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<document id="6038A2AA19D2B86F65CB1D78A979CBDD" ID-DOI="10.5281/zenodo.181480" ID-GBIF-Dataset="a312fd93-3766-46f6-90d4-7b689ccd3479" ID-ISSN="1175-5326" ID-Zenodo-Dep="181480" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459834491155" checkinUser="plazi" docAuthor="Sokolov, Igor M. &amp; Carlton, Christopher E." docDate="2008" docId="270587E8F40BFFDA74CEF0EC5216FD21" docLanguage="en" docName="zt01740p044.pdf" docOrigin="Zootaxa 1740" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Serranillus septentrionis Sokolov and Carlton, new species" docType="treatment" docVersion="9" lastPageNumber="40" masterDocId="DB3CFF90F40AFFD97459F46E5602FFB3" masterDocTitle="Two new species of blind, forest litter-inhabiting ground beetles from the subtribe Anillina (Carabidae: Trechinae: Bembidiini) from eastern U. S. A." masterLastPageNumber="44" masterPageNumber="37" pageNumber="38" updateTime="1698223317936" updateUser="plazi">
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<mods:title id="1BCDA48C2196BBB463D5AD512E9ECA11">Two new species of blind, forest litter-inhabiting ground beetles from the subtribe Anillina (Carabidae: Trechinae: Bembidiini) from eastern U. S. A.</mods:title>
</mods:titleInfo>
<mods:name id="5EA7FD54A16E95042FAE61B1D845F0EF" type="personal">
<mods:role id="31CAE8C0DA3281A2789C0AE2F6C3E496">
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</mods:role>
<mods:namePart id="0B21F9CC7456253210769F837E5F5AA2">Sokolov, Igor M.</mods:namePart>
</mods:name>
<mods:name id="AC03A12D8430EBE48246397A84B28926" type="personal">
<mods:role id="B4DF5F38318B56A5842588DD2AA7A2D8">
<mods:roleTerm id="60A96B395261FE360F5A468B8CF907B0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4F4D544CD82C27F00312D0FEE260E53C">Carlton, Christopher E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C9E44827376E25F49C1FD1CFC22934F9">text</mods:typeOfResource>
<mods:relatedItem id="DBCAF884018965582A8DE96D9526E68C" type="host">
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<mods:title id="8909C1C880878F742CA040356E71C749">Zootaxa</mods:title>
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<mods:part id="5364E37B6A758991CEA34FBB768680A0">
<mods:date id="BA982963E5AF9792937777F80947C758">2008</mods:date>
<mods:detail id="77CAD2C5850BA97AE5C3EC8C42C4D256" type="volume">
<mods:number id="48C0CCC765ACBF7FEE590C0B39A47F4C">1740</mods:number>
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<mods:classification id="5528E82C3DDA86E7EFEFC68380148BDE">journal article</mods:classification>
<mods:identifier id="CB7402527883EC081D764D91DCD1457D" type="DOI">10.5281/zenodo.181480</mods:identifier>
<mods:identifier id="A30CFF2CD5165E056DBA400B0375D3A4" type="GBIF-Dataset">a312fd93-3766-46f6-90d4-7b689ccd3479</mods:identifier>
<mods:identifier id="D079FD01180F9D4C794FB8A7C4172FF3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="46DD0BCA53750B839AC6087371B4F3B2" type="Zenodo-Dep">181480</mods:identifier>
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<treatment id="270587E8F40BFFDA74CEF0EC5216FD21" ID-DOI="http://doi.org/10.5281/zenodo.6233715" ID-GBIF-Taxon="119358520" ID-Zenodo-Dep="6233715" LSID="urn:lsid:plazi:treatment:270587E8F40BFFDA74CEF0EC5216FD21" httpUri="http://treatment.plazi.org/id/270587E8F40BFFDA74CEF0EC5216FD21" lastPageId="3" lastPageNumber="40" pageId="1" pageNumber="38">
<subSubSection id="E7B66575F40BFFD874CEF0EC556AFB2F" box="[151,872,1154,1180]" pageId="1" pageNumber="38" type="nomenclature">
<paragraph id="AF1336FEF40BFFD874CEF0EC556AFB2F" blockId="1.[151,872,1154,1180]" box="[151,872,1154,1180]" pageId="1" pageNumber="38">
<heading id="F45B8192F40BFFD874CEF0EC556AFB2F" bold="true" box="[151,872,1154,1180]" fontSize="11" level="1" pageId="1" pageNumber="38" reason="1">
<emphasis id="9DD8EAECF40BFFD874CEF0EC556AFB2F" bold="true" box="[151,872,1154,1180]" pageId="1" pageNumber="38">
<taxonomicName id="68AC4D7DF40BFFD874CEF0EC54C9FB2F" ID-CoL="6YKV9" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" box="[151,715,1154,1180]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="septentrionis" status="sp. nov.">
<emphasis id="9DD8EAECF40BFFD874CEF0EC57C2FB2F" bold="true" box="[151,448,1154,1180]" italics="true" pageId="1" pageNumber="38">Serranillus septentrionis</emphasis>
Sokolov and Carlton
</taxonomicName>
,
<taxonomicNameLabel id="86EB5797F40BFFD87681F0EC556AFB2F" box="[728,872,1154,1180]" pageId="1" pageNumber="38" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E7B66575F40BFFD874CEF0A35773F9FC" pageId="1" pageNumber="38" type="materials_examined">
<paragraph id="AF1336FEF40BFFD874CEF0A35221FA84" blockId="1.[151,1437,1229,2015]" pageId="1" pageNumber="38">
<emphasis id="9DD8EAECF40BFFD874CEF0A35712FB54" bold="true" box="[151,272,1229,1255]" pageId="1" pageNumber="38">
<typeStatus id="7017885CF40BFFD874CEF0A35709FB54" box="[151,267,1229,1255]" pageId="1" pageNumber="38" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male labeled / VIRGINIA: Botetourt Co., Blue Ridge Pwy., Black Horse Gap,
<geoCoordinate id="CA985039F40BFFD8715CF0A3539EFB54" box="[1285,1436,1229,1255]" direction="north" orientation="latitude" pageId="1" pageNumber="38" precision="9" value="37.422165">37°25.33 N</geoCoordinate>
<geoCoordinate id="CA985039F40BFFD874CEF09B5731FABC" box="[151,307,1269,1295]" direction="west" orientation="longitude" pageId="1" pageNumber="38" precision="9" value="-79.757164">79°45.43 W</geoCoordinate>
,
<specimenCount id="B9AAFD77F40BFFD8751BF09B578FFABC" box="[322,397,1269,1295]" pageId="1" pageNumber="38" type="generic">
<quantity id="68549B1BF40BFFD8751BF09B578BFABC" box="[322,393,1269,1295]" metricMagnitude="2" metricUnit="m" metricValue="7.45" pageId="1" pageNumber="38" unit="m" value="745.0">745m</quantity>
.
</specimenCount>
Litter sifting,
<date id="DB12103EF40BFFD8761BF09B54DAFABC" box="[578,728,1269,1295]" pageId="1" pageNumber="38" value="2006-05-02">2 May 2006</date>
A.K.Tishechkin / /
<typeStatus id="7017885CF40BFFD87796F09B526FFABC" box="[975,1133,1269,1295]" pageId="1" pageNumber="38" type="holotype">HOLOTYPE</typeStatus>
,
<taxonomicName id="68AC4D7DF40BFFD87025F09B578EFA84" authority="Sokolov and Carlton" authorityName="Sokolov and Carlton" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="septentrionis">Serranillus septentrionis Sokolov and Carlton</taxonomicName>
, des. 2007/. Deposited
<collectingCountry id="D7BB766EF40BFFD876C4F17354D3FA84" box="[669,721,1309,1335]" name="United States of America" pageId="1" pageNumber="38">U.S.</collectingCountry>
National Museum (
<collectionCode id="C9BDAE3BF40BFFD877E2F1735215FA84" LSID="urn:lsid:biocol.org:col:34871" box="[955,1047,1309,1335]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="38">USNM</collectionCode>
).
</paragraph>
<paragraph id="AF1336FEF40BFFD8749CF12B538DFAEC" blockId="1.[151,1437,1229,2015]" box="[197,1423,1349,1375]" pageId="1" pageNumber="38">
<emphasis id="9DD8EAECF40BFFD8749CF12B5769FAEC" bold="true" box="[197,363,1349,1375]" pageId="1" pageNumber="38">
<typeStatus id="7017885CF40BFFD8749CF12B5700FAEC" box="[197,258,1349,1375]" pageId="1" pageNumber="38">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="D7BB766EF40BFFD8752BF12B57A6FAEC" box="[370,420,1349,1375]" name="United States of America" pageId="1" pageNumber="38">U.S.</collectingCountry>
A, VA, Botetourt Co., Blue Ridge Pwy., Black Horse Gap,
<geoCoordinate id="CA985039F40BFFD87003F12B52E9FAEC" box="[1114,1259,1349,1375]" direction="north" orientation="latitude" pageId="1" pageNumber="38" precision="9" value="37.422165">37°25.33 N</geoCoordinate>
<geoCoordinate id="CA985039F40BFFD870ABF12B5389FAEC" box="[1266,1419,1349,1375]" direction="west" orientation="longitude" pageId="1" pageNumber="38" precision="9" value="-79.757164">79°45.43 W</geoCoordinate>
.
</paragraph>
<paragraph id="AF1336FEF40BFFD8749CF1035773F9FC" blockId="1.[151,1437,1229,2015]" pageId="1" pageNumber="38">
<typeStatus id="7017885CF40BFFD8749CF1035740FA34" box="[197,322,1389,1415]" pageId="1" pageNumber="38" type="paratype">
<emphasis id="9DD8EAECF40BFFD8749CF1035740FA34" bold="true" box="[197,322,1389,1415]" pageId="1" pageNumber="38">Paratypes</emphasis>
</typeStatus>
(6). One male and
<specimenCount id="B9AAFD77F40BFFD8767EF103548FFA34" box="[551,653,1389,1415]" pageId="1" pageNumber="38" type="female">1 female</specimenCount>
with same data as
<typeStatus id="7017885CF40BFFD87734F10355D5FA34" box="[877,983,1389,1415]" pageId="1" pageNumber="38" type="holotype">holotype</typeStatus>
;
<specimenCount id="B9AAFD77F40BFFD877BAF1035230FA34" box="[995,1074,1389,1415]" pageId="1" pageNumber="38" type="male">1 male</specimenCount>
labeled /
<collectingCountry id="D7BB766EF40BFFD870FFF10352E3FA34" box="[1190,1249,1389,1415]" name="United States of America" pageId="1" pageNumber="38">USA</collectingCountry>
VA: Giles Co., Jefferson NF, Flat Top Mt., at
<geoCoordinate id="CA985039F40BFFD875ACF1FB5475FA1C" box="[501,631,1429,1455]" direction="north" orientation="latitude" pageId="1" pageNumber="38" precision="92" value="37.243332">37°14.6 N</geoCoordinate>
<geoCoordinate id="CA985039F40BFFD87627F1FB5504FA1C" box="[638,774,1429,1455]" direction="west" orientation="longitude" pageId="1" pageNumber="38" precision="92" value="-80.86667">80°52.0 W</geoCoordinate>
, forest litter,
<quantity id="68549B1BF40BFFD877C6F1FB55F0FA1C" box="[927,1010,1429,1455]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="1" pageNumber="38" unit="m" value="1120.0">1120m</quantity>
, C.E.Carlton
<date id="DB12103EF40BFFD870C9F1FB532CFA1C" box="[1168,1326,1429,1455]" pageId="1" pageNumber="38" value="2006-05-25">25 May 2006</date>
/;
<specimenCount id="B9AAFD77F40BFFD8711BF1FB539EFA1C" box="[1346,1436,1429,1455]" pageId="1" pageNumber="38" type="male">2 males</specimenCount>
labeled /
<collectingCountry id="D7BB766EF40BFFD87551F1D35741FA64" box="[264,323,1469,1495]" name="United States of America" pageId="1" pageNumber="38">USA</collectingCountry>
VA: Giles Co., Jefferson NF, Flat Top Mt., at
<geoCoordinate id="CA985039F40BFFD877DAF1D3520AFA64" box="[899,1032,1469,1495]" direction="north" orientation="latitude" pageId="1" pageNumber="38" precision="92" value="37.233334">37°14.0 N</geoCoordinate>
<geoCoordinate id="CA985039F40BFFD8704AF1D352A2FA64" box="[1043,1184,1469,1495]" direction="west" orientation="longitude" pageId="1" pageNumber="38" precision="92" value="-80.875">80°52.5 W</geoCoordinate>
,
<quantity id="68549B1BF40BFFD870F7F1D35300FA64" box="[1198,1282,1469,1495]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="1" pageNumber="38" unit="m" value="1080.0">1080m</quantity>
, forest litter, C.E.Carlton
<date id="DB12103EF40BFFD87573F18B57C8FA4C" box="[298,458,1509,1535]" pageId="1" pageNumber="38" value="2006-05-25">25 May 2006</date>
/;
<specimenCount id="B9AAFD77F40BFFD87586F18B5447FA4C" box="[479,581,1509,1535]" pageId="1" pageNumber="38" type="female">1 female</specimenCount>
labeled /
<collectingCountry id="D7BB766EF40BFFD876E1F18B54F1FA4C" box="[696,755,1509,1535]" name="United States of America" pageId="1" pageNumber="38">USA</collectingCountry>
VA: Giles Co., Jefferson NF, Flat Top Mt., at
<geoCoordinate id="CA985039F40BFFD87140F18B539EFA4C" box="[1305,1436,1509,1535]" direction="north" orientation="latitude" pageId="1" pageNumber="38" precision="92" value="37.23">37°13.8 N</geoCoordinate>
<geoCoordinate id="CA985039F40BFFD874CEF2635724F994" box="[151,294,1549,1575]" direction="west" orientation="longitude" pageId="1" pageNumber="38" precision="92" value="-80.87334">80°52.4 W</geoCoordinate>
, hardwood berlese,
<quantity id="68549B1BF40BFFD87678F2635465F994" box="[545,615,1549,1575]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="38" unit="m" value="400.0">400m</quantity>
, C.E.Carlton
<date id="DB12103EF40BFFD8774BF26355BEF994" box="[786,956,1549,1575]" pageId="1" pageNumber="38" value="2006-05-25">25 May 2006</date>
/. Deposited Louisiana State Arthropod Museum (
<collectionCode id="C9BDAE3BF40BFFD87554F25B5767F9FC" LSID="urn:lsid:biocol.org:col:33726" box="[269,357,1589,1615]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33726" name="Louisiana State Arthropod Museum" pageId="1" pageNumber="38">LSAM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="E7B66575F40BFFD8749CF2335761F974" pageId="1" pageNumber="38" type="etymology">
<paragraph id="AF1336FEF40BFFD8749CF2335761F974" blockId="1.[151,1437,1229,2015]" pageId="1" pageNumber="38">
<emphasis id="9DD8EAECF40BFFD8749CF2335750F9C4" bold="true" box="[197,338,1629,1655]" pageId="1" pageNumber="38">Etymology.</emphasis>
The specific epithet &quot;
<taxonomicName id="68AC4D7DF40BFFD8760CF23354F0F9C4" box="[597,754,1629,1655]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="species" species="septentrionis">septentrionis</taxonomicName>
&quot; (Latin, “the northern regions, the northern countries, the north”) is an adjective referring to the geographic range of the species, the most northern among known species of the genus.
</paragraph>
</subSubSection>
<subSubSection id="E7B66575F40BFFDA749CF2BB55D5FEE1" lastPageId="3" lastPageNumber="40" pageId="1" pageNumber="38" type="description">
<paragraph id="AF1336FEF40BFFD8749CF2BB530BF86C" blockId="1.[151,1437,1229,2015]" pageId="1" pageNumber="38">
<emphasis id="9DD8EAECF40BFFD8749CF2BB5757F95C" bold="true" box="[197,341,1749,1775]" pageId="1" pageNumber="38">Description</emphasis>
. Large for genus (ABL range 2.00
<quantity id="68549B1BF40BFFD8775AF2BB5567F95C" box="[771,869,1749,1775]" metricMagnitude="-3" metricUnit="m" metricValue="2.14" pageId="1" pageNumber="38" unit="mm" value="2.14">2.14mm</quantity>
, mean 2.07±
<quantity id="68549B1BF40BFFD877A4F2BB527AF95C" box="[1021,1144,1749,1775]" metricMagnitude="-5" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="38" unit="mm" value="0.053">0.053 mm</quantity>
, n=7). Sexes of approximately equal size: size of males ABL range 2.00
<quantity id="68549B1BF40BFFD876B1F2935548F8A4" box="[744,842,1789,1815]" metricMagnitude="-3" metricUnit="m" metricValue="2.14" pageId="1" pageNumber="38" unit="mm" value="2.14">2.14mm</quantity>
, mean 2.06±
<quantity id="68549B1BF40BFFD877BBF293525CF8A4" box="[994,1118,1789,1815]" metricMagnitude="-5" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="38" unit="mm" value="0.055">0.055 mm</quantity>
, n=5, size of females ABL range
<quantity id="68549B1BF40BFFD874BEF34B578FF88C" box="[231,397,1829,1855]" metricMagnitude="-3" metricUnit="m" metricValue="2.085" metricValueMax="2.13" metricValueMin="2.04" pageId="1" pageNumber="38" unit="mm" value="2.085" valueMax="2.13" valueMin="2.04">2.042.13mm</quantity>
, mean 2.09±
<quantity id="68549B1BF40BFFD8766BF34B54B7F88C" box="[562,693,1829,1855]" metricMagnitude="-5" metricUnit="m" metricValue="6.4" pageId="1" pageNumber="38" unit="mm" value="0.064">0.064 mm</quantity>
, n=2. Habitus (
<figureCitation id="37972A7BF40BFFD877D9F34B55CEF88C" box="[896,972,1829,1855]" captionStart="FIGURES 1 2" captionStartId="2.[151,269,1700,1724]" captionTargetBox="[168,1428,672,1675]" captionTargetId="figure@2.[151,1436,665,1678]" captionTargetPageId="2" captionText="FIGURES 1 2. Habitus of (1) Serranillus septentrionis, new species (VA, Botetourt Co., Black Horse Gap), and (2) Anillinus cherokee, new species (TN, Blount Co., Great Smoky Mountains National Park), dorsal aspect, males." httpUri="https://zenodo.org/record/181481/files/figure.png" pageId="1" pageNumber="38">Fig. 1</figureCitation>
) markedly convex, ovoid (WE/ABL 0.39+0.010), head normally proportioned for genus (WH/WPm 0.72+0.017), pronotum narrow compared to elytra (WPm/WE 0.79+0.011). Body color dark, from dark brunneous to brunneorufous, appendages much lighter, testaceous. Microsculpture fine but distinct across dorsal surface of pronotum and most of head. On head only the vertex lacks microsculpture. Elytra with well-developed polygonal microsculpture.
</paragraph>
<paragraph id="AF1336FEF408FFDB749CF4F654DAFE99" blockId="2.[151,1436,152,618]" pageId="2" pageNumber="39">Pronotum moderately convex and comparatively strongly transverse (WPm/LP 1.33+0.043), with margins rectilinear and moderately constricted posteriad (WPm/WPp 1.31+0.032). Anterior angles evident, slightly prominent. Posterior angles slightly obtuse (105110°). Width between posterior angles slightly greater than between anterior angles (WPa/WPp 0.97+0.027).</paragraph>
<paragraph id="AF1336FEF408FFDB749CF5565289FE41" blockId="2.[151,1436,152,618]" pageId="2" pageNumber="39">Elytra markedly convex, slightly depressed along suture, of normal length for genus (LE/ABL 0.58+0.015), with traces of 12 interneurs. Humeri moderately prominent, in outline forming an obtuse angle with longitudinal axis of body, dorsally moderately rounded, gradually enlarged to middle. Margins subparallel, slightly divergent in basal half, evenly rounded to apex, maximum width of elytra behind midpoint. Elytra without subapical sinuation. Vestiture of elytra long (about half length of discal setae).</paragraph>
<paragraph id="AF1336FEF408FFDB749CF66E54B3FDF1" blockId="2.[151,1436,152,618]" pageId="2" pageNumber="39">Prothoracic leg of males with strongly dilated tarsomere 1 and moderately dilated tarsomere 2. Profemur moderately swollen. Metafemora unmodified.</paragraph>
<paragraph id="AF1336FEF408FFDB749CF63E5572FDD9" blockId="2.[151,1436,152,618]" box="[197,880,592,618]" pageId="2" pageNumber="39">Males with three denticles on the apex of the sternum VII.</paragraph>
<caption id="FBD36676F408FFDB74CEF2CA5343F96D" httpUri="https://zenodo.org/record/181481/files/figure.png" pageId="2" pageNumber="39" targetBox="[168,1428,672,1675]" targetPageId="2">
<paragraph id="AF1336FEF408FFDB74CEF2CA5343F96D" blockId="2.[151,1436,1700,1758]" pageId="2" pageNumber="39">
<emphasis id="9DD8EAECF408FFDB74CEF2CA5747F90F" bold="true" box="[151,325,1700,1724]" pageId="2" pageNumber="39">FIGURES 12.</emphasis>
Habitus of (1)
<taxonomicName id="68AC4D7DF408FFDB75AAF2CA54FEF908" box="[499,764,1700,1723]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="septentrionis" status="sp. nov.">
<emphasis id="9DD8EAECF408FFDB75AAF2CA54FEF908" box="[499,764,1700,1723]" italics="true" pageId="2" pageNumber="39">Serranillus septentrionis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="86EB5797F408FFDB7755F2CA558DF908" box="[780,911,1700,1723]" pageId="2" pageNumber="39" rank="species">new species</taxonomicNameLabel>
(VA, Botetourt Co., Black Horse Gap), and (2)
<emphasis id="9DD8EAECF408FFDB74CEF2A95766F96D" box="[151,356,1735,1758]" italics="true" pageId="2" pageNumber="39">
<taxonomicName id="68AC4D7DF408FFDB74CEF2A95762F96D" box="[151,352,1735,1758]" class="Insecta" family="Carabidae" genus="Anillinus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="cherokee" status="sp. nov.">Anillinus cherokee</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="86EB5797F408FFDB7533F2A957E8F96D" box="[362,490,1735,1758]" pageId="2" pageNumber="39" rank="species">new species</taxonomicNameLabel>
(TN, Blount Co., Great Smoky Mountains National Park), dorsal aspect, males.
</paragraph>
</caption>
<paragraph id="AF1336FEF408FFDB749CF37C5318F817" blockId="2.[151,1437,1810,2036]" pageId="2" pageNumber="39">
Median lobe (
<figureCitation id="37972A7BF408FFDB753EF37C57AEF89F" box="[359,428,1810,1836]" captionStart="FIGURES 3 6" captionStartId="4.[151,269,1573,1597]" captionTargetBox="[252,1331,268,1544]" captionTargetId="figure@4.[248,1341,262,1547]" captionTargetPageId="4" captionText="FIGURES 3 6. Serranillus septentrionis, new species, (VA, Giles Co., Jefferson National Forest). Male aedeagus: 3 median lobe, 4 left paramere; female genitalia: 5 spermatheca, 6 right stylomere and sternum IX. Scale = 100 μm." httpUri="https://zenodo.org/record/181482/files/figure.png" pageId="2" pageNumber="39">Fig. 3</figureCitation>
) nearly straight, with elongate, gradually narrowing and somewhat acute apex, ventral margin with obvious striation and few poriferous canals. Canals absent on walls of median lobe itself. Copulatory sclerites represented by two weakly sclerotized plates, subquadrate in form and one after the other in position. Walls of invagination of apical orifice obviously sclerotized and forming a wavy contour.
</paragraph>
<paragraph id="AF1336FEF408FFDB749CF3DC52CFF847" blockId="2.[151,1437,1810,2036]" pageId="2" pageNumber="39">
Left paramere (
<figureCitation id="37972A7BF408FFDB7524F3DC57C7F87F" box="[381,453,1970,1996]" captionStart="FIGURES 3 6" captionStartId="4.[151,269,1573,1597]" captionTargetBox="[252,1331,268,1544]" captionTargetId="figure@4.[248,1341,262,1547]" captionTargetPageId="4" captionText="FIGURES 3 6. Serranillus septentrionis, new species, (VA, Giles Co., Jefferson National Forest). Male aedeagus: 3 median lobe, 4 left paramere; female genitalia: 5 spermatheca, 6 right stylomere and sternum IX. Scale = 100 μm." httpUri="https://zenodo.org/record/181482/files/figure.png" pageId="2" pageNumber="39">Fig. 4</figureCitation>
) with enlarged base, which is distinctly separated by a wide border from the distal part of the paramere; paramere apex without visible setae (at X400). Right paramere absent.
</paragraph>
<paragraph id="AF1336FEF409FFDA749CF4F655C8FE99" blockId="3.[151,1436,152,658]" pageId="3" pageNumber="40">
Spermatheca (
<figureCitation id="37972A7BF409FFDA7536F4F657BAFF01" box="[367,440,152,178]" captionStart="FIGURES 3 6" captionStartId="4.[151,269,1573,1597]" captionTargetBox="[252,1331,268,1544]" captionTargetId="figure@4.[248,1341,262,1547]" captionTargetPageId="4" captionText="FIGURES 3 6. Serranillus septentrionis, new species, (VA, Giles Co., Jefferson National Forest). Male aedeagus: 3 median lobe, 4 left paramere; female genitalia: 5 spermatheca, 6 right stylomere and sternum IX. Scale = 100 μm." httpUri="https://zenodo.org/record/181482/files/figure.png" pageId="3" pageNumber="40">Fig. 5</figureCitation>
) moderately sclerotized, with three well-developed parts. Distal part, the cornu, is sclerotized and bean-like in form, more than 2 times longer than wide. Proximal part of cornu unsclerotized and straight, with traces of coils. Ramus well-differentiated, sclerotized, ovoid. Nodulus short. Spermathecal duct comparatively short, more or less straight, without obvious coils.
</paragraph>
<paragraph id="AF1336FEF409FFDA749CF55655D5FEE1" blockId="3.[151,1436,152,658]" box="[197,983,312,338]" pageId="3" pageNumber="40">
Stylomeres and sternum IX as in
<figureCitation id="37972A7BF409FFDA7612F55654B3FEE1" box="[587,689,312,338]" captionStart="FIGURES 3 6" captionStartId="4.[151,269,1573,1597]" captionTargetBox="[252,1331,268,1544]" captionTargetId="figure@4.[248,1341,262,1547]" captionTargetPageId="4" captionText="FIGURES 3 6. Serranillus septentrionis, new species, (VA, Giles Co., Jefferson National Forest). Male aedeagus: 3 median lobe, 4 left paramere; female genitalia: 5 spermatheca, 6 right stylomere and sternum IX. Scale = 100 μm." httpUri="https://zenodo.org/record/181482/files/figure.png" pageId="3" pageNumber="40">Figure 6</figureCitation>
. Sternum with 89 setae.
</paragraph>
</subSubSection>
<subSubSection id="E7B66575F409FFDA749CF531573FFE79" pageId="3" pageNumber="40" type="distribution">
<paragraph id="AF1336FEF409FFDA749CF53155D4FEC9" blockId="3.[151,1436,152,658]" box="[197,982,351,378]" pageId="3" pageNumber="40">
<emphasis id="9DD8EAECF409FFDA749CF5315764FECA" bold="true" box="[197,358,351,377]" pageId="3" pageNumber="40">Distribution.</emphasis>
Known from Giles and Botetourt Counties, Virginia.
</paragraph>
<paragraph id="AF1336FEF409FFDA749CF5E9573FFE79" blockId="3.[151,1436,152,658]" pageId="3" pageNumber="40">
<emphasis id="9DD8EAECF409FFDA749CF5E9572FFE12" bold="true" box="[197,301,391,417]" pageId="3" pageNumber="40">Habitat.</emphasis>
Beetles were collected in litter of deciduous hardwood forest from a wide range of altitudes (
<quantity id="68549B1BF409FFDA74C6F5DE5733FE79" box="[159,305,432,458]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="11.0" metricValueMin="4.0" pageId="3" pageNumber="40" unit="m" value="750.0" valueMax="1100.0" valueMin="400.0">4001100m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="E7B66575F409FFDA749CF5B95216FD21" pageId="3" pageNumber="40" type="description">
<paragraph id="AF1336FEF409FFDA749CF5B95216FD21" blockId="3.[151,1436,152,658]" pageId="3" pageNumber="40">
<emphasis id="9DD8EAECF409FFDA749CF5B957D5FE42" bold="true" box="[197,471,471,497]" pageId="3" pageNumber="40">Differential diagnosis.</emphasis>
<taxonomicName id="68AC4D7DF409FFDA7587F5B654FDFE42" box="[478,767,472,497]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="septentrionis">
<emphasis id="9DD8EAECF409FFDA7587F5B654FDFE42" box="[478,767,472,497]" italics="true" pageId="3" pageNumber="40">Serranillus septentrionis</emphasis>
</taxonomicName>
in general outline is very similar to
<taxonomicName id="68AC4D7DF409FFDA70FBF5B6531FFE42" box="[1186,1309,472,497]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="jeanneli">
<emphasis id="9DD8EAECF409FFDA70FBF5B6531FFE42" box="[1186,1309,472,497]" italics="true" pageId="3" pageNumber="40">S. jeanneli</emphasis>
</taxonomicName>
, but easily distinguished from the latter by the form of the apex of the median lobe. The latter species has comparatively shorter and subparallel apex of median lobe with a rounded tip. In comparison with
<taxonomicName id="68AC4D7DF409FFDA702DF64652FDFDF2" box="[1140,1279,552,577]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="dunavani">
<emphasis id="9DD8EAECF409FFDA702DF64652FDFDF2" box="[1140,1279,552,577]" italics="true" pageId="3" pageNumber="40">S. dunavani</emphasis>
</taxonomicName>
the new species is noticeably larger and has a comparatively smaller head. Additionally, the range of the new species is far north of the ranges of both previously known species of
<taxonomicName id="68AC4D7DF409FFDA7775F61655ACFD22" box="[812,942,632,657]" class="Insecta" family="Carabidae" genus="Serranillus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="9DD8EAECF409FFDA7775F61655ACFD22" box="[812,942,632,657]" italics="true" pageId="3" pageNumber="40">Serranillus</emphasis>
</taxonomicName>
(
<figureCitation id="37972A7BF409FFDA77E7F6165207FD21" box="[958,1029,632,658]" captionStart="FIGURE 7" captionStartId="5.[151,255,1880,1904]" captionTargetBox="[151,1435,986,1859]" captionTargetId="figure@5.[151,1436,986,1860]" captionTargetPageId="5" captionText="FIGURE 7. Schematic distribution of the Serranillus species. 1 S. jeanneli Barr; 2 S. dunavani (Jeannel); 3 S. septentrionis, new species. From Barr (1995) and original data." httpUri="https://zenodo.org/record/181483/files/figure.png" pageId="3" pageNumber="40">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>