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<document id="A9E0F0389D6AAEF89F0F80925BF0D571" ID-DOI="10.1649/0010-065X-72.mo4.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5102840" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626277672172" checkinUser="carolina" docAuthor="Mullen, Logan J., Campbell, J. M. &amp; Sikes, Derek S." docDate="2018" docId="03971078FFD0BC65FD14FA6EFCD9CCB6" docLanguage="en" docName="TheColeopBull.mo.72.16.1-54.pdf" docOrigin="The Coleopterists Bulletin 72" docSource="http://dx.doi.org/10.1649/0010-065x-72.mo4.1" docStyle="DocumentStyle:A431C8F32F3D86E670C18C13DA9B5F2B.4:TheColeopBull.2000-2009.monograph" docStyleId="A431C8F32F3D86E670C18C13DA9B5F2B" docStyleName="TheColeopBull.2000-2009.monograph" docStyleVersion="4" docTitle="Phlaeopterus hatchi Mullen and Campbell 2018, new species" docType="treatment" docUuid="FF1FC2B7-035B-4479-88D4-6E3B9C93B407" docUuidSource="ZooBank" docVersion="3" lastPageNumber="34" masterDocId="FFAE6800FFF0BC44FFDAFFA6FFBDCE36" masterDocTitle="Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="54" masterPageNumber="1" pageNumber="33" updateTime="1699007718751" updateUser="plazi">
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<mods:title id="CF345F93DE2ADF1EB3C9BCCB887548B9">Taxonomic Revision of the Rove Beetle Genus Phlaeopterus Motschulsky, 1853 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<mods:namePart id="32B67C0A2E94FD6B11F7B52DDB7F5448">Mullen, Logan J.</mods:namePart>
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<mods:namePart id="2D4C8E02E583F24611BBBB7524474BC3">Campbell, J. M.</mods:namePart>
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<mods:namePart id="30754D163EB5C7A2C496BDE32C0A42DB">Sikes, Derek S.</mods:namePart>
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<mods:title id="F53E6E30A1651BB98F25E13A67694DA7">The Coleopterists Bulletin</mods:title>
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<mods:date id="763573BFD12992F324AA03A03F20EA6E">2018</mods:date>
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<mods:number id="4D813C115BA8C263850CDF62CD64A515">2018-12-28</mods:number>
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<paragraph id="8B81A16EFFD0BC64FD14FA6EFC4DCBCE" blockId="32.[718,1182,1480,1608]" pageId="32" pageNumber="33">
<heading id="D0C91602FFD0BC64FD14FA6EFC4DCBCE" centered="true" fontSize="10" level="2" pageId="32" pageNumber="33" reason="2">
<emphasis id="B94A7D7CFFD0BC64FD14FA6EFC4DCBCE" bold="true" pageId="32" pageNumber="33">
9.
<taxonomicName id="4C3EDAEDFFD0BC64FD3DFA6EFB24CBEB" ID-CoL="8Y8CW" authority="Mullen and Campbell" authorityName="Mullen and Campbell" authorityYear="2018" box="[743,1177,1480,1501]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="hatchi" status="sp. nov.">
<emphasis id="B94A7D7CFFD0BC64FD3DFA6EFC12CBEB" bold="true" box="[743,943,1480,1501]" italics="true" pageId="32" pageNumber="33">Phlaeopterus hatchi</emphasis>
Mullen and Campbell
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,
<taxonomicNameLabel id="A279C007FFD0BC64FCA1FA45FC4DCBCE" box="[891,1008,1507,1528]" pageId="32" pageNumber="33" rank="species">new species</taxonomicNameLabel>
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<paragraph id="8B81A16EFFD0BC64FD2BFA58FB2BC87E" blockId="32.[718,1182,1480,1608]" pageId="32" pageNumber="33">
<emphasis id="B94A7D7CFFD0BC64FD2BFA58FD44C825" bold="true" box="[753,761,1534,1555]" pageId="32" pageNumber="33">(</emphasis>
<figureCitation id="1305BDEBFFD0BC64FD23FA58FCF3C825" box="[761,846,1534,1555]" captionStart="Fig" captionStartId="12.[197,228,1727,1746]" captionTargetBox="[177,1188,196,1710]" captionTargetId="figure-0@12.[175,1190,194,1712]" captionTargetPageId="12" captionText="Fig. 3. Phlaeopterus species, habitus. A) P. hatchi, male (UAMObs:Ento:232429), B) P. houkae, male (UAMObs:" figureDoi="http://doi.org/10.5281/zenodo.5102847" httpUri="https://zenodo.org/record/5102847/files/figure.png" pageId="32" pageNumber="33">Figs. 3A</figureCitation>
,
<figureCitation id="1305BDEBFFD0BC64FC80FA58FC38C825" box="[858,901,1534,1555]" captionStart="Fig" captionStartId="17.[709,740,1029,1048]" captionTargetBox="[154,1169,195,1012]" captionTargetPageId="17" captionText="Fig. 10. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. hatchi, B) P. houkae. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected." figureDoi="http://doi.org/10.5281/zenodo.5102859" httpUri="https://zenodo.org/record/5102859/files/figure.png" pageId="32" pageNumber="33">10A</figureCitation>
,
<figureCitation id="1305BDEBFFD0BC64FC4BFA58FC01C825" box="[913,956,1534,1555]" captionStart="Fig" captionStartId="23.[175,206,1547,1566]" captionTargetBox="[168,1155,197,1527]" captionTargetId="figure-55@23.[164,1159,194,1532]" captionTargetPageId="23" captionText="Fig. 17. Phlaeopterus species. Pronota, dorsal view, of A) P. fusconiger, B) P. frosti, C) P. filicornis, D) P. hatchi, E) P. elongatus. Abdominal segments 46, dorsal view, of F) P. houkae, G) P. obsoletus, H) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5102867" httpUri="https://zenodo.org/record/5102867/files/figure.png" pageId="32" pageNumber="33">17D</figureCitation>
,
<figureCitation id="1305BDEBFFD0BC64FC1DFA58FC4CC825" box="[967,1009,1534,1555]" captionStart="Fig" captionStartId="30.[197,228,1342,1361]" captionTargetBox="[177,1188,196,1324]" captionTargetId="figure-145@30.[175,1190,194,1326]" captionTargetPageId="30" captionText="Fig. 25. Phlaeopterus species, aedeagi (left = ventral view; right = dorsal view of median lobe). A) Phlaeopterus frosti, B) P. hatchi, C) P. filicornis, D) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5102881" httpUri="https://zenodo.org/record/5102881/files/figure.png" pageId="32" pageNumber="33">25B</figureCitation>
,
<figureCitation id="1305BDEBFFD0BC64FC27FA58FB9EC825" box="[1021,1059,1534,1555]" captionStart="Fig" captionStartId="36.[731,762,1027,1046]" captionTargetBox="[710,1190,195,1011]" captionTargetId="figure-267@36.[709,1191,194,1012]" captionTargetPageId="36" captionText="Fig. 29. Phlaeopterus species. Right mandibles, ventral view, of A) P. obsoletus, B) P. bakerensis, C) P. elongatus. D) P. bakerensis, humeral angle of elytron, dorsal view. Apices of elytra of E) P. loganensis, F) P. hatchi, G) P. filicornis female." figureDoi="http://doi.org/10.5281/zenodo.5102887" httpUri="https://zenodo.org/record/5102887/files/figure.png" pageId="32" pageNumber="33">29F</figureCitation>
,
<figureCitation id="1305BDEBFFD0BC64FBF4FA58FBC9C825" box="[1070,1140,1533,1555]" captionStart="Fig" captionStartId="42.[197,228,1196,1215]" captionTargetBox="[176,656,195,1179]" captionTargetId="figure-217@42.[175,657,194,1180]" captionTargetPageId="42" captionText="Fig. 35. Heads, dorsal and dorsal oblique views, of AB) Phlaeopterus kavanaughi, CD Phlaeopterus hatchi. Eyes, dorsal view, of E) Lesteva longoelytrata, F) Unamis sp. undescribed." figureDoi="http://doi.org/10.5281/zenodo.5102899" httpUri="https://zenodo.org/record/5102899/files/figure.png" pageId="32" pageNumber="33">35CD</figureCitation>
) Zoobank.org/
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<paragraph id="8B81A16EFFD0BC64FD19F9C6FB19C843" blockId="32.[707,1188,1632,1653]" box="[707,1188,1632,1653]" pageId="32" pageNumber="33">
<bibRefCitation id="EFAFDC9FFFD0BC64FD19F9C6FCE9C843" author="Hatch" box="[707,852,1632,1653]" pageId="32" pageNumber="33" refId="ref28862" refString="Hatch, M. H. 1957. The Beetles of the Pacific Northwest. Part II: Staphyliniformia. University of Washington Publications in Biology, Seattle, WA." type="book" year="1957">Hatch 1957: 59</bibRefCitation>
[misidentification as
<taxonomicName id="4C3EDAEDFFD0BC64FBFDF9C6FB2AC843" authorityName="Casey" authorityYear="1893" box="[1063,1175,1632,1653]" class="Insecta" family="Staphylinidae" genus="Tilea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="rufitarsis">
<emphasis id="B94A7D7CFFD0BC64FBFDF9C6FB2AC843" box="[1063,1175,1632,1653]" italics="true" pageId="32" pageNumber="33">P. rufitarsis</emphasis>
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].
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<materialsCitation id="3B56AB33FFD0BC64FD01F934FB18C891" box="[731,1189,1682,1703]" location="Mt. Begbie" pageId="32" pageNumber="33" specimenCount="1" stateProvince="British Columbia" typeStatus="holotype">
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<typeStatus id="54851FCCFFD0BC64FD01F934FCB0C891" box="[731,781,1682,1703]" pageId="32" pageNumber="33">Type</typeStatus>
Locality.
</emphasis>
<location id="8EE1F7B5FFD0BC64FCA0F934FC51C891" LSID="urn:lsid:plazi:treatment:03971078FFD0BC65FD14FA6EFCD9CCB6:8EE1F7B5FFD0BC64FCA0F934FC51C891" box="[890,1004,1682,1703]" name="Mt. Begbie" pageId="32" pageNumber="33" stateProvince="British Columbia">Mt. Begbie</location>
,
<collectingRegion id="49FA6F8CFFD0BC64FC20F934FB18C891" box="[1018,1189,1682,1703]" country="Canada" name="British Columbia" pageId="32" pageNumber="33">British Columbia</collectingRegion>
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,
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<collectingCountry id="F329E1FEFFD0BC64FD19F90BFCB0C8F4" box="[707,781,1709,1730]" name="Canada" pageId="32" pageNumber="33">Canada</collectingCountry>
.
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<subSubSection id="C324F2E5FFD1BC65FF75FF64FE57CB7B" pageId="33" pageNumber="34" type="description">
<paragraph id="8B81A16EFFD1BC65FF75FF64FE57CB7B" blockId="33.[151,639,193,1729]" pageId="33" pageNumber="34">
<emphasis id="B94A7D7CFFD1BC65FF75FF64FE3BCEE1" bold="true" box="[175,390,194,215]" pageId="33" pageNumber="34">Description. Habitus:</emphasis>
Length
<quantity id="4CC60C8BFFD1BC65FE09FF64FDFACEE1" box="[467,583,193,215]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" metricValueMax="6.9" metricValueMin="5.5" pageId="33" pageNumber="34" unit="mm" value="6.2" valueMax="6.9" valueMin="5.5">5.56.9 mm</quantity>
. Dark brown, almost black; palpi, elytra, and glabrous apices of tibiae lighter (
<figureCitation id="1305BDEBFFD1BC65FEA9FF51FE03CF3A" box="[371,446,247,268]" captionStart="Fig" captionStartId="12.[197,228,1727,1746]" captionTargetBox="[177,1188,196,1710]" captionTargetId="figure-0@12.[175,1190,194,1712]" captionTargetPageId="12" captionText="Fig. 3. Phlaeopterus species, habitus. A) P. hatchi, male (UAMObs:Ento:232429), B) P. houkae, male (UAMObs:" figureDoi="http://doi.org/10.5281/zenodo.5102847" httpUri="https://zenodo.org/record/5102847/files/figure.png" pageId="33" pageNumber="34">Fig. 3A</figureCitation>
).
<emphasis id="B94A7D7CFFD1BC65FE14FF51FDB6CF3A" bold="true" box="[462,523,247,268]" pageId="33" pageNumber="34">Head:</emphasis>
Moderately narrow, ratio of width across eyes subequal to head length. Interantennal groove shallowly impressed (
<figureCitation id="1305BDEBFFD1BC65FF44FEE0FEA7CF6D" box="[158,282,326,348]" captionStart="Fig" captionStartId="42.[197,228,1196,1215]" captionTargetBox="[176,656,195,1179]" captionTargetId="figure-217@42.[175,657,194,1180]" captionTargetPageId="42" captionText="Fig. 35. Heads, dorsal and dorsal oblique views, of AB) Phlaeopterus kavanaughi, CD Phlaeopterus hatchi. Eyes, dorsal view, of E) Lesteva longoelytrata, F) Unamis sp. undescribed." figureDoi="http://doi.org/10.5281/zenodo.5102899" httpUri="https://zenodo.org/record/5102899/files/figure.png" pageId="33" pageNumber="34">Fig. 35CD</figureCitation>
). Anteocellar foveae large, deeply impressed. Eyes glabrous or with less than 10 scattered setae near ventral margin. Antennomeres 510 at least 2 times longer than wide; antennomeres 411 each with many sensory pits with papilliform projections. Ocelli present. Nuchal constriction vague. Mandibular molar area with Lshaped row of setae. Labrum with sensory pores along entire surface. Labial palpi with 3
<superScript id="7C4B0C26FFD1BC65FDDDFDB0FDABCC12" attach="left" box="[519,534,534,548]" fontSize="6" pageId="33" pageNumber="34">rd</superScript>
palpomere 1.61.8 times longer than palpomere 2.
<emphasis id="B94A7D7CFFD1BC65FDF0FD93FDC0CC7C" bold="true" box="[554,637,565,586]" pageId="33" pageNumber="34">Thorax:</emphasis>
Pronotum moderately narrow (
<figureCitation id="1305BDEBFFD1BC65FE64FDF6FDABCC53" box="[446,534,592,613]" captionStart="Fig" captionStartId="23.[175,206,1547,1566]" captionTargetBox="[168,1155,197,1527]" captionTargetId="figure-55@23.[164,1159,194,1532]" captionTargetPageId="23" captionText="Fig. 17. Phlaeopterus species. Pronota, dorsal view, of A) P. fusconiger, B) P. frosti, C) P. filicornis, D) P. hatchi, E) P. elongatus. Abdominal segments 46, dorsal view, of F) P. houkae, G) P. obsoletus, H) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5102867" httpUri="https://zenodo.org/record/5102867/files/figure.png" pageId="33" pageNumber="34">Fig. 17D</figureCitation>
), length to width ratio = 0.780.84; ratio of pronotal width to head width = 1.301.39; maximum width less than width of elytra at bases; punctures on dorsal surface separated by average distance equal to greater than diameter of single puncture; without vague impression near midline on ventral surface (as in
<taxonomicName id="4C3EDAEDFFD1BC65FDB6FD56FF50CD29" authorityName="Casey" authorityYear="1886" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="filicornis">
<emphasis id="B94A7D7CFFD1BC65FDB6FD56FF50CD29" italics="true" pageId="33" pageNumber="34">P. filicornis</emphasis>
</taxonomicName>
); lateral margins deflexed posterad and anterad lateral foveae; lateral foveae moderately impressed. Elytra with humeral angles convex; epipleural carina not projecting; 2.43.0 times longer than pronotum; apical margins not sexually dimorphic, broadly convex and not diverging in both sexes (
<figureCitation id="1305BDEBFFD1BC65FEDEFC0CFEEBCD89" box="[260,342,938,959]" captionStart="Fig" captionStartId="36.[731,762,1027,1046]" captionTargetBox="[710,1190,195,1011]" captionTargetId="figure-267@36.[709,1191,194,1012]" captionTargetPageId="36" captionText="Fig. 29. Phlaeopterus species. Right mandibles, ventral view, of A) P. obsoletus, B) P. bakerensis, C) P. elongatus. D) P. bakerensis, humeral angle of elytron, dorsal view. Apices of elytra of E) P. loganensis, F) P. hatchi, G) P. filicornis female." figureDoi="http://doi.org/10.5281/zenodo.5102887" httpUri="https://zenodo.org/record/5102887/files/figure.png" pageId="33" pageNumber="34">Fig. 29F</figureCitation>
). Wings fully developed in most individuals, brachypterous in very few. Mesosternum with projecting tooth; longitudinal carina along midline of mesosternum complete.
<emphasis id="B94A7D7CFFD1BC65FE3FFC5FFDA6CA38" bold="true" box="[485,539,1017,1038]" pageId="33" pageNumber="34">Legs:</emphasis>
Apices of all tibiae glabrous, length of subglabrous apex of metatibia as ratio to metatibial length = 4.47.8. Metatrochanter without tooth on apical margin.
<emphasis id="B94A7D7CFFD1BC65FF4DFBC5FEBDCA4E" bold="true" box="[151,256,1123,1144]" pageId="33" pageNumber="34">Abdomen:</emphasis>
Wing-folding spicules on tergites IV and V broad, transverse, nearly contiguous; tergite VI lacking wing-folding patches.
<emphasis id="B94A7D7CFFD1BC65FE1DFB3FFD92CA98" bold="true" box="[455,559,1177,1198]" pageId="33" pageNumber="34">Aedeagus:</emphasis>
Length
<quantity id="4CC60C8BFFD1BC65FF42FB15FE91CAFE" box="[152,300,1203,1225]" metricMagnitude="-3" metricUnit="m" metricValue="1.3399999999999999" metricValueMax="1.4" metricValueMin="1.28" pageId="33" pageNumber="34" unit="mm" value="1.3399999999999999" valueMax="1.4" valueMin="1.28">1.281.40 mm</quantity>
. Median lobe short, with lateral margins subparallel then acutely triangular nearing apex (
<figureCitation id="1305BDEBFFD1BC65FF0AFB4EFE99CACB" box="[208,292,1256,1277]" captionStart="Fig" captionStartId="30.[197,228,1342,1361]" captionTargetBox="[177,1188,196,1324]" captionTargetId="figure-145@30.[175,1190,194,1326]" captionTargetPageId="30" captionText="Fig. 25. Phlaeopterus species, aedeagi (left = ventral view; right = dorsal view of median lobe). A) Phlaeopterus frosti, B) P. hatchi, C) P. filicornis, D) P. elongatus." figureDoi="http://doi.org/10.5281/zenodo.5102881" httpUri="https://zenodo.org/record/5102881/files/figure.png" pageId="33" pageNumber="34">Fig. 25B</figureCitation>
). Internal sac elongate, nearly as long as median lobe, moderately heavily sclerotized, covered with microspinules but with distinctly larger microspinules in patches near apex.
</paragraph>
</subSubSection>
<subSubSection id="C324F2E5FFD1BC65FF75FAF5FD9DC80B" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="8B81A16EFFD1BC65FF75FAF5FD9DC80B" blockId="33.[151,639,193,1729]" pageId="33" pageNumber="34">
<emphasis id="B94A7D7CFFD1BC65FF75FAF5FEDDCB5E" bold="true" box="[175,352,1363,1384]" pageId="33" pageNumber="34">Type Specimens.</emphasis>
<materialsCitation id="3B56AB33FFD1BC65FEB4FAF5FE48CB8E" collectingDate="1971-08-30" collectorName="B. C., Mt. Begbie &amp; J. M. Campbell" location="Mt. Begbie" pageId="33" pageNumber="34" specimenCount="2" typeStatus="holotype">
<typeStatus id="54851FCCFFD1BC65FEB4FAF5FE7BCB5E" box="[366,454,1363,1384]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
male (UAMObs: Ento:235832) and
<typeStatus id="54851FCCFFD1BC65FE9DFAC8FE2ECBB5" box="[327,403,1390,1411]" pageId="33" pageNumber="34" type="allotype">allotype</typeStatus>
female (UAMObs:Ento: 235833) labeled as follows:
<collectorName id="26CBC4B8FFD1BC65FE1FFA2EFDC7CBAB" box="[453,634,1416,1437]" pageId="33" pageNumber="34">
B.C.,
<location id="8EE1F7B5FFD1BC65FDDFFA2EFDC7CBAB" LSID="urn:lsid:plazi:treatment:03971078FFD0BC65FD14FA6EFCD9CCB6:8EE1F7B5FFD1BC65FDDFFA2EFDC7CBAB" box="[517,634,1416,1437]" name="Mt. Begbie" pageId="33" pageNumber="34">Mt. Begbie</location>
</collectorName>
, 6300,
<date id="FF8087AEFFD1BC65FF03FA05FEDECB8E" box="[217,355,1442,1464]" pageId="33" pageNumber="34" value="1971-08-30">
<collectingDate id="EFC47E46FFD1BC65FF03FA05FEDECB8E" box="[217,355,1442,1464]" pageId="33" pageNumber="34" value="1971-08-30">VIII301971</collectingDate>
</date>
,
<collectorName id="26CBC4B8FFD1BC65FEB1FA05FE48CB8E" box="[363,501,1443,1464]" pageId="33" pageNumber="34">J.M. Campbell</collectorName>
</materialsCitation>
/
<typeStatus id="54851FCCFFD1BC65FDDAFA05FDC0CB8E" box="[512,637,1443,1464]" pageId="33" pageNumber="34" type="holotype">HOLOTYPE</typeStatus>
³ (or
<typeStatus id="54851FCCFFD1BC65FF0FFA1BFEF2CBE4" box="[213,335,1469,1490]" pageId="33" pageNumber="34" type="allotype">ALLOTYPE</typeStatus>
<specimenCount id="9D386AE7FFD1BC65FE83FA66FEDBCBE4" box="[345,358,1472,1490]" pageId="33" pageNumber="34" type="female"></specimenCount>
)
<taxonomicName id="4C3EDAEDFFD1BC65FEA2FA1BFD85CBE4" authorityName="Mullen and Campbell" authorityYear="2018" box="[376,568,1469,1490]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="hatchi">Phlaeopterus hatchi</taxonomicName>
, desig. L.J. Mullen and J.M. Campbell, CNC No. 18463 (red label). Both specimens are deposited in the CNC.
<typeStatus id="54851FCCFFD1BC65FF42F9ABFF4DC814" box="[152,240,1549,1570]" pageId="33" pageNumber="34" type="paratype">Paratypes</typeStatus>
(
<emphasis id="B94A7D7CFFD1BC65FF21F9ABFEBAC814" box="[251,263,1549,1570]" italics="true" pageId="33" pageNumber="34">n</emphasis>
= 73) are deposited in the AMNH, CAS, CNC, CSCA, FMNH, MCZ, and USNM.
</paragraph>
</subSubSection>
<subSubSection id="C324F2E5FFD1BC65FF75F9E4FD40CF9D" pageId="33" pageNumber="34" type="distribution">
<paragraph id="8B81A16EFFD1BC65FF75F9E4FC6ACEC7" blockId="33.[151,639,193,1729]" lastBlockId="33.[686,1173,194,640]" pageId="33" pageNumber="34">
<emphasis id="B94A7D7CFFD1BC65FF75F9E4FE8CC861" bold="true" box="[175,305,1602,1623]" pageId="33" pageNumber="34">Distribution.</emphasis>
<taxonomicName id="4C3EDAEDFFD1BC65FEEFF9E4FE4FC861" authorityName="Mullen and Campbell" authorityYear="2018" box="[309,498,1602,1623]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="hatchi">
<emphasis id="B94A7D7CFFD1BC65FEEFF9E4FE4FC861" box="[309,498,1602,1623]" italics="true" pageId="33" pageNumber="34">Phlaeopterus hatchi</emphasis>
</taxonomicName>
is known from northern
<collectingRegion id="49FA6F8CFFD1BC65FF2DF9FBFEEBC844" box="[247,342,1629,1650]" country="United States of America" name="California" pageId="33" pageNumber="34">California</collectingRegion>
north through the Cascade Range, the Mission Mountains of
<collectingRegion id="49FA6F8CFFD1BC65FE20F9D1FDECC8BA" box="[506,593,1655,1676]" country="United States of America" name="Montana" pageId="33" pageNumber="34">Montana</collectingRegion>
, the Rocky Mountains of
<collectingRegion id="49FA6F8CFFD1BC65FEA0F934FD94C891" box="[378,553,1682,1703]" country="Canada" name="British Columbia" pageId="33" pageNumber="34">British Columbia</collectingRegion>
, and in
<collectingRegion id="49FA6F8CFFD1BC65FF42F90AFF67C8F7" box="[152,218,1708,1729]" country="United States of America" name="Alaska" pageId="33" pageNumber="34">Alaska</collectingRegion>
by a single specimen (UAMObs:Ento: 232742) collected by A. Ord in 2012 between Haines and Juneau (
<figureCitation id="1305BDEBFFD1BC65FCABFF7AFC71CEC7" box="[881,972,220,241]" captionStart="Fig" captionStartId="17.[709,740,1029,1048]" captionTargetBox="[154,1169,195,1012]" captionTargetPageId="17" captionText="Fig. 10. Distributions of Phlaeopterus species based on all georeferenced specimen records. A) P. hatchi, B) P. houkae. Grey dots represent localities where at least one Phlaeopterus species, but not the species of interest, was collected. Red dots represent localities where the species of interest was collected." figureDoi="http://doi.org/10.5281/zenodo.5102859" httpUri="https://zenodo.org/record/5102859/files/figure.png" pageId="33" pageNumber="34">Fig. 10A</figureCitation>
).
</paragraph>
<paragraph id="8B81A16EFFD1BC65FD1FFF51FD40CF9D" blockId="33.[686,1173,194,640]" pageId="33" pageNumber="34">
<emphasis id="B94A7D7CFFD1BC65FD1FFF51FC88CF3A" bold="true" box="[709,821,247,268]" pageId="33" pageNumber="34">Bionomics.</emphasis>
Adults have been collected at
<quantity id="4CC60C8BFFD1BC65FD74FEB7FCF4CF10" box="[686,841,273,295]" metricMagnitude="3" metricUnit="m" metricValue="1.6149999999999998" metricValueMax="2.13" metricValueMin="1.1" pageId="33" pageNumber="34" unit="m" value="1615.0" valueMax="2130.0" valueMin="1100.0">
<geoCoordinate id="EE0AC7A9FFD1BC65FD74FEB7FD59CF10" box="[686,740,273,294]" degrees="1,100" direction="north" orientation="latitude" pageId="33" pageNumber="34" precision="55" value="1.1">1,100</geoCoordinate>
<geoCoordinate id="EE0AC7A9FFD1BC65FD39FEB7FC98CF10" box="[739,805,273,295]" degrees="2,130" direction="west" orientation="longitude" pageId="33" pageNumber="34" precision="55" value="-2.13">2,130</geoCoordinate>
m
</quantity>
elevation during JulyAugust. Adults were mostly collected from under stones at the edges of small, cold streams, but sometimes were taken under rocks near snowfields or in splash zones of waterfalls and moss along the edges of streams.
</paragraph>
</subSubSection>
<subSubSection id="C324F2E5FFD1BC65FD1FFE17FC5BCC7C" pageId="33" pageNumber="34" type="discussion">
<paragraph id="8B81A16EFFD1BC65FD1FFE17FC5BCC7C" blockId="33.[686,1173,194,640]" pageId="33" pageNumber="34">
<emphasis id="B94A7D7CFFD1BC65FD1FFE17FC9ACFF0" bold="true" box="[709,807,433,454]" pageId="33" pageNumber="34">Remarks.</emphasis>
<taxonomicName id="4C3EDAEDFFD1BC65FCEEFE17FC47CFF0" authorityName="Mullen and Campbell" authorityYear="2018" box="[820,1018,433,454]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="hatchi">
<emphasis id="B94A7D7CFFD1BC65FCEEFE17FC47CFF0" box="[820,1018,433,454]" italics="true" pageId="33" pageNumber="34">Phlaeopterus hatchi</emphasis>
</taxonomicName>
can be distinguished from all other
<taxonomicName id="4C3EDAEDFFD1BC65FC52FE6DFBECCFD6" box="[904,1105,459,480]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B94A7D7CFFD1BC65FC52FE6DFBBBCFD6" box="[904,1030,459,480]" italics="true" pageId="33" pageNumber="34">Phlaeopterus</emphasis>
species
</taxonomicName>
by the shape of the pronotum, the glabrous apices of the tibiae, and the aedeagus. It is similar to
<taxonomicName id="4C3EDAEDFFD1BC65FBF8FDA6FB33CC23" authorityName="Casey" authorityYear="1886" box="[1058,1166,512,533]" class="Insecta" family="Staphylinidae" genus="Phlaeopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="filicornis">
<emphasis id="B94A7D7CFFD1BC65FBF8FDA6FB33CC23" box="[1058,1166,512,533]" italics="true" pageId="33" pageNumber="34">P. filicornis</emphasis>
</taxonomicName>
, from which it differs by characters discussed in the Remarks section of that species.
</paragraph>
</subSubSection>
<subSubSection id="C324F2E5FFD1BC65FD1FFDF6FCD9CCB6" pageId="33" pageNumber="34" type="etymology">
<paragraph id="8B81A16EFFD1BC65FD1FFDF6FCD9CCB6" blockId="33.[686,1173,194,640]" pageId="33" pageNumber="34">
<emphasis id="B94A7D7CFFD1BC65FD1FFDF6FC85CC53" bold="true" box="[709,824,592,613]" pageId="33" pageNumber="34">Etymology.</emphasis>
This species is named in honor of Dr. Melville H. Hatch.
</paragraph>
</subSubSection>
</treatment>
</document>