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<mods:title id="9E915B096DFB04E8546B44C5304C159F">The two southernmost species of the Andean genus Acrographinotus Holmgren (Arachnida, Opiliones, Gonyleptidae) described from Bolivia and Argentina</mods:title>
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<taxonomicName id="92F69E849F6AFF8F70AFFF28FE5BFF1C" authorityName="Acosta" authorityYear="2024" box="[151,475,151,178]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="556" phylum="Arthropoda" rank="species" species="calilegua" status="sp. nov.">
<emphasis id="678239159F6AFF8F70AFFF28FE5BFF1C" bold="true" box="[151,475,151,178]" italics="true" pageId="9" pageNumber="556">Acrographinotus calilegua</emphasis>
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<figureCitation id="CDCDF9829F6AFF8F70AFFF60FE96FF54" box="[151,278,223,250]" captionStart="FIGURE 5" captionStartId="9.[151,250,1745,1769]" captionTargetBox="[155,1432,374,1717]" captionTargetId="figure-45@9.[151,1435,370,1721]" captionTargetPageId="9" captionText="FIGURE 5. Acrographinotus calilegua sp. nov., HOlOtYpe ♂ (CDA 000.912). A. DORsal scutum, fRee teRgites (alsO shOwing VAP process from above), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, ventral anal plate, right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. Scale lines: 2 mm (A–E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783069" httpUri="https://zenodo.org/record/12783069/files/figure.png" pageId="9" pageNumber="556">Figs. 5A–F</figureCitation>
, 
<figureCitation id="CDCDF9829F6AFF8F7119FF60FEE4FF57" box="[289,356,223,249]" captionStart="FIGURE 6" captionStartId="11.[151,250,1298,1322]" captionTargetBox="[290,1297,185,1271]" captionTargetId="figure-15@11.[286,1301,181,1275]" captionTargetPageId="11" captionText="FIGURE 6. Acrographinotus calilegua sp. nov., distal end Of penis. A–B. PaRatYpe ♂ (MACN-AR 45699): A: dORsal view; B: lateRal view (membRanOus cRest is bent); C. HOlOtYpe ♂ (CDA 000.912), detail Of stYlus and ventRal pROcess. Scale lines: 0.1 mm (A, B), 0.05 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.12783071" httpUri="https://zenodo.org/record/12783071/files/figure.png" pageId="9" pageNumber="556">6A–C</figureCitation>
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<taxonomicName id="92F69E849F6AFF8F70AFFE9BFEC2FE92" ID-CoL="7NMYV" authorityName="Holmgren" authorityYear="1917" box="[151,322,292,316]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="556" phylum="Arthropoda" rank="genus">
<emphasis id="678239159F6AFF8F70AFFE9BFEC2FE92" box="[151,322,292,316]" italics="true" pageId="9" pageNumber="556">Acrographinotus</emphasis>
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[sp. from NW 
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] 
<treatmentCitation id="D457C3169F6AFF8F7265FE9BFC03FE95" author="Acosta, L. E. &amp; Maury, E. A." box="[605,899,291,316]" page="573" pageId="9" pageNumber="556" year="1998">
<bibRefCitation id="316798F69F6AFF8F7265FE9BFC03FE95" author="Acosta, L. E. &amp; Maury, E. A." box="[605,899,291,316]" pageId="9" pageNumber="556" pagination="569 - 580" refId="ref10832" refString="Acosta, L. E. &amp; Maury, E. A. (1998) Opiliones. In: Morrone, J. J. &amp; Coscaron, S. (Dirs.), Biodiversidad de Artropodos argentinos. Una perspectiva biotaxonomica. Ediciones Sur, La Plata, pp. 569 - 580." type="book chapter" year="1998">Acosta &amp; Maury 1998: 573</bibRefCitation>
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; 
<treatmentCitation id="D457C3169F6AFF8F73B7FE9BFBBEFE95" author="Acosta, L. E." box="[911,1086,291,316]" page="58" pageId="9" pageNumber="556" year="2001">
<bibRefCitation id="316798F69F6AFF8F73B7FE9BFBBEFE95" author="Acosta, L. E." box="[911,1086,291,316]" pageId="9" pageNumber="556" pagination="58 - 66" refId="ref10602" refString="Acosta, L. E. (2001) The identity of Acrographinotus erectispina, with a review of the generic diagnosis, and the description of a new species (Opiliones, Gonyleptidae, Pachylinae). Bulletin of the British Arachnological Society, 12 (2), 58 - 66." type="journal article" year="2001">Acosta 2001: 58</bibRefCitation>
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, 60; 2002b: 78; 
<treatmentCitation id="D457C3169F6AFF8F74D6FE9BFE16FEF4" author="Acosta, L. E. &amp; Perez-Gonzalez, A." page="326" pageId="9" pageNumber="556" year="2023">
<bibRefCitation id="316798F69F6AFF8F74D6FE9BFE16FEF4" author="Acosta, L. E. &amp; Perez-Gonzalez, A." pageId="9" pageNumber="556" refId="ref10889" refString="Acosta, L. E. &amp; Perez-Gonzalez, A. (2023) Opiliones. In: Claps, L. E., Roig-Junent, S. A. &amp; Morrone, J. J. (Dirs.), Biodiversidad de Artropodos Argentinos. Vol. 5. Ed. INSUE UNT, San Miguel de Tucuman, pp. 307 - 326." type="book" year="2023">Acosta &amp; Pérez-González 2023: 326</bibRefCitation>
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.
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<emphasis id="678239159F6AFF8F70AFF96EFE95F944" bold="true" box="[151,277,1745,1770]" pageId="9" pageNumber="556">FIGURE 5.</emphasis>
<taxonomicName id="92F69E849F6AFF8F7125F96DFDAEF944" authorityName="Acosta" authorityYear="2024" box="[285,558,1746,1770]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="556" phylum="Arthropoda" rank="species" species="calilegua" status="sp. nov.">
<emphasis id="678239159F6AFF8F7125F96DFDAEF944" box="[285,558,1746,1770]" italics="true" pageId="9" pageNumber="556">Acrographinotus calilegua</emphasis>
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, HOlOtYpe ♂ (CDA 000.912). A. DORsal scutum, fRee teRgites (alsO shOwing VAP process from above), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, ventral anal plate, right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. Scale lines: 2 mm (A–E), 1 mm (F).
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<emphasis id="678239159F69FF8C70AFFF29FEA7FF1E" bold="true" box="[151,295,150,176]" pageId="10" pageNumber="557">Type series.</emphasis>
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, 
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<collectingRegion id="97322BE59F69FF8C71D1FF28FD15FF1E" box="[489,661,151,176]" country="Argentina" name="Jujuy" pageId="10" pageNumber="557">Jujuy Province</collectingRegion>
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. 
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<specimenCount id="43F02E8E9F69FF8C7324FF28FCB1FF1F" box="[796,817,151,177]" count="1" pageId="10" pageNumber="557" type="male">♂</specimenCount>
(
<collectionCode id="33E77DC29F69FF8C737CFF29FC00FF1E" box="[836,896,150,176]" country="Canada" httpUri="http://grbio.org/cool/89m4-4i9r" name="Canadian Department of Agriculture" pageId="10" pageNumber="557">CDA</collectionCode>
000.912),
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<materialsCitation id="E59EEF5A9F69FF8C743AFF29FAE2FF1E" box="[1026,1378,150,177]" collectingDate="1994-12-23" collectingDateMax="1997-02-25" collectingDateMin="1994-12-23" collectionCode="CDA, LEA" collectorName="L. Acosta &amp; M. Acosta &amp; M. Ramirez &amp; P. Goloboff &amp; C. Szumik &amp; Mesada &amp; C. Grismado" country="Argentina" county="Calilegua National Park" elevation="1698" latitude="-23.625816" location="Calilegua NatiOnal PaRk" longLatPrecision="1" longitude="-64.93485" municipality="Monolito'" pageId="10" pageNumber="557" specimenCode="MACN-AR 45699, MACN-AR 45703, MACN-AR 45701" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="Jujuy" typeStatus="holotype">
<specimenCount id="43F02E8E9F69FF8C743AFF29FBB0FF1F" box="[1026,1072,150,177]" count="1" pageId="10" pageNumber="557" type="female">1 ♀</specimenCount>
<typeStatus id="8A4D5BA59F69FF8C7404FF28FB1CFF1F" box="[1084,1180,151,177]" pageId="10" pageNumber="557" type="paratype">paRatYpe</typeStatus>
(
<collectionCode id="33E77DC29F69FF8C7488FF29FB6CFF1E" box="[1200,1260,150,176]" country="Canada" httpUri="http://grbio.org/cool/89m4-4i9r" name="Canadian Department of Agriculture" pageId="10" pageNumber="557">CDA</collectionCode>
000.913),
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<materialsCitation id="E59EEF5A9F69FF8C7556FF29FA1CFF7B" collectingDate="1994-12-23" collectingDateMax="1997-02-25" collectingDateMin="1994-12-23" collectionCode="CDA, LEA" collectorName="L. Acosta &amp; M. Acosta &amp; M. Ramirez &amp; P. Goloboff &amp; C. Szumik &amp; Mesada &amp; C. Grismado" country="Argentina" county="Calilegua National Park" elevation="1698" latitude="-23.625816" location="Calilegua NatiOnal PaRk" longLatPrecision="1" longitude="-64.93485" municipality="Monolito'" pageId="10" pageNumber="557" specimenCode="MACN-AR 45699, MACN-AR 45703, MACN-AR 45701" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="Jujuy" typeStatus="holotype">
<specimenCount id="43F02E8E9F69FF8C7556FF29FA1CFF1F" box="[1390,1436,150,177]" count="1" pageId="10" pageNumber="557" type="female">1 ♀</specimenCount>
<typeStatus id="8A4D5BA59F69FF8C70AFFF04FF77FF7B" box="[151,247,187,213]" pageId="10" pageNumber="557" type="paratype">paratype</typeStatus>
(
<collectionCode id="33E77DC29F69FF8C7132FF05FEC1FF7A" box="[266,321,186,212]" pageId="10" pageNumber="557">LEA</collectionCode>
000.125): 
<collectingCounty id="BC289D8B9F69FF8C71FAFF05FD59FF7A" box="[450,729,186,212]" pageId="10" pageNumber="557">Calilegua National Park</collectingCounty>
, ‘ 
<collectingMunicipality id="B52D7F7D9F69FF8C72D4FF05FCE5FF7A" box="[748,869,186,212]" pageId="10" pageNumber="557">Monolito’</collectingMunicipality>
, 
<date id="2148C3C79F69FF8C734DFF05FBC6FF7A" box="[885,1094,186,213]" pageId="10" pageNumber="557" value="1997-02-25">
<collectingDate id="310C3A2F9F69FF8C734DFF05FBC6FF7A" box="[885,1094,186,213]" pageId="10" pageNumber="557" value="1997-02-25">25 February 1997</collectingDate>
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, 
<collectorName id="F80380D19F69FF8C746DFF04FB47FF7A" box="[1109,1223,186,213]" pageId="10" pageNumber="557">L. Acosta</collectorName>
, 
<collectorName id="F80380D19F69FF8C74EEFF04FACDFF7A" box="[1238,1357,186,213]" pageId="10" pageNumber="557">M. Acosta</collectorName>
<emphasis id="678239159F69FF8C756FFF05FA00FF7A" box="[1367,1408,186,212]" italics="true" pageId="10" pageNumber="557">leg.</emphasis>
—
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<materialsCitation id="E59EEF5A9F69FF8C70AFFF61FA13FEB2" collectingDate="1994-12-23" collectingDateMax="1997-02-25" collectingDateMin="1994-12-23" collectionCode="CDA, LEA" collectorName="L. Acosta &amp; M. Acosta &amp; M. Ramirez &amp; P. Goloboff &amp; C. Szumik &amp; Mesada &amp; C. Grismado" country="Argentina" county="Calilegua National Park" elevation="1698" latitude="-23.625816" location="Calilegua NatiOnal PaRk" longLatPrecision="1" longitude="-64.93485" municipality="Monolito'" pageId="10" pageNumber="557" specimenCode="MACN-AR 45699, MACN-AR 45703, MACN-AR 45701" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="Jujuy" typeStatus="holotype">
<specimenCount id="43F02E8E9F69FF8C70AFFF61FF47FF57" box="[151,199,222,249]" count="1" pageId="10" pageNumber="557" type="male">1 ♂</specimenCount>
, 
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(
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), 
<location id="5029B3DC9F69FF8C7240FF61FC11FF56" LSID="urn:lsid:plazi:treatment:DD5F54119F6AFF8B70AFFF28FC46FECB:5029B3DC9F69FF8C7240FF61FC11FF56" box="[632,913,222,248]" country="Argentina" county="Calilegua National Park" latitude="-23.625816" longLatPrecision="1" longitude="-64.93485" municipality="Monolito'" name="Calilegua NatiOnal PaRk" pageId="10" pageNumber="557" stateProvince="Jujuy">Calilegua NatiOnal PaRk</location>
, 
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fROm the entRance 
<location id="5029B3DC9F69FF8C74DBFF61FB7AFF56" LSID="urn:lsid:plazi:treatment:DD5F54119F6AFF8B70AFFF28FC46FECB:5029B3DC9F69FF8C74DBFF61FB7AFF56" box="[1251,1274,222,248]" country="Argentina" county="Calilegua National Park" latitude="-23.625816" longLatPrecision="1" longitude="-64.93485" municipality="Monolito'" name="Of" pageId="10" pageNumber="557" stateProvince="Jujuy">Of</location>
the 
<location id="5029B3DC9F69FF8C750BFF61FAE6FF56" LSID="urn:lsid:plazi:treatment:DD5F54119F6AFF8B70AFFF28FC46FECB:5029B3DC9F69FF8C750BFF61FAE6FF56" box="[1331,1382,222,248]" country="Argentina" county="Calilegua National Park" latitude="-23.625816" longLatPrecision="1" longitude="-64.93485" municipality="Monolito'" name="PaRk" pageId="10" pageNumber="557" stateProvince="Jujuy">PaRk</location>
[
<emphasis id="678239159F69FF8C7543FF60FA1CFF56" box="[1403,1436,223,248]" italics="true" pageId="10" pageNumber="557">ca.</emphasis>
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<geoCoordinate id="30C283C09F69FF8C7115FEBDFE44FEB3" box="[301,452,258,285]" degrees="64" direction="west" minutes="54.022" orientation="longitude" pageId="10" pageNumber="557" precision="1" value="-64.90037">64°54.022’W</geoCoordinate>
, ~ 
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<quantity id="920E48E29F69FF8C71D8FEBDFDB7FEB3" box="[480,567,258,285]" metricMagnitude="3" metricUnit="m" metricValue="1.698" pageId="10" pageNumber="557" unit="m" value="1698.0">1698 m</quantity>
a.s.l.
</elevation>
], 
<date id="2148C3C79F69FF8C72B4FEBDFC15FEB3" box="[652,917,258,285]" pageId="10" pageNumber="557" value="1995-09-23" valueMax="1995-09-24" valueMin="1995-09-23">
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</date>
, 
<collectorName id="F80380D19F69FF8C7399FEBCFBACFEB2" box="[929,1068,258,285]" pageId="10" pageNumber="557">M. Ramírez</collectorName>
, 
<collectorName id="F80380D19F69FF8C7400FEBCFB3DFEB2" box="[1080,1213,258,285]" pageId="10" pageNumber="557">P. Goloboff</collectorName>
, 
<collectorName id="F80380D19F69FF8C74F1FEBDFAC1FEB2" box="[1225,1345,258,284]" pageId="10" pageNumber="557">C. Szumik</collectorName>
<emphasis id="678239159F69FF8C7571FEBDFAF2FEB2" box="[1353,1394,258,284]" italics="true" pageId="10" pageNumber="557">leg.</emphasis>
—
</materialsCitation>
<materialsCitation id="E59EEF5A9F69FF8C75ABFEBDFB95FECA" collectingDate="1994-12-23" collectingDateMax="1997-02-25" collectingDateMin="1994-12-23" collectionCode="CDA, LEA" collectorName="L. Acosta &amp; M. Acosta &amp; M. Ramirez &amp; P. Goloboff &amp; C. Szumik &amp; Mesada &amp; C. Grismado" country="Argentina" county="Calilegua National Park" elevation="1698" latitude="-23.625816" location="Calilegua NatiOnal PaRk" longLatPrecision="1" longitude="-64.93485" municipality="Monolito'" pageId="10" pageNumber="557" specimenCode="MACN-AR 45699, MACN-AR 45703, MACN-AR 45701" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="Jujuy" typeStatus="holotype">
<specimenCount id="43F02E8E9F69FF8C75ABFEBDFF2CFEEF" count="1" pageId="10" pageNumber="557" type="female">1 ♀</specimenCount>
<typeStatus id="8A4D5BA59F69FF8C708CFE98FE94FEEF" box="[180,276,295,321]" pageId="10" pageNumber="557" type="paratype">paRatYpe</typeStatus>
(
<specimenCode id="05504D7C9F69FF8C711AFE99FE71FEEF" box="[290,497,294,321]" collectionCode="MACN-AR" pageId="10" pageNumber="557">MACN-AR 45703</specimenCode>
), 
<location id="5029B3DC9F69FF8C723DFE99FC97FEEE" LSID="urn:lsid:plazi:treatment:DD5F54119F6AFF8B70AFFF28FC46FECB:5029B3DC9F69FF8C723DFE99FC97FEEE" box="[517,791,294,320]" country="Argentina" county="Calilegua National Park" latitude="-23.625816" longLatPrecision="1" longitude="-64.93485" municipality="Monolito'" name="Calilegua NatiOnal PaRk" pageId="10" pageNumber="557" stateProvince="Jujuy">Calilegua NatiOnal PaRk</location>
, 
<quantity id="920E48E29F69FF8C7319FE99FCDBFEEE" box="[801,859,294,320]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="557" unit="km" value="8.0">8 km</quantity>
W fROm 
<collectorName id="F80380D19F69FF8C73F9FE99FB98FEEE" box="[961,1048,294,320]" pageId="10" pageNumber="557">Mesada</collectorName>
de las COlmenas, 
<emphasis id="678239159F69FF8C74DEFE98FA88FEEE" box="[1254,1288,295,320]" italics="true" pageId="10" pageNumber="557">ca.</emphasis>
<geoCoordinate id="30C283C09F69FF8C7537FE99FA1CFEEF" box="[1295,1436,294,321]" degrees="23" direction="south" minutes="41.155" orientation="latitude" pageId="10" pageNumber="557" precision="1" value="-23.685917">23°41.155’S</geoCoordinate>
<geoCoordinate id="30C283C09F69FF8C70AFFEF5FEAFFECB" box="[151,303,330,357]" degrees="64" direction="west" minutes="54.032" orientation="longitude" pageId="10" pageNumber="557" precision="1" value="-64.900536">64°54.032’W</geoCoordinate>
, ~ 
<elevation id="DEDB02349F69FF8C7171FEF5FE5DFECA" box="[329,477,330,357]" metricMagnitude="3" metricUnit="m" metricValue="1.66" pageId="10" pageNumber="557" unit="m" value="1660.0">
<quantity id="920E48E29F69FF8C7171FEF5FE20FECB" box="[329,416,330,357]" metricMagnitude="3" metricUnit="m" metricValue="1.66" pageId="10" pageNumber="557" unit="m" value="1660.0">1660 m</quantity>
a.s.l.
</elevation>
, 
<date id="2148C3C79F69FF8C71D3FEF5FD25FECB" box="[491,677,330,357]" pageId="10" pageNumber="557" value="1995-09">
<collectingDate id="310C3A2F9F69FF8C71D3FEF5FD25FECB" box="[491,677,330,357]" pageId="10" pageNumber="557" value="1995-09">September 1995</collectingDate>
</date>
, 
<collectorName id="F80380D19F69FF8C7289FEF4FCB5FECA" box="[689,821,330,357]" pageId="10" pageNumber="557">P. Goloboff</collectorName>
, 
<collectorName id="F80380D19F69FF8C7379FEF4FC48FECA" box="[833,968,330,357]" pageId="10" pageNumber="557">M. Ramírez</collectorName>
<emphasis id="678239159F69FF8C73F7FEF5FC79FECA" box="[975,1017,330,356]" italics="true" pageId="10" pageNumber="557">leg.</emphasis>
—
</materialsCitation>
<materialsCitation id="E59EEF5A9F69FF8C742DFEF5FAD2FE26" collectingDate="1994-12-23" collectingDateMax="1997-02-25" collectingDateMin="1994-12-23" collectionCode="CDA, LEA" collectorName="L. Acosta &amp; M. Acosta &amp; M. Ramirez &amp; P. Goloboff &amp; C. Szumik &amp; Mesada &amp; C. Grismado" country="Argentina" county="Calilegua National Park" elevation="1698" latitude="-23.625816" location="Calilegua NatiOnal PaRk" longLatPrecision="1" longitude="-64.93485" municipality="Monolito'" pageId="10" pageNumber="557" specimenCode="MACN-AR 45699, MACN-AR 45703, MACN-AR 45701" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="Jujuy" typeStatus="holotype">
<specimenCount id="43F02E8E9F69FF8C742DFEF5FBBFFECB" box="[1045,1087,330,357]" count="2" pageId="10" pageNumber="557" type="female">2 ♀</specimenCount>
<typeStatus id="8A4D5BA59F69FF8C747FFEF4FB32FECB" box="[1095,1202,331,357]" pageId="10" pageNumber="557" type="paratype">paRatYpes</typeStatus>
(
<specimenCode id="05504D7C9F69FF8C74F9FEF5FA10FECB" box="[1217,1424,330,357]" collectionCode="MACN-AR" pageId="10" pageNumber="557">MACN-AR 45701</specimenCode>
), 
<location id="5029B3DC9F69FF8C70AFFED1FE27FE26" LSID="urn:lsid:plazi:treatment:DD5F54119F6AFF8B70AFFF28FC46FECB:5029B3DC9F69FF8C70AFFED1FE27FE26" box="[151,423,366,392]" country="Argentina" county="Calilegua National Park" latitude="-23.625816" longLatPrecision="1" longitude="-64.93485" municipality="Monolito'" name="Calilegua National Park" pageId="10" pageNumber="557" stateProvince="Jujuy">Calilegua National Park</location>
, 
<emphasis id="678239159F69FF8C718AFED0FE54FE26" box="[434,468,367,392]" italics="true" pageId="10" pageNumber="557">ca.</emphasis>
<geoCoordinate id="30C283C09F69FF8C71E3FED1FDE7FE27" box="[475,615,366,393]" degrees="23" direction="south" minutes="37.549" orientation="latitude" pageId="10" pageNumber="557" precision="1" value="-23.625816">23°37.549’S</geoCoordinate>
<geoCoordinate id="30C283C09F69FF8C7257FED1FC87FE27" box="[623,775,366,393]" degrees="64" direction="west" minutes="56.091" orientation="longitude" pageId="10" pageNumber="557" precision="1" value="-64.93485">64°56.091’W</geoCoordinate>
, 
<elevation id="DEDB02349F69FF8C732AFED1FC24FE26" box="[786,932,366,393]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="10" pageNumber="557" unit="m" value="2100.0">
<quantity id="920E48E29F69FF8C732AFED1FCE7FE27" box="[786,871,366,393]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="10" pageNumber="557" unit="m" value="2100.0">2100 m</quantity>
a.s.l.
</elevation>
, 
<date id="2148C3C79F69FF8C7388FED1FB08FE26" box="[944,1160,366,392]" pageId="10" pageNumber="557" value="1994-12-23">
<collectingDate id="310C3A2F9F69FF8C7388FED1FB08FE26" box="[944,1160,366,392]" pageId="10" pageNumber="557" value="1994-12-23">23 December 1994</collectingDate>
</date>
, 
<collectorName id="F80380D19F69FF8C74ABFED1FAA2FE26" box="[1171,1314,366,392]" pageId="10" pageNumber="557">C. Grismado</collectorName>
<emphasis id="678239159F69FF8C7511FED1FAD2FE26" box="[1321,1362,366,392]" italics="true" pageId="10" pageNumber="557">leg.</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="5549E5079F69FF8C70FFFE2DFE8EFE7E" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="557">
<materialsCitation id="E59EEF5A9F69FF8C70FFFE2DFE8AFE7E" country="Argentina" elevation="1705" latitude="-23.681116" location="Calilegua National Park" longLatPrecision="1" longitude="-64.90147" municipality="Monolito'" pageId="10" pageNumber="557" specimenCount="1" stateProvince="Jujuy" typeStatus="holotype">
<emphasis id="678239159F69FF8C70FFFE2DFEE4FE02" bold="true" box="[199,356,402,428]" pageId="10" pageNumber="557">
<typeStatus id="8A4D5BA59F69FF8C70FFFE2DFE82FE02" box="[199,258,402,428]" pageId="10" pageNumber="557">Type</typeStatus>
locality.
</emphasis>
<location id="5029B3DC9F69FF8C7152FE2DFDF7FE02" LSID="urn:lsid:plazi:treatment:DD5F54119F6AFF8B70AFFF28FC46FECB:5029B3DC9F69FF8C7152FE2DFDF7FE02" box="[362,631,402,428]" country="Argentina" latitude="-23.681116" longLatPrecision="1" longitude="-64.90147" municipality="Monolito'" name="Calilegua National Park" pageId="10" pageNumber="557" stateProvince="Jujuy">Calilegua National Park</location>
, ‘ 
<collectingMunicipality id="B52D7F7D9F69FF8C72BDFE2DFD7AFE02" box="[645,762,402,428]" pageId="10" pageNumber="557">Monolito’</collectingMunicipality>
(
<geoCoordinate id="30C283C09F69FF8C733FFE2DFC15FE02" box="[775,917,402,428]" degrees="23" direction="south" minutes="40.867" orientation="latitude" pageId="10" pageNumber="557" precision="1" value="-23.681116">23°40.867’S</geoCoordinate>
<geoCoordinate id="30C283C09F69FF8C73A3FE2DFBB3FE03" box="[923,1075,402,429]" degrees="64" direction="west" minutes="54.088" orientation="longitude" pageId="10" pageNumber="557" precision="1" value="-64.90147">64°54.088’W</geoCoordinate>
, ~ 
<elevation id="DEDB02349F69FF8C7473FE2DFB5AFE02" box="[1099,1242,402,429]" metricMagnitude="3" metricUnit="m" metricValue="1.705" pageId="10" pageNumber="557" unit="m" value="1705.0">
<quantity id="920E48E29F69FF8C7473FE2DFB20FE03" box="[1099,1184,402,429]" metricMagnitude="3" metricUnit="m" metricValue="1.705" pageId="10" pageNumber="557" unit="m" value="1705.0">1705 m</quantity>
a.s.l.
</elevation>
), 
<collectingRegion id="97322BE59F69FF8C74D7FE2DFA18FE02" box="[1263,1432,402,428]" country="Argentina" name="Jujuy" pageId="10" pageNumber="557">Jujuy Province</collectingRegion>
, 
<collectingCountry id="2DE1A5979F69FF8C70AFFE09FE8AFE7E" box="[151,266,438,464]" name="Argentina" pageId="10" pageNumber="557">Argentina</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="5549E5079F69FF8C70FFFE65FB63FDCE" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="557">
<emphasis id="678239159F69FF8C70FFFE65FE37FE5A" bold="true" box="[199,439,474,500]" pageId="10" pageNumber="557">Additional material.</emphasis>
<materialsCitation id="E59EEF5A9F69FF8C7181FE65FB60FDCE" collectingDate="1997-03-21" collectionCode="LEA" collectorName="Guardaparques Nicolossi &amp; Zalazar &amp; Remarks &amp; This" country="Argentina" elevation="2400" latitude="-23.624317" location="Alto Calilegua" longLatPrecision="1" longitude="-64.9227" municipality="San" pageId="10" pageNumber="557" specimenCount="1" specimenCount-female="1" stateProvince="Jujuy">
<collectingCountry id="2DE1A5979F69FF8C7181FE65FDDFFE5A" box="[441,607,474,500]" name="Argentina" pageId="10" pageNumber="557">ARGENTINA</collectingCountry>
, 
<emphasis id="678239159F69FF8C725FFE64FC8BFE5A" box="[615,779,475,500]" italics="true" pageId="10" pageNumber="557">
<collectingRegion id="97322BE59F69FF8C725FFE64FC8BFE5A" box="[615,779,475,500]" country="Argentina" name="Jujuy" pageId="10" pageNumber="557">Jujuy Province</collectingRegion>
</emphasis>
. 
<specimenCount id="43F02E8E9F69FF8C732EFE65FCBDFE5B" box="[790,829,474,501]" count="1" pageId="10" pageNumber="557" type="female">1 ♀</specimenCount>
(
<collectionCode id="33E77DC29F69FF8C7370FE65FCFFFE5A" box="[840,895,474,500]" pageId="10" pageNumber="557">LEA</collectionCode>
000.148), DuRaznillO,between 
<collectingMunicipality id="B52D7F7D9F69FF8C74EDFE65FB7FFE5A" box="[1237,1279,474,500]" pageId="10" pageNumber="557">San</collectingMunicipality>
FRanciscO and 
<location id="5029B3DC9F69FF8C70AFFE41FEBAFDB6" LSID="urn:lsid:plazi:treatment:DD5F54119F6AFF8B70AFFF28FC46FECB:5029B3DC9F69FF8C70AFFE41FEBAFDB6" box="[151,314,510,536]" country="Argentina" latitude="-23.624317" longLatPrecision="1" longitude="-64.9227" municipality="San" name="Alto Calilegua" pageId="10" pageNumber="557" stateProvince="Jujuy">Alto Calilegua</location>
(Depto. Valle Grande), 
<emphasis id="678239159F69FF8C727CFE40FDE5FDB6" box="[580,613,511,536]" italics="true" pageId="10" pageNumber="557">ca.</emphasis>
<geoCoordinate id="30C283C09F69FF8C7253FE41FD78FDB7" box="[619,760,510,537]" degrees="23" direction="south" minutes="37.459" orientation="latitude" pageId="10" pageNumber="557" precision="1" value="-23.624317">23°37.459’S</geoCoordinate>
<geoCoordinate id="30C283C09F69FF8C72C6FE41FC16FDB7" box="[766,918,510,537]" degrees="64" direction="west" minutes="55.362" orientation="longitude" pageId="10" pageNumber="557" precision="1" value="-64.9227">64°55.362’W</geoCoordinate>
, 
<elevation id="DEDB02349F69FF8C7399FE41FBB1FDB6" box="[929,1073,510,537]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="557" unit="m" value="2400.0">
<quantity id="920E48E29F69FF8C7399FE41FC75FDB7" box="[929,1013,510,537]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="557" unit="m" value="2400.0">2400 m</quantity>
a.s.l.
</elevation>
, 
<date id="2148C3C79F69FF8C7404FE41FB69FDB6" box="[1084,1257,510,536]" pageId="10" pageNumber="557" value="1997-03-21">
<collectingDate id="310C3A2F9F69FF8C7404FE41FB69FDB6" box="[1084,1257,510,536]" pageId="10" pageNumber="557" value="1997-03-21">21 March 1997</collectingDate>
</date>
, 
<collectorName id="F80380D19F69FF8C74CBFE41FE81FD92" pageId="10" pageNumber="557">Guardaparques Nicolossi</collectorName>
&amp; 
<collectorName id="F80380D19F69FF8C711DFD9DFEF9FD92" box="[293,377,546,572]" pageId="10" pageNumber="557">Zalazar</collectorName>
<emphasis id="678239159F69FF8C71B8FD9DFE2AFD92" box="[384,426,546,572]" italics="true" pageId="10" pageNumber="557">leg.</emphasis>
<collectorName id="F80380D19F69FF8C7189FD9DFD99FD92" box="[433,537,546,572]" pageId="10" pageNumber="557">Remarks</collectorName>
: 
<collectorName id="F80380D19F69FF8C721BFD9DFDD5FD92" box="[547,597,546,572]" pageId="10" pageNumber="557">This</collectorName>
female largely oversizes all remaining specimens (DS length 
<quantity id="920E48E29F69FF8C7534FD9DFAE4FD93" box="[1292,1380,546,573]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="10" pageNumber="557" unit="mm" value="7.6">7.6 mm</quantity>
, 
<figureCitation id="CDCDF9829F69FF8C7548FD9DFF27FDCE" captionStart="FIGURE 2" captionStartId="2.[151,250,1241,1265]" captionTargetBox="[155,1431,509,1214]" captionTargetId="figure-169@2.[151,1435,505,1218]" captionTargetPageId="2" captionText="FIGURE 2. Comparative morphometrics of Acrographinotus tariquiae sp. nov. and A. calilegua sp. nov., displayed as box-plots with data points (internal lines, dashed: mean, solid: median). Left: DS length (in mm) for males and females; the single outlying female from El Duraznillo (star), tentatively assigned to A. calilegua but not used in calculations, is shown for reference. Right: FeIV/DS ratio, males. In box-plots of male A. tariquiae, small aRROws fROm the left identifY the twO examined β-males." figureDoi="http://doi.org/10.5281/zenodo.12783062" httpUri="https://zenodo.org/record/12783062/files/figure.png" pageId="10" pageNumber="557">Fig. 2</figureCitation>
), so it is assigned to 
<taxonomicName id="92F69E849F69FF8C71A9FDF8FD98FDCE" authorityName="Acosta" authorityYear="2024" box="[401,536,582,609]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="557" phylum="Arthropoda" rank="species" species="calilegua" status="sp. nov.">
<emphasis id="678239159F69FF8C71A9FDF8FD98FDCE" box="[401,536,582,609]" italics="true" pageId="10" pageNumber="557">A. calilegua</emphasis>
</taxonomicName>
<emphasis id="678239159F69FF8C7227FDF9FDF9FDCE" bold="true" box="[543,633,582,608]" pageId="10" pageNumber="557">
<taxonomicNameLabel id="7CB1846E9F69FF8C7227FDF9FDF9FDCE" box="[543,633,582,608]" pageId="10" pageNumber="557" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
only tentatively, until additional samples are available
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="1DECB68C9F69FF8C70FFFDD5FD13FD06" pageId="10" pageNumber="557" type="etymology">
<paragraph id="5549E5079F69FF8C70FFFDD5FD13FD06" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="557">
<emphasis id="678239159F69FF8C70FFFDD5FECEFD2A" bold="true" box="[199,334,618,644]" pageId="10" pageNumber="557">Etymology.</emphasis>
The specific name is a noun in apposition, referring to the Calilegua National Park (
<collectingRegion id="97322BE59F69FF8C74D5FDD5FA18FD2A" box="[1261,1432,618,644]" country="Argentina" name="Jujuy" pageId="10" pageNumber="557">Jujuy Province</collectingRegion>
, 
<collectingCountry id="2DE1A5979F69FF8C70AFFD31FE8BFD06" box="[151,267,654,680]" name="Argentina" pageId="10" pageNumber="557">Argentina</collectingCountry>
), where most 
<typeStatus id="8A4D5BA59F69FF8C719FFD30FE62FD07" box="[423,482,655,681]" pageId="10" pageNumber="557">types</typeStatus>
were collected.
</paragraph>
</subSubSection>
<subSubSection id="1DECB68C9F69FF8D70FFFD0DFCE2F9E5" lastPageId="11" lastPageNumber="558" pageId="10" pageNumber="557" type="description">
<paragraph id="5549E5079F69FF8C70FFFD0DFB4AFCBA" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="557">
<emphasis id="678239159F69FF8C70FFFD0DFED9FD62" bold="true" box="[199,345,690,716]" pageId="10" pageNumber="557">Description.</emphasis>
<emphasis id="678239159F69FF8C715AFD0CFD2DFD62" box="[354,685,690,716]" italics="true" pageId="10" pageNumber="557">Measurements and meristics.</emphasis>
DS length: males 
<quantity id="920E48E29F69FF8C73BCFD0DFB8DFD63" box="[900,1037,690,717]" metricMagnitude="-3" metricUnit="m" metricValue="5.95" metricValueMax="6.2" metricValueMin="5.7" pageId="10" pageNumber="557" unit="mm" value="5.95" valueMax="6.2" valueMin="5.7">5.7–6.2 mm</quantity>
(n=2), females 
<quantity id="920E48E29F69FF8C74FCFD0DFACEFD63" box="[1220,1358,690,717]" metricMagnitude="-3" metricUnit="m" metricValue="6.45" metricValueMax="6.7" metricValueMin="6.2" pageId="10" pageNumber="557" unit="mm" value="6.45" valueMax="6.7" valueMin="6.2">6.2–6.7 mm</quantity>
(mean 
<quantity id="920E48E29F69FF8C70AFFD69FF70FD5F" box="[151,240,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="10" pageNumber="557" unit="mm" value="6.4">6.4 mm</quantity>
, n=6). FeIV/DS ratio (males): 0.84–0.89 (n=2). Detailed measurements of the 
<typeStatus id="8A4D5BA59F69FF8C745DFD69FB47FD5E" box="[1125,1223,726,752]" pageId="10" pageNumber="557" type="holotype">holotype</typeStatus>
male and a female 
<typeStatus id="8A4D5BA59F69FF8C70AFFD44FF77FCBB" box="[151,247,763,789]" pageId="10" pageNumber="557" type="paratype">paratype</typeStatus>
in 
<tableCitation id="1874D0BC9F69FF8C7123FD45FEEFFCBA" box="[283,367,762,788]" captionStart="TABLE 1" captionStartId="5.[151,239,831,855]" captionTargetId="graphics-230@5.[151,1436,937,1819]" captionTargetPageId="5" captionText="TABLE 1. Measurements (mm) of male holotype and a female paratype of Acrographinotus tariquiae sp. nov. and Acrographinotus calilegua sp. nov.." pageId="10" pageNumber="557">Table 1</tableCitation>
. Tarsal formula: males 6:7–8:7:7 (
<typeStatus id="8A4D5BA59F69FF8C72C8FD45FCD4FCBA" box="[752,852,762,788]" pageId="10" pageNumber="557" type="holotype">holotype</typeStatus>
with 6:7:7:7), females 6:7–9:7:7.
</paragraph>
<paragraph id="5549E5079F69FF8C70FFFCA1FCCDFC2E" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="557">
<emphasis id="678239159F69FF8C70FFFCA1FE7DFC96" box="[199,509,798,824]" italics="true" pageId="10" pageNumber="557">Coloration in ethanol 70%.</emphasis>
General color light cinnamon, with chelicera, pedipalps, legs I–III and metatarsus IV lighter; faint reticulated pigment on carapace and appendages; leg IV (coxa to tibia), free tergites (including apophysis) and venter of male darker (towards a reddish hue).
</paragraph>
<paragraph id="5549E5079F69FF8C70FFFC35FA08FB9A" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="557">
<emphasis id="678239159F69FF8C70FFFC35FE98FC0A" bold="true" box="[199,280,906,932]" pageId="10" pageNumber="557">Males:</emphasis>
<emphasis id="678239159F69FF8C7126FC34FE00FC0A" box="[286,384,907,932]" italics="true" pageId="10" pageNumber="557">Dorsum.</emphasis>
DS tYpe λ (lambda, cOda shORt, caRapace with subpaRallel sides; 
<figureCitation id="CDCDF9829F69FF8C747CFC35FB1BFC0B" box="[1092,1179,906,933]" captionStart="FIGURE 5" captionStartId="9.[151,250,1745,1769]" captionTargetBox="[155,1432,374,1717]" captionTargetId="figure-45@9.[151,1435,370,1721]" captionTargetPageId="9" captionText="FIGURE 5. Acrographinotus calilegua sp. nov., HOlOtYpe ♂ (CDA 000.912). A. DORsal scutum, fRee teRgites (alsO shOwing VAP process from above), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, ventral anal plate, right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. Scale lines: 2 mm (A–E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783069" httpUri="https://zenodo.org/record/12783069/files/figure.png" pageId="10" pageNumber="557">Fig. 5A</figureCitation>
), unaRmed, gRanulatiOn very unconspicuous. Anterior margin of carapace smooth, frontal hump shallow. Ocular mound low, with a blunt small median apophysis (
<figureCitation id="CDCDF9829F69FF8C7186FC6DFD97FC43" box="[446,535,978,1005]" captionStart="FIGURE 5" captionStartId="9.[151,250,1745,1769]" captionTargetBox="[155,1432,374,1717]" captionTargetId="figure-45@9.[151,1435,370,1721]" captionTargetPageId="9" captionText="FIGURE 5. Acrographinotus calilegua sp. nov., HOlOtYpe ♂ (CDA 000.912). A. DORsal scutum, fRee teRgites (alsO shOwing VAP process from above), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, ventral anal plate, right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. Scale lines: 2 mm (A–E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783069" httpUri="https://zenodo.org/record/12783069/files/figure.png" pageId="10" pageNumber="557">Fig. 5F</figureCitation>
). DS area I divided; areas I–IV entire, almost smooth, except for sparse, tiny granules in a row. Area V, lateral areas and free tergites with a row of small granules. FT-III with an inconspicuous row of scarce granules and a strong median apophysis, curved (
<typeStatus id="8A4D5BA59F69FF8C7365FBA5FC42FB9A" box="[861,962,1050,1076]" pageId="10" pageNumber="557" type="holotype">holotype</typeStatus>
) or horizontal (
<typeStatus id="8A4D5BA59F69FF8C7457FBA5FB53FB9A" box="[1135,1235,1050,1076]" pageId="10" pageNumber="557" type="paratype">paratype</typeStatus>
) in lateral view.
</paragraph>
<paragraph id="5549E5079F69FF8C70FFFB80FD54FB0E" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="557">
<emphasis id="678239159F69FF8C70FFFB80FE93FBF6" box="[199,275,1087,1112]" italics="true" pageId="10" pageNumber="557">Venter.</emphasis>
Posterior margin of stigmatic segment like a flat bell curve. Free sternites with a row of minute granules. VAP bears a small ‘shelf-like’ projection; typical apophyses are lacking but they are replaced by a pair of small ventral, subapically shifted tubercles (
<figureCitation id="CDCDF9829F69FF8C7279FB39FD47FB0E" box="[577,711,1158,1185]" captionStart="FIGURE 5" captionStartId="9.[151,250,1745,1769]" captionTargetBox="[155,1432,374,1717]" captionTargetId="figure-45@9.[151,1435,370,1721]" captionTargetPageId="9" captionText="FIGURE 5. Acrographinotus calilegua sp. nov., HOlOtYpe ♂ (CDA 000.912). A. DORsal scutum, fRee teRgites (alsO shOwing VAP process from above), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, ventral anal plate, right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. Scale lines: 2 mm (A–E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783069" httpUri="https://zenodo.org/record/12783069/files/figure.png" pageId="10" pageNumber="557">Figs. 5B, D</figureCitation>
).
</paragraph>
<paragraph id="5549E5079F69FF8C70FFFB15FD23FB6A" blockId="10.[151,1437,150,1977]" box="[199,675,1194,1221]" pageId="10" pageNumber="557">
<emphasis id="678239159F69FF8C70FFFB15FECAFB6A" box="[199,330,1194,1220]" italics="true" pageId="10" pageNumber="557">Chelicerae.</emphasis>
Subtly rugous, bulla unarmed.
</paragraph>
<paragraph id="5549E5079F69FF8C70FFFB71FA83FAA2" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="557">
<emphasis id="678239159F69FF8C70FFFB71FEBFFB46" box="[199,319,1230,1256]" italics="true" pageId="10" pageNumber="557">Pedipalps.</emphasis>
Feeble, tegument finely rugous; blunt ventral setigerous tubercles on trochanter and femur; no medial subapical spine on femur. Pedipalp spination: LAT: Ti i_[Ii•], Ta iiii or •iii – MED: Ti Iiii or IiIi, Ta IIi.
</paragraph>
<paragraph id="5549E5079F69FF8C70FFFAA8FCAEFA9F" blockId="10.[151,1437,150,1977]" box="[199,814,1302,1329]" pageId="10" pageNumber="557">
<emphasis id="678239159F69FF8C70FFFAA8FEBEFA9E" box="[199,318,1303,1329]" italics="true" pageId="10" pageNumber="557">Legs I–III.</emphasis>
Unarmed, finely granulous on all segments.
</paragraph>
<paragraph id="5549E5079F69FF8C70FFFA84FD8FFA33" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="557">
<emphasis id="678239159F69FF8C70FFFA84FE9AFAFA" box="[199,282,1339,1364]" italics="true" pageId="10" pageNumber="557">Leg IV.</emphasis>
Coxa IV smooth, with a diagonal prolateral apophysis, short and blunt, and a minuscule retrolateral one, perceptible from ventral and mostly hidden between trochanter and stigmatic segment. Coxa-trochanter joint oblique in ventral view (
<figureCitation id="CDCDF9829F69FF8C7191FA3DFD82FA33" box="[425,514,1410,1437]" captionStart="FIGURE 5" captionStartId="9.[151,250,1745,1769]" captionTargetBox="[155,1432,374,1717]" captionTargetId="figure-45@9.[151,1435,370,1721]" captionTargetPageId="9" captionText="FIGURE 5. Acrographinotus calilegua sp. nov., HOlOtYpe ♂ (CDA 000.912). A. DORsal scutum, fRee teRgites (alsO shOwing VAP process from above), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, ventral anal plate, right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. Scale lines: 2 mm (A–E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783069" httpUri="https://zenodo.org/record/12783069/files/figure.png" pageId="10" pageNumber="557">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="5549E5079F69FF8C70FFFA19FB59FA4A" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="557">Trochanter IV wide, asymmetric-subtrapezoidal; armature modest: a blunt prodorsal tubercle near the apical border, opposite to the coxal apophysis, and a small conic tubercle on the retrodorsal apical edge.</paragraph>
<paragraph id="5549E5079F69FF8C70FFFA50FC6AF912" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="557">
FemuR IV shORteR than DS and gentlY aRched, it aRticulates diagOnallY sidewaYs (diveRging bY ⁓45° fROm bOdY axis; 
<figureCitation id="CDCDF9829F69FF8C70EDF9ADFEDBF983" box="[213,347,1554,1581]" captionStart="FIGURE 5" captionStartId="9.[151,250,1745,1769]" captionTargetBox="[155,1432,374,1717]" captionTargetId="figure-45@9.[151,1435,370,1721]" captionTargetPageId="9" captionText="FIGURE 5. Acrographinotus calilegua sp. nov., HOlOtYpe ♂ (CDA 000.912). A. DORsal scutum, fRee teRgites (alsO shOwing VAP process from above), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, ventral anal plate, right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. Scale lines: 2 mm (A–E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783069" httpUri="https://zenodo.org/record/12783069/files/figure.png" pageId="10" pageNumber="557">Figs. 5A–B</figureCitation>
); it is simple, with longitudinal rows of granules or small tubercles and subdistal narrowing not evident; dorsal and retrodorsal rows of blunt tubercles or low apophyses, the retrodorsal ends in a small conic subapical apophysis, pointing upwards (
<figureCitation id="CDCDF9829F69FF8C7261F9E5FD5CF9DA" box="[601,732,1626,1653]" captionStart="FIGURE 5" captionStartId="9.[151,250,1745,1769]" captionTargetBox="[155,1432,374,1717]" captionTargetId="figure-45@9.[151,1435,370,1721]" captionTargetPageId="9" captionText="FIGURE 5. Acrographinotus calilegua sp. nov., HOlOtYpe ♂ (CDA 000.912). A. DORsal scutum, fRee teRgites (alsO shOwing VAP process from above), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, ventral anal plate, right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. Scale lines: 2 mm (A–E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783069" httpUri="https://zenodo.org/record/12783069/files/figure.png" pageId="10" pageNumber="557">Figs. 5A, E</figureCitation>
); pro- and retrolateral rows of small, sparse granules, the latter ends in a conic apical aphophysis pointing medially (
<figureCitation id="CDCDF9829F69FF8C733BF9C1FCDFF937" box="[771,863,1662,1689]" captionStart="FIGURE 5" captionStartId="9.[151,250,1745,1769]" captionTargetBox="[155,1432,374,1717]" captionTargetId="figure-45@9.[151,1435,370,1721]" captionTargetPageId="9" captionText="FIGURE 5. Acrographinotus calilegua sp. nov., HOlOtYpe ♂ (CDA 000.912). A. DORsal scutum, fRee teRgites (alsO shOwing VAP process from above), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, ventral anal plate, right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. Scale lines: 2 mm (A–E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783069" httpUri="https://zenodo.org/record/12783069/files/figure.png" pageId="10" pageNumber="557">Fig. 5B</figureCitation>
); pro- and retroventral rows incomplete, granules larger and more acute distally, they end in a small unciform apophysis each.
</paragraph>
<paragraph id="5549E5079F69FF8C70FFF979FC6EF94E" blockId="10.[151,1437,150,1977]" box="[199,1006,1734,1761]" pageId="10" pageNumber="557">Patella IV gently granulous, with 1–2 small acute retroventral tubercles.</paragraph>
<paragraph id="5549E5079F69FF8C70FFF955FE94F886" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="557">Tibia IV with rows of conic granules, progressively larger on the retroventral row to end as small acute apophyses.</paragraph>
<paragraph id="5549E5079F69FF8C70FFF88CFAAFF817" blockId="10.[151,1437,150,1977]" pageId="10" pageNumber="557">
<emphasis id="678239159F69FF8C70FFF88CFE85F8E2" box="[199,261,1843,1868]" italics="true" pageId="10" pageNumber="557">Penis</emphasis>
(
<figureCitation id="CDCDF9829F69FF8C712FF88DFE1FF8E2" box="[279,415,1842,1868]" captionStart="FIGURE 6" captionStartId="11.[151,250,1298,1322]" captionTargetBox="[290,1297,185,1271]" captionTargetId="figure-15@11.[286,1301,181,1275]" captionTargetPageId="11" captionText="FIGURE 6. Acrographinotus calilegua sp. nov., distal end Of penis. A–B. PaRatYpe ♂ (MACN-AR 45699): A: dORsal view; B: lateRal view (membRanOus cRest is bent); C. HOlOtYpe ♂ (CDA 000.912), detail Of stYlus and ventRal pROcess. Scale lines: 0.1 mm (A, B), 0.05 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.12783071" httpUri="https://zenodo.org/record/12783071/files/figure.png" pageId="10" pageNumber="557">Figs. 6A–C</figureCitation>
). Distal end higher than wide, hourglass shaped in dorsal view due to VP sides markedly concave; front border of VP incurvate. Basal macrosetae (A1–A3 + B) in a transversal row, shifted more basally than the truncus-glans joint. Stylus tip abruptly inflected; VPS has the ‘ibis head with crest’ look typical for the genus, curvature of ‘beak-like’ process mild, while ‘veins’ in the membranous expansion are extremely tenuous.
</paragraph>
<caption id="0189B58F9F68FF8D70AFFAADFE1BFADD" ID-DOI="http://doi.org/10.5281/zenodo.12783071" ID-Zenodo-Dep="12783071" httpUri="https://zenodo.org/record/12783071/files/figure.png" pageId="11" pageNumber="558" startId="11.[151,250,1298,1322]" targetBox="[290,1297,185,1271]" targetPageId="11" targetType="figure">
<paragraph id="5549E5079F68FF8D70AFFAADFE1BFADD" blockId="11.[151,1437,1298,1395]" pageId="11" pageNumber="558">
<emphasis id="678239159F68FF8D70AFFAADFE93FA84" bold="true" box="[151,275,1298,1322]" pageId="11" pageNumber="558">FIGURE 6.</emphasis>
<taxonomicName id="92F69E849F68FF8D7123FAACFDAAFA85" authorityName="Acosta" authorityYear="2024" box="[283,554,1299,1323]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="558" phylum="Arthropoda" rank="species" species="calilegua" status="sp. nov.">
<emphasis id="678239159F68FF8D7123FAACFDAAFA85" box="[283,554,1299,1323]" italics="true" pageId="11" pageNumber="558">Acrographinotus calilegua</emphasis>
</taxonomicName>
<emphasis id="678239159F68FF8D7209FAACFD02FA85" bold="true" box="[561,642,1299,1323]" pageId="11" pageNumber="558">
<taxonomicNameLabel id="7CB1846E9F68FF8D7209FAACFD02FA85" box="[561,642,1299,1323]" pageId="11" pageNumber="558" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, distal end Of penis. A–B. PaRatYpe ♂ (MACN-AR 45699): A: dORsal view; B: lateRal view (membRanOus cRest is bent); C. HOlOtYpe ♂ (CDA 000.912), detail Of stYlus and ventRal pROcess. Scale lines: 0.1 mm (A, B), 0.05 mm (C).
</paragraph>
</caption>
<paragraph id="5549E5079F68FF8D70FFFA1EFCE2F9E5" blockId="11.[151,1437,1441,2007]" pageId="11" pageNumber="558">
<emphasis id="678239159F68FF8D70FFFA1EFEB2FA15" bold="true" box="[199,306,1441,1467]" pageId="11" pageNumber="558">Females:</emphasis>
DS 
<typeStatus id="8A4D5BA59F68FF8D715CFA1EFE14FA15" box="[356,404,1441,1467]" pageId="11" pageNumber="558">tYpe</typeStatus>
α-K (alpha-keYhOle, cOda mOdeRatelY lOng with diveRging sides), unaRmed, with gRanulatiOn subtly more conspicuous than males. Ocular mound slightly higher than males. FT-III with a row of acute granules and a small median apophysis. Margin of VAP with a pair of flat tubercles, as vestiges of the ‘shelf’ of males. Coxa IV: small acute prolateral apophysis. FeIV slender, slightly curved at its base, then sub-straight, with rows of granules; retroapical apophysis very small, pointing backwards.
</paragraph>
</subSubSection>
<subSubSection id="1DECB68C9F68FF8A70FFF9EAFD6CF8E5" lastPageId="12" lastPageNumber="559" pageId="11" pageNumber="558" type="diagnosis">
<paragraph id="5549E5079F68FF8D70FFF9EAFE19F879" blockId="11.[151,1437,1441,2007]" pageId="11" pageNumber="558">
<emphasis id="678239159F68FF8D70FFF9EAFD9CF9C1" bold="true" box="[199,540,1621,1647]" pageId="11" pageNumber="558">Comparisons and diagnosis.</emphasis>
This species is the smallest, less robust and with the simplest morphology in the genus. FeIV of male is slightly arched and articulates decidedly sideways (
<figureCitation id="CDCDF9829F68FF8D7435F9C6FBE7F93A" box="[1037,1127,1657,1684]" captionStart="FIGURE 5" captionStartId="9.[151,250,1745,1769]" captionTargetBox="[155,1432,374,1717]" captionTargetId="figure-45@9.[151,1435,370,1721]" captionTargetPageId="9" captionText="FIGURE 5. Acrographinotus calilegua sp. nov., HOlOtYpe ♂ (CDA 000.912). A. DORsal scutum, fRee teRgites (alsO shOwing VAP process from above), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, ventral anal plate, right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. Scale lines: 2 mm (A–E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783069" httpUri="https://zenodo.org/record/12783069/files/figure.png" pageId="11" pageNumber="558">Fig. 5A</figureCitation>
); it is shorter than DS (
<figureCitation id="CDCDF9829F68FF8D7557F9C6FF27F919" captionStart="FIGURE 2" captionStartId="2.[151,250,1241,1265]" captionTargetBox="[155,1431,509,1214]" captionTargetId="figure-169@2.[151,1435,505,1218]" captionTargetPageId="2" captionText="FIGURE 2. Comparative morphometrics of Acrographinotus tariquiae sp. nov. and A. calilegua sp. nov., displayed as box-plots with data points (internal lines, dashed: mean, solid: median). Left: DS length (in mm) for males and females; the single outlying female from El Duraznillo (star), tentatively assigned to A. calilegua but not used in calculations, is shown for reference. Right: FeIV/DS ratio, males. In box-plots of male A. tariquiae, small aRROws fROm the left identifY the twO examined β-males." figureDoi="http://doi.org/10.5281/zenodo.12783062" httpUri="https://zenodo.org/record/12783062/files/figure.png" pageId="11" pageNumber="558">Fig. 2</figureCitation>
), while the ‘subapical narrowing’, typical for many species in the genus (like 
<emphasis id="678239159F68FF8D742FF921FBBFF919" box="[1047,1087,1694,1719]" italics="true" pageId="11" pageNumber="558">e.g.</emphasis>
, 
<taxonomicName id="92F69E849F68FF8D7476F922FB4EF919" authorityName="Acosta" authorityYear="2001" box="[1102,1230,1693,1719]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="558" phylum="Arthropoda" rank="species" species="niawpaq">
<emphasis id="678239159F68FF8D7476F922FB4EF919" box="[1102,1230,1693,1719]" italics="true" pageId="11" pageNumber="558">A. niawpaq</emphasis>
</taxonomicName>
or 
<taxonomicName id="92F69E849F68FF8D74CCF922FAF4F919" authorityName="Acosta" authorityYear="2024" box="[1268,1396,1693,1719]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="558" phylum="Arthropoda" rank="species" species="tariquiae" status="sp. nov.">
<emphasis id="678239159F68FF8D74CCF922FAF4F919" box="[1268,1396,1693,1719]" italics="true" pageId="11" pageNumber="558">A tariquiae</emphasis>
</taxonomicName>
<emphasis id="678239159F68FF8D7543F922FF48F975" bold="true" pageId="11" pageNumber="558">
<taxonomicNameLabel id="7CB1846E9F68FF8D7543F922FF48F975" pageId="11" pageNumber="558" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) is hardly perceptible. In fact, most features present in other congeners (namely those species from the couple 5 onwards in the key above), including apophyses and rows of tubercles of FeIV, can be identified though much more attenuated. Like in 
<taxonomicName id="92F69E849F68FF8D7185F8B6FDBFF88D" authorityName="Acosta" authorityYear="2001" box="[445,575,1801,1827]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="558" phylum="Arthropoda" rank="species" species="niawpaq">
<emphasis id="678239159F68FF8D7185F8B6FDBFF88D" box="[445,575,1801,1827]" italics="true" pageId="11" pageNumber="558">A. niawpaq</emphasis>
</taxonomicName>
and 
<taxonomicName id="92F69E849F68FF8D7244F8B6FC86F88D" authorityName="Acosta" authorityYear="2024" box="[636,774,1801,1827]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="558" phylum="Arthropoda" rank="species" species="tariquiae" status="sp. nov.">
<emphasis id="678239159F68FF8D7244F8B6FC86F88D" box="[636,774,1801,1827]" italics="true" pageId="11" pageNumber="558">A. tariquiae</emphasis>
</taxonomicName>
<emphasis id="678239159F68FF8D7328F8B6FCEDF88D" bold="true" box="[784,877,1801,1827]" pageId="11" pageNumber="558">
<taxonomicNameLabel id="7CB1846E9F68FF8D7328F8B6FCEDF88D" box="[784,877,1801,1827]" pageId="11" pageNumber="558" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, VAP in 
<taxonomicName id="92F69E849F68FF8D73E3F8B6FBE5F88D" authorityName="Acosta" authorityYear="2024" box="[987,1125,1801,1827]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="558" phylum="Arthropoda" rank="species" species="calilegua" status="sp. nov.">
<emphasis id="678239159F68FF8D73E3F8B6FBE5F88D" box="[987,1125,1801,1827]" italics="true" pageId="11" pageNumber="558">A. calilegua</emphasis>
</taxonomicName>
<emphasis id="678239159F68FF8D7448F8B6FB4CF88D" bold="true" box="[1136,1228,1801,1827]" pageId="11" pageNumber="558">
<taxonomicNameLabel id="7CB1846E9F68FF8D7448F8B6FB4CF88D" box="[1136,1228,1801,1827]" pageId="11" pageNumber="558" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
bears a shelf-like projection (quadrangular in ventral view), but it is not flanked by large conical apophyses; instead, these appear to be represented by conic tubercles shifted subdistally in the ‘shelf’ (
<figureCitation id="CDCDF9829F68FF8D73ABF8EEFC6CF8C2" box="[915,1004,1873,1900]" captionStart="FIGURE 5" captionStartId="9.[151,250,1745,1769]" captionTargetBox="[155,1432,374,1717]" captionTargetId="figure-45@9.[151,1435,370,1721]" captionTargetPageId="9" captionText="FIGURE 5. Acrographinotus calilegua sp. nov., HOlOtYpe ♂ (CDA 000.912). A. DORsal scutum, fRee teRgites (alsO shOwing VAP process from above), coxae IV, right trochanter and femur IV, dorsal view. B. Venter, with coxae IV, genital and stigmatic segments, free sternites, ventral anal plate, right trochanter and femur IV. C. Right coxa, trochanter, femur, patella and tibia IV, prolateral view. D. Detail of free tergites, dorsal and ventral anal operculum, lateral view. E. Right femur and patella IV, retrolateral view. F. Ocular mound, posterior view. Scale lines: 2 mm (A–E), 1 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.12783069" httpUri="https://zenodo.org/record/12783069/files/figure.png" pageId="11" pageNumber="558">Fig. 5B</figureCitation>
). These differences remain valid even when compared to smaller 
<taxonomicName id="92F69E849F68FF8D71F2F8CAFDD1F821" authorityName="Acosta" authorityYear="2024" box="[458,593,1909,1935]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="558" phylum="Arthropoda" rank="species" species="tariquiae">
<emphasis id="678239159F68FF8D71F2F8CAFDD1F821" box="[458,593,1909,1935]" italics="true" pageId="11" pageNumber="558">A. tariquiae</emphasis>
</taxonomicName>
males, which still displaY an α-tYpe lOOk in spite Of theiR Reduced size and weakeR aRmatuRe. MORe challenging is tO identifY β-males Of 
<taxonomicName id="92F69E849F68FF8D737BF826FC49F81D" authorityName="Acosta" authorityYear="2024" box="[835,969,1945,1971]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="558" phylum="Arthropoda" rank="species" species="tariquiae">
<emphasis id="678239159F68FF8D737BF826FC49F81D" box="[835,969,1945,1971]" italics="true" pageId="11" pageNumber="558">A. tariquiae</emphasis>
</taxonomicName>
, for which a suite of subtle details can be considered, as follows:
</paragraph>
<paragraph id="5549E5079F6FFF8A7204FF1CFAB8FDCE" pageId="12" pageNumber="559">
<table id="27F617A79F6F0079709BFF1CFA0DFDCE" box="[163,1421,163,608]" gridcols="5" gridrows="7" pageId="12" pageNumber="559">
<tr id="EBC6E7459F6F0079709BFF1CFA0DFF71" box="[163,1421,163,223]" gridrow="0" pageId="12" pageNumber="559" rowspan-0="1" rowspan-1="1">
<th id="A8178E399F6F00797204FF1CFCA8FF71" box="[572,808,163,223]" gridcol="2" gridrow="0" pageId="12" pageNumber="559">
<taxonomicName id="92F69E849F6FFF8A7204FF1CFD37FF15" authorityName="Acosta" authorityYear="2024" box="[572,695,163,187]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="559" phylum="Arthropoda" rank="species" species="calilegua" status="sp. nov.">
<emphasis id="678239159F6FFF8A7204FF1CFD37FF15" box="[572,695,163,187]" italics="true" pageId="12" pageNumber="559">A. calilegua</emphasis>
</taxonomicName>
<emphasis id="678239159F6FFF8A7285FF1CFC8EFF15" bold="true" box="[701,782,163,187]" pageId="12" pageNumber="559">
<taxonomicNameLabel id="7CB1846E9F6FFF8A7285FF1CFC8EFF15" box="[701,782,163,187]" pageId="12" pageNumber="559" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
♂ (n=2)
</th>
<th id="A8178E399F6F00797367FF1CFBCFFF71" box="[863,1103,163,223]" gridcol="3" gridrow="0" pageId="12" pageNumber="559">
<taxonomicName id="92F69E849F6FFF8A7357FF1CFC68FF14" authorityName="Acosta" authorityYear="2024" box="[879,1000,163,186]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="559" phylum="Arthropoda" rank="species" species="tariquiae" status="sp. nov.">
<emphasis id="678239159F6FFF8A7357FF1CFC68FF14" box="[879,1000,163,186]" italics="true" pageId="12" pageNumber="559">A. tariquiae</emphasis>
</taxonomicName>
<emphasis id="678239159F6FFF8A73D6FF1CFBBFFF15" bold="true" box="[1006,1087,163,187]" pageId="12" pageNumber="559">
<taxonomicNameLabel id="7CB1846E9F6FFF8A73D6FF1CFBBFFF15" box="[1006,1087,163,187]" pageId="12" pageNumber="559" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
– small α ♂ (n=3)
</th>
<th id="A8178E399F6F00797445FF1CFA0DFF71" box="[1149,1421,163,223]" gridcol="4" gridrow="0" pageId="12" pageNumber="559">
<taxonomicName id="92F69E849F6FFF8A7445FF1CFB77FF14" authorityName="Acosta" authorityYear="2024" box="[1149,1271,163,186]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="559" phylum="Arthropoda" rank="species" species="tariquiae" status="sp. nov.">
<emphasis id="678239159F6FFF8A7445FF1CFB77FF14" box="[1149,1271,163,186]" italics="true" pageId="12" pageNumber="559">A. tariquiae</emphasis>
</taxonomicName>
<emphasis id="678239159F6FFF8A74C5FF1CFACDFF15" bold="true" box="[1277,1357,163,187]" pageId="12" pageNumber="559">
<taxonomicNameLabel id="7CB1846E9F6FFF8A74C5FF1CFACDFF15" box="[1277,1357,163,187]" pageId="12" pageNumber="559" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
– β ♂ (n=2)
</th>
</tr>
<tr id="EBC6E7459F6F0079709BFF50FA0DFEA9" box="[163,1421,239,263]" gridrow="1" pageId="12" pageNumber="559">
<th id="A8178E399F6F0079709BFF50FD98FEA9" box="[163,536,239,263]" colspan="2" colspanRight="1" gridcol="0" gridrow="1" pageId="12" pageNumber="559">VAP: ‘shelf’-like projection</th>
<td id="A8178E399F6F00797204FF50FCA8FEA9" box="[572,808,239,263]" gridcol="2" gridrow="1" pageId="12" pageNumber="559">present (Figs. 5B, D)</td>
<td id="A8178E399F6F00797367FF50FBCFFEA9" box="[863,1103,239,263]" gridcol="3" gridrow="1" pageId="12" pageNumber="559">present (Figs. 3G–H)</td>
<td id="A8178E399F6F00797445FF50FA0DFEA9" box="[1149,1421,239,263]" gridcol="4" gridrow="1" pageId="12" pageNumber="559">absent (Figs. 3I–J)</td>
</tr>
<tr id="EBC6E7459F6F0079709BFEA8FA0DFEFD" box="[163,1421,279,339]" gridrow="2" pageId="12" pageNumber="559" rowspan-1="1">
<th id="A8178E399F6F0079709BFEA8FE24FEFD" box="[163,420,279,339]" gridcol="0" gridrow="2" pageId="12" pageNumber="559">VAP: flanking apophyses</th>
<td id="A8178E399F6F00797204FEA8FCA8FEFD" box="[572,808,279,339]" gridcol="2" gridrow="2" pageId="12" pageNumber="559">absent</td>
<td id="A8178E399F6F00797367FEA8FBCFFEFD" box="[863,1103,279,339]" gridcol="3" gridrow="2" pageId="12" pageNumber="559">present, may be weak (Figs. 3G–H)</td>
<td id="A8178E399F6F00797445FEA8FA0DFEFD" box="[1149,1421,279,339]" gridcol="4" gridrow="2" pageId="12" pageNumber="559">replaced by button-like tubercles (Figs. 3I–J)</td>
</tr>
<tr id="EBC6E7459F6F0079709BFEDCFA0DFE31" box="[163,1421,355,415]" gridrow="3" pageId="12" pageNumber="559">
<th id="A8178E399F6F0079709BFEDCFE24FE31" box="[163,420,355,415]" gridcol="0" gridrow="3" pageId="12" pageNumber="559">VAP: subapical conic beneath ‘shelf’</th>
<td id="A8178E399F6F00797185FEDCFD98FE31" box="[445,536,355,415]" gridcol="1" gridrow="3" pageId="12" pageNumber="559">tubercles</td>
<td id="A8178E399F6F00797204FEDCFCA8FE31" box="[572,808,355,415]" gridcol="2" gridrow="3" pageId="12" pageNumber="559">present (Figs. 5B, D)</td>
<td id="A8178E399F6F00797367FEDCFBCFFE31" box="[863,1103,355,415]" gridcol="3" gridrow="3" pageId="12" pageNumber="559">absent</td>
<td id="A8178E399F6F00797445FEDCFA0DFE31" box="[1149,1421,355,415]" gridcol="4" gridrow="3" pageId="12" pageNumber="559">NA</td>
</tr>
<tr id="EBC6E7459F6F0079709BFE0FFA0DFE42" box="[163,1421,432,492]" gridrow="4" pageId="12" pageNumber="559">
<th id="A8178E399F6F0079709BFE0FFD98FE42" box="[163,536,432,492]" colspan="2" colspanRight="1" gridcol="0" gridrow="4" pageId="12" pageNumber="559">FeIV: shape and articulation</th>
<td id="A8178E399F6F00797204FE0FFCA8FE42" box="[572,808,432,492]" gridcol="2" gridrow="4" pageId="12" pageNumber="559">arched, articulated sideways (Fig. 5A)</td>
<td id="A8178E399F6F00797367FE0FFBCFFE42" box="[863,1103,432,492]" gridcol="3" gridrow="4" pageId="12" pageNumber="559">sub-straight, articulated diagonally</td>
<td id="A8178E399F6F00797445FE0FFA0DFE42" box="[1149,1421,432,492]" gridcol="4" gridrow="4" pageId="12" pageNumber="559">female-type</td>
</tr>
<tr id="EBC6E7459F6F0079709BFE43FA0DFD96" box="[163,1421,508,568]" gridrow="5" pageId="12" pageNumber="559">
<th id="A8178E399F6F0079709BFE43FD98FD96" box="[163,536,508,568]" colspan="2" colspanRight="1" gridcol="0" gridrow="5" pageId="12" pageNumber="559">FeIV: retrodorsal row of tubercles</th>
<td id="A8178E399F6F00797204FE43FCA8FD96" box="[572,808,508,568]" gridcol="2" gridrow="5" pageId="12" pageNumber="559">moderate but distinct (Fig. 5A, E)</td>
<td id="A8178E399F6F00797367FE43FBCFFD96" box="[863,1103,508,568]" gridcol="3" gridrow="5" pageId="12" pageNumber="559">conspicuous, truncate tubercles</td>
<td id="A8178E399F6F00797445FE43FA0DFD96" box="[1149,1421,508,568]" gridcol="4" gridrow="5" pageId="12" pageNumber="559">little differentiated</td>
</tr>
<tr id="EBC6E7459F6F0079709BFDF7FA0DFDCE" box="[163,1421,584,608]" gridrow="6" pageId="12" pageNumber="559">
<th id="A8178E399F6F0079709BFDF7FD98FDCE" box="[163,536,584,608]" colspan="2" colspanRight="1" gridcol="0" gridrow="6" pageId="12" pageNumber="559">FeIV/DS ratio (relative size)</th>
<td id="A8178E399F6F00797204FDF7FCA8FDCE" box="[572,808,584,608]" gridcol="2" gridrow="6" pageId="12" pageNumber="559">0.84–0.89</td>
<td id="A8178E399F6F00797367FDF7FBCFFDCE" box="[863,1103,584,608]" gridcol="3" gridrow="6" pageId="12" pageNumber="559">0.94–0.95–1.09</td>
<td id="A8178E399F6F00797445FDF7FA0DFDCE" box="[1149,1421,584,608]" gridcol="4" gridrow="6" pageId="12" pageNumber="559">0.86–1.02</td>
</tr>
</table>
</paragraph>
<caption id="0189B58F9F6FFF8A70AFF9E5FDBFF970" ID-DOI="http://doi.org/10.5281/zenodo.12783073" ID-Zenodo-Dep="12783073" httpUri="https://zenodo.org/record/12783073/files/figure.png" pageId="12" pageNumber="559" startId="12.[151,250,1626,1650]" targetBox="[151,1435,634,1602]" targetPageId="12" targetType="figure">
<paragraph id="5549E5079F6FFF8A70AFF9E5FDBFF970" blockId="12.[151,1436,1626,1758]" pageId="12" pageNumber="559">
<emphasis id="678239159F6FFF8A70AFF9E5FE93F9DC" bold="true" box="[151,275,1626,1650]" pageId="12" pageNumber="559">FIGURE 7.</emphasis>
A: Records of 
<taxonomicName id="92F69E849F6FFF8A7194F9E5FD3AF9DC" authorityName="Acosta" authorityYear="2024" box="[428,698,1626,1650]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="559" phylum="Arthropoda" rank="species" species="tariquiae" status="sp. nov.">
<emphasis id="678239159F6FFF8A7194F9E5FD3AF9DC" box="[428,698,1626,1650]" italics="true" pageId="12" pageNumber="559">Acrographinotus tariquiae</emphasis>
</taxonomicName>
<emphasis id="678239159F6FFF8A72F8F9E5FC90F9DC" bold="true" box="[704,784,1626,1650]" pageId="12" pageNumber="559">
<taxonomicNameLabel id="7CB1846E9F6FFF8A72F8F9E5FC90F9DC" box="[704,784,1626,1650]" pageId="12" pageNumber="559" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in Tarija Department, Bolivia (yellow dots) and 
<taxonomicName id="92F69E849F6FFF8A74C5F9E5FAF7F9DC" authorityName="Acosta" authorityYear="2024" box="[1277,1399,1626,1650]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="559" phylum="Arthropoda" rank="species" species="calilegua" status="sp. nov.">
<emphasis id="678239159F6FFF8A74C5F9E5FAF7F9DC" box="[1277,1399,1626,1650]" italics="true" pageId="12" pageNumber="559">A. calilegua</emphasis>
</taxonomicName>
<emphasis id="678239159F6FFF8A7546F9E5FF43F938" bold="true" pageId="12" pageNumber="559">
<taxonomicNameLabel id="7CB1846E9F6FFF8A7546F9E5FF43F938" pageId="12" pageNumber="559" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in Jujuy Province, Argentina (red dots); blue contours: protected areas (1: Tariquía Flora and Fauna National Reserve; 2: Calilegua National Park). B: Type locality of 
<taxonomicName id="92F69E849F6FFF8A7257F91DFD6BF914" authorityName="Acosta" authorityYear="2024" box="[623,747,1698,1722]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="559" phylum="Arthropoda" rank="species" species="calilegua" status="sp. nov.">
<emphasis id="678239159F6FFF8A7257F91DFD6BF914" box="[623,747,1698,1722]" italics="true" pageId="12" pageNumber="559">A. calilegua</emphasis>
</taxonomicName>
<emphasis id="678239159F6FFF8A72CBF91DFCC6F914" bold="true" box="[755,838,1698,1722]" pageId="12" pageNumber="559">
<taxonomicNameLabel id="7CB1846E9F6FFF8A72CBF91DFCC6F914" box="[755,838,1698,1722]" pageId="12" pageNumber="559" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: cloud forests at ‘Monolito’ (
<emphasis id="678239159F6FFF8A7441F91CFB18F914" box="[1145,1176,1699,1722]" italics="true" pageId="12" pageNumber="559">ca.</emphasis>
1700 m a.s.l.), Calilegua National Park, Jujuy Province, Argentina.
</paragraph>
</caption>
<paragraph id="5549E5079F6FFF8A70FFF8B3FD6CF8E5" blockId="12.[151,1437,1804,2011]" pageId="12" pageNumber="559">
Regarding females, DS of 
<taxonomicName id="92F69E849F6FFF8A71CBF8B2FDFAF888" authorityName="Acosta" authorityYear="2024" box="[499,634,1804,1831]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="559" phylum="Arthropoda" rank="species" species="calilegua" status="sp. nov.">
<emphasis id="678239159F6FFF8A71CBF8B2FDFAF888" box="[499,634,1804,1831]" italics="true" pageId="12" pageNumber="559">A. calilegua</emphasis>
</taxonomicName>
<emphasis id="678239159F6FFF8A72BAF8B3FD5CF888" bold="true" box="[642,732,1804,1830]" pageId="12" pageNumber="559">
<taxonomicNameLabel id="7CB1846E9F6FFF8A72BAF8B3FD5CF888" box="[642,732,1804,1830]" pageId="12" pageNumber="559" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
looks slightly wider, with constriction 2 less marked and midbulge less differentiated than in 
<taxonomicName id="92F69E849F6FFF8A71C7F88EFD05F8E4" authorityName="Acosta" authorityYear="2024" box="[511,645,1841,1867]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="559" phylum="Arthropoda" rank="species" species="tariquiae" status="sp. nov.">
<emphasis id="678239159F6FFF8A71C7F88EFD05F8E4" box="[511,645,1841,1867]" italics="true" pageId="12" pageNumber="559">A. tariquiae</emphasis>
</taxonomicName>
<emphasis id="678239159F6FFF8A72B3F88FFD65F8E4" bold="true" box="[651,741,1840,1866]" pageId="12" pageNumber="559">
<taxonomicNameLabel id="7CB1846E9F6FFF8A72B3F88FFD65F8E4" box="[651,741,1840,1866]" pageId="12" pageNumber="559" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="1DECB68C9F6FFF8B70FFF8EBFC46FECB" lastPageId="13" lastPageNumber="560" pageId="12" pageNumber="559" type="distribution">
<paragraph id="5549E5079F6FFF8B70FFF8EBFC46FECB" blockId="12.[151,1437,1804,2011]" lastBlockId="13.[151,1437,150,357]" lastPageId="13" lastPageNumber="560" pageId="12" pageNumber="559">
<emphasis id="678239159F6FFF8A70FFF8EBFE74F8C0" bold="true" box="[199,500,1876,1902]" pageId="12" pageNumber="559">Distribution and habitat.</emphasis>
<taxonomicName id="92F69E849F6FFF8A71C4F8EBFCA9F8C0" authorityName="Acosta" authorityYear="2024" box="[508,809,1876,1902]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="559" phylum="Arthropoda" rank="species" species="calilegua" status="sp. nov.">
<emphasis id="678239159F6FFF8A71C4F8EBFCA9F8C0" box="[508,809,1876,1902]" italics="true" pageId="12" pageNumber="559">Acrographinotus calilegua</emphasis>
</taxonomicName>
<emphasis id="678239159F6FFF8A730AF8EBFC0CF8C0" bold="true" box="[818,908,1876,1902]" pageId="12" pageNumber="559">
<taxonomicNameLabel id="7CB1846E9F6FFF8A730AF8EBFC0CF8C0" box="[818,908,1876,1902]" pageId="12" pageNumber="559" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was recorded from a few sites in and near the Calilegua National Park, 
<collectingRegion id="97322BE59F6FFF8A718AF8C7FDDCF83C" box="[434,604,1912,1938]" country="Argentina" name="Jujuy" pageId="12" pageNumber="559">Jujuy Province</collectingRegion>
, 
<collectingCountry id="2DE1A5979F6FFF8A725EF8C7FD58F83C" box="[614,728,1912,1938]" name="Argentina" pageId="12" pageNumber="559">Argentina</collectingCountry>
, a renowned biodiversity hotspot in the Yungas ecoregion (
<figureCitation id="CDCDF9829F6FFF8A7557F8C7FF3DF818" captionStart="FIGURE 7" captionStartId="12.[151,250,1626,1650]" captionTargetBox="[151,1435,634,1602]" captionTargetId="figure-132@12.[151,1435,634,1602]" captionTargetPageId="12" captionText="FIGURE 7. A: Records of Acrographinotus tariquiae sp. nov. in Tarija Department, Bolivia (yellow dots) and A. calilegua sp. nov. in Jujuy Province, Argentina (red dots); blue contours: protected areas (1: Tariquía Flora and Fauna National Reserve; 2: Calilegua National Park). B: Type locality of A. calilegua sp. nov.: cloud forests at ‘Monolito’ (ca. 1700 m a.s.l.), Calilegua National Park, Jujuy Province, Argentina." figureDoi="http://doi.org/10.5281/zenodo.12783073" httpUri="https://zenodo.org/record/12783073/files/figure.png" pageId="12" pageNumber="559">Fig. 7A</figureCitation>
). This protected area is accessible by Provincial Route 83, which ascends on the east-faced slope across different vegetation altitudinal belts: a piedmont-montane transition (
<quantity id="920E48E29F6FFF8A7317F87FFC2EF875" box="[815,942,1984,2011]" metricMagnitude="2" metricUnit="m" metricValue="6.75" metricValueMax="8.5" metricValueMin="5.0" pageId="12" pageNumber="559" unit="m" value="675.0" valueMax="850.0" valueMin="500.0">500–850 m</quantity>
a.s.l.), the montane rainforests (the belt with the highest precipitation, 
<quantity id="920E48E29F6EFF8B7180FF29FDC6FF1F" box="[440,582,150,177]" metricMagnitude="3" metricUnit="m" metricValue="1.075" metricValueMax="1.3" metricValueMin="0.85" pageId="13" pageNumber="560" unit="m" value="1075.0" valueMax="1300.0" valueMin="850.0">850–1300 m</quantity>
a.s.l.), and a montane rainforests-montane forests transition (
<quantity id="920E48E29F6EFF8B74C6FF28FA1CFF1F" box="[1278,1436,151,177]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="13" pageNumber="560" unit="m" value="1450.0" valueMax="1600.0" valueMin="1300.0">1300–1600 m</quantity>
a.s.l.). Beyond the National Park itself, the road further leads to cloud forests (
<quantity id="920E48E29F6EFF8B73C6FF04FB1AFF7B" box="[1022,1178,187,213]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="2.4" metricValueMin="1.6" pageId="13" pageNumber="560" unit="m" value="2000.0" valueMax="2400.0" valueMin="1600.0">1600–2400 m</quantity>
a.s.l.), which give way to high altitude grasslands above (
<bibRefCitation id="316798F69F6EFF8B7228FF60FD7DFF57" author="Politi, N. &amp; Rivera, L. O." box="[528,765,222,249]" pageId="13" pageNumber="560" pagination="43 - 52" refId="ref11821" refString="Politi, N. &amp; Rivera, L. O. (2005) Abundance and distribution of parrots along the elevational gradient of Calilegua National Park, Argentina. Ornitologia Neotropical, 16, 43 - 52." type="journal article" year="2005">Politi &amp; Rivera 2005</bibRefCitation>
; 
<bibRefCitation id="316798F69F6EFF8B7330FF60FC5EFF57" author="Malizia, L. &amp; Pacheco, S. &amp; Blundo, C. &amp; Brown, A. D." box="[776,990,223,249]" pageId="13" pageNumber="560" pagination="53 - 73" refId="ref11679" refString="Malizia, L., Pacheco, S., Blundo, C. &amp; Brown, A. D. (2012) Caracterizacion altitudinal, uso y conservacion de las Yungas Subtropicales de Argentina. Ecosistemas, 21 (1 - 2), 53 - 73." type="journal article" year="2012">
Malizia 
<emphasis id="678239159F6EFF8B735BFF60FC1BFF57" box="[867,923,223,249]" italics="true" pageId="13" pageNumber="560">et al.</emphasis>
2012
</bibRefCitation>
). The elevation range of 
<taxonomicName id="92F69E849F6EFF8B74D6FF60FAF4FF57" authorityName="Acosta" authorityYear="2024" box="[1262,1396,223,249]" class="Arachnida" family="Gonyleptidae" genus="Acrographinotus" kingdom="Animalia" order="Opiliones" pageId="13" pageNumber="560" phylum="Arthropoda" rank="species" species="calilegua" status="sp. nov.">
<emphasis id="678239159F6EFF8B74D6FF60FAF4FF57" box="[1262,1396,223,249]" italics="true" pageId="13" pageNumber="560">A. calilegua</emphasis>
</taxonomicName>
<emphasis id="678239159F6EFF8B7543FF60FF48FEB3" bold="true" pageId="13" pageNumber="560">
<taxonomicNameLabel id="7CB1846E9F6EFF8B7543FF60FF48FEB3" pageId="13" pageNumber="560" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<quantity id="920E48E29F6EFF8B70EFFEBCFEF5FEB3" box="[215,373,259,285]" metricMagnitude="3" metricUnit="m" metricValue="2.03" metricValueMax="2.4" metricValueMin="1.66" pageId="13" pageNumber="560" unit="m" value="2030.0" valueMax="2400.0" valueMin="1660.0">1660–2400 m</quantity>
a.s.l.) places this species in the upper forest belt (cloud forests: 
<figureCitation id="CDCDF9829F6EFF8B747EFEBCFB1BFEB3" box="[1094,1179,259,285]" captionStart="FIGURE 7" captionStartId="12.[151,250,1626,1650]" captionTargetBox="[151,1435,634,1602]" captionTargetId="figure-132@12.[151,1435,634,1602]" captionTargetPageId="12" captionText="FIGURE 7. A: Records of Acrographinotus tariquiae sp. nov. in Tarija Department, Bolivia (yellow dots) and A. calilegua sp. nov. in Jujuy Province, Argentina (red dots); blue contours: protected areas (1: Tariquía Flora and Fauna National Reserve; 2: Calilegua National Park). B: Type locality of A. calilegua sp. nov.: cloud forests at ‘Monolito’ (ca. 1700 m a.s.l.), Calilegua National Park, Jujuy Province, Argentina." figureDoi="http://doi.org/10.5281/zenodo.12783073" httpUri="https://zenodo.org/record/12783073/files/figure.png" pageId="13" pageNumber="560">Fig. 7B</figureCitation>
) and some transitional sites. In lower vegetation belts of Calilegua, especially in montane rainforests, samples are dominated by another gonyleptid, 
<taxonomicName id="92F69E849F6EFF8B7125FEF4FD3BFECB" authority="(Roewer, 1925)" baseAuthorityName="Roewer" baseAuthorityYear="1925" box="[285,699,330,357]" class="Arachnida" family="Gonyleptidae" genus="Pachyloides" kingdom="Animalia" order="Opiliones" pageId="13" pageNumber="560" phylum="Arthropoda" rank="species" species="sicarius">
<emphasis id="678239159F6EFF8B7125FEF4FD86FECA" box="[285,518,331,357]" italics="true" pageId="13" pageNumber="560">Pachyloides sicarius</emphasis>
(
<bibRefCitation id="316798F69F6EFF8B722CFEF4FD33FECB" author="Roewer, C. F." box="[532,691,330,357]" pageId="13" pageNumber="560" pagination="1 - 34" refId="ref11926" refString="Roewer, C. F. (1925) Opilioniden aus Sud-Amerika. Bolletino dei Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino, New Series, 40 (34), 1 - 34, pls. 5 - 6." type="journal article" year="1925">Roewer, 1925</bibRefCitation>
)
</taxonomicName>
(LEA unpubl. records).
</paragraph>
</subSubSection>
</treatment>
</document>