treatments-xml/data/03/93/FD/0393FD7CFFBDF760FF0511ADFEC3C3EB.xml
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<document id="94EAE66D56B296CBA2957FBDE8DDB2CA" ID-DOI="10.11646/zootaxa.1949.1.4" ID-ISSN="1175­5334" ID-Zenodo-Dep="5240414" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629588191494" checkinUser="felipe" docAuthor="Southern, Phillip Sterling" docDate="2008" docId="0393FD7CFFBDF760FF0511ADFEC3C3EB" docLanguage="en" docName="zt01949p062.pdf" docOrigin="Zootaxa 1949 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1949.1.4" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Empoasca coofa Southern 2008, n. sp." docType="treatment" docVersion="3" lastPageNumber="60" masterDocId="FFAA8504FFBBF769FF921752FFB1C014" masterDocTitle="New species and color forms of Empoasca (Hemiptera: Cicadellidae: Typhlocybinae: Empoascini) from South America" masterLastPageNumber="62" masterPageNumber="51" pageNumber="57" updateTime="1699081674479" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="68A7AAB2ADA07CB0950A34E2AE68758C">New species and color forms of Empoasca (Hemiptera: Cicadellidae: Typhlocybinae: Empoascini) from South America</mods:title>
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<mods:name id="BCD4A8847D796496C1D4A3A0E730070C" type="personal">
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<mods:namePart id="26099D3CC66AFE3EB40A6502648CFEF0">Southern, Phillip Sterling</mods:namePart>
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<mods:title id="C2EEE22EF0214671A3BD2C73BBBB76D7">Zootaxa</mods:title>
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<mods:part id="846AF25CA56B187EF041948D636D4120">
<mods:date id="D196DB57CD362834E48B5BE35751CF5E">2008</mods:date>
<mods:detail id="B8EB1B098F3F90923045DD8B956E5A8D" type="pubDate">
<mods:number id="0460671ACED7D9A1567E70DBF23928C9">2008-12-03</mods:number>
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<mods:number id="9D6E741560CF3D387BA81368361D130E">1949</mods:number>
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<mods:number id="2F64A4FDC2773C56C04281508E9A85E9">1</mods:number>
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<mods:url id="199D1B7ACB34FE0DC96E3C5846A2264C">https://biotaxa.org/Zootaxa/article/view/zootaxa.1949.1.4</mods:url>
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<mods:classification id="4D63106D374E79EAFF278068DF1E2245">journal article</mods:classification>
<mods:identifier id="5B5D13EEFA874C63DBEDE2081FAE7915" type="DOI">10.11646/zootaxa.1949.1.4</mods:identifier>
<mods:identifier id="B5D413D82F73A3403041A1B8BEB574F7" type="ISSN">1175­5334</mods:identifier>
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<treatment id="0393FD7CFFBDF760FF0511ADFEC3C3EB" ID-DOI="http://doi.org/10.5281/zenodo.5240876" ID-Zenodo-Dep="5240876" LSID="urn:lsid:plazi:treatment:0393FD7CFFBDF760FF0511ADFEC3C3EB" httpUri="http://treatment.plazi.org/id/0393FD7CFFBDF760FF0511ADFEC3C3EB" lastPageId="9" lastPageNumber="60" pageId="6" pageNumber="57">
<subSubSection id="C3201FE1FFBDF76FFF0511ADFE19C70D" box="[151,424,1791,1817]" pageId="6" pageNumber="57" type="nomenclature">
<paragraph id="8B854C6AFFBDF76FFF0511ADFE19C70D" blockId="6.[151,424,1791,1852]" box="[151,424,1791,1817]" pageId="6" pageNumber="57">
<heading id="D0CDFB06FFBDF76FFF0511ADFE19C70D" bold="true" box="[151,424,1791,1817]" fontSize="11" level="1" pageId="6" pageNumber="57" reason="1">
<emphasis id="B94E9078FFBDF76FFF0511ADFE19C70D" bold="true" box="[151,424,1791,1817]" pageId="6" pageNumber="57">
<taxonomicName id="4C3A37E9FFBDF76FFF0511ADFEECC70D" ID-CoL="e3783a63-a149-44b6-a4f7-3a2badca9aef" authority="Southern, 2008" authorityName="Southern" authorityYear="2008" box="[151,349,1791,1817]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="coofa" status="sp. nov.">
<emphasis id="B94E9078FFBDF76FFF0511ADFEECC70D" bold="true" box="[151,349,1791,1817]" italics="true" pageId="6" pageNumber="57">Empoasca coofa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D2D03FFBDF76FFEF711ADFE19C70D" box="[357,424,1791,1817]" pageId="6" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3201FE1FFBDF76EFF051070FB24C77E" lastPageId="7" lastPageNumber="58" pageId="6" pageNumber="57" type="description">
<paragraph id="8B854C6AFFBDF76FFF051070FE95C728" blockId="6.[151,424,1791,1852]" box="[151,292,1826,1852]" pageId="6" pageNumber="57">
(
<figureCitation id="130150EFFFBDF76FFF0D1070FF4EC728" box="[159,255,1826,1852]" captionStart="FIGURE 1" captionStartId="1.[151,255,1143,1167]" captionTargetBox="[173,1418,505,1107]" captionTargetId="figure-378@1.[151,1436,493,1117]" captionTargetPageId="1" captionText="FIGURE 1. Empoasca spp. dorsal aspect (abdomen removed). a: Empoasca bartletti n. sp. b: Empoasca concava n. sp. c: Empoasca coofa n. sp. d: Empoasca longibrachiata Southern—common color pattern. e: Empoasca longibrachiata Southern—variant color pattern." figureDoi="http://doi.org/10.5281/zenodo.5240416" httpUri="https://zenodo.org/record/5240416/files/figure.png" pageId="6" pageNumber="57">Figs. 1c</figureCitation>
,
<figureCitation id="130150EFFFBDF76FFE9E1070FEADC728" box="[268,284,1826,1852]" captionStart="FIGURE 4" captionStartId="8.[151,255,1790,1814]" captionTargetBox="[244,1342,189,1763]" captionTargetId="figure-139@8.[244,1346,176,1822]" captionTargetPageId="8" captionText="FIGURE 4. Empoasca coofa n. sp. a. genital capsule, lateral view (broken line = outline of aedeagus); b. genital capsule, ventral view (broken line = outline of aedeagus); c1. aedeagus, lateral view; c2. aedeagus, variation in apical portion of shaft, lateral view; d. aedeagus, left anal hook and dorsal aedeagal support, posteroventral view; e. connective, ventral view; g. style, ventral view; h. pygofer process, lateral view of apical portion; i. pygofer process, ventral view of apical portion; k. pattern of sclerotization of dorsal surface of genital capsule, dorsal view; m. first sternal complex, anterior view; n. base of abdomen and second sternal apodemes, ventral view; p. second tergal apodemes, anterior view; q. base of abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5240424" httpUri="https://zenodo.org/record/5240424/files/figure.png" pageId="6" pageNumber="57">4</figureCitation>
)
</paragraph>
<paragraph id="8B854C6AFFBDF76EFF051020FD17C256" blockId="6.[151,1436,1906,2012]" lastBlockId="7.[151,1437,152,2018]" lastPageId="7" lastPageNumber="58" pageId="6" pageNumber="57">
<emphasis id="B94E9078FFBDF76FFF051020FE8DC798" bold="true" box="[151,316,1906,1932]" pageId="6" pageNumber="57">Body (male):</emphasis>
Length:
<quantity id="4CC2E18FFFBDF76FFE381020FD8CC798" box="[426,573,1906,1932]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="3.9" metricValueMin="3.5" pageId="6" pageNumber="57" unit="mm" value="3.7" valueMax="3.9" valueMin="3.5">3.53.9 mm</quantity>
; crown length:
<quantity id="4CC2E18FFFBDF76FFD681020FC1AC798" box="[762,939,1906,1932]" metricMagnitude="-4" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="6" pageNumber="57" unit="mm" value="0.245" valueMax="0.26" valueMin="0.23">0.230.26 mm</quantity>
; crown projection:
<quantity id="4CC2E18FFFBDF76FFB071020FAF5C798" box="[1173,1348,1906,1932]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" metricValueMax="11.0" metricValueMin="7" pageId="6" pageNumber="57" unit="mm" value="0.09" valueMax="0.11" valueMin="0.07">0.070.11 mm</quantity>
; crown projection ratio: 0.300.45; head length (including eyes):
<quantity id="4CC2E18FFFBDF76FFCDE10C8FC4AC7A0" box="[844,1019,1946,1972]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="4.8" metricValueMin="4.2" pageId="6" pageNumber="57" unit="mm" value="0.44999999999999996" valueMax="0.48" valueMin="0.42">0.420.48 mm</quantity>
; interocular width:
<quantity id="4CC2E18FFFBDF76FFB7510C8FA26C7A0" box="[1255,1431,1946,1972]" metricMagnitude="-4" metricUnit="m" metricValue="3.95" metricValueMax="4.1" metricValueMin="3.8" pageId="6" pageNumber="57" unit="mm" value="0.395" valueMax="0.41" valueMin="0.38">0.380.41 mm</quantity>
; head width (including eyes):
<quantity id="4CC2E18FFFBDF76FFE7C1090FD2CC7C8" box="[494,669,1986,2012]" metricMagnitude="-4" metricUnit="m" metricValue="8.200000000000001" metricValueMax="8.299999999999999" metricValueMin="8.100000000000001" pageId="6" pageNumber="57" unit="mm" value="0.82" valueMax="0.83" valueMin="0.81">0.810.83 mm</quantity>
; face length:
<quantity id="4CC2E18FFFBDF76FFCA91090FC58C7C8" box="[827,1001,1986,2012]" metricMagnitude="-4" metricUnit="m" metricValue="8.85" metricValueMax="9.3" metricValueMin="8.4" pageId="6" pageNumber="57" unit="mm" value="0.885" valueMax="0.93" valueMin="0.84">0.840.93 mm</quantity>
; interantennal width:
<quantity id="4CC2E18FFFBDF76FFB781090FA29C7C8" box="[1258,1432,1986,2012]" metricMagnitude="-4" metricUnit="m" metricValue="3.15" metricValueMax="3.3" metricValueMin="3.0" pageId="6" pageNumber="57" unit="mm" value="0.315" valueMax="0.33" valueMin="0.3">0.300.33 mm</quantity>
. pronotal width:
<quantity id="4CC2E18FFFBCF76EFEC617CAFDB3C0A6" box="[340,514,152,178]" metricMagnitude="-4" metricUnit="m" metricValue="8.2" metricValueMax="8.5" metricValueMin="7.9" pageId="7" pageNumber="58" unit="mm" value="0.82" valueMax="0.85" valueMin="0.79">0.790.85 mm</quantity>
; pronotal length:
<quantity id="4CC2E18FFFBCF76EFD4017CAFC30C0A6" box="[722,897,152,178]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" metricValueMax="4.6" metricValueMin="4.0" pageId="7" pageNumber="58" unit="mm" value="0.43" valueMax="0.46" valueMin="0.4">0.400.46 mm</quantity>
; hind tibia length:
<quantity id="4CC2E18FFFBCF76EFBF217CAFABFC0A6" box="[1120,1294,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="1.605" metricValueMax="1.63" metricValueMin="1.58" pageId="7" pageNumber="58" unit="mm" value="1.605" valueMax="1.63" valueMin="1.58">1.581.63 mm</quantity>
; hind tarsus length:
<quantity id="4CC2E18FFFBCF76EFF621792FE2FC0CE" box="[240,414,192,218]" metricMagnitude="-4" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.1" metricValueMin="5.8" pageId="7" pageNumber="58" unit="mm" value="0.595" valueMax="0.61" valueMin="0.58">0.580.61 mm</quantity>
; hind tarsomere I length:
<quantity id="4CC2E18FFFBCF76EFD5F1792FCCAC0CE" box="[717,891,192,218]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" metricValueMax="4.4" metricValueMin="3.8" pageId="7" pageNumber="58" unit="mm" value="0.41" valueMax="0.44" valueMin="0.38">0.380.44 mm</quantity>
; hind tarsomere I/hind tarsus ratio: 0.620.76. Body generally pale yellow-brown with cream markings (all specimens available appear to have been collected or stored in alcohol, possibly affecting color). Crown (
<figureCitation id="130150EFFFBCF76EFCCF1642FC1FC13E" box="[861,942,272,298]" captionStart="FIGURE 1" captionStartId="1.[151,255,1143,1167]" captionTargetBox="[173,1418,505,1107]" captionTargetId="figure-378@1.[151,1436,493,1117]" captionTargetPageId="1" captionText="FIGURE 1. Empoasca spp. dorsal aspect (abdomen removed). a: Empoasca bartletti n. sp. b: Empoasca concava n. sp. c: Empoasca coofa n. sp. d: Empoasca longibrachiata Southern—common color pattern. e: Empoasca longibrachiata Southern—variant color pattern." figureDoi="http://doi.org/10.5281/zenodo.5240416" httpUri="https://zenodo.org/record/5240416/files/figure.png" pageId="7" pageNumber="58">Fig. 1c</figureCitation>
) pale yellow-brown with large subhyaline spot on either side of midline. Coronal suture approaching but not reaching anterior margin. Face yellowbrown with cream markings; frontal suture ending below ocelli, nearer eye than midline. Pronotum yellowbrown along anterior margin, with cream spots, the majority of pronotum subhyaline. Scutellum yellowbrown with cream markings on midline and lateral margins. Forewings yellowish subhyaline in basal 2/3, hyaline, brownish in apical 1/3; apical veins R2 and
<collectingRegion id="49FE8288FFBCF76EFC9B168AFC86C1E6" box="[777,823,472,498]" country="Peru" name="Lima" pageId="7" pageNumber="58">RM</collectingRegion>
confluent basally (second apical cell triangular and stalked). Hind wing venation similar to
<taxonomicName id="4C3A37E9FFBCF76EFDE51552FC8DC20D" baseAuthorityName="Harris" baseAuthorityYear="1841" box="[631,828,512,537]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="fabae">
<emphasis id="B94E9078FFBCF76EFDE51552FC8DC20D" box="[631,828,512,537]" italics="true" pageId="7" pageNumber="58">Empoasca fabae</emphasis>
</taxonomicName>
.
<emphasis id="B94E9078FFBCF76EFCDF16ADFBB9C20D" bold="true" box="[845,1032,511,537]" pageId="7" pageNumber="58">Body (female):</emphasis>
Length:
<quantity id="4CC2E18FFFBCF76EFBE51552FABBC20E" box="[1143,1290,512,538]" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.2" metricValueMin="3.9" pageId="7" pageNumber="58" unit="mm" value="4.05" valueMax="4.2" valueMin="3.9">3.94.2 mm</quantity>
. Coloration, markings and wings similar to those of male.
</paragraph>
<paragraph id="8B854C6AFFBCF76EFF54151DFEBDC646" blockId="7.[151,1437,152,2018]" pageId="7" pageNumber="58">
<emphasis id="B94E9078FFBCF76EFF54151DFE99C27D" bold="true" box="[198,296,591,617]" pageId="7" pageNumber="58">Pygofer</emphasis>
tapered on both dorsal and ventral margins to narrowly rounded apex (
<figureCitation id="130150EFFFBCF76EFBF91502FB73C27E" box="[1131,1218,592,618]" captionStart="FIGURE 4" captionStartId="8.[151,255,1790,1814]" captionTargetBox="[244,1342,189,1763]" captionTargetId="figure-139@8.[244,1346,176,1822]" captionTargetPageId="8" captionText="FIGURE 4. Empoasca coofa n. sp. a. genital capsule, lateral view (broken line = outline of aedeagus); b. genital capsule, ventral view (broken line = outline of aedeagus); c1. aedeagus, lateral view; c2. aedeagus, variation in apical portion of shaft, lateral view; d. aedeagus, left anal hook and dorsal aedeagal support, posteroventral view; e. connective, ventral view; g. style, ventral view; h. pygofer process, lateral view of apical portion; i. pygofer process, ventral view of apical portion; k. pattern of sclerotization of dorsal surface of genital capsule, dorsal view; m. first sternal complex, anterior view; n. base of abdomen and second sternal apodemes, ventral view; p. second tergal apodemes, anterior view; q. base of abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5240424" httpUri="https://zenodo.org/record/5240424/files/figure.png" pageId="7" pageNumber="58">Fig. 4a</figureCitation>
); stout macrosetae (1118) along dorsal margin and in rough diagonal line across outer face of pygofer; 34 filamentous setae scattered on outer face.
<emphasis id="B94E9078FFBCF76EFE2115CDFE43C2AD" bold="true" box="[435,498,671,697]" pageId="7" pageNumber="58">Plate</emphasis>
in lateral view expanded dorsolaterally near base, thence subparallel-margined to rounded apex, curving evenly upward but not expanded in apical 1/3; in ventral view, expanded laterally near base. A-group setae (23); B-group setae (1624) relatively uniform in distribution, roughly uniseriate along dorsolateral margin to apex of plate; C-group setae (1821) biseriate near base of plate, uniseriate apically (ca. last 6); D-group setae (2735) roughly bi- or tri-seriate, extended almost to anterior margin of plate.
<emphasis id="B94E9078FFBCF76EFF051435FF62C395" bold="true" box="[151,211,871,897]" pageId="7" pageNumber="58">Style</emphasis>
in lateral view parallel margined to dentifer, bowed slightly dorsad at body/dentifer junction, ca. 4 setal bases or sensory pits laterally at apex of body, teeth (57) low, ridgelike, widely spaced (spacing decreasing apically), apex rounded; in ventral view (
<figureCitation id="130150EFFFBCF76EFD3614EAFCB1C3C6" box="[676,768,952,978]" captionStart="FIGURE 4" captionStartId="8.[151,255,1790,1814]" captionTargetBox="[244,1342,189,1763]" captionTargetId="figure-139@8.[244,1346,176,1822]" captionTargetPageId="8" captionText="FIGURE 4. Empoasca coofa n. sp. a. genital capsule, lateral view (broken line = outline of aedeagus); b. genital capsule, ventral view (broken line = outline of aedeagus); c1. aedeagus, lateral view; c2. aedeagus, variation in apical portion of shaft, lateral view; d. aedeagus, left anal hook and dorsal aedeagal support, posteroventral view; e. connective, ventral view; g. style, ventral view; h. pygofer process, lateral view of apical portion; i. pygofer process, ventral view of apical portion; k. pattern of sclerotization of dorsal surface of genital capsule, dorsal view; m. first sternal complex, anterior view; n. base of abdomen and second sternal apodemes, ventral view; p. second tergal apodemes, anterior view; q. base of abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5240424" httpUri="https://zenodo.org/record/5240424/files/figure.png" pageId="7" pageNumber="58">
Fig.
<quantity id="4CC2E18FFFBCF76EFD7014EAFCB1C3C6" box="[738,768,952,978]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="7" pageNumber="58" unit="g" value="4.0">4g</quantity>
</figureCitation>
) body bowed mediad, dentifer angled posterolaterad.
<emphasis id="B94E9078FFBCF76EFF05148DFEEDC3ED" bold="true" box="[151,348,991,1017]" pageId="7" pageNumber="58">Pygofer process</emphasis>
in lateral view angled slightly dorsad near base thence relatively straight, tapered slightly to apex, long, nearly reaching apex of plate, apically flattened dorsoventrally and spoon-shaped (
<figureCitation id="130150EFFFBCF76EFA83135AFADCC436" box="[1297,1389,1032,1058]" captionStart="FIGURE 4" captionStartId="8.[151,255,1790,1814]" captionTargetBox="[244,1342,189,1763]" captionTargetId="figure-139@8.[244,1346,176,1822]" captionTargetPageId="8" captionText="FIGURE 4. Empoasca coofa n. sp. a. genital capsule, lateral view (broken line = outline of aedeagus); b. genital capsule, ventral view (broken line = outline of aedeagus); c1. aedeagus, lateral view; c2. aedeagus, variation in apical portion of shaft, lateral view; d. aedeagus, left anal hook and dorsal aedeagal support, posteroventral view; e. connective, ventral view; g. style, ventral view; h. pygofer process, lateral view of apical portion; i. pygofer process, ventral view of apical portion; k. pattern of sclerotization of dorsal surface of genital capsule, dorsal view; m. first sternal complex, anterior view; n. base of abdomen and second sternal apodemes, ventral view; p. second tergal apodemes, anterior view; q. base of abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5240424" httpUri="https://zenodo.org/record/5240424/files/figure.png" pageId="7" pageNumber="58">Fig. 4h</figureCitation>
), in some specimens a few aculeae are scattered on dorsal surface near apex (difficult to see); in ventral view angled toward midline from near base, straight or very slightly bowed inward, apically with very fine serrations on medial margin (
<figureCitation id="130150EFFFBCF76EFE2A13D2FDBBC48E" box="[440,522,1152,1178]" captionStart="FIGURE 4" captionStartId="8.[151,255,1790,1814]" captionTargetBox="[244,1342,189,1763]" captionTargetId="figure-139@8.[244,1346,176,1822]" captionTargetPageId="8" captionText="FIGURE 4. Empoasca coofa n. sp. a. genital capsule, lateral view (broken line = outline of aedeagus); b. genital capsule, ventral view (broken line = outline of aedeagus); c1. aedeagus, lateral view; c2. aedeagus, variation in apical portion of shaft, lateral view; d. aedeagus, left anal hook and dorsal aedeagal support, posteroventral view; e. connective, ventral view; g. style, ventral view; h. pygofer process, lateral view of apical portion; i. pygofer process, ventral view of apical portion; k. pattern of sclerotization of dorsal surface of genital capsule, dorsal view; m. first sternal complex, anterior view; n. base of abdomen and second sternal apodemes, ventral view; p. second tergal apodemes, anterior view; q. base of abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5240424" httpUri="https://zenodo.org/record/5240424/files/figure.png" pageId="7" pageNumber="58">Fig. 4i</figureCitation>
).
<emphasis id="B94E9078FFBCF76EFDB2132DFD13C48D" bold="true" box="[544,674,1151,1177]" pageId="7" pageNumber="58">Anal hook</emphasis>
in lateral view (
<figureCitation id="130150EFFFBCF76EFCF613D2FC05C48E" box="[868,948,1152,1178]" captionStart="FIGURE 4" captionStartId="8.[151,255,1790,1814]" captionTargetBox="[244,1342,189,1763]" captionTargetId="figure-139@8.[244,1346,176,1822]" captionTargetPageId="8" captionText="FIGURE 4. Empoasca coofa n. sp. a. genital capsule, lateral view (broken line = outline of aedeagus); b. genital capsule, ventral view (broken line = outline of aedeagus); c1. aedeagus, lateral view; c2. aedeagus, variation in apical portion of shaft, lateral view; d. aedeagus, left anal hook and dorsal aedeagal support, posteroventral view; e. connective, ventral view; g. style, ventral view; h. pygofer process, lateral view of apical portion; i. pygofer process, ventral view of apical portion; k. pattern of sclerotization of dorsal surface of genital capsule, dorsal view; m. first sternal complex, anterior view; n. base of abdomen and second sternal apodemes, ventral view; p. second tergal apodemes, anterior view; q. base of abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5240424" httpUri="https://zenodo.org/record/5240424/files/figure.png" pageId="7" pageNumber="58">Fig. 4j</figureCitation>
) directed ventrad, reaching ca. mid-point of pygofer, slightly sinuate, tapered to sharp apex; in caudoventral view (
<figureCitation id="130150EFFFBCF76EFC7A13FAFB8DC4D6" box="[1000,1084,1192,1218]" captionStart="FIGURE 4" captionStartId="8.[151,255,1790,1814]" captionTargetBox="[244,1342,189,1763]" captionTargetId="figure-139@8.[244,1346,176,1822]" captionTargetPageId="8" captionText="FIGURE 4. Empoasca coofa n. sp. a. genital capsule, lateral view (broken line = outline of aedeagus); b. genital capsule, ventral view (broken line = outline of aedeagus); c1. aedeagus, lateral view; c2. aedeagus, variation in apical portion of shaft, lateral view; d. aedeagus, left anal hook and dorsal aedeagal support, posteroventral view; e. connective, ventral view; g. style, ventral view; h. pygofer process, lateral view of apical portion; i. pygofer process, ventral view of apical portion; k. pattern of sclerotization of dorsal surface of genital capsule, dorsal view; m. first sternal complex, anterior view; n. base of abdomen and second sternal apodemes, ventral view; p. second tergal apodemes, anterior view; q. base of abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5240424" httpUri="https://zenodo.org/record/5240424/files/figure.png" pageId="7" pageNumber="58">Fig. 4d</figureCitation>
) bifurcate in transverse plane, outer ramus curved mediad, inner ramus a variably developed lobe (may be broader and more distinct than in
<figureCitation id="130150EFFFBCF76EFF0513AAFF5CC506" box="[151,237,1272,1298]" captionStart="FIGURE 4" captionStartId="8.[151,255,1790,1814]" captionTargetBox="[244,1342,189,1763]" captionTargetId="figure-139@8.[244,1346,176,1822]" captionTargetPageId="8" captionText="FIGURE 4. Empoasca coofa n. sp. a. genital capsule, lateral view (broken line = outline of aedeagus); b. genital capsule, ventral view (broken line = outline of aedeagus); c1. aedeagus, lateral view; c2. aedeagus, variation in apical portion of shaft, lateral view; d. aedeagus, left anal hook and dorsal aedeagal support, posteroventral view; e. connective, ventral view; g. style, ventral view; h. pygofer process, lateral view of apical portion; i. pygofer process, ventral view of apical portion; k. pattern of sclerotization of dorsal surface of genital capsule, dorsal view; m. first sternal complex, anterior view; n. base of abdomen and second sternal apodemes, ventral view; p. second tergal apodemes, anterior view; q. base of abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5240424" httpUri="https://zenodo.org/record/5240424/files/figure.png" pageId="7" pageNumber="58">Fig. 4d</figureCitation>
) directed mediad.
<emphasis id="B94E9078FFBCF76EFE5C13A5FDF7C505" bold="true" box="[462,582,1271,1297]" pageId="7" pageNumber="58">Aedeagus</emphasis>
in lateral view (
<figureCitation id="130150EFFFBCF76EFC9D13AAFCD9C506" box="[783,872,1272,1298]" captionStart="FIGURE 4" captionStartId="8.[151,255,1790,1814]" captionTargetBox="[244,1342,189,1763]" captionTargetId="figure-139@8.[244,1346,176,1822]" captionTargetPageId="8" captionText="FIGURE 4. Empoasca coofa n. sp. a. genital capsule, lateral view (broken line = outline of aedeagus); b. genital capsule, ventral view (broken line = outline of aedeagus); c1. aedeagus, lateral view; c2. aedeagus, variation in apical portion of shaft, lateral view; d. aedeagus, left anal hook and dorsal aedeagal support, posteroventral view; e. connective, ventral view; g. style, ventral view; h. pygofer process, lateral view of apical portion; i. pygofer process, ventral view of apical portion; k. pattern of sclerotization of dorsal surface of genital capsule, dorsal view; m. first sternal complex, anterior view; n. base of abdomen and second sternal apodemes, ventral view; p. second tergal apodemes, anterior view; q. base of abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5240424" httpUri="https://zenodo.org/record/5240424/files/figure.png" pageId="7" pageNumber="58">Fig. 4c</figureCitation>
<figureCitation id="130150EFFFBCF76EFCFA13AAFCC2C506" box="[872,883,1272,1298]" captionStart="FIGURE 1" captionStartId="1.[151,255,1143,1167]" captionTargetBox="[173,1418,505,1107]" captionTargetId="figure-378@1.[151,1436,493,1117]" captionTargetPageId="1" captionText="FIGURE 1. Empoasca spp. dorsal aspect (abdomen removed). a: Empoasca bartletti n. sp. b: Empoasca concava n. sp. c: Empoasca coofa n. sp. d: Empoasca longibrachiata Southern—common color pattern. e: Empoasca longibrachiata Southern—variant color pattern." figureDoi="http://doi.org/10.5281/zenodo.5240416" httpUri="https://zenodo.org/record/5240416/files/figure.png" pageId="7" pageNumber="58">1</figureCitation>
) with preatrium bowed caudad at base thence straight, shaft diverging from atrium at near right angle thence curved dorsad, nearly as long as preatrium, tapered apically to acute apex, each side of shaft with a very small process at about mid-length, closely appressed to the shaft and often very difficult to see (apparently absent in
<specimenCount id="9D3C87E3FFBCF76EFB8A1222FB0EC59E" box="[1048,1215,1392,1418]" pageId="7" pageNumber="58" type="generic">one specimen</specimenCount>
), dorsal margin of shaft with triangular or hook-shaped (
<figureCitation id="130150EFFFBCF76EFDC912CAFD01C5A6" box="[603,688,1432,1458]" captionStart="FIGURE 4" captionStartId="8.[151,255,1790,1814]" captionTargetBox="[244,1342,189,1763]" captionTargetId="figure-139@8.[244,1346,176,1822]" captionTargetPageId="8" captionText="FIGURE 4. Empoasca coofa n. sp. a. genital capsule, lateral view (broken line = outline of aedeagus); b. genital capsule, ventral view (broken line = outline of aedeagus); c1. aedeagus, lateral view; c2. aedeagus, variation in apical portion of shaft, lateral view; d. aedeagus, left anal hook and dorsal aedeagal support, posteroventral view; e. connective, ventral view; g. style, ventral view; h. pygofer process, lateral view of apical portion; i. pygofer process, ventral view of apical portion; k. pattern of sclerotization of dorsal surface of genital capsule, dorsal view; m. first sternal complex, anterior view; n. base of abdomen and second sternal apodemes, ventral view; p. second tergal apodemes, anterior view; q. base of abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5240424" httpUri="https://zenodo.org/record/5240424/files/figure.png" pageId="7" pageNumber="58">Fig. 4c</figureCitation>
<figureCitation id="130150EFFFBCF76EFD2212CAFD0EC5A6" box="[688,703,1432,1458]" captionStart="FIGURE 2" captionStartId="3.[151,255,1834,1858]" captionTargetBox="[186,1405,188,1761]" captionTargetId="figure-124@3.[181,1406,183,1848]" captionTargetPageId="3" captionText="FIGURE 2. Empoasca bartletti n. sp. a. genital capsule, lateral view (broken line = outline of aedeagus); b. genital capsule, ventral view (broken line = outline of aedeagus); c1. aedeagus, lateral view; c2. aedeagus, variation in processes, lateral view; d. aedeagus, left anal hook and dorsal aedeagal support, posteroventral view; e. connective, ventral view; g. style, ventral view; h. pygofer process, lateral view of apical portion; i. pygofer process, ventral view; k. pattern of sclerotization of dorsal surface of genital capsule, dorsal view; m. first sternal complex, anterior view; n. base of abdomen and second sternal apodemes, ventral view; q. base of abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5240418" httpUri="https://zenodo.org/record/5240418/files/figure.png" pageId="7" pageNumber="58">2</figureCitation>
) expansion just beyond mid-length; in ventral view preatium narrow basally, expanding to broad atrium, shaft parallel-sided to truncate apex, gonopore appearing terminal or subterminal on ventral side.
<emphasis id="B94E9078FFBCF76EFD9C12B5FD2BC615" bold="true" box="[526,666,1511,1537]" pageId="7" pageNumber="58">Connective</emphasis>
(
<figureCitation id="130150EFFFBCF76EFD3E12BAFCB0C616" box="[684,769,1512,1538]" captionStart="FIGURE 4" captionStartId="8.[151,255,1790,1814]" captionTargetBox="[244,1342,189,1763]" captionTargetId="figure-139@8.[244,1346,176,1822]" captionTargetPageId="8" captionText="FIGURE 4. Empoasca coofa n. sp. a. genital capsule, lateral view (broken line = outline of aedeagus); b. genital capsule, ventral view (broken line = outline of aedeagus); c1. aedeagus, lateral view; c2. aedeagus, variation in apical portion of shaft, lateral view; d. aedeagus, left anal hook and dorsal aedeagal support, posteroventral view; e. connective, ventral view; g. style, ventral view; h. pygofer process, lateral view of apical portion; i. pygofer process, ventral view of apical portion; k. pattern of sclerotization of dorsal surface of genital capsule, dorsal view; m. first sternal complex, anterior view; n. base of abdomen and second sternal apodemes, ventral view; p. second tergal apodemes, anterior view; q. base of abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5240424" httpUri="https://zenodo.org/record/5240424/files/figure.png" pageId="7" pageNumber="58">Fig. 4e</figureCitation>
) broader than long, anterior bar well sclerotized, apical notch broad and deep.
<emphasis id="B94E9078FFBCF76EFE36115DFD5DC63D" bold="true" box="[420,748,1551,1577]" pageId="7" pageNumber="58">Dorsum of genital capsule</emphasis>
(
<figureCitation id="130150EFFFBCF76EFD6F1142FCE2C63E" box="[765,851,1552,1578]" captionStart="FIGURE 4" captionStartId="8.[151,255,1790,1814]" captionTargetBox="[244,1342,189,1763]" captionTargetId="figure-139@8.[244,1346,176,1822]" captionTargetPageId="8" captionText="FIGURE 4. Empoasca coofa n. sp. a. genital capsule, lateral view (broken line = outline of aedeagus); b. genital capsule, ventral view (broken line = outline of aedeagus); c1. aedeagus, lateral view; c2. aedeagus, variation in apical portion of shaft, lateral view; d. aedeagus, left anal hook and dorsal aedeagal support, posteroventral view; e. connective, ventral view; g. style, ventral view; h. pygofer process, lateral view of apical portion; i. pygofer process, ventral view of apical portion; k. pattern of sclerotization of dorsal surface of genital capsule, dorsal view; m. first sternal complex, anterior view; n. base of abdomen and second sternal apodemes, ventral view; p. second tergal apodemes, anterior view; q. base of abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5240424" httpUri="https://zenodo.org/record/5240424/files/figure.png" pageId="7" pageNumber="58">Fig. 4k</figureCitation>
) with horns distinct and widely separated, bridge indistinct.
</paragraph>
<paragraph id="8B854C6AFFBCF76EFF54110DFB24C77E" blockId="7.[151,1437,152,2018]" pageId="7" pageNumber="58">
<emphasis id="B94E9078FFBCF76EFF54110DFE61C66D" bold="true" box="[198,464,1631,1657]" pageId="7" pageNumber="58">First sternal complex</emphasis>
(
<figureCitation id="130150EFFFBCF76EFE721132FD8FC66E" box="[480,574,1632,1658]" captionStart="FIGURE 4" captionStartId="8.[151,255,1790,1814]" captionTargetBox="[244,1342,189,1763]" captionTargetId="figure-139@8.[244,1346,176,1822]" captionTargetPageId="8" captionText="FIGURE 4. Empoasca coofa n. sp. a. genital capsule, lateral view (broken line = outline of aedeagus); b. genital capsule, ventral view (broken line = outline of aedeagus); c1. aedeagus, lateral view; c2. aedeagus, variation in apical portion of shaft, lateral view; d. aedeagus, left anal hook and dorsal aedeagal support, posteroventral view; e. connective, ventral view; g. style, ventral view; h. pygofer process, lateral view of apical portion; i. pygofer process, ventral view of apical portion; k. pattern of sclerotization of dorsal surface of genital capsule, dorsal view; m. first sternal complex, anterior view; n. base of abdomen and second sternal apodemes, ventral view; p. second tergal apodemes, anterior view; q. base of abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5240424" httpUri="https://zenodo.org/record/5240424/files/figure.png" pageId="7" pageNumber="58">
Fig.
<quantity id="4CC2E18FFFBCF76EFD8A1132FD8FC66E" box="[536,574,1632,1658]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="58" unit="m" value="4.0">4m</quantity>
</figureCitation>
) with dorsal apodeme well developed, medial notch variable but in some specimens constricted above base forming a two-step taper.
<emphasis id="B94E9078FFBCF76EFCC311D5FB38C6B5" bold="true" box="[849,1161,1671,1697]" pageId="7" pageNumber="58">Second sternal apodemes</emphasis>
(
<figureCitation id="130150EFFFBCF76EFB0B11DAFB5CC6B6" box="[1177,1261,1672,1698]" captionStart="FIGURE 4" captionStartId="8.[151,255,1790,1814]" captionTargetBox="[244,1342,189,1763]" captionTargetId="figure-139@8.[244,1346,176,1822]" captionTargetPageId="8" captionText="FIGURE 4. Empoasca coofa n. sp. a. genital capsule, lateral view (broken line = outline of aedeagus); b. genital capsule, ventral view (broken line = outline of aedeagus); c1. aedeagus, lateral view; c2. aedeagus, variation in apical portion of shaft, lateral view; d. aedeagus, left anal hook and dorsal aedeagal support, posteroventral view; e. connective, ventral view; g. style, ventral view; h. pygofer process, lateral view of apical portion; i. pygofer process, ventral view of apical portion; k. pattern of sclerotization of dorsal surface of genital capsule, dorsal view; m. first sternal complex, anterior view; n. base of abdomen and second sternal apodemes, ventral view; p. second tergal apodemes, anterior view; q. base of abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5240424" httpUri="https://zenodo.org/record/5240424/files/figure.png" pageId="7" pageNumber="58">Fig. 4n</figureCitation>
) reaching 4.0 4.3 (length:
<quantity id="4CC2E18FFFBCF76EFEB411E2FE62C6DE" box="[294,467,1712,1738]" metricMagnitude="-4" metricUnit="m" metricValue="2.0500000000000003" metricValueMax="2.3000000000000003" metricValueMin="1.7999999999999998" pageId="7" pageNumber="58" unit="mm" value="0.205" valueMax="0.23" valueMin="0.18">0.180.23 mm</quantity>
; width:
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), well separated medially, subparallel-sided to rounded apicies.
<emphasis id="B94E9078FFBCF76EFF461185FDB5C6E5" bold="true" box="[212,516,1751,1777]" pageId="7" pageNumber="58">Second tergal apodemes</emphasis>
developed as vertical plates extended from dorsolateral surface towards midline, flattened at medial end in sagital plane (
<figureCitation id="130150EFFFBCF76EFD3A1052FC99C70E" box="[680,808,1792,1818]" captionStart="FIGURE 4" captionStartId="8.[151,255,1790,1814]" captionTargetBox="[244,1342,189,1763]" captionTargetId="figure-139@8.[244,1346,176,1822]" captionTargetPageId="8" captionText="FIGURE 4. Empoasca coofa n. sp. a. genital capsule, lateral view (broken line = outline of aedeagus); b. genital capsule, ventral view (broken line = outline of aedeagus); c1. aedeagus, lateral view; c2. aedeagus, variation in apical portion of shaft, lateral view; d. aedeagus, left anal hook and dorsal aedeagal support, posteroventral view; e. connective, ventral view; g. style, ventral view; h. pygofer process, lateral view of apical portion; i. pygofer process, ventral view of apical portion; k. pattern of sclerotization of dorsal surface of genital capsule, dorsal view; m. first sternal complex, anterior view; n. base of abdomen and second sternal apodemes, ventral view; p. second tergal apodemes, anterior view; q. base of abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5240424" httpUri="https://zenodo.org/record/5240424/files/figure.png" pageId="7" pageNumber="58">Figs. 4p, q</figureCitation>
), apodemes closely appressed but separate at midline (sometimes slightly more widely separated and concave medially than in
<figureCitation id="130150EFFFBCF76EFBB5107AFB32C756" box="[1063,1155,1832,1858]" captionStart="FIGURE 4" captionStartId="8.[151,255,1790,1814]" captionTargetBox="[244,1342,189,1763]" captionTargetId="figure-139@8.[244,1346,176,1822]" captionTargetPageId="8" captionText="FIGURE 4. Empoasca coofa n. sp. a. genital capsule, lateral view (broken line = outline of aedeagus); b. genital capsule, ventral view (broken line = outline of aedeagus); c1. aedeagus, lateral view; c2. aedeagus, variation in apical portion of shaft, lateral view; d. aedeagus, left anal hook and dorsal aedeagal support, posteroventral view; e. connective, ventral view; g. style, ventral view; h. pygofer process, lateral view of apical portion; i. pygofer process, ventral view of apical portion; k. pattern of sclerotization of dorsal surface of genital capsule, dorsal view; m. first sternal complex, anterior view; n. base of abdomen and second sternal apodemes, ventral view; p. second tergal apodemes, anterior view; q. base of abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5240424" httpUri="https://zenodo.org/record/5240424/files/figure.png" pageId="7" pageNumber="58">Fig. 4q</figureCitation>
).
<emphasis id="B94E9078FFBCF76EFB0E1075FE7BC77D" bold="true" pageId="7" pageNumber="58">Posterior margin of abdominal sternum VIII</emphasis>
nearly straight with very small, inconspicuous medial notch.
</paragraph>
</subSubSection>
<subSubSection id="C3201FE1FFBCF760FF541025FEC3C3EB" lastPageId="9" lastPageNumber="60" pageId="7" pageNumber="58" type="materials_examined">
<paragraph id="8B854C6AFFBCF76EFF541025FCE0C7F6" blockId="7.[151,1437,152,2018]" pageId="7" pageNumber="58">
<emphasis id="B94E9078FFBCF76EFF541025FECDC785" bold="true" box="[198,380,1911,1937]" pageId="7" pageNumber="58">
<typeStatus id="5481F2C8FFBCF76EFF541025FE89C785" box="[198,312,1911,1937]" pageId="7" pageNumber="58" type="holotype">Holotype</typeStatus>
male
</emphasis>
:
<collectingCountry id="F32D0CFAFFBCF76EFE1E102AFE4EC786" box="[396,511,1912,1938]" name="Colombia" pageId="7" pageNumber="58">Colombia</collectingCountry>
/ Boy Sogamoso /
<date id="FF846AAAFFBCF76EFD73102AFCCFC786" box="[737,894,1912,1938]" pageId="7" pageNumber="58" value="1946-05-29">29 May 1946</date>
// EA Chapin / Sta 46-23 // PSS RESEARCH / 06-241-01 //
<typeStatus id="5481F2C8FFBCF76EFED510F2FE53C7AE" box="[327,482,1952,1978]" pageId="7" pageNumber="58" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C3A37E9FFBCF76EFE6F10F2FDC6C7AD" authorityName="Walsh" authorityYear="1862" box="[509,631,1952,1977]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="B94E9078FFBCF76EFE6F10F2FDC6C7AD" box="[509,631,1952,1977]" italics="true" pageId="7" pageNumber="58">Empoasca</emphasis>
</taxonomicName>
/
<taxonomicName id="4C3A37E9FFBCF76EFD0110F2FD65C7AD" authority="Southern, 2008" authorityName="Southern" authorityYear="2008" box="[659,724,1952,1977]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="coofa" status="sp. nov.">
<emphasis id="B94E9078FFBCF76EFD0110F2FD65C7AD" box="[659,724,1952,1977]" italics="true" pageId="7" pageNumber="58">coofa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D2D03FFBCF76EFD4F10F2FCA4C7AE" box="[733,789,1952,1978]" pageId="7" pageNumber="58" rank="species">n.sp.</taxonomicNameLabel>
/ P. S. Southern 08 (
<collectionCode id="ED2BD4AFFFBCF76EFB8110F2FBD9C7AE" box="[1043,1128,1952,1978]" country="USA" httpUri="http://grbio.org/cool/ij62-iybb" name="North Carolina State University Insect Museum" pageId="7" pageNumber="58" type="Museum">NCSU</collectionCode>
).
<typeStatus id="5481F2C8FFBCF76EFBEC10F2FB45C7AE" box="[1150,1268,1952,1978]" pageId="7" pageNumber="58" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D3C87E3FFBCF76EFA9010F2FAC1C7AE" box="[1282,1392,1952,1978]" pageId="7" pageNumber="58" type="male">13 males</specimenCount>
,
<specimenCount id="9D3C87E3FFBCF76EFAEC10F2FF44C7F6" pageId="7" pageNumber="58" type="female">13 females</specimenCount>
, same collection data as
<typeStatus id="5481F2C8FFBCF76EFD84109AFDCDC7F6" box="[534,636,1992,2018]" pageId="7" pageNumber="58" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED2BD4AFFFBCF76EFD1F109AFD51C7F6" box="[653,736,1992,2018]" country="USA" httpUri="http://grbio.org/cool/ij62-iybb" name="North Carolina State University Insect Museum" pageId="7" pageNumber="58" type="Museum">NCSU</collectionCode>
,
<collectionCode id="ED2BD4AFFFBCF76EFD79109AFCF4C7F6" box="[747,837,1992,2018]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="58" type="Museum">USNM</collectionCode>
).
</paragraph>
<caption id="DF451CE2FFB3F761FF0511ACFE15C7F1" ID-DOI="http://doi.org/10.5281/zenodo.5240424" ID-Zenodo-Dep="5240424" httpUri="https://zenodo.org/record/5240424/files/figure.png" pageId="8" pageNumber="59" startId="8.[151,255,1790,1814]" targetBox="[244,1342,189,1763]" targetPageId="8">
<paragraph id="8B854C6AFFB3F761FF0511ACFE15C7F1" blockId="8.[151,1436,1790,2021]" pageId="8" pageNumber="59">
<emphasis id="B94E9078FFB3F761FF0511ACFEA9C702" bold="true" box="[151,280,1790,1814]" pageId="8" pageNumber="59">FIGURE 4.</emphasis>
<taxonomicName id="4C3A37E9FFB3F761FE8C11ADFE7EC702" authority="Southern, 2008" authorityName="Southern" authorityYear="2008" box="[286,463,1791,1814]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="59" phylum="Arthropoda" rank="species" species="coofa" status="sp. nov.">
<emphasis id="B94E9078FFB3F761FE8C11ADFE7EC702" box="[286,463,1791,1814]" italics="true" pageId="8" pageNumber="59">Empoasca coofa</emphasis>
</taxonomicName>
<emphasis id="B94E9078FFB3F761FE4711ACFDA1C702" bold="true" box="[469,528,1790,1814]" pageId="8" pageNumber="59">
<taxonomicNameLabel id="A27D2D03FFB3F761FE4711ACFDA1C702" box="[469,528,1790,1814]" pageId="8" pageNumber="59" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
a. genital capsule, lateral view (broken line = outline of aedeagus); b. genital capsule, ventral view (broken line = outline of aedeagus); c1. aedeagus, lateral view; c2. aedeagus, variation in apical portion of shaft, lateral view; d. aedeagus, left anal hook and dorsal aedeagal support, posteroventral view; e. connective, ventral view; g. style, ventral view; h. pygofer process, lateral view of apical portion; i. pygofer process, ventral view of apical portion; k. pattern of sclerotization of dorsal surface of genital capsule, dorsal view; m. first sternal complex, anterior view; n. base of abdomen and second sternal apodemes, ventral view; p. second tergal apodemes, anterior view; q. base of abdomen, dorsal view.
</paragraph>
</caption>
<paragraph id="8B854C6AFFB2F760FF5417CAFACAC0A6" blockId="9.[151,1436,152,1023]" box="[198,1403,152,178]" pageId="9" pageNumber="60">
The specific name
<taxonomicName id="4C3A37E9FFB2F760FE3617CAFE54C0A5" authorityName="Southern" authorityYear="2008" box="[420,485,152,177]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="coofa">
<emphasis id="B94E9078FFB2F760FE3617CAFE54C0A5" box="[420,485,152,177]" italics="true" pageId="9" pageNumber="60">coofa</emphasis>
</taxonomicName>
is an arbitrary combination of letters treated as a feminine noun in apposition.
</paragraph>
<paragraph id="8B854C6AFFB2F760FF541792FEC3C3EB" blockId="9.[151,1436,152,1023]" pageId="9" pageNumber="60">
In the key to
<taxonomicName id="4C3A37E9FFB2F760FEF21792FE6BC0CD" authorityName="Walsh" authorityYear="1862" box="[352,474,192,217]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="B94E9078FFB2F760FEF21792FE6BC0CD" box="[352,474,192,217]" italics="true" pageId="9" pageNumber="60">Empoasca</emphasis>
</taxonomicName>
of eastern
<collectingCountry id="F32D0CFAFFB2F760FDC81792FD21C0CE" box="[602,656,192,218]" name="Peru" pageId="9" pageNumber="60">Peru</collectingCountry>
(
<bibRefCitation id="EFAB319BFFB2F760FD0D1792FCE6C0CE" author="Southern P. S." box="[671,855,192,218]" pageId="9" pageNumber="60" refId="ref6784" refString="Southern P. S. (1982) A taxonomic study of the leafhopper genus Empoasca (Homoptera: Cidadellidae in eastern Peru. Technical Bulletin 272. North Carolina State University, Raleigh, N. C., 194 pp." type="book" year="1982">Southern, 1982</bibRefCitation>
),
<taxonomicName id="4C3A37E9FFB2F760FCFE1792FC7CC0CD" authorityName="Southern" authorityYear="2008" box="[876,973,192,217]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="coofa">
<emphasis id="B94E9078FFB2F760FCFE1792FC7CC0CD" box="[876,973,192,217]" italics="true" pageId="9" pageNumber="60">E. coofa</emphasis>
</taxonomicName>
keys to couplet 17 (if the aedeagal processes are present and judged to be at least 1/10
<superScript id="7C4FE122FFB2F760FD4817BAFD57C0E2" attach="left" box="[730,742,232,246]" fontSize="6" pageId="9" pageNumber="60">th</superScript>
the length of the shaft) and
<taxonomicName id="4C3A37E9FFB2F760FBAD17B9FF7AC138" authority="Southern. This" authorityName="Southern. This" authorityYear="1982" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="trindula">
<emphasis id="B94E9078FFB2F760FBAD17B9FA90C110" box="[1087,1313,235,260]" italics="true" pageId="9" pageNumber="60">Empoasca trindula</emphasis>
Southern. This
</taxonomicName>
species differs from
<taxonomicName id="4C3A37E9FFB2F760FE521641FD8FC138" authorityName="Southern. This" authorityYear="1982" box="[448,574,275,300]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="trindula">
<emphasis id="B94E9078FFB2F760FE521641FD8FC138" box="[448,574,275,300]" italics="true" pageId="9" pageNumber="60">E. trindula</emphasis>
</taxonomicName>
in having the aedeagal processes (if present) arising from near the middle of the shaft rather than from the base, the unique structure of the second tergal apodemes flattened in a sagital plane medially, the length of the second sternal apodemes which extend no further than 4.3, and the broad and deep medial notch in the dorsal apodeme of the first sternal complex. If the aedeagal processes are absent, not seen, or judged to be less than 1/10
<superScript id="7C4FE122FFB2F760FD1616E1FD21C1D5" attach="left" box="[644,656,435,449]" fontSize="6" pageId="9" pageNumber="60">th</superScript>
the length of the shaft,
<taxonomicName id="4C3A37E9FFB2F760FC5316E4FB99C1DB" authorityName="Southern" authorityYear="2008" box="[961,1064,438,463]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="coofa">
<emphasis id="B94E9078FFB2F760FC5316E4FB99C1DB" box="[961,1064,438,463]" italics="true" pageId="9" pageNumber="60">E. coofa</emphasis>
</taxonomicName>
keys to couplet 42 and either
<taxonomicName id="4C3A37E9FFB2F760FF05168CFE75C1E3" authority="Southern" authorityName="Southern" authorityYear="1982" box="[151,452,477,503]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="celta">
<emphasis id="B94E9078FFB2F760FF05168CFEE3C1E3" box="[151,338,478,503]" italics="true" pageId="9" pageNumber="60">Empoasca celta</emphasis>
Southern
</taxonomicName>
or
<taxonomicName id="4C3A37E9FFB2F760FE7C168CFC29C1E3" authority="Southern. However" authorityName="Southern. However" authorityYear="1982" box="[494,920,477,503]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="lecta">
<emphasis id="B94E9078FFB2F760FE7C168CFD18C1E3" box="[494,681,478,503]" italics="true" pageId="9" pageNumber="60">Empoasca lecta</emphasis>
Southern. However
</taxonomicName>
,
<taxonomicName id="4C3A37E9FFB2F760FC37168CFBB9C1E3" authorityName="Southern" authorityYear="2008" box="[933,1032,478,503]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="coofa">
<emphasis id="B94E9078FFB2F760FC37168CFBB9C1E3" box="[933,1032,478,503]" italics="true" pageId="9" pageNumber="60">E. coofa</emphasis>
</taxonomicName>
differs from both these species in external coloration, the anal hook much less deeply bifid in caudoventral view, the triangular or hooklike expansion on the dorsal margin of the aedeagal shaft, the form of the second tergal apodemes, and in the much shorter second sternal apodemes.
<taxonomicName id="4C3A37E9FFB2F760FDB71504FD39C27B" authorityName="Southern" authorityYear="2008" box="[549,648,598,623]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="coofa">
<emphasis id="B94E9078FFB2F760FDB71504FD39C27B" box="[549,648,598,623]" italics="true" pageId="9" pageNumber="60">E. coofa</emphasis>
</taxonomicName>
appears most closely related to
<taxonomicName id="4C3A37E9FFB2F760FB941504FF61C283" authority="Southern. Both" authorityName="Southern. Both" authorityYear="2006" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="ecuadorensis">
<emphasis id="B94E9078FFB2F760FB941504FA93C27B" box="[1030,1314,598,623]" italics="true" pageId="9" pageNumber="60">Empoasca ecuadorensis</emphasis>
Southern. Both
</taxonomicName>
species have relatively small processes arising from the shaft of the aedeagus, transversely bifurcate anal hooks, the aedeagal shaft initially diverging from the line of the atrial complex but then curving dorsad, short second sternal apodemes, and second tergal apodemes and flattened on the medial ends into plate-like structures which are closely opposed but are not fused. However, the aedeagal shaft of
<taxonomicName id="4C3A37E9FFB2F760FBF615A4FB76C31B" authorityName="Southern" authorityYear="2008" box="[1124,1223,758,783]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="coofa">
<emphasis id="B94E9078FFB2F760FBF615A4FB76C31B" box="[1124,1223,758,783]" italics="true" pageId="9" pageNumber="60">E. coofa</emphasis>
</taxonomicName>
is longer and that of
<taxonomicName id="4C3A37E9FFB2F760FF24144CFEDEC323" authorityName="Southern. Both" authorityYear="2006" box="[182,367,798,823]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="ecuadorensis">
<emphasis id="B94E9078FFB2F760FF24144CFEDEC323" box="[182,367,798,823]" italics="true" pageId="9" pageNumber="60">E. ecuadorensis</emphasis>
</taxonomicName>
lacks a dorsal expansion. The outer ramus of the anal hook of
<taxonomicName id="4C3A37E9FFB2F760FBD8144CFAB2C323" authorityName="Southern. Both" authorityYear="2006" box="[1098,1283,798,823]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="ecuadorensis">
<emphasis id="B94E9078FFB2F760FBD8144CFAB2C323" box="[1098,1283,798,823]" italics="true" pageId="9" pageNumber="60">E. ecuadorensis</emphasis>
</taxonomicName>
is much narrower and is bowed anteriorly with the apex directed caudad. Though the second tergal apodemes of both are flattened sagitally at the midline, the apodemes of
<taxonomicName id="4C3A37E9FFB2F760FD73143CFCF3C393" authorityName="Southern" authorityYear="2008" box="[737,834,878,903]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="coofa">
<emphasis id="B94E9078FFB2F760FD73143CFCF3C393" box="[737,834,878,903]" italics="true" pageId="9" pageNumber="60">E. coofa</emphasis>
</taxonomicName>
appear to be separated from the dorsum for most of their length (at most there may be an obscure membranous connection), while those of
<taxonomicName id="4C3A37E9FFB2F760FB2014C4FADFC3BB" authorityName="Southern. Both" authorityYear="2006" box="[1202,1390,918,943]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="ecuadorensis">
<emphasis id="B94E9078FFB2F760FB2014C4FADFC3BB" box="[1202,1390,918,943]" italics="true" pageId="9" pageNumber="60">E. ecuadorensis</emphasis>
</taxonomicName>
are clearly connected to the dorsum except at the midline.
<taxonomicName id="4C3A37E9FFB2F760FCBE14ECFC21C3C3" authorityName="Southern" authorityYear="2008" box="[812,912,958,983]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="coofa">
<emphasis id="B94E9078FFB2F760FCBE14ECFC21C3C3" box="[812,912,958,983]" italics="true" pageId="9" pageNumber="60">E. coofa</emphasis>
</taxonomicName>
is considered to be a member of the
<taxonomicName id="4C3A37E9FFB2F760FAC614ECFA2AC3C3" authorityName="Ruppel &amp; DeLong" authorityYear="1956" box="[1364,1435,958,983]" class="Insecta" family="Cicadellidae" genus="Empoasca" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="papae">
<emphasis id="B94E9078FFB2F760FAC614ECFA2AC3C3" box="[1364,1435,958,983]" italics="true" pageId="9" pageNumber="60">papae</emphasis>
</taxonomicName>
group (see below).
</paragraph>
</subSubSection>
</treatment>
</document>