year="1893">
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<taxonomicName id="4C3EDAEDFFF9BC4DFD74F9FBFC70C844" ID-CoL="56WMW" authority="Casey 1893: 403" authorityName="Casey" authorityPageNumber="403" authorityYear="1893" box="[686,973,1629,1650]" class="Insecta" family="Staphylinidae" genus="Tilea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="castanea">
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<emphasis id="B94A7D7CFFF9BC4DFD74F9FBFC8DC844" box="[686,816,1629,1650]" italics="true" pageId="9" pageNumber="10">Tilea castanea</emphasis>
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<bibRefCitation id="EFAFDC9FFFF9BC4DFCEFF9FBFC70C844" author="Casey" box="[821,973,1629,1650]" pageId="9" pageNumber="10" pagination="281 - 624" refId="ref28683" refString="Casey, T. L. 1893. Coleopterological Notices V. Annals of the New York Academy of Sciences 7 (1): 281 - 624." type="journal article" year="1893">Casey 1893: 403</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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[original description].
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B81A16EFFF9BC4DFD77F9DEFC02C8F4" blockId="9.[685,1173,1629,1730]" pageId="9" pageNumber="10">
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<treatmentCitationGroup id="AB2E8640FFF9BC4DFD77F9DEFC02C8F4" pageId="9" pageNumber="10">
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<taxonomicName id="4C3EDAEDFFF9BC4DFD77F9DEFB9DC8BB" ID-CoL="76ZZS" authority="(Casey, 1893)" authorityName="" baseAuthorityName="Casey" baseAuthorityYear="1893" box="[685,1056,1656,1677]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="castaneus">
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<emphasis id="B94A7D7CFFF9BC4DFD77F9DEFC2CC8BB" box="[685,913,1656,1677]" italics="true" pageId="9" pageNumber="10">Phlaeopterus castaneus</emphasis>
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(
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<bibRefCitation id="EFAFDC9FFFF9BC4DFC45F9DEFBA4C8BB" author="Casey" box="[927,1049,1656,1677]" pageId="9" pageNumber="10" pagination="281 - 624" refId="ref28683" refString="Casey, T. L. 1893. Coleopterological Notices V. Annals of the New York Academy of Sciences 7 (1): 281 - 624." type="journal article" year="1893">Casey, 1893</bibRefCitation>
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||
)
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||
</taxonomicName>
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. Moore and Legner 1975: 208; Campbell and Davies 1991: 5;
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<treatmentCitation id="0A9F877FFFF9BC4DFD12F90BFC06C8F4" author="Bousquet" box="[712,955,1709,1730]" page="89" pageId="9" pageNumber="10" year="2013">
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||
<bibRefCitation id="EFAFDC9FFFF9BC4DFD12F90BFC06C8F4" author="Bousquet" box="[712,955,1709,1730]" pageId="9" pageNumber="10" refId="ref28340" refString="Bousquet, Y., P. Bouchard, A. E. Davies, and D. S. Sikes. 2013. Checklist of Beetles (Coleoptera) of Canada and Alaska. Second Edition. Pensoft, Sofia, Bulgaria." type="book" year="2013">
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Bousquet
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<emphasis id="B94A7D7CFFF9BC4DFCF0F90BFCE6C8F4" box="[810,859,1709,1730]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
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2013: 89
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</bibRefCitation>
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</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<caption id="DF41F1E6FFFABC4EFF1FF965FB14C8E0" ID-DOI="http://doi.org/10.5281/zenodo.5102842" ID-Zenodo-Dep="5102842" box="[197,1193,1731,1750]" httpUri="https://zenodo.org/record/5102842/files/figure.png" pageId="10" pageNumber="11" startId="10.[197,228,1731,1750]" targetBox="[177,1188,196,1713]" targetPageId="10">
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||
<paragraph id="8B81A16EFFFABC4EFF1FF965FB14C8E0" blockId="10.[197,1193,1731,1750]" box="[197,1193,1731,1750]" pageId="10" pageNumber="11">
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||
<emphasis id="B94A7D7CFFFABC4EFF1FF965FF42C8E0" bold="true" box="[197,255,1731,1750]" pageId="10" pageNumber="11">Fig. 1.</emphasis>
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<taxonomicName id="4C3EDAEDFFFABC4EFECEF965FE76C8E0" box="[276,459,1731,1750]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B94A7D7CFFFABC4EFECEF965FE39C8E0" box="[276,388,1731,1750]" italics="true" pageId="10" pageNumber="11">Phlaeopterus</emphasis>
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species
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</taxonomicName>
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, habitus. A)
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<taxonomicName id="4C3EDAEDFFFABC4EFDE7F965FD10C8E0" ID-CoL="8Y8CT" authorityName="Mullen & Campbell & Sikes" authorityYear="2018" box="[573,685,1731,1750]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bakerensis">
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||
<emphasis id="B94A7D7CFFFABC4EFDE7F965FD10C8E0" box="[573,685,1731,1750]" italics="true" pageId="10" pageNumber="11">P. bakerensis</emphasis>
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</taxonomicName>
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, female (UAMObs:Ento:233271), B)
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<taxonomicName id="4C3EDAEDFFFABC4EFC23F965FBDEC8E0" ID-CoL="76ZZS" authorityName="" baseAuthorityName="Casey" baseAuthorityYear="1893" box="[1017,1123,1731,1750]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="castaneus">
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<emphasis id="B94A7D7CFFFABC4EFC23F965FBDEC8E0" box="[1017,1123,1731,1750]" italics="true" pageId="10" pageNumber="11">P. castaneus</emphasis>
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</taxonomicName>
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, female
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</paragraph>
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</caption>
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<paragraph id="8B81A16EFFFABC4EFF77F97DFB1DC8D8" blockId="10.[173,311,1755,1774]" box="[173,1184,1755,1774]" lastBlockId="10.[1061,1184,1755,1774]" pageId="10" pageNumber="11">
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||
(UAMObs:Ento bars =
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||
<quantity id="4CC60C8BFFFABC4EFBB2F97DFB21C8D8" box="[1128,1180,1755,1774]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="mm" value="1.0">1 mm</quantity>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C324F2E5FFFABC4AFEEDF97DFD81CDDE" lastPageId="14" lastPageNumber="15" pageId="10" pageNumber="11" type="description">
|
||
<paragraph id="8B81A16EFFFABC4FFEEDF97DFE10C8DF" blockId="10.[311,1054,1755,1774]" lastBlockId="11.[151,429,1750,1793]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
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||
:234040), C)
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||
<taxonomicName id="4C3EDAEDFFFABC4EFE72F97DFDB2C8D8" box="[424,527,1755,1774]" class="Insecta" family="Staphylinidae" genus="Tilea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="cavicollis">
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||
<emphasis id="B94A7D7CFFFABC4EFE72F97DFDB2C8D8" box="[424,527,1755,1774]" italics="true" pageId="10" pageNumber="11">P. cavicollis</emphasis>
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||
</taxonomicName>
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||
, male (UAMObs:Ento:235795), D)
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||
<taxonomicName id="4C3EDAEDFFFABC4EFC9DF97DFC1DC8D8" baseAuthorityName="Shavrin" baseAuthorityYear="2001" box="[839,928,1755,1774]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="czerskyi">
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||
<emphasis id="B94A7D7CFFFABC4EFC9DF97DFC1DC8D8" box="[839,928,1755,1774]" italics="true" pageId="10" pageNumber="11">P. czerskyi</emphasis>
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||
</taxonomicName>
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||
, female. Scale (UAMObs:Ento:232406), C)
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||
<emphasis id="B94A7D7CFFFBBC4FFE7AF970FE10C8DF" box="[416,429,1750,1769]" italics="true" pageId="11" pageNumber="12">P</emphasis>
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||
</paragraph>
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||
<caption id="DF41F1E6FFFBBC4FFF75F918FB29C8E7" ID-DOI="http://doi.org/10.5281/zenodo.5102845" ID-Zenodo-Dep="5102845" box="[175,1172,1726,1745]" httpUri="https://zenodo.org/record/5102845/files/figure.png" pageId="11" pageNumber="12" startId="11.[175,206,1726,1745]" targetBox="[156,1167,196,1708]" targetPageId="11">
|
||
<paragraph id="8B81A16EFFFBBC4FFF75F918FB29C8E7" blockId="11.[175,1172,1726,1745]" box="[175,1172,1726,1745]" pageId="11" pageNumber="12">
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||
<emphasis id="B94A7D7CFFFBBC4FFF75F918FF51C8E7" bold="true" box="[175,236,1726,1745]" pageId="11" pageNumber="12">Fig. 2.</emphasis>
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||
<taxonomicName id="4C3EDAEDFFFBBC4FFEDBF918FE06C8E7" box="[257,443,1726,1745]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="undetermined">
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||
<emphasis id="B94A7D7CFFFBBC4FFEDBF918FECCC8E7" box="[257,369,1726,1745]" italics="true" pageId="11" pageNumber="12">Phlaeopterus</emphasis>
|
||
species
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||
</taxonomicName>
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||
, habitus. A)
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<taxonomicName id="4C3EDAEDFFFBBC4FFDEFF918FD1CC8E7" authorityName="Mullen and Campbell" authorityYear="2018" box="[565,673,1726,1745]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="elongatus">
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||
<emphasis id="B94A7D7CFFFBBC4FFDEFF918FD1CC8E7" box="[565,673,1726,1745]" italics="true" pageId="11" pageNumber="12">P. elongatus</emphasis>
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||
</taxonomicName>
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, male (UAMObs:Ento:232543), B)
|
||
<taxonomicName id="4C3EDAEDFFFBBC4FFC3CF918FBF1C8E7" authorityName="Casey" authorityYear="1886" box="[998,1100,1726,1745]" class="Insecta" family="Cleridae" genus="Phloeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="filicornis">
|
||
<emphasis id="B94A7D7CFFFBBC4FFC3CF918FBF1C8E7" box="[998,1100,1726,1745]" italics="true" pageId="11" pageNumber="12">P. filicornis</emphasis>
|
||
</taxonomicName>
|
||
, female
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B81A16EFFFBBC4FFF4DF948FE24C937" blockId="11.[151,429,1750,1793]" box="[151,409,1774,1793]" pageId="11" pageNumber="12">
|
||
235496). Scale bars =
|
||
<quantity id="4CC60C8BFFFBBC4FFEBAF948FE29C937" box="[352,404,1774,1793]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="12" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B81A16EFFFBBC48FBD6F970FE9BC8DC" blockId="11.[1036,1173,1750,1769]" lastBlockId="12.[291,1070,1751,1794]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||
UAMObs:Ento:.
|
||
<taxonomicName id="4C3EDAEDFFFBBC4FFE66F970FE5AC8DF" authorityName="Hatch" authorityYear="1957" box="[444,487,1750,1769]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="frosti">
|
||
<emphasis id="B94A7D7CFFFBBC4FFE66F970FE5AC8DF" box="[444,487,1750,1769]" italics="true" pageId="11" pageNumber="12">frosti</emphasis>
|
||
</taxonomicName>
|
||
, male (UAMObs:Ento:235798), D)
|
||
<taxonomicName id="4C3EDAEDFFFBBC4FFCE3F970FC13C8DF" box="[825,942,1750,1769]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fusconiger">
|
||
<emphasis id="B94A7D7CFFFBBC4FFCE3F970FC13C8DF" box="[825,942,1750,1769]" italics="true" pageId="11" pageNumber="12">P. fusconiger</emphasis>
|
||
</taxonomicName>
|
||
, female (Ento:234115),
|
||
</paragraph>
|
||
<caption id="DF41F1E6FFFCBC48FF1FF919FB17C8E4" ID-DOI="http://doi.org/10.5281/zenodo.5102847" ID-Zenodo-Dep="5102847" box="[197,1194,1727,1746]" httpUri="https://zenodo.org/record/5102847/files/figure.png" pageId="12" pageNumber="13" startId="12.[197,228,1727,1746]" targetBox="[177,1188,196,1710]" targetPageId="12">
|
||
<paragraph id="8B81A16EFFFCBC48FF1FF919FB17C8E4" blockId="12.[197,1194,1727,1746]" box="[197,1194,1727,1746]" pageId="12" pageNumber="13">
|
||
<emphasis id="B94A7D7CFFFCBC48FF1FF919FF40C8E4" bold="true" box="[197,253,1727,1746]" pageId="12" pageNumber="13">Fig. 3.</emphasis>
|
||
<taxonomicName id="4C3EDAEDFFFCBC48FEC9F919FE74C8E4" box="[275,457,1727,1746]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B94A7D7CFFFCBC48FEC9F919FE3EC8E4" box="[275,387,1727,1746]" italics="true" pageId="12" pageNumber="13">Phlaeopterus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, habitus. A)
|
||
<taxonomicName id="4C3EDAEDFFFCBC48FDE2F919FD39C8E4" authorityName="Mullen and Campbell" authorityYear="2018" box="[568,644,1727,1746]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="hatchi">
|
||
<emphasis id="B94A7D7CFFFCBC48FDE2F919FD39C8E4" box="[568,644,1727,1746]" italics="true" pageId="12" pageNumber="13">P. hatchi</emphasis>
|
||
</taxonomicName>
|
||
, male (UAMObs:Ento:232429), B)
|
||
<taxonomicName id="4C3EDAEDFFFCBC48FC60F919FBB0C8E4" authorityName="Hatch" authorityYear="1957" box="[954,1037,1727,1746]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="houkae">
|
||
<emphasis id="B94A7D7CFFFCBC48FC60F919FBB0C8E4" box="[954,1037,1727,1746]" italics="true" pageId="12" pageNumber="13">P. houkae</emphasis>
|
||
</taxonomicName>
|
||
, male (UAMObs:
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B81A16EFFFCBC48FF77F949FEE1C934" blockId="12.[173,294,1751,1794]" lastBlockId="12.[291,1070,1751,1794]" pageId="12" pageNumber="13">
|
||
Scale bars =:Ento:234056). C)
|
||
<taxonomicName id="4C3EDAEDFFFCBC48FE92F971FE7CC8DC" authorityName="Mullen and Campbell" authorityYear="2018" box="[328,449,1751,1770]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="kavanaughi">
|
||
<emphasis id="B94A7D7CFFFCBC48FE92F971FE7CC8DC" box="[328,449,1751,1770]" italics="true" pageId="12" pageNumber="13">P. kavanaughi</emphasis>
|
||
</taxonomicName>
|
||
, female (UAMObs:Ento:232423), D)
|
||
<taxonomicName id="4C3EDAEDFFFCBC48FCD3F971FC39C8DC" authorityName="Hatch" authorityYear="1957" box="[777,900,1751,1770]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="lagrandeuri">
|
||
<emphasis id="B94A7D7CFFFCBC48FCD3F971FC39C8DC" box="[777,900,1751,1770]" italics="true" pageId="12" pageNumber="13">P. lagrandeuri</emphasis>
|
||
</taxonomicName>
|
||
, female (UAMObs
|
||
<quantity id="4CC60C8BFFFCBC48FEF9F949FEEAC934" box="[291,343,1775,1794]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="13" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF41F1E6FFFDBC49FF75F968FB29C8D7" ID-DOI="http://doi.org/10.5281/zenodo.5102849" ID-Zenodo-Dep="5102849" box="[175,1172,1742,1761]" httpUri="https://zenodo.org/record/5102849/files/figure.png" pageId="13" pageNumber="14" startId="13.[175,206,1742,1761]" targetBox="[157,1166,196,1725]" targetPageId="13">
|
||
<paragraph id="8B81A16EFFFDBC49FF75F968FB29C8D7" blockId="13.[175,1172,1742,1761]" box="[175,1172,1742,1761]" pageId="13" pageNumber="14">
|
||
<emphasis id="B94A7D7CFFFDBC49FF75F968FF54C8D7" bold="true" box="[175,233,1742,1761]" pageId="13" pageNumber="14">Fig. 4.</emphasis>
|
||
<taxonomicName id="4C3EDAEDFFFDBC49FF25F968FE0BC8D7" box="[255,438,1742,1761]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B94A7D7CFFFDBC49FF25F968FED2C8D7" box="[255,367,1742,1761]" italics="true" pageId="13" pageNumber="14">Phlaeopterus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, habitus. A)
|
||
<taxonomicName id="4C3EDAEDFFFDBC49FDF2F968FD24C8D7" authorityName="Hatch" authorityYear="1957" box="[552,665,1742,1761]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="loganensis">
|
||
<emphasis id="B94A7D7CFFFDBC49FDF2F968FD24C8D7" box="[552,665,1742,1761]" italics="true" pageId="13" pageNumber="14">P. loganensis</emphasis>
|
||
</taxonomicName>
|
||
, female (UAMObs:Ento:233072), B)
|
||
<taxonomicName id="4C3EDAEDFFFDBC49FC3EF968FBE3C8D7" authorityName="Mullen & Campbell & Sikes" authorityYear="2018" baseAuthorityName="Casey" baseAuthorityYear="1885" box="[996,1118,1742,1761]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="longipennis">
|
||
<emphasis id="B94A7D7CFFFDBC49FC3EF968FBE3C8D7" box="[996,1118,1742,1761]" italics="true" pageId="13" pageNumber="14">P. longipennis</emphasis>
|
||
</taxonomicName>
|
||
, male
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B81A16EFFFDBC49FF42F940FE1AC8CF" blockId="13.[152,423,1766,1809]" box="[152,423,1766,1785]" pageId="13" pageNumber="14">
|
||
(UAMObs:Ento:235802), C)
|
||
<emphasis id="B94A7D7CFFFDBC49FE4CF940FE1AC8CF" box="[406,423,1766,1785]" italics="true" pageId="13" pageNumber="14">P.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B81A16EFFFDBC49FF42F958FE24C927" blockId="13.[152,423,1766,1809]" box="[152,409,1790,1809]" pageId="13" pageNumber="14">
|
||
233445). Scale bars =
|
||
<quantity id="4CC60C8BFFFDBC49FEBAF958FE28C927" box="[352,405,1790,1809]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B81A16EFFFDBC4AFBD6F940FD81CDDE" blockId="13.[1036,1173,1766,1785]" lastBlockId="14.[173,1193,957,1000]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||
UAMObs:Ento: (UAMObs:Ento:233301). Scale bars =
|
||
<quantity id="4CC60C8BFFFEBC4AFDD9FC73FD8ACDDE" box="[515,567,981,1000]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF41F1E6FFFEBC4AFF1FFC1BFB14CDE6" ID-DOI="http://doi.org/10.5281/zenodo.5102853" ID-Zenodo-Dep="5102853" box="[197,1193,957,976]" httpUri="https://zenodo.org/record/5102853/files/figure.png" pageId="14" pageNumber="15" startId="14.[197,228,957,976]" targetBox="[177,1188,196,940]" targetPageId="14">
|
||
<paragraph id="8B81A16EFFFEBC4AFF1FFC1BFB14CDE6" blockId="14.[173,1193,957,1000]" box="[197,1193,957,976]" pageId="14" pageNumber="15">
|
||
<emphasis id="B94A7D7CFFFEBC4AFF1FFC1BFEBCCDE6" bold="true" box="[197,257,957,976]" pageId="14" pageNumber="15">Fig. 5.</emphasis>
|
||
<taxonomicName id="4C3EDAEDFFFEBC4AFECCFC1BFE72CDE6" box="[278,463,957,976]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B94A7D7CFFFEBC4AFECCFC1BFE3BCDE6" box="[278,390,957,976]" italics="true" pageId="14" pageNumber="15">Phlaeopterus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, habitus. A)
|
||
<taxonomicName id="4C3EDAEDFFFEBC4AFD92FC1BFD0ACDE6" authorityName="Mullen and Campbell" authorityYear="2018" box="[584,695,957,976]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="olympicus">
|
||
<emphasis id="B94A7D7CFFFEBC4AFD92FC1BFD0ACDE6" box="[584,695,957,976]" italics="true" pageId="14" pageNumber="15">P. olympicus</emphasis>
|
||
</taxonomicName>
|
||
, female (UAMObs:Ento:232960), B)
|
||
<taxonomicName id="4C3EDAEDFFFEBC4AFBD1FC1BFBCCCDE6" authorityName="Mullen and Campbell" authorityYear="2018" box="[1035,1137,957,976]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="smetanai">
|
||
<emphasis id="B94A7D7CFFFEBC4AFBD1FC1BFBCCCDE6" box="[1035,1137,957,976]" italics="true" pageId="14" pageNumber="15">P. smetanai</emphasis>
|
||
</taxonomicName>
|
||
, male
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C324F2E5FFFEBC4AFF77FB89FD8CCA4F" pageId="14" pageNumber="15" type="reference_group">
|
||
<paragraph id="8B81A16EFFFEBC4AFF77FB89FD8CCA4F" blockId="14.[173,659,1071,1145]" pageId="14" pageNumber="15">
|
||
<treatmentCitationGroup id="AB2E8640FFFEBC4AFF77FB89FD8CCA4F" pageId="14" pageNumber="15">
|
||
<treatmentCitation id="0A9F877FFFFEBC4AFF77FB89FD80CA72" author="Hatch" box="[173,573,1071,1092]" page="59" pageId="14" pageNumber="15" year="1957">
|
||
<taxonomicName id="4C3EDAEDFFFEBC4AFF77FB89FD80CA72" ID-CoL="76ZZR" authority="Hatch 1957: 59" authorityName="Hatch" authorityPageNumber="59" authorityYear="1957" box="[173,573,1071,1092]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="cascadiensis">
|
||
<emphasis id="B94A7D7CFFFEBC4AFF77FB89FE15CA72" box="[173,424,1071,1092]" italics="true" pageId="14" pageNumber="15">Phlaeopterus cascadiensis</emphasis>
|
||
<bibRefCitation id="EFAFDC9FFFFEBC4AFE77FB89FD80CA72" author="Hatch" box="[429,573,1071,1092]" pageId="14" pageNumber="15" refId="ref28862" refString="Hatch, M. H. 1957. The Beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology, Seattle, WA." type="book" year="1957">Hatch 1957: 59</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
[original description]. Campbell and Davies 1991: 5.
|
||
<emphasis id="B94A7D7CFFFEBC4AFDBCFBEFFD8CCA4F" bold="true" pageId="14" pageNumber="15">New status as subspecies and synonymy.</emphasis>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFFEBC4AFF1FFB3FFE47CA98" box="[197,506,1177,1198]" pageId="14" pageNumber="15" type="materials_examined">
|
||
<paragraph id="8B81A16EFFFEBC4AFF1FFB3FFE47CA98" blockId="14.[173,660,1177,1730]" box="[197,506,1177,1198]" pageId="14" pageNumber="15">
|
||
<materialsCitation id="3B56AB33FFFEBC4AFF1FFB3FFE4BCA98" box="[197,502,1177,1198]" country="United States of America" location="Colorado" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Colorado" typeStatus="holotype">
|
||
<emphasis id="B94A7D7CFFFEBC4AFF1FFB3FFEEBCA98" bold="true" box="[197,342,1177,1198]" pageId="14" pageNumber="15">
|
||
<typeStatus id="54851FCCFFFEBC4AFF1FFB3FFF4ACA98" box="[197,247,1177,1198]" pageId="14" pageNumber="15">Type</typeStatus>
|
||
Locality.
|
||
</emphasis>
|
||
<collectingRegion id="49FA6F8CFFFEBC4AFE87FB3FFE05CA98" box="[349,440,1177,1198]" country="United States of America" name="Colorado" pageId="14" pageNumber="15">Colorado</collectingRegion>
|
||
,
|
||
<collectingCountry id="F329E1FEFFFEBC4AFE19FB3FFE4BCA98" box="[451,502,1177,1198]" name="United States of America" pageId="14" pageNumber="15">USA</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFFEBC4BFF1FFB12FE3ACF11" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
|
||
<paragraph id="8B81A16EFFFEBC4BFF1FFB12FE3ACF11" blockId="14.[173,660,1177,1730]" lastBlockId="15.[151,639,194,1730]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
|
||
<emphasis id="B94A7D7CFFFEBC4AFF1FFB12FE05CAFF" bold="true" box="[197,440,1204,1225]" pageId="14" pageNumber="15">Redescription. Habitus:</emphasis>
|
||
Length
|
||
<quantity id="4CC60C8BFFFEBC4AFDC8FB12FD2DCAFF" box="[530,656,1203,1225]" metricMagnitude="-3" metricUnit="m" metricValue="6.65" metricValueMax="7.7" metricValueMin="5.6" pageId="14" pageNumber="15" unit="mm" value="6.65" valueMax="7.7" valueMin="5.6">5.6–7.7 mm</quantity>
|
||
. Dark brown to reddish brown; lateral margins of pronotum often lighter (
|
||
<figureCitation id="1305BDEBFFFEBC4AFE70FB4FFE46CAC8" box="[426,507,1257,1278]" captionStart="Fig" captionStartId="10.[197,228,1731,1750]" captionTargetBox="[177,1188,196,1713]" captionTargetId="figure-0@10.[175,1190,194,1715]" captionTargetPageId="10" captionText="Fig. 1. Phlaeopterus species, habitus. A) P. bakerensis, female (UAMObs:Ento:233271), B) P. castaneus, female" figureDoi="http://doi.org/10.5281/zenodo.5102842" httpUri="https://zenodo.org/record/5102842/files/figure.png" pageId="14" pageNumber="15">Fig. 1B</figureCitation>
|
||
); head, elytra, palpi, and antennae sometimes lighter.
|
||
<emphasis id="B94A7D7CFFFEBC4AFD89FAA5FD2ECB2E" bold="true" box="[595,659,1283,1304]" pageId="14" pageNumber="15">Head:</emphasis>
|
||
Moderately broad, ratio of width across eyes to length of head = 1.0–1.3. Interantennal groove broadly and deeply impressed. Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered setae near ventral margin. Antennomeres 5–10 at least 2 times longer than wide; antennomeres 4–11 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with L-shaped row of setae. Labrum with sensory pores along entire surface. Labial palpi with 3
|
||
<superScript id="7C4B0C26FFFEBC4AFE18F99BFE6CC87D" attach="left" box="[450,465,1597,1611]" fontSize="6" pageId="14" pageNumber="15">rd</superScript>
|
||
palpomere 1.6–1.8 times longer than palpomere 2.
|
||
<emphasis id="B94A7D7CFFFEBC4AFE2BF9FBFDF5C844" bold="true" box="[497,584,1629,1650]" pageId="14" pageNumber="15">Thorax:</emphasis>
|
||
Pronotum broad (Fig. 16B), length to width ratio = 0.63–0.73; ratio of pronotal width to head width 1.5–1.61; maximum width subequal to elytral width at humeral angles; punctures on dorsal surface separated by average distance equal to twice diameter of a puncture; lateral margins explanate posterad lateral foveae, narrowly explanate (
|
||
<taxonomicName id="4C3EDAEDFFFEBC4AFBB3FBD9FCD8CA98" authority="castaneus" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subSpecies" species="castaneus" subSpecies="castaneus">
|
||
<emphasis id="B94A7D7CFFFEBC4AFBB3FBD9FCD8CA98" italics="true" pageId="14" pageNumber="15">P. castaneus castaneus</emphasis>
|
||
</taxonomicName>
|
||
) to not explanate (
|
||
<taxonomicName id="4C3EDAEDFFFEBC4AFBEAFB3FFC84CAFF" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subSpecies" species="castaneus" subSpecies="cascadiensis">
|
||
<emphasis id="B94A7D7CFFFEBC4AFBEAFB3FFC84CAFF" italics="true" pageId="14" pageNumber="15">P. castaneus cascadiensis</emphasis>
|
||
</taxonomicName>
|
||
) anterad lateral foveae; lateral foveae deeply impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.2–2.5 times longer than pronotum; apical margins broadly convex, uniformly dark reddish brown (
|
||
<taxonomicName id="4C3EDAEDFFFEBC4AFC14FAB8FB1CCB05" authority="castaneus" box="[974,1185,1310,1331]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subSpecies" species="castaneus" subSpecies="castaneus">
|
||
<emphasis id="B94A7D7CFFFEBC4AFC14FAB8FB1CCB05" box="[974,1185,1310,1331]" italics="true" pageId="14" pageNumber="15">P. castaneus castaneus</emphasis>
|
||
</taxonomicName>
|
||
) or with sutural area lighter reddish brown than remainder of elytral surface (
|
||
<taxonomicName id="4C3EDAEDFFFEBC4AFBF6FAF5FC86CBB5" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subSpecies" species="castaneus" subSpecies="cascadiensis">
|
||
<emphasis id="B94A7D7CFFFEBC4AFBF6FAF5FC86CBB5" italics="true" pageId="14" pageNumber="15">P. castaneus cascadiensis</emphasis>
|
||
</taxonomicName>
|
||
). Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum complete but not strongly carinate.
|
||
<emphasis id="B94A7D7CFFFEBC4AFCC4FA7EFCE9CBDB" bold="true" box="[798,852,1496,1517]" pageId="14" pageNumber="15">Legs:</emphasis>
|
||
All tibiae with dense pubescence from base to apex. Metatrochanter without tooth on apical margin.
|
||
<emphasis id="B94A7D7CFFFEBC4AFCA1F9ABFC5BC814" bold="true" box="[891,998,1549,1570]" pageId="14" pageNumber="15">Abdomen:</emphasis>
|
||
Wing-folding spicules on tergites IV and V broad, transverse, narrowly separated; tergite VI lacking wing-folding patches.
|
||
<emphasis id="B94A7D7CFFFEBC4AFCC5F9FBFC37C844" bold="true" box="[799,906,1629,1650]" pageId="14" pageNumber="15">Aedeagus:</emphasis>
|
||
Length
|
||
<quantity id="4CC60C8BFFFEBC4AFC31F9FBFBC2C844" box="[1003,1151,1628,1650]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" metricValueMax="1.36" metricValueMin="1.2" pageId="14" pageNumber="15" unit="mm" value="1.28" valueMax="1.36" valueMin="1.2">1.20–1.36 mm</quantity>
|
||
for
|
||
<taxonomicName id="4C3EDAEDFFFEBC4AFD19F9D1FC0DC8BA" box="[707,944,1655,1676]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subSpecies" species="castaneus" subSpecies="cascadiensis">
|
||
<emphasis id="B94A7D7CFFFEBC4AFD19F9D1FC0DC8BA" box="[707,944,1655,1676]" italics="true" pageId="14" pageNumber="15">P. castaneus cascadiensis</emphasis>
|
||
</taxonomicName>
|
||
(Fig. 21F),
|
||
<quantity id="4CC60C8BFFFEBC4AFBFAF9D1FB14C8BA" box="[1056,1193,1655,1677]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.48" metricValueMin="1.32" pageId="14" pageNumber="15" unit="mm" value="1.4" valueMax="1.48" valueMin="1.32">1.32–1.48 mm</quantity>
|
||
for
|
||
<taxonomicName id="4C3EDAEDFFFEBC4AFD31F934FC68C891" authority="castaneus" box="[747,981,1682,1703]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subSpecies" species="castaneus" subSpecies="castaneus">
|
||
<emphasis id="B94A7D7CFFFEBC4AFD31F934FC68C891" box="[747,981,1682,1703]" italics="true" pageId="14" pageNumber="15">P. castaneus castaneus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1305BDEBFFFEBC4AFC32F934FBF4C891" box="[1000,1097,1682,1703]" captionStart="Fig" captionStartId="27.[175,206,1390,1409]" captionTargetBox="[156,1167,196,1372]" captionTargetId="figure-125@27.[154,1169,194,1374]" captionTargetPageId="27" captionText="Fig. 22. Phlaeopterus species, aedeagi (left = ventral view; right = dorsal view of median lobe). A) P. castaneus castaneus, B–D) variation in P. cavicollis." figureDoi="http://doi.org/10.5281/zenodo.5102875" httpUri="https://zenodo.org/record/5102875/files/figure.png" pageId="14" pageNumber="15">Fig. 22A</figureCitation>
|
||
). Median lobe narrowed abruptly just before apex, with longitudinal carina at apex. Parameres narrow. Internal sac variable; 2/3 to half length of median lobe; covered in microspinules; with or without subapical transverse fold.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFFFBC4BFF75FE8AFDA0CD0C" pageId="15" pageNumber="16" type="materials_examined">
|
||
<paragraph id="8B81A16EFFFFBC4BFF75FE8AFE2FCCB6" blockId="15.[151,639,194,1730]" pageId="15" pageNumber="16">
|
||
<emphasis id="B94A7D7CFFFFBC4BFF75FE8AFDB0CF77" bold="true" box="[175,525,300,321]" pageId="15" pageNumber="16">
|
||
Type Specimens.
|
||
<taxonomicName id="4C3EDAEDFFFFBC4BFEB4FE8AFDB8CF77" authorityName="Casey" authorityYear="1893" box="[366,517,300,321]" class="Insecta" family="Staphylinidae" genus="Tilea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="castanea">
|
||
<emphasis id="B94A7D7CFFFFBC4BFEB4FE8AFDB8CF77" bold="true" box="[366,517,300,321]" italics="true" pageId="15" pageNumber="16">Tilea castanea</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
</emphasis>
|
||
<typeStatus id="54851FCCFFFFBC4BFDC1FE8AFDC0CF77" box="[539,637,300,321]" pageId="15" pageNumber="16" type="lectotype">Lectotype</typeStatus>
|
||
male (here designated, UAMObs:Ento:235783) labeled as follows: Col./³/CASEY bequest 1925/ TYPE USNM 48100/[
|
||
<taxonomicName id="4C3EDAEDFFFFBC4BFE55FEDAFE79CFA7" authorityName="Fauvel" authorityYear="1878" box="[399,452,380,401]" class="Insecta" family="Staphylinidae" genus="Tilea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="33" phylum="Arthropoda" rank="genus">Tilea</taxonomicName>
|
||
] Castanea [sic]/
|
||
<typeStatus id="54851FCCFFFFBC4BFF4DFE30FE99CF9D" box="[151,292,406,427]" pageId="15" pageNumber="16" type="lectotype">LECTOTYPE</typeStatus>
|
||
³
|
||
<taxonomicName id="4C3EDAEDFFFFBC4BFE96FE30FDC3CF9D" authority="(Casey)" baseAuthorityName="Casey" box="[332,638,406,427]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="castanea">Phlaeopterus castanea (Casey)</taxonomicName>
|
||
des. 1984, J.M. Campbell.
|
||
<typeStatus id="54851FCCFFFFBC4BFE4EFE17FDABCFF0" box="[404,534,433,454]" pageId="15" pageNumber="16" type="paralectotype">Paralectotype</typeStatus>
|
||
male (here designated) labeled as follows: CASEY bequest 1925/castanea
|
||
<typeStatus id="54851FCCFFFFBC4BFEFDFE40FE27CFCD" box="[295,410,486,507]" pageId="15" pageNumber="16" type="paratype">PARATYPE</typeStatus>
|
||
USNM 48110. We designated these specimens as
|
||
<typeStatus id="54851FCCFFFFBC4BFE69FDA7FDB1CC20" box="[435,524,513,534]" pageId="15" pageNumber="16" type="lectotype">lectotype</typeStatus>
|
||
and
|
||
<typeStatus id="54851FCCFFFFBC4BFD9CFDA7FF54CC06" pageId="15" pageNumber="16" type="paralectotype">paralectotype</typeStatus>
|
||
to clarify the application of this name to this taxon and because a
|
||
<typeStatus id="54851FCCFFFFBC4BFE48FD90FE5ACC7D" box="[402,487,566,587]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
was not designated in the series examined by
|
||
<bibRefCitation id="EFAFDC9FFFFFBC4BFE64FDF6FD81CC53" author="Casey" box="[446,572,592,613]" pageId="15" pageNumber="16" pagination="281 - 624" refId="ref28683" refString="Casey, T. L. 1893. Coleopterological Notices V. Annals of the New York Academy of Sciences 7 (1): 281 - 624." type="journal article" year="1893">Casey (1893)</bibRefCitation>
|
||
, which comprised
|
||
<specimenCount id="9D386AE7FFFFBC4BFF25FDCDFE33CCB6" box="[255,398,619,640]" pageId="15" pageNumber="16" type="generic">two specimens</specimenCount>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B81A16EFFFFBC4BFF75FD23FE6ECD16" blockId="15.[151,639,194,1730]" pageId="15" pageNumber="16">
|
||
<emphasis id="B94A7D7CFFFFBC4BFF75FD23FE6ECCAC" bold="true" box="[175,467,645,666]" pageId="15" pageNumber="16">
|
||
<taxonomicName id="4C3EDAEDFFFFBC4BFF75FD23FE77CCAC" authorityName="Hatch" authorityYear="1957" box="[175,458,645,666]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cascadiensis">
|
||
<emphasis id="B94A7D7CFFFFBC4BFF75FD23FE77CCAC" bold="true" box="[175,458,645,666]" italics="true" pageId="15" pageNumber="16">Phlaeopterus cascadiensis</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
</emphasis>
|
||
<typeStatus id="54851FCCFFFFBC4BFE38FD23FD82CCAC" box="[482,575,645,666]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
|
||
male (UAMObs:Ento:235219) and
|
||
<typeStatus id="54851FCCFFFFBC4BFE02FD06FD94CC83" box="[472,553,672,693]" pageId="15" pageNumber="16" type="allotype">allotype</typeStatus>
|
||
female (UAMObs:Ento:235777) labeled as follows: Mt. Rainier, WASH., Tipsoo Lake,
|
||
<date id="FF8087AEFFFFBC4BFE1BFD73FDFCCCDC" box="[449,577,725,746]" pageId="15" pageNumber="16" value="1938-08-02">Aug. 2, 1938</date>
|
||
, M.H. Hatch/TYPE ³ (or
|
||
<typeStatus id="54851FCCFFFFBC4BFE85FD56FE65CD33" box="[351,472,752,773]" pageId="15" pageNumber="16" type="allotype">ALLOTYPE</typeStatus>
|
||
<specimenCount id="9D386AE7FFFFBC4BFE3EFD54FE4CCD32" box="[484,497,754,772]" pageId="15" pageNumber="16" type="female">♀</specimenCount>
|
||
)
|
||
<taxonomicName id="4C3EDAEDFFFFBC4BFDDCFD56FEFDCD16" authorityName="Hatch" authorityYear="1957" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cascadiensis">Phlaeopterus cascadiensis 1951</taxonomicName>
|
||
– M.H. Hatch.
|
||
</paragraph>
|
||
<paragraph id="8B81A16EFFFFBC4BFF75FC83FDA0CD0C" blockId="15.[151,639,194,1730]" box="[175,541,805,826]" pageId="15" pageNumber="16">
|
||
All
|
||
<specimenCount id="9D386AE7FFFFBC4BFF0FFC83FEDACD0C" box="[213,359,805,826]" pageId="15" pageNumber="16" type="generic">four specimens</specimenCount>
|
||
are in the USNM.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFFFBC4BFF75FCE6FE7ACAD2" pageId="15" pageNumber="16" type="distribution">
|
||
<paragraph id="8B81A16EFFFFBC4BFF75FCE6FE26CA4F" blockId="15.[151,639,194,1730]" pageId="15" pageNumber="16">
|
||
<emphasis id="B94A7D7CFFFFBC4BFF75FCE6FE93CD63" bold="true" box="[175,302,832,853]" pageId="15" pageNumber="16">Distribution.</emphasis>
|
||
<taxonomicName id="4C3EDAEDFFFFBC4BFEE6FCE6FF79CD59" authority="castaneus" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="subSpecies" species="castaneus" subSpecies="castaneus">
|
||
<emphasis id="B94A7D7CFFFFBC4BFEE6FCE6FF79CD59" italics="true" pageId="15" pageNumber="16">Phlaeopterus castaneus castaneus</emphasis>
|
||
</taxonomicName>
|
||
is known from the Rocky Mountains of
|
||
<collectingRegion id="49FA6F8CFFFFBC4BFF4DFCD3FF5CCDBC" box="[151,225,885,906]" country="Canada" name="Alberta" pageId="15" pageNumber="16">Alberta</collectingRegion>
|
||
south to
|
||
<collectingRegion id="49FA6F8CFFFFBC4BFE96FCD3FE15CDBC" box="[332,424,885,906]" country="United States of America" name="Colorado" pageId="15" pageNumber="16">Colorado</collectingRegion>
|
||
and west to eastern
|
||
<collectingRegion id="49FA6F8CFFFFBC4BFF4DFC36FF59CD93" box="[151,228,912,933]" country="United States of America" name="Oregon" pageId="15" pageNumber="16">Oregon</collectingRegion>
|
||
, eastern
|
||
<collectingRegion id="49FA6F8CFFFFBC4BFE8DFC36FE6DCD93" box="[343,464,912,933]" country="United States of America" name="Washington" pageId="15" pageNumber="16">Washington</collectingRegion>
|
||
, and Garibaldi Provincial Park,
|
||
<collectingRegion id="49FA6F8CFFFFBC4BFE8AFC0CFDBECD89" box="[336,515,938,959]" country="Canada" name="British Columbia" pageId="15" pageNumber="16">British Columbia</collectingRegion>
|
||
(
|
||
<figureCitation id="1305BDEBFFFFBC4BFDC1FC0CFDCDCD89" box="[539,624,938,959]" captionStart="Fig" captionStartId="15.[709,740,1495,1514]" captionTargetBox="[690,1168,666,1478]" captionTargetId="figure-473@15.[688,1170,664,1480]" captionTargetPageId="15" captionText="Fig. 6. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. bakerensis, B) P. castaneus. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected. In B, red triangles represent localities of P. castaneus castaneus, red squares represent localities of P. castaneus cascadiensis, and green stars represent areas of overlap of the two subspecies." figureDoi="http://doi.org/10.5281/zenodo.5102855" httpUri="https://zenodo.org/record/5102855/files/figure.png" pageId="15" pageNumber="16">Fig. 6B</figureCitation>
|
||
).
|
||
<taxonomicName id="4C3EDAEDFFFFBC4BFF4DFC63FDAECDEC" box="[151,531,965,986]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="subSpecies" species="castaneus" subSpecies="cascadiensis">
|
||
<emphasis id="B94A7D7CFFFFBC4BFF4DFC63FDAECDEC" box="[151,531,965,986]" italics="true" pageId="15" pageNumber="16">Phlaeopterus castaneus cascadiensis</emphasis>
|
||
</taxonomicName>
|
||
is known from the Aleutian Islands and southeastern
|
||
<collectingRegion id="49FA6F8CFFFFBC4BFDEEFC79FDC7CDC2" box="[564,634,991,1012]" country="United States of America" name="Alaska" pageId="15" pageNumber="16">Alaska</collectingRegion>
|
||
, and the Pacific Coast and Cascade Ranges of
|
||
<collectingRegion id="49FA6F8CFFFFBC4BFF4DFBB3FEFBCA1C" box="[151,326,1045,1066]" country="Canada" name="British Columbia" pageId="15" pageNumber="16">British Columbia</collectingRegion>
|
||
south to
|
||
<collectingRegion id="49FA6F8CFFFFBC4BFE79FBB3FE53CA1C" box="[419,494,1045,1066]" country="United States of America" name="Oregon" pageId="15" pageNumber="16">Oregon</collectingRegion>
|
||
(
|
||
<figureCitation id="1305BDEBFFFFBC4BFE26FBB3FDF6CA1C" box="[508,587,1045,1066]" captionStart="Fig" captionStartId="15.[709,740,1495,1514]" captionTargetBox="[690,1168,666,1478]" captionTargetId="figure-473@15.[688,1170,664,1480]" captionTargetPageId="15" captionText="Fig. 6. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. bakerensis, B) P. castaneus. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected. In B, red triangles represent localities of P. castaneus castaneus, red squares represent localities of P. castaneus cascadiensis, and green stars represent areas of overlap of the two subspecies." figureDoi="http://doi.org/10.5281/zenodo.5102855" httpUri="https://zenodo.org/record/5102855/files/figure.png" pageId="15" pageNumber="16">Fig. 6B</figureCitation>
|
||
). We confirm the surprising occurrence of this species on Unalaska Island from a single specimen (UAMObs:Ento:233345).
|
||
</paragraph>
|
||
<paragraph id="8B81A16EFFFFBC4BFF75FBD9FE7ACAD2" blockId="15.[151,639,194,1730]" pageId="15" pageNumber="16">
|
||
<emphasis id="B94A7D7CFFFFBC4BFF75FBD9FEA2CAA2" bold="true" box="[175,287,1151,1172]" pageId="15" pageNumber="16">Bionomics.</emphasis>
|
||
This species has been collected at elevations of
|
||
<quantity id="4CC60C8BFFFFBC4BFECCFB3FFE2ECA98" box="[278,403,1177,1199]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="2.5" metricValueMin="0.5" pageId="15" pageNumber="16" unit="m" value="1500.0" valueMax="2500.0" valueMin="500.0">500–2,500 m</quantity>
|
||
at the edges of cold, fast streams and sometimes at the edges of snowfields and lakes fed by melting snow.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324F2E5FFFFBC4BFF75FB4FFC15CF3A" pageId="15" pageNumber="16" type="discussion">
|
||
<paragraph id="8B81A16EFFFFBC4BFF75FB4FFC15CF3A" blockId="15.[151,639,194,1730]" lastBlockId="15.[685,1172,194,268]" pageId="15" pageNumber="16">
|
||
<emphasis id="B94A7D7CFFFFBC4BFF75FB4FFEACCAC8" bold="true" box="[175,273,1257,1278]" pageId="15" pageNumber="16">Remarks.</emphasis>
|
||
<taxonomicName id="4C3EDAEDFFFFBC4BFEFEFB4FFDA3CAC8" authorityName="" baseAuthorityName="Casey" baseAuthorityYear="1893" box="[292,542,1257,1278]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="castaneus">
|
||
<emphasis id="B94A7D7CFFFFBC4BFEFEFB4FFDA3CAC8" box="[292,542,1257,1278]" italics="true" pageId="15" pageNumber="16">Phlaeopterus castaneus</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from all other
|
||
<taxonomicName id="4C3EDAEDFFFFBC4BFE76FAA2FDC3CB2F" box="[428,638,1284,1305]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B94A7D7CFFFFBC4BFE76FAA2FD8CCB2F" box="[428,561,1284,1305]" italics="true" pageId="15" pageNumber="16">Phlaeopterus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
by the shape of the pronotum, tibiae evenly pubescent to apex, mesosternum weakly but completely carinate, and the form of the aedeagus. We chose to classify
|
||
<taxonomicName id="4C3EDAEDFFFFBC4BFE9CFAC8FE63CBB5" authorityName="Hatch" authorityYear="1957" box="[326,478,1390,1411]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cascadiensis">
|
||
<emphasis id="B94A7D7CFFFFBC4BFE9CFAC8FE63CBB5" box="[326,478,1390,1411]" italics="true" pageId="15" pageNumber="16">P. cascadiensis</emphasis>
|
||
</taxonomicName>
|
||
as a subspecies of
|
||
<taxonomicName id="4C3EDAEDFFFFBC4BFF6CFA2FFE88CBA8" authorityName="" baseAuthorityName="Casey" baseAuthorityYear="1893" box="[182,309,1417,1438]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="castaneus">
|
||
<emphasis id="B94A7D7CFFFFBC4BFF6CFA2FFE88CBA8" box="[182,309,1417,1438]" italics="true" pageId="15" pageNumber="16">P. castaneus</emphasis>
|
||
</taxonomicName>
|
||
primarily because of the abundance of intermediate forms at the zone of overlap of the two taxa. The two subspecies can be distinguished throughout most of their range by the length and microspinules of the internal sac of the aedeagus, the explanate lateral margins of the pronotum, and, to some extent, the coloration of the elytra. However, in Garibaldi and Manning Provincial Parks, intermediate states of microspinules of the internal sac and the explanate lateral margins of the pronotum can be found. The two subspecies are diagnosable throughout their range except in the zone of overlap in Garibaldi and Manning Provincial Parks,
|
||
<collectingRegion id="49FA6F8CFFFFBC4BFD29FF51FC1ECF3A" box="[755,931,247,268]" country="Canada" name="British Columbia" pageId="15" pageNumber="16">British Columbia</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |