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<document id="DC979CC764E1E52FE9FCEE4041A91DF8" ID-DOI="10.1649/0010-065X-72.mo4.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5102840" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626277672172" checkinUser="carolina" docAuthor="Mullen, Logan J., Campbell, J. M. &amp; Sikes, Derek S." docDate="2018" docId="03971078FFD3BC60FD17FF65FCFAC8F4" docLanguage="en" docName="TheColeopBull.mo.72.16.1-54.pdf" docOrigin="The Coleopterists Bulletin 72" docSource="http://dx.doi.org/10.1649/0010-065x-72.mo4.1" docStyle="DocumentStyle:A431C8F32F3D86E670C18C13DA9B5F2B.4:TheColeopBull.2000-2009.monograph" docStyleId="A431C8F32F3D86E670C18C13DA9B5F2B" docStyleName="TheColeopBull.2000-2009.monograph" docStyleVersion="4" docTitle="Phlaeopterus lagrandeuri Hatch 1957" docType="treatment" docVersion="3" lastPageNumber="37" masterDocId="FFAE6800FFF0BC44FFDAFFA6FFBDCE36" masterDocTitle="Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="54" masterPageNumber="1" pageNumber="36" updateTime="1699007718751" updateUser="plazi">
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<mods:title id="5290C163EC03C5BD8C9F3CA03B8537BA">Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<mods:namePart id="0518AA21C32D06D92B4B330B365B4D6B">Mullen, Logan J.</mods:namePart>
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<mods:namePart id="A606CB1400E61E92DBF8D4B48F9F6026">Sikes, Derek S.</mods:namePart>
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<mods:title id="2AB4F839551FFC5646553849C9F4ADA7">The Coleopterists Bulletin</mods:title>
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<heading id="D0C91602FFD3BC67FD17FF65FBC9CEEE" box="[717,1140,195,216]" centered="true" fontSize="10" level="2" pageId="35" pageNumber="36" reason="2">
<emphasis id="B94A7D7CFFD3BC67FD17FF65FBC9CEEE" bold="true" box="[717,1140,195,216]" pageId="35" pageNumber="36">
12.
<taxonomicName id="4C3EDAEDFFD3BC67FD29FF65FBC9CEEE" ID-CoL="76ZM2" authority="Hatch, 1957" authorityName="Hatch" authorityYear="1957" box="[755,1140,195,216]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lagrandeuri">
<emphasis id="B94A7D7CFFD3BC67FD29FF65FC4CCEEE" bold="true" box="[755,1009,195,216]" italics="true" pageId="35" pageNumber="36">Phlaeopterus lagrandeuri</emphasis>
<bibRefCitation id="EFAFDC9FFFD3BC67FC23FF65FBC9CEEE" author="Hatch" box="[1017,1140,195,216]" pageId="35" pageNumber="36" refId="ref28862" refString="Hatch, M. H. 1957. The Beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology, Seattle, WA." type="book" year="1957">Hatch, 1957</bibRefCitation>
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</emphasis>
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</paragraph>
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<subSubSection id="C324F2E5FFD3BC67FD62FF78FBE5CF3B" pageId="35" pageNumber="36" type="description">
<paragraph id="8B81A16EFFD3BC67FD62FF78FBE5CF3B" blockId="35.[696,1161,195,269]" pageId="35" pageNumber="36">
(
<figureCitation id="1305BDEBFFD3BC67FD65FF78FCABCEC5" box="[703,790,222,243]" captionStart="Fig" captionStartId="12.[197,228,1727,1746]" captionTargetBox="[177,1188,196,1710]" captionTargetId="figure-0@12.[175,1190,194,1712]" captionTargetPageId="12" captionText="Fig. 3. Phlaeopterus species, habitus. A) P. hatchi, male (UAMObs:Ento:232429), B) P. houkae, male (UAMObs:" figureDoi="http://doi.org/10.5281/zenodo.5102847" httpUri="https://zenodo.org/record/5102847/files/figure.png" pageId="35" pageNumber="36">Figs. 3D</figureCitation>
,
<figureCitation id="1305BDEBFFD3BC67FCFBFF78FCF7CEC5" box="[801,842,222,243]" captionStart="Fig" captionStartId="18.[197,228,1028,1047]" captionText="Fig. 11. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. kavanaughi, B) P. lagrandeuri. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected." pageId="35" pageNumber="36">11B</figureCitation>
,
<figureCitation id="1305BDEBFFD3BC67FC8CFF78FCC3CEC5" box="[854,894,222,243]" captionStart="Fig" captionStartId="21.[175,206,1672,1691]" captionTargetBox="[157,1166,199,1650]" captionTargetId="figure-1@21.[154,1169,194,1656]" captionTargetPageId="21" captionText="Fig. 15. Pronota, dorsal view, of A) Lesteva pallipes LeConte, 1863, B) Lesteva longoelytrata (Goeze, 1777), C) Unamis sp. undescribed, D) Unamis sp., E) Phlaeopterus lagrandeuri, F) Phlaeopterus houkae, G) Phlaeopterus longipennis, H) Phlaeopterus obsoletus." figureDoi="http://doi.org/10.5281/zenodo.5102865" httpUri="https://zenodo.org/record/5102865/files/figure.png" pageId="35" pageNumber="36">15E</figureCitation>
,
<figureCitation id="1305BDEBFFD3BC67FC50FF78FC0ECEC5" box="[906,947,222,243]" captionStart="Fig" captionStartId="25.[175,206,1235,1254]" captionTargetBox="[154,634,195,1217]" captionTargetId="figure-213@25.[153,635,194,1219]" captionTargetPageId="25" captionText="Fig. 19. Mesosterna, ventral oblique view, of A) Lesteva pallipes, B) Lesteva longoelytrata, C) Unamis sp. undescribed, D) Unamis sp., E) Phlaeopterus lagrandeuri, F) Phlaeopterus houkae." figureDoi="http://doi.org/10.5281/zenodo.5102873" httpUri="https://zenodo.org/record/5102873/files/figure.png" pageId="35" pageNumber="36">19E</figureCitation>
, 21A, 28E, 31E, 33E, 34AB, 36A, 38EF, 40E, 41D, 42C)
</paragraph>
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<subSubSection id="C324F2E5FFD3BC67FD77FE85FE69CBB5" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="8B81A16EFFD3BC67FD77FE85FC45CFB1" blockId="35.[685,1172,291,391]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="AB2E8640FFD3BC67FD77FE85FC45CFB1" pageId="35" pageNumber="36">
<treatmentCitation id="0A9F877FFFD3BC67FD77FE85FB80CF0E" author="Hatch" box="[685,1085,291,312]" page="58" pageId="35" pageNumber="36" year="1957">
<taxonomicName id="4C3EDAEDFFD3BC67FD77FE85FB80CF0E" ID-CoL="76ZM2" authority="Hatch 1957: 58" authorityName="Hatch" authorityPageNumber="58" authorityYear="1957" box="[685,1085,291,312]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lagrandeuri">
<emphasis id="B94A7D7CFFD3BC67FD77FE85FC19CF0E" box="[685,932,291,312]" italics="true" pageId="35" pageNumber="36">Phlaeopterus lagrandeuri</emphasis>
<bibRefCitation id="EFAFDC9FFFD3BC67FC70FE85FB80CF0E" author="Hatch" box="[938,1085,291,312]" pageId="35" pageNumber="36" refId="ref28862" refString="Hatch, M. H. 1957. The Beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology, Seattle, WA." type="book" year="1957">Hatch 1957: 58</bibRefCitation>
</taxonomicName>
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[original description]. Campbell and Davies 1991: 5;
<treatmentCitation id="0A9F877FFFD3BC67FD12FEFEFC3BCF5B" author="Herman" box="[712,902,344,365]" page="377" pageId="35" pageNumber="36" year="2001">
<bibRefCitation id="EFAFDC9FFFD3BC67FD12FEFEFC3BCF5B" author="Herman" box="[712,902,344,365]" pageId="35" pageNumber="36" pagination="1 - 650" refId="ref28893" refString="Herman, L. H. 2001. Catalog of the Staphylinidae (Insecta, Coleoptera): 1758 to the end of the Second Millennium. I. Introduction, history, biogeographical sketches, and Omaliine group. Bulletin of the American Museum of Natural History 265: 1 - 650." type="journal article" year="2001">Herman 2001: 377</bibRefCitation>
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;
<treatmentCitation id="0A9F877FFFD3BC67FC49FEFEFB32CF5B" author="Bousquet" box="[915,1167,344,365]" page="89" pageId="35" pageNumber="36" year="2013">
<bibRefCitation id="EFAFDC9FFFD3BC67FC49FEFEFB32CF5B" author="Bousquet" box="[915,1167,344,365]" pageId="35" pageNumber="36" refId="ref28340" refString="Bousquet, Y., P. Bouchard, A. E. Davies, and D. S. Sikes. 2013. Checklist of Beetles (Coleoptera) of Canada and Alaska. Second Edition. Pensoft, Sofia, Bulgaria." type="book" year="2013">
Bousquet
<emphasis id="B94A7D7CFFD3BC67FC2DFEFEFB91CF5B" box="[1015,1068,344,365]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
2013: 89
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; Shavrin and Mullen 2015: 121.
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</paragraph>
<caption id="DF41F1E6FFD3BC67FF75FB75FE9ACB20" ID-DOI="http://doi.org/10.5281/zenodo.5102885" ID-Zenodo-Dep="5102885" httpUri="https://zenodo.org/record/5102885/files/figure.png" pageId="35" pageNumber="36" startId="35.[175,206,1235,1254]" targetBox="[154,634,195,1218]" targetPageId="35">
<paragraph id="8B81A16EFFD3BC67FF75FB75FE9ACB20" blockId="35.[151,639,1235,1302]" pageId="35" pageNumber="36">
<emphasis id="B94A7D7CFFD3BC67FF75FB75FF48CAD0" bold="true" box="[175,245,1235,1254]" pageId="35" pageNumber="36">Fig. 27.</emphasis>
Heads, dorsal and dorsal oblique views, of AB)
<taxonomicName id="4C3EDAEDFFD3BC67FF14FB4DFED4CAC8" ID-CoL="6Q33B" authorityName="Mannerheim" authorityYear="1830" box="[206,361,1259,1278]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="B94A7D7CFFD3BC67FF14FB4DFED4CAC8" box="[206,361,1259,1278]" italics="true" pageId="35" pageNumber="36">Lesteva pubescens</emphasis>
</taxonomicName>
, CD)
<taxonomicName id="4C3EDAEDFFD3BC67FE77FB4DFDB3CAC8" box="[429,526,1259,1278]" class="Insecta" family="Staphylinidae" genus="Unamis" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B94A7D7CFFD3BC67FE77FB4DFE53CAC8" box="[429,494,1259,1278]" italics="true" pageId="35" pageNumber="36">Unamis</emphasis>
sp.
</taxonomicName>
undescribed, EF)
<taxonomicName id="4C3EDAEDFFD3BC67FF12FAA5FE9ACB20" box="[200,295,1283,1302]" class="Insecta" family="Staphylinidae" genus="Unamis" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B94A7D7CFFD3BC67FF12FAA5FEB5CB20" box="[200,264,1283,1302]" italics="true" pageId="35" pageNumber="36">Unamis</emphasis>
sp.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="8B81A16EFFD3BC67FF4DFAF2FE69CBB5" blockId="35.[151,639,1364,1730]" pageId="35" pageNumber="36">they were observed feeding on windblown arthropods on the surface of the snow.</paragraph>
</subSubSection>
<subSubSection id="C324F2E5FFD3BC67FF75FA2FFEC9C844" pageId="35" pageNumber="36" type="discussion">
<paragraph id="8B81A16EFFD3BC67FF75FA2FFEC9C844" blockId="35.[151,639,1364,1730]" pageId="35" pageNumber="36">
<emphasis id="B94A7D7CFFD3BC67FF75FA2FFEACCBA8" bold="true" box="[175,273,1417,1438]" pageId="35" pageNumber="36">Remarks.</emphasis>
<taxonomicName id="4C3EDAEDFFD3BC67FEF9FA2FFD9ECBA8" authorityName="Mullen and Campbell" authorityYear="2018" box="[291,547,1417,1438]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="kavanaughi">
<emphasis id="B94A7D7CFFD3BC67FEF9FA2FFD9ECBA8" box="[291,547,1417,1438]" italics="true" pageId="35" pageNumber="36">Phlaeopterus kavanaughi</emphasis>
</taxonomicName>
can be distinguished from all other
<taxonomicName id="4C3EDAEDFFD3BC67FE6BFA05FDC0CB8E" box="[433,637,1443,1464]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B94A7D7CFFD3BC67FE6BFA05FD92CB8E" box="[433,559,1443,1464]" italics="true" pageId="35" pageNumber="36">Phlaeopterus</emphasis>
species
</taxonomicName>
by the tibiae pubescent to the apices, the shape of the pronotal lateral margins, the ventral longitudinal carina of the median lobe of the aedeagus, the shape and median transverse fold of the internal sac of the aedeagus, and the usually yellowish brown elytra and pronotal lateral margins that are paler than remainder of the body.
</paragraph>
</subSubSection>
<subSubSection id="C324F2E5FFD3BC67FF75F9DEFDAFC8F4" pageId="35" pageNumber="36" type="etymology">
<paragraph id="8B81A16EFFD3BC67FF75F9DEFDAFC8F4" blockId="35.[151,639,1364,1730]" pageId="35" pageNumber="36">
<emphasis id="B94A7D7CFFD3BC67FF75F9DEFE9FC8BB" bold="true" box="[175,290,1656,1677]" pageId="35" pageNumber="36">Etymology.</emphasis>
This species is named in honor of Dr. David H. Kavanaugh,
<collectingRegion id="49FA6F8CFFD3BC67FEA8F934FE6CC891" box="[370,465,1682,1703]" country="United States of America" name="California" pageId="35" pageNumber="36">California</collectingRegion>
Academy of Sciences, San Francisco,
<collectingRegion id="49FA6F8CFFD3BC67FEB4F90BFE6DC8F4" box="[366,464,1709,1730]" country="United States of America" name="California" pageId="35" pageNumber="36">California</collectingRegion>
,
<collectingCountry id="F329E1FEFFD3BC67FE06F90BFDB2C8F4" box="[476,527,1709,1730]" name="United States of America" pageId="35" pageNumber="36">USA</collectingCountry>
.
</paragraph>
</subSubSection>
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<paragraph id="8B81A16EFFD3BC67FD1FFE14FC93CFD7" blockId="35.[685,1173,434,1730]" pageId="35" pageNumber="36">
<materialsCitation id="3B56AB33FFD3BC67FD1FFE14FC96CFD7" country="United States of America" location="Carkeek Park" municipality="Seattle" pageId="35" pageNumber="36" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
<emphasis id="B94A7D7CFFD3BC67FD1FFE14FCE1CFF1" bold="true" box="[709,860,434,455]" pageId="35" pageNumber="36">
<typeStatus id="54851FCCFFD3BC67FD1FFE14FD4ACFF1" box="[709,759,434,455]" pageId="35" pageNumber="36">Type</typeStatus>
Locality.
</emphasis>
<location id="8EE1F7B5FFD3BC67FCB3FE14FC49CFF1" LSID="urn:lsid:plazi:treatment:03971078FFD3BC60FD17FF65FCFAC8F4:8EE1F7B5FFD3BC67FCB3FE14FC49CFF1" box="[873,1012,434,455]" country="United States of America" municipality="Seattle" name="Carkeek Park" pageId="35" pageNumber="36" stateProvince="Washington">Carkeek Park</location>
,
<collectingMunicipality id="6BE53B14FFD3BC67FBDEFE14FBFBCFF1" box="[1028,1094,434,455]" pageId="35" pageNumber="36">Seattle</collectingMunicipality>
,
<collectingRegion id="49FA6F8CFFD3BC67FB82FE14FD51CFD7" country="United States of America" name="Washington" pageId="35" pageNumber="36">Washington</collectingRegion>
,
<collectingCountry id="F329E1FEFFD3BC67FD22FE6AFC96CFD7" box="[760,811,460,481]" name="United States of America" pageId="35" pageNumber="36">USA</collectingCountry>
</materialsCitation>
.
</paragraph>
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<subSubSection id="C324F2E5FFD3BC60FD1FFE41FEC2CBB5" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="description">
<paragraph id="8B81A16EFFD3BC60FD1FFE41FEC2CBB5" blockId="35.[685,1173,434,1730]" lastBlockId="36.[173,660,1337,1730]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<emphasis id="B94A7D7CFFD3BC67FD1FFE41FC04CFCA" bold="true" box="[709,953,487,508]" pageId="35" pageNumber="36">Redescription. Habitus:</emphasis>
Length
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. Dark brown to reddish brown; legs, mouthparts, antennae, and elytra usually lighter (
<figureCitation id="1305BDEBFFD3BC67FBE9FDBAFB35CC07" box="[1075,1160,540,561]" captionStart="Fig" captionStartId="12.[197,228,1727,1746]" captionTargetBox="[177,1188,196,1710]" captionTargetId="figure-0@12.[175,1190,194,1712]" captionTargetPageId="12" captionText="Fig. 3. Phlaeopterus species, habitus. A) P. hatchi, male (UAMObs:Ento:232429), B) P. houkae, male (UAMObs:" figureDoi="http://doi.org/10.5281/zenodo.5102847" httpUri="https://zenodo.org/record/5102847/files/figure.png" pageId="35" pageNumber="36">Fig. 3D</figureCitation>
); elytra with distinctive yellow to reddish yellow area ranging from small spot on humeri to nearly covering elytral disc.
<emphasis id="B94A7D7CFFD3BC67FC52FDCAFC75CCB7" bold="true" box="[904,968,620,641]" pageId="35" pageNumber="36">Head:</emphasis>
Moderately narrow, width across eyes to head length slightly wider than long (
<figureCitation id="1305BDEBFFD3BC67FCFCFD07FC18CC80" box="[806,933,672,694]" captionStart="Fig" captionStartId="41.[175,206,1236,1255]" captionTargetBox="[154,634,195,1220]" captionTargetId="figure-493@41.[153,635,194,1221]" captionTargetPageId="41" captionText="Fig. 34. Phlaeopterus species, head, dorsal and dorsal oblique views. AB) P. lagrandeuri, CD) P. houkae, EF) P. obsoletus." figureDoi="http://doi.org/10.5281/zenodo.5102897" httpUri="https://zenodo.org/record/5102897/files/figure.png" pageId="35" pageNumber="36">Fig. 34AB</figureCitation>
). Interantennal groove large, deeply impressed. Anteocellar foveae small, shallowly impressed. Eyes moderately pubescent on ventral half, with more than 10 setae (
<figureCitation id="1305BDEBFFD3BC67FD6FFCADFCA5CD16" box="[693,792,779,800]" captionStart="Fig" captionStartId="42.[731,762,1272,1291]" captionTargetBox="[711,1189,196,1255]" captionTargetId="figure-251@42.[709,1191,194,1257]" captionTargetPageId="42" captionText="Fig. 36. Phlaeopterus species, eyes, dorsal view. A) P. lagrandeuri, B) P. houkae, C) P. longipennis, D) P. frosti, E) P. bakerensis, F) P. olympicus." figureDoi="http://doi.org/10.5281/zenodo.5102901" httpUri="https://zenodo.org/record/5102901/files/figure.png" pageId="35" pageNumber="36">Fig. 36A</figureCitation>
). Antennomeres 510 1.21.8 times longer than wide; antennomeres 411 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with L-shaped row of setae and additional oblique row of setae (
<figureCitation id="1305BDEBFFD3BC67FBB1FC36FD64CD89" captionStart="Fig" captionStartId="36.[197,228,1167,1186]" captionTargetBox="[176,656,195,1150]" captionTargetId="figure-220@36.[175,657,194,1151]" captionTargetPageId="36" captionText="Fig. 28. Ventral view of A) Left mandible of Lesteva longoelytrata, B) Right mandible of Lesteva pallipes, C) Right mandible of Unamis sp. undescribed, D) Right mandible of Unamis sp., E) Left mandible of Phlaeopterus lagrandeuri, F) Left mandible of Phlaeopterus houkae." figureDoi="http://doi.org/10.5281/zenodo.5102889" httpUri="https://zenodo.org/record/5102889/files/figure.png" pageId="35" pageNumber="36">Fig. 28E</figureCitation>
). Labrum with sensory pores along anterior margin only (
<figureCitation id="1305BDEBFFD3BC67FCEEFC63FC32CDEC" box="[820,911,965,986]" captionStart="Fig" captionStartId="38.[197,228,1598,1617]" captionTargetBox="[177,1188,196,1581]" captionTargetId="figure-0@38.[175,1190,194,1583]" captionTargetPageId="38" captionText="Fig. 31. Labra of A) Lesteva pallipes, B) Lesteva pubescens, C) Unamis sp. undescribed, D) Unamis sp., E) Phlaeopterus lagrandeuri, F) Phlaeopterus houkae, G) Phlaeopterus obsoletus, H) Phlaeopterus filicornis. White arrows indicate labrum." figureDoi="http://doi.org/10.5281/zenodo.5102893" httpUri="https://zenodo.org/record/5102893/files/figure.png" pageId="35" pageNumber="36">Fig. 31E</figureCitation>
). Maxilla with lacinia and galea as in
<figureCitation id="1305BDEBFFD3BC67FCC2FC46FC3ACDC3" box="[792,903,991,1013]" captionStart="Fig" captionStartId="44.[197,228,1311,1330]" captionTargetBox="[177,655,196,1293]" captionTargetId="figure-155@44.[175,657,194,1295]" captionTargetPageId="44" captionText="Fig. 38. Maxillae, dorsal and ventral views, of AB) Lesteva longoelytrata, CD) Unamis sp. undescribed, EF) Phlaeopterus lagrandeuri." figureDoi="http://doi.org/10.5281/zenodo.5102907" httpUri="https://zenodo.org/record/5102907/files/figure.png" pageId="35" pageNumber="36">Fig. 38EF</figureCitation>
. Epipharynx as in Fig. 33E. Hypopharynx as in
<figureCitation id="1305BDEBFFD3BC67FCB6FC5CFC79CA39" box="[876,964,1018,1039]" captionStart="Fig" captionStartId="45.[175,206,1177,1196]" captionText="Fig. 40. Hypopharynges, dorsal view, of A) Lesteva pallipes, B) Lesteva longoelytrata, C) Unamis sp. undescribed, D) Unamis sp., E) Phlaeopterus lagrandeuri, F) Phlaeopterus houkae." pageId="35" pageNumber="36">Fig. 40E</figureCitation>
. Labial palpi with 3
<superScript id="7C4B0C26FFD3BC67FB5EFC53FB2ECA35" attach="none" box="[1156,1171,1013,1027]" fontSize="6" pageId="35" pageNumber="36">rd</superScript>
palpomere 1.11.3 times longer than 2
<superScript id="7C4B0C26FFD3BC67FBE3FBA9FBF6CA2B" attach="left" box="[1081,1099,1039,1053]" fontSize="6" pageId="35" pageNumber="36">nd</superScript>
palpomere. Gula as in
<figureCitation id="1305BDEBFFD3BC67FCBCFB89FC79CA72" box="[870,964,1071,1092]" captionStart="Fig" captionStartId="46.[197,228,1197,1216]" captionTargetBox="[176,656,195,1180]" captionTargetId="figure-522@46.[175,657,194,1181]" captionTargetPageId="46" captionText="Fig. 42. Heads, ventral view, of A) Unamis sp. undescribed, B) Unamis sp., C) Phlaeopterus lagrandeuri, D) Phlaeopterus houkae, E) Phlaeopterus longipennis, F) Phlaeopterus elongatus." figureDoi="http://doi.org/10.5281/zenodo.5102911" httpUri="https://zenodo.org/record/5102911/files/figure.png" pageId="35" pageNumber="36">Fig. 42C</figureCitation>
.
<emphasis id="B94A7D7CFFD3BC67FC0BFB89FB95CA72" bold="true" box="[977,1064,1071,1092]" pageId="35" pageNumber="36">Thorax:</emphasis>
Pronotum moderately narrow (
<figureCitation id="1305BDEBFFD3BC67FC5AFBECFC63CA69" box="[896,990,1098,1119]" captionStart="Fig" captionStartId="21.[175,206,1672,1691]" captionTargetBox="[157,1166,199,1650]" captionTargetId="figure-1@21.[154,1169,194,1656]" captionTargetPageId="21" captionText="Fig. 15. Pronota, dorsal view, of A) Lesteva pallipes LeConte, 1863, B) Lesteva longoelytrata (Goeze, 1777), C) Unamis sp. undescribed, D) Unamis sp., E) Phlaeopterus lagrandeuri, F) Phlaeopterus houkae, G) Phlaeopterus longipennis, H) Phlaeopterus obsoletus." figureDoi="http://doi.org/10.5281/zenodo.5102865" httpUri="https://zenodo.org/record/5102865/files/figure.png" pageId="35" pageNumber="36">Fig. 15E</figureCitation>
), length to width ratio = 0.680.72; ratio of pronotal width to head width = 1.541.67; maximum width less than elytral width at bases; dorsal surface coarsely, sparsely punctate, punctures separated by average distance slightly less than diameter of a puncture; lateral margins narrowly explanate posterad lateral fovea, not explanate anterad lateral foveae; lateral foveae moderately impressed, contiguous with lateral margins. Elytra with humeral angles convex; epipleural carina not projecting; 2.22.5 times longer than pronotum; apical margins convex. Wings fully developed. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum absent or vague.
<emphasis id="B94A7D7CFFD3BC67FB80FA18FB29CBE5" bold="true" box="[1114,1172,1470,1491]" pageId="35" pageNumber="36">Legs:</emphasis>
All tibiae evenly pubescent to apex. Metatrochanter with tooth on apical margin (
<figureCitation id="1305BDEBFFD3BC67FBF0FA55FB34C83E" box="[1066,1161,1523,1544]" captionStart="Fig" captionStartId="45.[709,740,1185,1204]" captionTargetBox="[154,1169,195,1168]" captionTargetPageId="45" captionText="Fig. 41. Phlaeopterus species, hypopharynges, dorsal view, of A) P. obsoletus, B) P. kavanaughi, C) P. elongatus. D) Phlaeopterus lagrandeuri, metatrochanter with apical tooth. Heads, ventral view, of E) Lesteva pallipes, F) Lesteva pubescens." figureDoi="http://doi.org/10.5281/zenodo.5102909" httpUri="https://zenodo.org/record/5102909/files/figure.png" pageId="35" pageNumber="36">Fig. 41D</figureCitation>
). Protibia slightly sinuate on medioventral margin in male.
<emphasis id="B94A7D7CFFD3BC67FCDDF98EFCC9C80B" bold="true" box="[775,884,1576,1597]" pageId="35" pageNumber="36">Abdomen:</emphasis>
Wing-folding spicules on tergites IV and V broad, transverse, nearly contiguous; tergite VI without wing-folding spicules. Apical palisade fringe on tergite VII absent.
<emphasis id="B94A7D7CFFD3BC67FBF6F9DEFB29C8BB" bold="true" box="[1068,1172,1656,1677]" pageId="35" pageNumber="36">Aedeagus:</emphasis>
Length
<quantity id="4CC60C8BFFD3BC67FD25F934FC29C891" box="[767,916,1682,1704]" metricMagnitude="-4" metricUnit="m" metricValue="6.65" metricValueMax="7.0" metricValueMin="6.3" pageId="35" pageNumber="36" unit="mm" value="0.665" valueMax="0.7" valueMin="0.63">0.630.70 mm</quantity>
. Median lobe triangular. Parameres broad, diverging until just past midline then converging towards apex of median lobe (Fig. 21A). Internal sac irregular in shape, with 2 apical patches of thick setae.
</paragraph>
</subSubSection>
<caption id="DF41F1E6FFD4BC60FD01FBA5FC75CA40" ID-DOI="http://doi.org/10.5281/zenodo.5102887" ID-Zenodo-Dep="5102887" httpUri="https://zenodo.org/record/5102887/files/figure.png" pageId="36" pageNumber="37" startId="36.[731,762,1027,1046]" targetBox="[710,1190,195,1011]" targetPageId="36">
<paragraph id="8B81A16EFFD4BC60FD01FBA5FC75CA40" blockId="36.[707,1194,1027,1142]" pageId="36" pageNumber="37">
<emphasis id="B94A7D7CFFD4BC60FD01FBA5FC95CA20" bold="true" box="[731,808,1027,1046]" pageId="36" pageNumber="37">Fig. 29.</emphasis>
<taxonomicName id="4C3EDAEDFFD4BC60FCE7FBA5FBBDCA20" box="[829,1024,1027,1046]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B94A7D7CFFD4BC60FCE7FBA5FC10CA20" box="[829,941,1027,1046]" italics="true" pageId="36" pageNumber="37">Phlaeopterus</emphasis>
species.
</taxonomicName>
Right mandibles, ventral view, of A)
<taxonomicName id="4C3EDAEDFFD4BC60FCAAFBBDFC6BCA18" authorityName="Mullen and Campbell" authorityYear="2018" box="[880,982,1051,1070]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="obsoletus">
<emphasis id="B94A7D7CFFD4BC60FCAAFBBDFC6BCA18" box="[880,982,1051,1070]" italics="true" pageId="36" pageNumber="37">P. obsoletus</emphasis>
</taxonomicName>
, B)
<taxonomicName id="4C3EDAEDFFD4BC60FBDAFBBDFBCDCA18" authorityName="Mullen &amp; Campbell &amp; Sikes" authorityYear="2018" box="[1024,1136,1051,1070]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="bakerensis">
<emphasis id="B94A7D7CFFD4BC60FBDAFBBDFBCDCA18" box="[1024,1136,1051,1070]" italics="true" pageId="36" pageNumber="37">P. bakerensis</emphasis>
</taxonomicName>
, C)
<taxonomicName id="4C3EDAEDFFD4BC60FB43FBBDFCA8CA70" authorityName="Mullen and Campbell" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="elongatus">
<emphasis id="B94A7D7CFFD4BC60FB43FBBDFCA8CA70" italics="true" pageId="36" pageNumber="37">P. elongatus</emphasis>
</taxonomicName>
. D)
<taxonomicName id="4C3EDAEDFFD4BC60FC9EFB95FC0ACA70" authorityName="Mullen &amp; Campbell &amp; Sikes" authorityYear="2018" box="[836,951,1075,1094]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="bakerensis">
<emphasis id="B94A7D7CFFD4BC60FC9EFB95FC0ACA70" box="[836,951,1075,1094]" italics="true" pageId="36" pageNumber="37">P. bakerensis</emphasis>
</taxonomicName>
, humeral angle of elytron, dorsal view. Apices of elytra of E)
<taxonomicName id="4C3EDAEDFFD4BC60FC25FBEDFBCDCA68" authorityName="Hatch" authorityYear="1957" box="[1023,1136,1099,1118]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="loganensis">
<emphasis id="B94A7D7CFFD4BC60FC25FBEDFBCDCA68" box="[1023,1136,1099,1118]" italics="true" pageId="36" pageNumber="37">P. loganensis</emphasis>
</taxonomicName>
, F)
<taxonomicName id="4C3EDAEDFFD4BC60FB43FBEDFD45CA40" authorityName="Mullen and Campbell" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="hatchi">
<emphasis id="B94A7D7CFFD4BC60FB43FBEDFD45CA40" italics="true" pageId="36" pageNumber="37">P. hatchi</emphasis>
</taxonomicName>
, G)
<taxonomicName id="4C3EDAEDFFD4BC60FCFAFBC5FC3FCA40" authorityName="Casey" authorityYear="1886" box="[800,898,1123,1142]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="filicornis">
<emphasis id="B94A7D7CFFD4BC60FCFAFBC5FC3FCA40" box="[800,898,1123,1142]" italics="true" pageId="36" pageNumber="37">P. filicornis</emphasis>
</taxonomicName>
female.
</paragraph>
</caption>
<caption id="DF41F1E6FFD4BC60FF1FFB29FEDBCB2F" ID-DOI="http://doi.org/10.5281/zenodo.5102889" ID-Zenodo-Dep="5102889" httpUri="https://zenodo.org/record/5102889/files/figure.png" pageId="36" pageNumber="37" startId="36.[197,228,1167,1186]" targetBox="[176,656,195,1150]" targetPageId="36">
<paragraph id="8B81A16EFFD4BC60FF1FFB29FEDBCB2F" blockId="36.[173,660,1167,1305]" pageId="36" pageNumber="37">
<emphasis id="B94A7D7CFFD4BC60FF1FFB29FEACCA94" bold="true" box="[197,273,1167,1186]" pageId="36" pageNumber="37">Fig. 28.</emphasis>
Ventral view of A) Left mandible of
<taxonomicName id="4C3EDAEDFFD4BC60FF77FB01FED7CA8C" baseAuthorityName="Goeze" baseAuthorityYear="1777" box="[173,362,1191,1210]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="longoelytrata">
<emphasis id="B94A7D7CFFD4BC60FF77FB01FED7CA8C" box="[173,362,1191,1210]" italics="true" pageId="36" pageNumber="37">Lesteva longoelytrata</emphasis>
</taxonomicName>
, B) Right mandible of
<taxonomicName id="4C3EDAEDFFD4BC60FD89FB01FF4DCAE4" authorityName="LeConte" authorityYear="1863" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B94A7D7CFFD4BC60FD89FB01FF4DCAE4" italics="true" pageId="36" pageNumber="37">Lesteva pallipes</emphasis>
</taxonomicName>
, C) Right mandible of
<taxonomicName id="4C3EDAEDFFD4BC60FE65FB19FD9CCAE4" box="[447,545,1215,1234]" class="Insecta" family="Staphylinidae" genus="Unamis" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B94A7D7CFFD4BC60FE65FB19FDBCCAE4" box="[447,513,1215,1234]" italics="true" pageId="36" pageNumber="37">Unamis</emphasis>
sp.
</taxonomicName>
undescribed, D) Right mandible of
<taxonomicName id="4C3EDAEDFFD4BC60FE5AFB70FE55CADF" box="[384,488,1238,1257]" class="Insecta" family="Staphylinidae" genus="Unamis" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B94A7D7CFFD4BC60FE5AFB70FE7FCADF" box="[384,450,1238,1257]" italics="true" pageId="36" pageNumber="37">Unamis</emphasis>
sp.
</taxonomicName>
, E) Left mandible of
<taxonomicName id="4C3EDAEDFFD4BC60FF15FB48FE08CB37" authorityName="Hatch" authorityYear="1957" box="[207,437,1262,1281]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lagrandeuri">
<emphasis id="B94A7D7CFFD4BC60FF15FB48FE08CB37" box="[207,437,1262,1281]" italics="true" pageId="36" pageNumber="37">Phlaeopterus lagrandeuri</emphasis>
</taxonomicName>
, F) Left mandible of
<taxonomicName id="4C3EDAEDFFD4BC60FF77FAA0FEDCCB2F" authorityName="Hatch" authorityYear="1957" box="[173,353,1286,1305]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="houkae">
<emphasis id="B94A7D7CFFD4BC60FF77FAA0FEDCCB2F" box="[173,353,1286,1305]" italics="true" pageId="36" pageNumber="37">Phlaeopterus houkae</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C324F2E5FFD4BC60FF1FFA2FFE3EC844" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="8B81A16EFFD4BC60FF1FFA2FFE3EC844" blockId="36.[173,660,1337,1730]" pageId="36" pageNumber="37">
<emphasis id="B94A7D7CFFD4BC60FF1FFA2FFEC8CBA8" bold="true" box="[197,373,1417,1438]" pageId="36" pageNumber="37">Type Specimens.</emphasis>
<materialsCitation id="3B56AB33FFD4BC60FE5EFA2FFE55CBDB" collectionCode="WASH, V" collectorName="Carkeek Park &amp; Bel. &amp; Type &amp; M. H. Hatch" pageId="36" pageNumber="37" specimenCount="1" typeStatus="holotype">
<typeStatus id="54851FCCFFD4BC60FE5EFA2FFE61CBA8" box="[388,476,1417,1438]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
male (UAMObs: Ento:235215) labeled as follows: Seattle,
<collectionCode id="ED2F39ABFFD4BC60FD99FA05FD31CB8E" box="[579,652,1443,1464]" pageId="36" pageNumber="37">WASH</collectionCode>
.,
<collectorName id="26CBC4B8FFD4BC60FF77FA18FE92CBE5" box="[173,303,1470,1491]" pageId="36" pageNumber="37">Carkeek Park</collectorName>
,
<collectorName id="26CBC4B8FFD4BC60FEEDFA18FEDCCBE5" box="[311,353,1470,1491]" pageId="36" pageNumber="37">Bel.</collectorName>
,
<collectionCode id="ED2F39ABFFD4BC60FEB3FA18FEC7CBE5" box="[361,378,1470,1491]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="36" pageNumber="37" type="Museum">V</collectionCode>
.3.1949/
<collectorName id="26CBC4B8FFD4BC60FE1FFA18FE48CBE5" box="[453,501,1470,1491]" pageId="36" pageNumber="37">Type</collectorName>
³,
<taxonomicName id="4C3EDAEDFFD4BC60FDCEFA18FEDCCBDB" authorityName="Hatch" authorityYear="1957" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lagrandeuri">
<emphasis id="B94A7D7CFFD4BC60FDCEFA18FEA2CBDB" italics="true" pageId="36" pageNumber="37">Phlaeopterus lagrandeuri</emphasis>
, 1951
</taxonomicName>
,
<collectorName id="26CBC4B8FFD4BC60FEB5FA7EFE55CBDB" box="[367,488,1496,1517]" pageId="36" pageNumber="37">M.H. Hatch</collectorName>
</materialsCitation>
.
<materialsCitation id="3B56AB33FFD4BC60FE2FFA7EFD31C815" collectionCode="ORE, VI" collectorName="DM Fender" pageId="36" pageNumber="37" specimenCount="1" typeStatus="allotype">
<typeStatus id="54851FCCFFD4BC60FE2FFA7EFDF5CBDB" box="[501,584,1496,1517]" pageId="36" pageNumber="37" type="allotype">Allotype</typeStatus>
female (UAMObs:Ento:235775) labeled as follows: Corvallis,
<collectionCode id="ED2F39ABFFD4BC60FF2CF9A8FE96C815" box="[246,299,1550,1571]" pageId="36" pageNumber="37">ORE</collectionCode>
.,
<collectionCode id="ED2F39ABFFD4BC60FE98F9A8FEE0C815" box="[322,349,1550,1571]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="36" pageNumber="37">VI</collectionCode>
.1.1946, KM &amp;
<collectorName id="26CBC4B8FFD4BC60FDCEF9A8FD31C815" box="[532,652,1550,1571]" pageId="36" pageNumber="37">DM Fender</collectorName>
</materialsCitation>
/
<typeStatus id="54851FCCFFD4BC60FF77F98EFE9AC80B" box="[173,295,1576,1597]" pageId="36" pageNumber="37" type="allotype">ALLOTYPE</typeStatus>
<specimenCount id="9D386AE7FFD4BC60FEE2F98CFEF8C80A" box="[312,325,1578,1596]" pageId="36" pageNumber="37" type="female"></specimenCount>
<taxonomicName id="4C3EDAEDFFD4BC60FE8CF98EFD32C80B" authorityName="Hatch" authorityYear="1957" box="[342,655,1576,1597]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lagrandeuri">Phlaeopterus lagrandeuri 1951</taxonomicName>
, M.H. Hatch/Marys Peak. Both specimens are deposited in the USNM.
</paragraph>
</subSubSection>
<subSubSection id="C324F2E5FFD4BC60FF1FF9DEFB17CB78" pageId="36" pageNumber="37" type="distribution">
<paragraph id="8B81A16EFFD4BC60FF1FF9DEFC89CB2F" blockId="36.[173,660,1337,1730]" lastBlockId="36.[707,1194,1204,1730]" pageId="36" pageNumber="37">
<emphasis id="B94A7D7CFFD4BC60FF1FF9DEFEF3C8BB" bold="true" box="[197,334,1656,1677]" pageId="36" pageNumber="37">Distribution.</emphasis>
<taxonomicName id="4C3EDAEDFFD4BC60FEBAF9DEFDCCC8BB" authorityName="Hatch" authorityYear="1957" box="[352,625,1656,1677]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lagrandeuri">
<emphasis id="B94A7D7CFFD4BC60FEBAF9DEFDCCC8BB" box="[352,625,1656,1677]" italics="true" pageId="36" pageNumber="37">Phlaeopterus lagrandeuri</emphasis>
</taxonomicName>
is widely distributed from the Kenai Peninsula of
<collectingRegion id="49FA6F8CFFD4BC60FF17F90BFEA8C8F4" box="[205,277,1709,1730]" country="United States of America" name="Alaska" pageId="36" pageNumber="37">Alaska</collectingRegion>
,
<collectingRegion id="49FA6F8CFFD4BC60FEFFF90BFE61C8F4" box="[293,476,1709,1730]" country="Canada" name="British Columbia" pageId="36" pageNumber="37">British Columbia</collectingRegion>
, including Haida Gwaii and Vancouver Island, south through
<collectingRegion id="49FA6F8CFFD4BC60FD19FB69FC86CAD2" box="[707,827,1231,1252]" country="United States of America" name="Washington" pageId="36" pageNumber="37">Washington</collectingRegion>
and
<collectingRegion id="49FA6F8CFFD4BC60FCB0FB69FC05CAD2" box="[874,952,1231,1252]" country="United States of America" name="Oregon" pageId="36" pageNumber="37">Oregon</collectingRegion>
, and
<specimenCount id="9D386AE7FFD4BC60FC31FB69FBC9CAD2" box="[1003,1140,1231,1252]" pageId="36" pageNumber="37" type="generic">one specimen</specimenCount>
from Calahan,
<collectingRegion id="49FA6F8CFFD4BC60FCEBFB4FFC2ACAC8" box="[817,919,1257,1278]" country="United States of America" name="California" pageId="36" pageNumber="37">California</collectingRegion>
(UAMObs:Ento:232633) (
<figureCitation id="1305BDEBFFD4BC60FD10FAA2FC95CB2F" box="[714,808,1284,1305]" captionStart="Fig" captionStartId="18.[197,228,1028,1047]" captionText="Fig. 11. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. kavanaughi, B) P. lagrandeuri. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected." pageId="36" pageNumber="37">Fig. 11B</figureCitation>
).
</paragraph>
<paragraph id="8B81A16EFFD4BC60FD01FAB8FB17CB78" blockId="36.[707,1194,1204,1730]" pageId="36" pageNumber="37">
<emphasis id="B94A7D7CFFD4BC60FD01FAB8FCF7CB05" bold="true" box="[731,842,1310,1331]" pageId="36" pageNumber="37">Bionomics.</emphasis>
Adults have been collected from near sea level to
<quantity id="4CC60C8BFFD4BC60FCE8FA9FFC3DCB78" box="[818,896,1337,1358]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="36" pageNumber="37" unit="m" value="2200.0">2,200 m</quantity>
elevation during MayAugust.
</paragraph>
</subSubSection>
<subSubSection id="C324F2E5FFD4BC60FD01FAF2FCFAC8F4" pageId="36" pageNumber="37" type="discussion">
<paragraph id="8B81A16EFFD4BC60FD01FAF2FCFAC8F4" blockId="36.[707,1194,1204,1730]" pageId="36" pageNumber="37">
<emphasis id="B94A7D7CFFD4BC60FD01FAF2FC80CB5F" bold="true" box="[731,829,1364,1385]" pageId="36" pageNumber="37">Remarks.</emphasis>
<taxonomicName id="4C3EDAEDFFD4BC60FC91FAF2FBF4CB5F" authorityName="Hatch" authorityYear="1957" box="[843,1097,1364,1385]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lagrandeuri">
<emphasis id="B94A7D7CFFD4BC60FC91FAF2FBF4CB5F" box="[843,1097,1364,1385]" italics="true" pageId="36" pageNumber="37">Phlaeopterus lagrandeuri</emphasis>
</taxonomicName>
is easily distinguished from all other
<taxonomicName id="4C3EDAEDFFD4BC60FC00FAC8FB14CBB5" box="[986,1193,1390,1411]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B94A7D7CFFD4BC60FC00FAC8FBE1CBB5" box="[986,1116,1390,1411]" italics="true" pageId="36" pageNumber="37">Phlaeopterus</emphasis>
species
</taxonomicName>
by the especially coarse, sparse punctation of the pronotum and elytra, yellow spot on each elytron (although occasionally vague or absent), tooth on the apical margin of the metatrochanter, form of the mandibles, and the unique structure of the internal sac of the aedeagus. This species most closely resembles Siberian
<taxonomicName id="4C3EDAEDFFD4BC60FC0BF98EFB84C80B" baseAuthorityName="Shavrin" baseAuthorityYear="2001" box="[977,1081,1576,1597]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="czerskyi">
<emphasis id="B94A7D7CFFD4BC60FC0BF98EFB84C80B" box="[977,1081,1576,1597]" italics="true" pageId="36" pageNumber="37">P. czerskyi</emphasis>
</taxonomicName>
but can be distinguished by many characters including the more shallowly impressed anteocellar foveae, less coarse punctation, lateral margins of the pronotum smooth, not crenulate, fully developed wings, and longer elytra.
</paragraph>
</subSubSection>
</treatment>
</document>