204 lines
37 KiB
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204 lines
37 KiB
XML
<document ID-CLB-Dataset="6710" ID-DOI="http://dx.doi.org/10.3897/dez.63.10241" ID-GBIF-Dataset="0a6965a7-b124-4871-95d4-9e35c49073de" ID-Pensoft-Pub="1860-1324-2-211" ID-Pensoft-UUID="51243BBBE9F158F58BBC9BF3EA964017" ID-Zenodo-Dep="167458" ID-ZooBank="45552C4EC6AE4F9499980C2D492333B3" ModsDocID="1860-1324-63-2-211" checkinTime="1621372567443" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2016" docId="F8E281ED3BCCB38DA82F3AEDEB19805A" docLanguage="en" docName="DeutEntomolZeit 63(2): 211-270" docOrigin="Deutsche Entomologische Zeitschrift 63 (2)" docPubDate="2016-11-18" docSource="http://dx.doi.org/10.3897/dez.63.10241" docTitle="Cyphocoleus globulicollis Fauvel 1882" docType="treatment" docVersion="5" id="51243BBBE9F158F58BBC9BF3EA964017" lastPageNumber="250" masterDocId="51243BBBE9F158F58BBC9BF3EA964017" masterDocTitle="Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia" masterLastPageNumber="270" masterPageNumber="211" pageNumber="249" updateTime="1691866435949" updateUser="ExternalLinkService">
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<mods:title id="AF1A713BCBD6C6B76DEDB3DEBD0FCE2C">Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia</mods:title>
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<mods:namePart id="89D8A68B55511B1D362E0B949DC3FD83">Liebherr, James K.</mods:namePart>
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<mods:affiliation id="F65D2A7BE4E2690ADF87FC70A1E02396">Cornell University Insect Collection, John H. and Anna B. Comstock Hall, Cornell University, Ithaca, NY 14853 - 2601, USA</mods:affiliation>
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<mods:nameIdentifier id="48BB205350194927ADB48FC216B98E6D" type="email">jkl5@cornell.edu</mods:nameIdentifier>
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<mods:title id="FDEF4E0C28C09559FCD76BE9533C63DA">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="AE8FF99D793C003A44E60E56C1026958">2016</mods:date>
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<mods:number id="5A8B886BC68E3C0F62AB74D8DC933271">2016-11-18</mods:number>
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<mods:number id="98E19A3AFAFAD404D222583DB134C592">63</mods:number>
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<mods:classification id="617F0E5F278FAE3C53667663E3F42BCF">journal article</mods:classification>
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<mods:identifier id="E5015646B93E2713AD23EB632A4310CD" type="DOI">http://dx.doi.org/10.3897/dez.63.10241</mods:identifier>
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<treatment id="F8E281ED3BCCB38DA82F3AEDEB19805A" ID-GBIF-Taxon="182223707" LSID="urn:lsid:plazi:treatment:F8E281ED3BCCB38DA82F3AEDEB19805A" httpUri="http://treatment.plazi.org/id/F8E281ED3BCCB38DA82F3AEDEB19805A" lastPageId="39" lastPageNumber="250" pageId="38" pageNumber="249">
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<subSubSection id="9E550943B4D7A339E5B4295AB9119291" pageId="38" pageNumber="249" type="nomenclature">
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<paragraph id="42099DB74A226912E035F7F2DB8F3FB7" pageId="38" pageNumber="249">
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18.
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<taxonomicName id="213E772E1850E301869BA7CB89F25200" ID-CoL="6BWYZ" authority="Fauvel, 1882" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus globulicollis" order="Coleoptera" pageId="38" pageNumber="249" phylum="Arthropoda" rank="species" species="globulicollis">Cyphocoleus globulicollis Fauvel, 1882</taxonomicName>
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<figureCitation id="A791C9F7D01D571389ABBBEF23DB4A33" captionStart="Figures 86–98" captionStartId="F16" captionText="Figures 86 - 98. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 86, Cyphocoleus burwelli, internal sac everted, Ningua Res.; 87, Cyphocoleus angustatus, dextroventral view, internal sac everted, Mt. Panie; 88, Cyphocoleus monteithi, Parque Prov. Riviere Bleue, Haute Pourina; 89, Cyphocoleus miricollis, Mt. Mou; 90, Cyphocoleus fasciatus, dextral view, sac partially everted, Mt. Panie; 91 - 98, Cyphocoleus cychroides; 91, internal sac everted, Mandjelia; 92 - 93, Mandjelia; 94, internal sac everted, Hienghene; 95, Touho; 96, Tionaka Tal; 97, Ateou; 98, Pic d'Amoa. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures86-98" httpUri="https://binary.pensoft.net/fig/111553" pageId="38" pageNumber="249">Figures 91-98</figureCitation>
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<figureCitation id="A3C8768E806B3D8DE0E5FFC025AE069F" captionStart="Figures 105–112" captionStartId="F18" captionText="Figures 105 - 112. Female reproductive tract, ventral view, for Cyphocoleus spp.; 105, Cyphocoleus mirabilis, Mt. Koghi; 106, Cyphocoleus fasciatus, Mt. Panie; 107, Cyphocoleus cychroides, Pic d'Amoa; 108, Cyphocoleus latipennis, Mt. Panie; 109, Cyphocoleus cardiopterus, Mt. Mou; 110, Cyphocoleus globulicollis, Col des Roussettes; 111, Cyphocoleus flavipes, Noumea; 112, Cyphocoleus subulatus, Foret de Thi. For abbreviations see Table 1." figureDoi="10.3897/dez.63.10241.figures105-112" httpUri="https://binary.pensoft.net/fig/111555" pageId="38" pageNumber="249">, 110</figureCitation>
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<figureCitation id="4C5E32DB1105BA781ED602992767C8DC" captionStart="Figures 113–120" captionStartId="F19" captionText="Figures 113 - 120. Female right gonocoxae, ventral view, for Cyphocoleus spp. illustrating apical fringe setae of basal gonocoxite 1, and two lateral ensiform setae, one dorsal ensiform seta, and two apical nematiform setae on apical gonocoxite 2; 113, Cyphocoleus fasciatus, Mt. Panie; 114, Cyphocoleus cychroides, Pic d'Amoa; 115, Cyphocoleus latipennis Mt. Panie; 116, Cyphocoleus cordatus, Foret Nord; 117, Cyphocoleus globulicollis, Col des Roussettes; 118, Cyphocoleus flavipes, Noumea; 119, Cyphocoleus bourailensis, Bourail; 120, Cyphocoleus subulatus, Foret de Thi. Scale bar, 0.10 mm." figureDoi="10.3897/dez.63.10241.figures113-120" httpUri="https://binary.pensoft.net/fig/111556" pageId="38" pageNumber="249">, 117</figureCitation>
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<figureCitation id="5708EEF68F89194A0A3C4F9F72A56EF2" captionStart="Figures 121–125" captionStartId="F20" captionText="Figures 121 - 125. Dorsal habitus photographs of Cyphocoleus spp., environmental patina removed: 121, Cyphocoleus cychroides male, Tionaki Tal; 122, Cyphocoleus latipennis male, Mt. Panie; 123, Cyphocoleus cordatus female, Foret Nord; 124, Cyphocoleus cardiopterus female, Mt. Mou; 125, Cyphocoleus globulicollis female, Col des Roussettes." figureDoi="10.3897/dez.63.10241.figures121-125" httpUri="https://binary.pensoft.net/fig/111557" pageId="38" pageNumber="249">, 125</figureCitation>
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<figureCitation id="9656427B48AE50F6FF7419A3AA586ECB" captionStart="Figures 137–140" captionStartId="F22" captionText="Figures 137 - 140. Distribution maps of Cyphocoleus spp.: 137, Cyphocoleus cychroides, Cyphocoleus cordatus, Cyphocoleus cardiopterus; 138, Cyphocoleus latipennis; 139, Cyphocoleus globulicollis; 140 Cyphocoleus flavipes, Cyphocoleus bourailensis." figureDoi="10.3897/dez.63.10241.figures137-140" httpUri="https://binary.pensoft.net/fig/111561" pageId="38" pageNumber="249">, 139</figureCitation>
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<figureCitation id="18B42CC92E2746CABB3FC9FC8835DBD5" captionStart="Figures 141–160" captionStartId="F23" captionText="Figures 141 - 160. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 141 - 152, Cyphocoleus globulicollis; 141, Aoupinie; 142, Me Maoya; 143, Col de Roussettes; 144, internal sac partially everted, Gelima; 145, Col d'Amieu; 146, Mt. Do; 147, internal sac everted, Ningua; 148 - 150, Mt. Dzumac Rd; 151 - 152, Mt. Koghi; 153, Cyphocoleus flavipes, Yahoue; 154, Cyphocoleus bourailensis, Bourail; 155 - 158, Cyphocoleus subulatus; 155, La Crouen; 156, Mt. Rembai; 157, Col d'Amieu; 158, Mt. Koghi; 159 - 160, Cyphocoleus iledespinsensis, Ile des Pins; 159, dextral view; 160, ventral view." figureDoi="10.3897/dez.63.10241.figures141-160" httpUri="https://binary.pensoft.net/fig/111562" pageId="38" pageNumber="249">, 141-152</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="9CD670F254D7C0202F6685331ECF3369" pageId="38" pageNumber="249" type="diagnosis">
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<paragraph id="FA73A65F308CABFFBD4B051417C1D09A" pageId="38" pageNumber="249">Diagnosis</paragraph>
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<paragraph id="4A986E7DD8D6B6FEE2AEB3302F808578" pageId="38" pageNumber="249">
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(n = 5). Beetles comprising this species (Fig.
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<figureCitation id="0EF33EF1D0AF4D12DD4DA1E002B5DCF4" captionStart="Figures 121–125" captionStartId="F20" captionText="Figures 121 - 125. Dorsal habitus photographs of Cyphocoleus spp., environmental patina removed: 121, Cyphocoleus cychroides male, Tionaki Tal; 122, Cyphocoleus latipennis male, Mt. Panie; 123, Cyphocoleus cordatus female, Foret Nord; 124, Cyphocoleus cardiopterus female, Mt. Mou; 125, Cyphocoleus globulicollis female, Col des Roussettes." figureDoi="10.3897/dez.63.10241.figures121-125" httpUri="https://binary.pensoft.net/fig/111557" pageId="38" pageNumber="249">125</figureCitation>
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) exhibit a pronotal disc with distinct transverse wrinkles bordered laterally by deep parasagittal impressions, and a body surface covered with a well-developed pelage of evident, elongate microsetae. These setae are especially visible on the vertex and frons, even occurring between the ommatidia on the surface of the slightly convex eyes: each eye with 12-13 ommatidia across the horizontal diameter. The elytra are broadly ovoid, with intervals 3-7 very convex, subcarinate basally. The elytral surface is glossy, with only very shallow transverse sculpticells visible over portions of the surface. Standardized body length 4.8-6.7 mm.
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</paragraph>
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</subSubSection>
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<subSubSection id="53CED72580F846080A1CDD3C1B50F5EA" pageId="38" pageNumber="249" type="description">
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<paragraph id="CCB1C73BD4127AE8CD3190B30DA7BCFE" pageId="38" pageNumber="249">Description.</paragraph>
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<paragraph id="37CE659D270C6B739FBDA928604A8851" pageId="38" pageNumber="249">
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Head capsule parallel-sided, appearing
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<normalizedToken id="D6E96CAD62B01E06110C203E022F68AF" originalValue="“reduviidoid”">"reduviidoid"</normalizedToken>
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, the small, little-convex eyes only slightly extended beyond the genal profile in dorsal view; neck only slightly depressed medially, genae only indistinctly narrowed laterally before pronotal front angles; frons with narrow median longitudinal crest surrounded by flattened, rugose, lateral areas that lie mesad the deep, sinuous frontal grooves; frontoclypeal suture reduced, difficult to trace, frontal grooves apparently extended onto clypeus; mandibles elongate, length 2.1
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<normalizedToken id="75D96DC3D724CE61B3A3614F0EF39B5F" originalValue="×">x</normalizedToken>
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distance from antennal articulatory socket to anterolateral margin of labrum; antennae moderately elongate, scape length 2.5
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<normalizedToken id="DF39195B44CD835729908846A9005EEC" originalValue="×">x</normalizedToken>
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maximal breadth; gena setose, subgenal seta present. Pronotum with front angles protruded, angulate, the angle right to slightly acute; lateral marginal carinae extended to base, slightly sinuate just before depressed median base, the marginal bead across base irregularly interrupted by convex basal wrinkles of disc; median longitudinal impression deep and broad, separating discal wrinkles, extended nearly to anterior margin but deepest at juncture with transverse anterior impression; anterior transverse impression a depressed oblique border defining an irregularly surfaced anterior callosity bearing distinct longitudinal wrinkles; proepisternum bulging, distinctly extended beyond marginal bead in dorsal view; proepisternal surface dimpled, the irregularities mostly short vertical wrinkles near dorsal margin with proepipleuron, short longitudinal wrinkles ventrally; prosternum deeply depressed medially from pronotal anterior marginal bead to prosternal process, process broadly, slightly depressed on ventral face, posterior face with depressed ventral triangle and rounded medial ridge above. Elytra broadly ovoid, parascutellar region depressed and inflexed, basal margin sinuously subangulate basad stria 4, then broadly extended and posteriorly curved; sutural intervals elevated and fused into median callosity on middle of disc, intervals 3, 5, and 7, and to a lesser degree intervals 4 and 6, subcarinate, very convex on elevated basal portion of disc immediately posterad apex of parascutellar striole; striae deep, not visibly punctate except in extremely clean specimens where very small elongate punctulae can be seen in the deepest portion of the striae, these elongate depressions separated from each other by twice their length or more; parascutellar seta present, articulatory socket distinctly papillate, very upraised; dorsal elytral setae absent; lateral elytral setae arranged as 6 + 6, the articulatory sockets distinctly papillate; both subapical and apical elytral seta present, the articulatory socket for apical seta much smaller, not papillate; subapical sinuation distinctly developed, lateral portion of concavity joined to depressed lateral margin in tightly rounded curve; elytral apices rounded, slightly separated by shallow invagination at fused suture. Mesepisternum impunctate; metepisternal dorsal length 1.1
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<normalizedToken id="8F77FB6B77595F163BEEB686B00B1A0B" originalValue="×">x</normalizedToken>
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diagonal width; metasternum with deep, circular median fossa at base of metasternal process. Abdomen with apical margin of apical ventrite shallowly concave medially in both males and females; males with one seta each side of apical margin of apical ventrite, females with two setae each side. Legs moderately elongate; metacoxae bisetose; mt1 length/tibial length = 0.24; mt4 length to apex of outer lobe 2.0
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<normalizedToken id="0430DFA212599078F13904E29BC90DCF" originalValue="×">x</normalizedToken>
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median tarsomere length, 3-4 ventrolateral setae each side. Microsculpture on vertex densely transverse, shallow, difficult to trace on the glossy surface; higher areas between pronotal transverse wrinkles glossy, depressions of wrinkles and lateral flanges outside parasagittal groove covered with granulate isodiametric sculpticells, the surface matte; pelage well developed, head capsule, pronotum and elytra with densely distributed, evident microsetae, the setae on each elytral interval arranged in two irregular, longitudinal rows; prosternum with elongate pelage setae, meso- and metasternum largely glabrous, and meso- and metacoxae, and abdominal ventrites 2-3 medially with very short microsetae. Coloration of body dorsum piceous; thoracic sternites and elytral epipleura piceous, abdominal ventrites dark rufous; femora and tibiae rufopiceous, their articulation rufoflavous, tarsi rufobrunneous.
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</paragraph>
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<paragraph id="E8BF2FD51336EC7C798FEEC3EB69BD4A" pageId="38" pageNumber="249">
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<emphasis id="3735DA52690C90D72C0D0CDD56FA35DB" italics="true" pageId="38" pageNumber="249">Male genitalia</emphasis>
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(n = 21). Male aedeagal median lobe variably robust, from gracile (Figs
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<figureCitation id="B290424213C0200C6C9FA1AE60F09012" captionStart="Figures 141–160" captionStartId="F23" captionText="Figures 141 - 160. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 141 - 152, Cyphocoleus globulicollis; 141, Aoupinie; 142, Me Maoya; 143, Col de Roussettes; 144, internal sac partially everted, Gelima; 145, Col d'Amieu; 146, Mt. Do; 147, internal sac everted, Ningua; 148 - 150, Mt. Dzumac Rd; 151 - 152, Mt. Koghi; 153, Cyphocoleus flavipes, Yahoue; 154, Cyphocoleus bourailensis, Bourail; 155 - 158, Cyphocoleus subulatus; 155, La Crouen; 156, Mt. Rembai; 157, Col d'Amieu; 158, Mt. Koghi; 159 - 160, Cyphocoleus iledespinsensis, Ile des Pins; 159, dextral view; 160, ventral view." figureDoi="10.3897/dez.63.10241.figures141-160" httpUri="https://binary.pensoft.net/fig/111562" pageId="38" pageNumber="249">144</figureCitation>
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,
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<figureCitation id="67990BB682F0F7223CED3AB59F336FE4" captionStart="Figures 126–136" captionStartId="F21" captionText="Figures 126 - 136. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 126 - 133, Cyphocoleus latipennis; 126, internal sac everted, Mandjelia; 127 - 128, Mandjelia; 129, Mt. Panie; 130, Pic d'Amoa; 131 - 132, Aoupinie; 133, Ouenarou; 134 - 135, Cyphocoleus cordatus; 134, Mt. Koghi; 135, Cyphocoleus cordatus, internal sac everted, Foret Nord; 136, Cyphocoleus cardiopterus, Dzumac Rd. jct." figureDoi="10.3897/dez.63.10241.figures126-136" httpUri="https://binary.pensoft.net/fig/111559" pageId="38" pageNumber="249">136</figureCitation>
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) to more robust (Figs
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<figureCitation id="39C5F5ACA635A118177715EB8331006F" captionStart="Figures 141–160" captionStartId="F23" captionText="Figures 141 - 160. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 141 - 152, Cyphocoleus globulicollis; 141, Aoupinie; 142, Me Maoya; 143, Col de Roussettes; 144, internal sac partially everted, Gelima; 145, Col d'Amieu; 146, Mt. Do; 147, internal sac everted, Ningua; 148 - 150, Mt. Dzumac Rd; 151 - 152, Mt. Koghi; 153, Cyphocoleus flavipes, Yahoue; 154, Cyphocoleus bourailensis, Bourail; 155 - 158, Cyphocoleus subulatus; 155, La Crouen; 156, Mt. Rembai; 157, Col d'Amieu; 158, Mt. Koghi; 159 - 160, Cyphocoleus iledespinsensis, Ile des Pins; 159, dextral view; 160, ventral view." figureDoi="10.3897/dez.63.10241.figures141-160" httpUri="https://binary.pensoft.net/fig/111562" pageId="38" pageNumber="249">147</figureCitation>
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,
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<figureCitation id="4DBAD4B67CA14DE8C7619B4B7F32AF26" captionStart="Figures 141–160" captionStartId="F23" captionText="Figures 141 - 160. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 141 - 152, Cyphocoleus globulicollis; 141, Aoupinie; 142, Me Maoya; 143, Col de Roussettes; 144, internal sac partially everted, Gelima; 145, Col d'Amieu; 146, Mt. Do; 147, internal sac everted, Ningua; 148 - 150, Mt. Dzumac Rd; 151 - 152, Mt. Koghi; 153, Cyphocoleus flavipes, Yahoue; 154, Cyphocoleus bourailensis, Bourail; 155 - 158, Cyphocoleus subulatus; 155, La Crouen; 156, Mt. Rembai; 157, Col d'Amieu; 158, Mt. Koghi; 159 - 160, Cyphocoleus iledespinsensis, Ile des Pins; 159, dextral view; 160, ventral view." figureDoi="10.3897/dez.63.10241.figures141-160" httpUri="https://binary.pensoft.net/fig/111562" pageId="38" pageNumber="249">151</figureCitation>
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,
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<figureCitation id="C704B45BB86210371A82C5384FCD30B1" captionStart="Figures 141–160" captionStartId="F23" captionText="Figures 141 - 160. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 141 - 152, Cyphocoleus globulicollis; 141, Aoupinie; 142, Me Maoya; 143, Col de Roussettes; 144, internal sac partially everted, Gelima; 145, Col d'Amieu; 146, Mt. Do; 147, internal sac everted, Ningua; 148 - 150, Mt. Dzumac Rd; 151 - 152, Mt. Koghi; 153, Cyphocoleus flavipes, Yahoue; 154, Cyphocoleus bourailensis, Bourail; 155 - 158, Cyphocoleus subulatus; 155, La Crouen; 156, Mt. Rembai; 157, Col d'Amieu; 158, Mt. Koghi; 159 - 160, Cyphocoleus iledespinsensis, Ile des Pins; 159, dextral view; 160, ventral view." figureDoi="10.3897/dez.63.10241.figures141-160" httpUri="https://binary.pensoft.net/fig/111562" pageId="38" pageNumber="249">152</figureCitation>
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), dorsoventral breadth at midlength ranging from 0.16-0.22
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<normalizedToken id="917D798DA024CC87AEF59E1C67D71C6B" originalValue="×">x</normalizedToken>
|
||
distance from tip to base of closed basal bulb (Figs
|
||
<figureCitation id="A734D66BFD10F2FBAD0819C296235C6C" captionStart="Figures 141–160" captionStartId="F23" captionText="Figures 141 - 160. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 141 - 152, Cyphocoleus globulicollis; 141, Aoupinie; 142, Me Maoya; 143, Col de Roussettes; 144, internal sac partially everted, Gelima; 145, Col d'Amieu; 146, Mt. Do; 147, internal sac everted, Ningua; 148 - 150, Mt. Dzumac Rd; 151 - 152, Mt. Koghi; 153, Cyphocoleus flavipes, Yahoue; 154, Cyphocoleus bourailensis, Bourail; 155 - 158, Cyphocoleus subulatus; 155, La Crouen; 156, Mt. Rembai; 157, Col d'Amieu; 158, Mt. Koghi; 159 - 160, Cyphocoleus iledespinsensis, Ile des Pins; 159, dextral view; 160, ventral view." figureDoi="10.3897/dez.63.10241.figures141-160" httpUri="https://binary.pensoft.net/fig/111562" pageId="38" pageNumber="249">141-152</figureCitation>
|
||
); lobe apex little extended beyond ostial opening, rounded; internal sac membranous without melanized microtrichia, length 1.5
|
||
<normalizedToken id="A5D44EBA02888F1F9F7C501E23F25AB6" originalValue="×">x</normalizedToken>
|
||
breadth (Fig.
|
||
<figureCitation id="70CD26DDCD6909C2E64269F121B750C2" captionStart="Figures 141–160" captionStartId="F23" captionText="Figures 141 - 160. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 141 - 152, Cyphocoleus globulicollis; 141, Aoupinie; 142, Me Maoya; 143, Col de Roussettes; 144, internal sac partially everted, Gelima; 145, Col d'Amieu; 146, Mt. Do; 147, internal sac everted, Ningua; 148 - 150, Mt. Dzumac Rd; 151 - 152, Mt. Koghi; 153, Cyphocoleus flavipes, Yahoue; 154, Cyphocoleus bourailensis, Bourail; 155 - 158, Cyphocoleus subulatus; 155, La Crouen; 156, Mt. Rembai; 157, Col d'Amieu; 158, Mt. Koghi; 159 - 160, Cyphocoleus iledespinsensis, Ile des Pins; 159, dextral view; 160, ventral view." figureDoi="10.3897/dez.63.10241.figures141-160" httpUri="https://binary.pensoft.net/fig/111562" pageId="38" pageNumber="249">147</figureCitation>
|
||
). The substantial variation in robustness of the aedeagi for males assigned to
|
||
<taxonomicName id="D55759E2CC6A48514D56046AC2A9F99D" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus globulicollis" order="Diptera" pageId="38" pageNumber="249" phylum="Arthropoda" rank="species" species="globulicollis">
|
||
<emphasis id="4CBE41FB53769766DE64477583169FE7" italics="true" pageId="38" pageNumber="249">Cyphocoleus globulicollis</emphasis>
|
||
</taxonomicName>
|
||
is not associated with geography, as a male from Mt. Dzumac Rd. in the southern portion of the range (Fig.
|
||
<figureCitation id="E74AC84082BB0D654F9657A63AE1191C" captionStart="Figures 141–160" captionStartId="F23" captionText="Figures 141 - 160. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 141 - 152, Cyphocoleus globulicollis; 141, Aoupinie; 142, Me Maoya; 143, Col de Roussettes; 144, internal sac partially everted, Gelima; 145, Col d'Amieu; 146, Mt. Do; 147, internal sac everted, Ningua; 148 - 150, Mt. Dzumac Rd; 151 - 152, Mt. Koghi; 153, Cyphocoleus flavipes, Yahoue; 154, Cyphocoleus bourailensis, Bourail; 155 - 158, Cyphocoleus subulatus; 155, La Crouen; 156, Mt. Rembai; 157, Col d'Amieu; 158, Mt. Koghi; 159 - 160, Cyphocoleus iledespinsensis, Ile des Pins; 159, dextral view; 160, ventral view." figureDoi="10.3897/dez.63.10241.figures141-160" httpUri="https://binary.pensoft.net/fig/111562" pageId="38" pageNumber="249">150</figureCitation>
|
||
) varies little from the male from
|
||
<normalizedToken id="4FB9BB1CD8D2BD21F04E66CA321F1252" originalValue="Aoupinié">Aoupinie</normalizedToken>
|
||
at the northern margin (Fig.
|
||
<figureCitation id="D45E6EF51775AFBA2F7BBF35BDC8A771" captionStart="Figures 141–160" captionStartId="F23" captionText="Figures 141 - 160. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 141 - 152, Cyphocoleus globulicollis; 141, Aoupinie; 142, Me Maoya; 143, Col de Roussettes; 144, internal sac partially everted, Gelima; 145, Col d'Amieu; 146, Mt. Do; 147, internal sac everted, Ningua; 148 - 150, Mt. Dzumac Rd; 151 - 152, Mt. Koghi; 153, Cyphocoleus flavipes, Yahoue; 154, Cyphocoleus bourailensis, Bourail; 155 - 158, Cyphocoleus subulatus; 155, La Crouen; 156, Mt. Rembai; 157, Col d'Amieu; 158, Mt. Koghi; 159 - 160, Cyphocoleus iledespinsensis, Ile des Pins; 159, dextral view; 160, ventral view." figureDoi="10.3897/dez.63.10241.figures141-160" httpUri="https://binary.pensoft.net/fig/111562" pageId="38" pageNumber="249">141</figureCitation>
|
||
). Also two males from Mt. Koghi vary substantially in robustness (Figs
|
||
<figureCitation id="54708551A076BC33B5E7C9D5D3020613" captionStart="Figures 141–160" captionStartId="F23" captionText="Figures 141 - 160. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 141 - 152, Cyphocoleus globulicollis; 141, Aoupinie; 142, Me Maoya; 143, Col de Roussettes; 144, internal sac partially everted, Gelima; 145, Col d'Amieu; 146, Mt. Do; 147, internal sac everted, Ningua; 148 - 150, Mt. Dzumac Rd; 151 - 152, Mt. Koghi; 153, Cyphocoleus flavipes, Yahoue; 154, Cyphocoleus bourailensis, Bourail; 155 - 158, Cyphocoleus subulatus; 155, La Crouen; 156, Mt. Rembai; 157, Col d'Amieu; 158, Mt. Koghi; 159 - 160, Cyphocoleus iledespinsensis, Ile des Pins; 159, dextral view; 160, ventral view." figureDoi="10.3897/dez.63.10241.figures141-160" httpUri="https://binary.pensoft.net/fig/111562" pageId="38" pageNumber="249">151</figureCitation>
|
||
,
|
||
<figureCitation id="54CA0DA2BADA64F042378D1EDB586E27" captionStart="Figures 141–160" captionStartId="F23" captionText="Figures 141 - 160. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 141 - 152, Cyphocoleus globulicollis; 141, Aoupinie; 142, Me Maoya; 143, Col de Roussettes; 144, internal sac partially everted, Gelima; 145, Col d'Amieu; 146, Mt. Do; 147, internal sac everted, Ningua; 148 - 150, Mt. Dzumac Rd; 151 - 152, Mt. Koghi; 153, Cyphocoleus flavipes, Yahoue; 154, Cyphocoleus bourailensis, Bourail; 155 - 158, Cyphocoleus subulatus; 155, La Crouen; 156, Mt. Rembai; 157, Col d'Amieu; 158, Mt. Koghi; 159 - 160, Cyphocoleus iledespinsensis, Ile des Pins; 159, dextral view; 160, ventral view." figureDoi="10.3897/dez.63.10241.figures141-160" httpUri="https://binary.pensoft.net/fig/111562" pageId="38" pageNumber="249">152</figureCitation>
|
||
). All of these specimens exhibit the glossy cuticle with well-developed pelage of microsetae characteristic of
|
||
<taxonomicName id="E2AA7CB982CAAFA4EC8444444BF0CBA0" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus globulicollis" order="Diptera" pageId="38" pageNumber="249" phylum="Arthropoda" rank="species" species="globulicollis">
|
||
<emphasis id="7DFD8E253B5283FC52C1063A684A3465" italics="true" pageId="38" pageNumber="249">Cyphocoleus globulicollis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="AFFE5DD53113C65FE7078B3D8FE7AEBD" pageId="38" pageNumber="249">
|
||
<emphasis id="43B813ACE6DD7F6D0F8AA5EC6203446E" italics="true" pageId="38" pageNumber="249">Female reproductive tract</emphasis>
|
||
(n = 3). Bursa copulatrix columnar, distance from base of r gonocoxites to spermathecal basal sclerite 3
|
||
<normalizedToken id="4CADE73D9CE473C7F569569D0E97DFBC" originalValue="×">x</normalizedToken>
|
||
maximum breadth (dissection compressed under cover slip) (Fig.
|
||
<figureCitation id="CF50086CE616691F1AD52129C8C45A74" captionStart="Figures 105–112" captionStartId="F18" captionText="Figures 105 - 112. Female reproductive tract, ventral view, for Cyphocoleus spp.; 105, Cyphocoleus mirabilis, Mt. Koghi; 106, Cyphocoleus fasciatus, Mt. Panie; 107, Cyphocoleus cychroides, Pic d'Amoa; 108, Cyphocoleus latipennis, Mt. Panie; 109, Cyphocoleus cardiopterus, Mt. Mou; 110, Cyphocoleus globulicollis, Col des Roussettes; 111, Cyphocoleus flavipes, Noumea; 112, Cyphocoleus subulatus, Foret de Thi. For abbreviations see Table 1." figureDoi="10.3897/dez.63.10241.figures105-112" httpUri="https://binary.pensoft.net/fig/111555" pageId="38" pageNumber="249">110</figureCitation>
|
||
); bursal walls thin, translucent, lumenal surface smooth except for thin band of very fine setae over part of surface at midlength; basal gonocoxite with apical fringe of five to six setae (Fig.
|
||
<figureCitation id="58D2543D264CE4A15CFFC46B315140FE" captionStart="Figures 113–120" captionStartId="F19" captionText="Figures 113 - 120. Female right gonocoxae, ventral view, for Cyphocoleus spp. illustrating apical fringe setae of basal gonocoxite 1, and two lateral ensiform setae, one dorsal ensiform seta, and two apical nematiform setae on apical gonocoxite 2; 113, Cyphocoleus fasciatus, Mt. Panie; 114, Cyphocoleus cychroides, Pic d'Amoa; 115, Cyphocoleus latipennis Mt. Panie; 116, Cyphocoleus cordatus, Foret Nord; 117, Cyphocoleus globulicollis, Col des Roussettes; 118, Cyphocoleus flavipes, Noumea; 119, Cyphocoleus bourailensis, Bourail; 120, Cyphocoleus subulatus, Foret de Thi. Scale bar, 0.10 mm." figureDoi="10.3897/dez.63.10241.figures113-120" httpUri="https://binary.pensoft.net/fig/111556" pageId="38" pageNumber="249">117</figureCitation>
|
||
); apical gonocoxite broadly triangular with pointed apex and two lateral ensiform setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4CF2150C4B0BEEF9B1F4431E2B906833" lastPageId="39" lastPageNumber="250" pageId="38" pageNumber="249" type="materials_examined">
|
||
<paragraph id="DE2872E7C1D3A5F772804C123BA639EA" pageId="38" pageNumber="249">
|
||
<typeStatus id="5DA28735987EBE14DC937A6AC367403A">Type</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="CDA7C6A23D5A76A1E9DB6A9541562C1C" lastPageId="39" lastPageNumber="250" pageId="38" pageNumber="249">
|
||
<materialsCitation id="96F95C707FD6D701E3CF459370639609" collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" collectorName="Coll. R. I. Sc. N. B., Coll. R. I. Sc., Fauvel, Deplanche" determinerName="A. Fauvel" location="Ile des Pins" municipality="Mt. Mou" specimenCount="1" typeStatus="Lectotype">
|
||
<typeStatus id="01DF352BCD9DBA9F605EB801A1F8AA4E">Lectotype</typeStatus>
|
||
female (IRSN) hereby designated:
|
||
<location id="AA765B4587673C1796C50E65012CF857" LSID="urn:lsid:plazi:treatment:F8E281ED3BCCB38DA82F3AEDEB19805A:AA765B4587673C1796C50E65012CF857" municipality="Mt. Mou" name="Ile des Pins">Ile des Pins</location>
|
||
[and]
|
||
<collectingMunicipality id="D22A243525B0A53D865DEC3E6641C965">Mt. Mou</collectingMunicipality>
|
||
(two separate white labels glued to larger pink label)
|
||
<collectorName id="1F2E867D6F97FAD124C42B80340CAC1A">
|
||
<collectorName id="F9D4161A88C587D706CBE69679646276">Coll. R. I. Sc.</collectorName>
|
||
N. B.
|
||
</collectorName>
|
||
/ Nou
|
||
<pageBreakToken id="E588BA494FE7C0C458493F4E5132B72E" pageId="39" pageNumber="250" start="start">velle</pageBreakToken>
|
||
<normalizedToken id="A50C2F01EF69E9EC7E88987C57AA04C0" originalValue="Calédonie">Caledonie</normalizedToken>
|
||
/ ex. coll.
|
||
<collectorName id="56CF2D14B36B16C3F9F6D6C27B5796E7">Fauvel</collectorName>
|
||
/ rec.
|
||
<collectorName id="CD6444CD045F3281D23E037C20063DFC">Deplanche</collectorName>
|
||
// type (pink label with black border) // Coll. et det.
|
||
<determinerName id="7E1E42C7BB283EA1537C3D19A1DF0D0D">A. Fauvel</determinerName>
|
||
/
|
||
<taxonomicName id="27E8E2279CC3D487CF957254B5BF48A2" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus" order="Diptera" pageId="39" pageNumber="250" phylum="Arthropoda" rank="genus">Cyphocoleus</taxonomicName>
|
||
1882 /
|
||
<taxonomicName id="E5C4FCC68A28CC68D074C628804D4032" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="globulicollis" order="Diptera" pageId="39" pageNumber="250" phylum="Arthropoda" rank="species" species="globulicollis">globulicollis</taxonomicName>
|
||
Fauv. //
|
||
<typeStatus id="6C9F92BAEE462F855A5071C2A6424E65">LECTOTYPE</typeStatus>
|
||
/
|
||
<taxonomicName id="056C5673B7721B947D1F47C804C73CF9" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus" order="Diptera" pageId="39" pageNumber="250" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4B50CE29287A0C2F138326080DF40972" italics="true" pageId="39" pageNumber="250">Cyphocoleus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="0DEC13C7EA72009F3AFBB1C2B6166448" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="globulicollis" order="Diptera" pageId="39" pageNumber="250" phylum="Arthropoda" rank="species" species="globulicollis">
|
||
<emphasis id="0E92405463AD39B0DA16A55A09A6C59F" italics="true" pageId="39" pageNumber="250">globulicollis</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<bibRefCitation id="0222A79A58818C9661D01321C5BE46A5" author="Fauvel, A" journalOrPublisher="Revue d'Entomologie" pageId="48" pageNumber="259" pagination="241 - 261" refId="B18" refString="Fauvel, A, 1882. Les Coleopteres de la Nouvelle-Caledonie. Revue d'Entomologie 1: 241 - 261" title="Les Coleopteres de la Nouvelle-Caledonie." volume="1" year="1882">Fauvel 1882</bibRefCitation>
|
||
(black-bordered red label.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="97C6FE8AA2F510EFD438942C1BCC92B6" pageId="39" pageNumber="250" type="nomenclatural note">
|
||
<paragraph id="88C1C372AD269E218FED4A1068D426CA" pageId="39" pageNumber="250">Nomenclatural note.</paragraph>
|
||
<paragraph id="C64EEC8CB4C8BC5F39F951B492A72136" pageId="39" pageNumber="250">
|
||
<bibRefCitation id="3C292F705458C5DAD376318E5B725878" author="Fauvel, A" journalOrPublisher="Revue d'Entomologie" pageId="48" pageNumber="259" pagination="241 - 261" refId="B18" refString="Fauvel, A, 1882. Les Coleopteres de la Nouvelle-Caledonie. Revue d'Entomologie 1: 241 - 261" title="Les Coleopteres de la Nouvelle-Caledonie." volume="1" year="1882">Fauvel (1882</bibRefCitation>
|
||
: 250) described this species listing specimens from Mt. Mou and
|
||
<normalizedToken id="DCCBC712C95E4975B3531B6CADF2DD00" originalValue="Île">Ile</normalizedToken>
|
||
des Pins. The lectotype specimen has two labels glued to the pink IRSN museum card (above), one specifying
|
||
<normalizedToken id="0D4F66300F72501C9BD5DC02EF779F93" originalValue="“Île">"Ile</normalizedToken>
|
||
des Pins," the second "Mt. Mou." As these two localities are impossible to reconcile geographically on a single specimen, and as numerous other specimens of this species are recorded from Mt. Mou and vicinity, Mt. Mou is designated type locality, with
|
||
<normalizedToken id="8EEFC3ADD62F81FA89281B9BC3F6C52C" originalValue="Île">Ile</normalizedToken>
|
||
des Pins considered to represent an unverified locality for this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FF2D7BA489D2A23CA6A4D17373D3E2B8" pageId="39" pageNumber="250" type="distribution">
|
||
<paragraph id="EE083464269713BB7D0D3C32890AAB50" pageId="39" pageNumber="250">Distribution and habitat.</paragraph>
|
||
<paragraph id="4FE82BA082D3CFDD37AF03B54B8E980C" pageId="39" pageNumber="250">
|
||
This species is distributed in the southern two-thirds of Grande Terre, being known from
|
||
<normalizedToken id="E1E5A617F8E09F3EFC01E2208DCB861A" originalValue="Aoupinié">Aoupinie</normalizedToken>
|
||
on the north to Montagne des Sources on the south (Fig.
|
||
<figureCitation id="C964D79294D50752329BCB84D4BF7D1A" captionStart="Figures 137–140" captionStartId="F22" captionText="Figures 137 - 140. Distribution maps of Cyphocoleus spp.: 137, Cyphocoleus cychroides, Cyphocoleus cordatus, Cyphocoleus cardiopterus; 138, Cyphocoleus latipennis; 139, Cyphocoleus globulicollis; 140 Cyphocoleus flavipes, Cyphocoleus bourailensis." figureDoi="10.3897/dez.63.10241.figures137-140" httpUri="https://binary.pensoft.net/fig/111561" pageId="39" pageNumber="250">139</figureCitation>
|
||
, Suppl. material 3). Beetles have been found in leaf litter, and via the application of pyrethrin fog to trunks and logs. The well-developed pelage of microsetae is associated with a deep layer of environmental patina developing on these beetles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |