370 lines
58 KiB
XML
370 lines
58 KiB
XML
<document id="D2D1FC38A3724E666BB96873A9DDA0AB" ID-DOI="10.37520/fi.2023.006" ID-ISSN="2533-4069" ID-Zenodo-Dep="10480778" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1704918079491" checkinUser="felipe" docAuthor="Bomfleur, Benjamin, Hedenäs, Lars, Friis, Else Marie, Crane, Peter R., Raunsgaard, Kaj, Pedersen, Mendes, Mário Miguel & Kvaček, Jiří" docDate="2023" docId="1A34ED3FF339FFA3FC32359BFB59EBBD" docLanguage="en" docName="FossilImprint.79.2.103-125.pdf" docOrigin="Fossil Imprint 79 (2)" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Polytrichastrum incurvum HEDENAS, BOMFLEUR et E. M. FRIIS 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="107" masterDocId="E60D9547F33BFFA7FFB43170FFC3ED45" masterDocTitle="Fossil Mosses From The Early Cretaceous Catefica Mesofossil Flora, Portugal - A Window Into The Mesozoic History Of Bryophytes" masterLastPageNumber="125" masterPageNumber="103" pageNumber="105" updateTime="1704923666188" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
|
||
<mods:mods id="5D4D50A71B3A598C75579BEECC9608C7" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="1C6F3C7CD964370535BFADC08B3E4A22">
|
||
<mods:title id="5681747D4745A3B3983748D03A5309D5">Fossil Mosses From The Early Cretaceous Catefica Mesofossil Flora, Portugal - A Window Into The Mesozoic History Of Bryophytes</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="F124BC714F15A73BE489BE7EC88CE83E" type="personal">
|
||
<mods:role id="187BA4B29A7BD3E7CD64A7A01E27F3C4">
|
||
<mods:roleTerm id="B45F7F2706D577E85938F5C9DCEEA823">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="6B907F50CAB254A514BCB938106B609C">Bomfleur, Benjamin</mods:namePart>
|
||
<mods:affiliation id="254C045A12178E18F2965EF50C12D4AC">Institute of Geology and Palaeontology, Palaeobotany, University of MÜnster, Heisenbergstr. 2, D- 48149 MÜnster, Germany; e-mail:</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="F1D03F5A2966097177CC51E2EB5BD01D" type="personal">
|
||
<mods:role id="029CC313A9295F7A58FEB8D0E623D888">
|
||
<mods:roleTerm id="E38C0E0B76FADC75789082DBF1B5DBAF">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="7614CAB7DA4DC46E4A3C129DF5635335">Hedenäs, Lars</mods:namePart>
|
||
<mods:affiliation id="4830550255E7383AFCBEA0012F8983A7">Department of Botany, Swedish Museum of Natural History, Box 50007, SE- 104 05 Stockholm, Sweden; e-mail: lars. hedenas @ nrm. se.</mods:affiliation>
|
||
<mods:nameIdentifier id="C0E30F907133A4A36E81D7BC46679DD2" type="email">lars.hedenas@nrm.se.</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="444F36601ECA9A6FDE9556BBE62992DE" type="personal">
|
||
<mods:role id="E1B527C9F7DD4B4C7F4A5F0DBBFE5CB6">
|
||
<mods:roleTerm id="55FF8FE31657ACC88C2E8B13143A5598">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2D8F7DDFF1E36FE6495D53045FB2FAEF">Friis, Else Marie</mods:namePart>
|
||
<mods:affiliation id="98772DE9463090F05E52F0056AF220C7">Department of Geoscience, University of Aarhus, HØegh-Guldbergs Gade 2, DK- 8000 Aarhus C Denmark. & Department of Palaeobiology, Swedish Museum of Natural History, Box 50007, SE- 104 05 Stockholm, Sweden.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="146B3D7FB31C409752B787711FF42C83" type="personal">
|
||
<mods:role id="67EE137179A14C8D8FD09002416E669B">
|
||
<mods:roleTerm id="8AA27FE372ABFB1A2D8E11FEACEBD492">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="FEBEFC0E1F3C825EFE61BAA25737556B">Crane, Peter R.</mods:namePart>
|
||
<mods:affiliation id="D0077C9A7B71E146C151BA5C64E3EC45">Oak Spring Garden Foundation, 1776 Loughborough Lane, Upperville, Virginia 20184 USA. & Yale School of Environment, Yale University, New Haven, Connecticut 06511 USA.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="E4CF7CC71061E5631E030C9576E03DEE" type="personal">
|
||
<mods:role id="8DDEC3C89AE1962412A4172A81694790">
|
||
<mods:roleTerm id="4A4468F67FA11326C7E386BD49D53DC9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8480C6D92F38309C5372EC8CF5FD54D0">Raunsgaard, Kaj</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="A66960118F0AC207CDC4E97EEE43EB77" type="personal">
|
||
<mods:role id="41D434AB761E90B343F0A179B1783434">
|
||
<mods:roleTerm id="33E8A96E4F5870B27AF0C94C20280079">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="9984EE25FCA3C4602DB2E0B93307C400">Pedersen</mods:namePart>
|
||
<mods:affiliation id="2345C2F6290403916BA5115E06893EE4">Department of Geoscience, University of Aarhus, HØegh-Guldbergs Gade 2, DK- 8000 Aarhus C Denmark.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="D7F7984951A488F501BABA9AA3D97E9F" type="personal">
|
||
<mods:role id="9C9547D1F040FD67640CE9A0F5C80F42">
|
||
<mods:roleTerm id="E91010B75B9A361A6295DC5AE65F478F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E985140024D03FEF0FB634D3B82293C1">Mendes, Mário Miguel</mods:namePart>
|
||
<mods:affiliation id="EBAE49A4F2BE9F59A372B2404C43841E">University of Coimbra, MARE - Marine and Environmental Sciences Centre / ARNET - Aquatic Research Network, Earth Sciences & Fernando Pessoa University, Praça 9 de Abril 349, Porto 4249 - 004, Portugal.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="6098C9F31EC6819AD2961BE72D4D2DBB" type="personal">
|
||
<mods:role id="EAF85245A0A2B94905438972E3F81CF6">
|
||
<mods:roleTerm id="EAB41F0A2929E55D14825B81523FB292">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B66390B1A6210F40BAE106728EEA4587">Kvaček, Jiří</mods:namePart>
|
||
<mods:affiliation id="3BED940FCB7D6190B03B48BD0212CCDE">National Museum Prague, Václavské nám. 68, 115 79 Praha 1, Czech Republic. * corresponding author</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="1183722DA8BA56A27D33855C5161263E">text</mods:typeOfResource>
|
||
<mods:relatedItem id="18977EA6C329E2E56EF7C21C011446F0" type="host">
|
||
<mods:titleInfo id="B49A16FEA84FACBEB85B38CE8CFAC691">
|
||
<mods:title id="3F9788AA6DCA5873B96A81B022F53BA8">Fossil Imprint</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="D6F5BE9B1E424C459AE41C8F16C7D3E1">
|
||
<mods:date id="D3010A89A56515D1E75E85729D1B6AA1">2023</mods:date>
|
||
<mods:detail id="3C00F96166FBC74F8FF510D95A1C222E" type="volume">
|
||
<mods:number id="57EEE5844F7F384DD6C382D0D61A2CAE">79</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="FC28E4B6D6D612FAED8AB36CF8F303A1" type="issue">
|
||
<mods:number id="A6A3C2F52015FC2BB34B9A744353C5AF">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="827DF69D1B68F196DFB0CB449AFCD3E6" unit="page">
|
||
<mods:start id="63F2DD7C51812F051789C13425AE10B8">103</mods:start>
|
||
<mods:end id="78181DC8CA37886CD9C35C0ADC505462">125</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="DCE1B92452342C33F09093423684C38A">journal article</mods:classification>
|
||
<mods:identifier id="96463B103DCC2E1316B2B63B8CC5B632" type="DOI">10.37520/fi.2023.006</mods:identifier>
|
||
<mods:identifier id="5D9C6F7EA9681054CD19868795A5C660" type="ISSN">2533-4069</mods:identifier>
|
||
<mods:identifier id="9877396C222BBE5FF14718FCB0EA6F07" type="Zenodo-Dep">10480778</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="1A34ED3FF339FFA3FC32359BFB59EBBD" LSID="urn:lsid:plazi:treatment:1A34ED3FF339FFA3FC32359BFB59EBBD" httpUri="http://treatment.plazi.org/id/1A34ED3FF339FFA3FC32359BFB59EBBD" lastPageId="4" lastPageNumber="107" pageId="2" pageNumber="105">
|
||
<subSubSection id="DA870FA2F339FFA5FC32359BFB3DE863" pageId="2" pageNumber="105" type="nomenclature">
|
||
<paragraph id="92225C29F339FFA5FC32359BFB3DE863" blockId="2.[902,1422,1259,1318]" pageId="2" pageNumber="105">
|
||
<heading id="C96AEB45F339FFA5FC32359BFB3DE863" centered="true" fontSize="8" level="2" pageId="2" pageNumber="105" reason="2">
|
||
<emphasis id="A0E9803BF339FFA5FC32359BFB3DE863" bold="true" pageId="2" pageNumber="105">
|
||
<taxonomicName id="559D27AAF339FFA5FC32359BFB62E861" authority="HEDENAS, BOMFLEUR et E.M.FRIIS" authorityName="HEDENAS, BOMFLEUR et E. M. FRIIS" authorityYear="2023" class="Bryopsida" family="Polytrichaceae" genus="Polytrichastrum" kingdom="Plantae" order="Polytrichales" pageId="2" pageNumber="105" phylum="Bryophyta" rank="species" species="incurvum" status="sp. nov.">
|
||
<emphasis id="A0E9803BF339FFA5FC32359BFB6AE846" bold="true" box="[902,1193,1259,1283]" italics="true" pageId="2" pageNumber="105">Polytrichastrum incurvum</emphasis>
|
||
<smallCapsWord id="94C4CAF5F339FFA5FB04359CFAD3E846" baselines="1278,1278" box="[1200,1296,1260,1284]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Hedenas" pageId="2" pageNumber="105">HEDENÄS</smallCapsWord>
|
||
,
|
||
<smallCapsWord id="94C4CAF5F339FFA5FAA9359CFA4DE847" baselines="1278,1278" box="[1309,1422,1260,1284]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Bomfleur" pageId="2" pageNumber="105">BOMFLEUR</smallCapsWord>
|
||
et E.M.
|
||
<smallCapsWord id="94C4CAF5F339FFA5FBDF347EFB62E861" baselines="1312,1312" box="[1131,1185,1294,1318]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Friis" pageId="2" pageNumber="105">FRIIS</smallCapsWord>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BBDA3D40F339FFA5FB1C347EFB3DE863" box="[1192,1278,1294,1318]" pageId="2" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DA870FA2F339FFA3FBA93441FB59EBBD" lastPageId="4" lastPageNumber="107" pageId="2" pageNumber="105" type="description">
|
||
<paragraph id="92225C29F339FFA5FBA93441FB34E80C" blockId="2.[1053,1271,1329,1353]" box="[1053,1271,1329,1353]" pageId="2" pageNumber="105">
|
||
<figureCitation id="0AA640ACF339FFA5FBA93441FB7AE80C" box="[1053,1209,1329,1353]" captionStart="Text-fig" captionStartId="5.[125,206,1270,1291]" captionTargetBox="[125,1439,151,1254]" captionTargetId="figure-267@5.[125,1439,151,1254]" captionTargetPageId="5" captionText="Text-fig. 3. Scanning electron microscope (SEM, a–e) and synchrotron radiation X-ray tomographic microscopy (SRXTM, f–g) images of a shoot and leaves of Polytrichastrum incurvum HEDENÄS, BOMFLEUR et E.M.FRIIS from the Early Cretaceous Catefica mesofossil flora, Almargem Formation, Portugal. a: Shoot apex showing several incurved leaves. b: Leaf showing ventral longitudinal lamellae (arrow) and incurved leaf margins. c: Detail of longitudinal lamellae on the ventral leaf surface (arrow) and the incurved leaf margin. d: Transverse fracture through the upper portion of leaf showing lamellae on the ventral surface (arrows). e: Detail of upper surface of three longitudinal lamellae showing outlines of individual cells. f: Transverse section (orthoslice xy0655) of shoot and several leaves at different levels. Note the lamellae on the ventral surface over the costa in sections in the distal part of the leaf (arrow) and the mostly unistratose leaf lamina with smooth cells (arrowheads) and lacking lamellae in the proximal part of the leaf. g: Longitudinal section (orthoslice yz1435) showing shoot with several leaves. Specimen, Catefica 49-S174251 (a–g; holotype). Scale bars = 1 mm (a), 200 µm (b, f, g), 100 µm (c), 50 µm (d), 10 µm (e)." figureDoi="http://doi.org/10.5281/zenodo.10480784" httpUri="https://zenodo.org/record/10480784/files/figure.png" pageId="2" pageNumber="105">Text-figs 3a–g</figureCitation>
|
||
,
|
||
<figureCitation id="0AA640ACF339FFA5FB773441FB34E80C" box="[1219,1271,1329,1353]" captionStart="Text-fig" captionStartId="6.[148,229,816,837]" captionTargetBox="[148,1462,151,801]" captionTargetId="figure-561@6.[148,1462,151,801]" captionTargetPageId="6" captionText="Text-fig. 4. Synchrotron radiation X-ray tomographic microscopy (SRXTM) images (volume renderings, a, b; orthoslices, c, d) of a shoot and leaves of Polytrichastrum incurvum HEDENÄS, BOMFLEUR et E.M.FRIIS from the Early Cretaceous Catefica mesofossil flora, Almargem Formation, Portugal. a, b: Shoot apex showing incurved leaves with distinctly incurved leaf margins. Note a section through the proximal part of a leaf (b) showing lamellae (arrow) over the costa and the unistratose leaf lamina (arrow head). c: Transverse section (orthoslice xy1440) of a shoot showing sections through the distal portions of six leaves showing the distinct lamellae (arrows). d: Transverse section (orthoslice xy2160) of a shoot showing sections through the proximal portions of several leaves showing the unistratose leaf lamina (arrow) with smooth cells. Specimen, Catefica 49-S266392 (a–d). Scale bars = 1 mm (a), 500 µm (b), 200 µm (c, d)." figureDoi="http://doi.org/10.5281/zenodo.10480786" httpUri="https://zenodo.org/record/10480786/files/figure.png" pageId="2" pageNumber="105">4a–d</figureCitation>
|
||
</paragraph>
|
||
<paragraph id="92225C29F339FFA5FCC53413FA09E83E" blockId="2.[881,1482,1379,1403]" box="[881,1482,1379,1403]" pageId="2" pageNumber="105">H o l o t y p e. S174251 (Catefica sample 49; figured</paragraph>
|
||
<paragraph id="92225C29F339FFA5FCFF34F6FC33E8DB" blockId="2.[843,1008,1414,1438]" box="[843,1008,1414,1438]" pageId="2" pageNumber="105">
|
||
<figureCitation id="0AA640ACF339FFA5FCFF34F6FC27E8DB" box="[843,996,1414,1438]" captionStart="Text-fig" captionStartId="5.[125,206,1270,1291]" captionTargetBox="[125,1439,151,1254]" captionTargetId="figure-267@5.[125,1439,151,1254]" captionTargetPageId="5" captionText="Text-fig. 3. Scanning electron microscope (SEM, a–e) and synchrotron radiation X-ray tomographic microscopy (SRXTM, f–g) images of a shoot and leaves of Polytrichastrum incurvum HEDENÄS, BOMFLEUR et E.M.FRIIS from the Early Cretaceous Catefica mesofossil flora, Almargem Formation, Portugal. a: Shoot apex showing several incurved leaves. b: Leaf showing ventral longitudinal lamellae (arrow) and incurved leaf margins. c: Detail of longitudinal lamellae on the ventral leaf surface (arrow) and the incurved leaf margin. d: Transverse fracture through the upper portion of leaf showing lamellae on the ventral surface (arrows). e: Detail of upper surface of three longitudinal lamellae showing outlines of individual cells. f: Transverse section (orthoslice xy0655) of shoot and several leaves at different levels. Note the lamellae on the ventral surface over the costa in sections in the distal part of the leaf (arrow) and the mostly unistratose leaf lamina with smooth cells (arrowheads) and lacking lamellae in the proximal part of the leaf. g: Longitudinal section (orthoslice yz1435) showing shoot with several leaves. Specimen, Catefica 49-S174251 (a–g; holotype). Scale bars = 1 mm (a), 200 µm (b, f, g), 100 µm (c), 50 µm (d), 10 µm (e)." figureDoi="http://doi.org/10.5281/zenodo.10480784" httpUri="https://zenodo.org/record/10480784/files/figure.png" pageId="2" pageNumber="105">Text-fig. 3a–g</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="92225C29F339FFA5FCC534C8FA09E88A" blockId="2.[843,1482,1464,1522]" box="[881,1482,1464,1488]" pageId="2" pageNumber="105">P l a n t F o s s i l N a m e s R e g i s t r y N u m b e r.</paragraph>
|
||
<paragraph id="92225C29F339FFA5FCFF34AAFC10E8B7" blockId="2.[843,1482,1464,1522]" box="[843,979,1498,1522]" pageId="2" pageNumber="105">PFN003145.</paragraph>
|
||
<paragraph id="92225C29F339FFA5FCC5377DFB71EB02" blockId="2.[843,1481,1548,1607]" pageId="2" pageNumber="105">
|
||
P a r a t y p e s. S266391, S266392 [Catefica sample 49; figured
|
||
<figureCitation id="0AA640ACF339FFA5FC29375FFBF1EB02" box="[925,1074,1583,1607]" captionStart="Text-fig" captionStartId="6.[148,229,816,837]" captionTargetBox="[148,1462,151,801]" captionTargetId="figure-561@6.[148,1462,151,801]" captionTargetPageId="6" captionText="Text-fig. 4. Synchrotron radiation X-ray tomographic microscopy (SRXTM) images (volume renderings, a, b; orthoslices, c, d) of a shoot and leaves of Polytrichastrum incurvum HEDENÄS, BOMFLEUR et E.M.FRIIS from the Early Cretaceous Catefica mesofossil flora, Almargem Formation, Portugal. a, b: Shoot apex showing incurved leaves with distinctly incurved leaf margins. Note a section through the proximal part of a leaf (b) showing lamellae (arrow) over the costa and the unistratose leaf lamina (arrow head). c: Transverse section (orthoslice xy1440) of a shoot showing sections through the distal portions of six leaves showing the distinct lamellae (arrows). d: Transverse section (orthoslice xy2160) of a shoot showing sections through the proximal portions of several leaves showing the unistratose leaf lamina (arrow) with smooth cells. Specimen, Catefica 49-S266392 (a–d). Scale bars = 1 mm (a), 500 µm (b), 200 µm (c, d)." figureDoi="http://doi.org/10.5281/zenodo.10480786" httpUri="https://zenodo.org/record/10480786/files/figure.png" pageId="2" pageNumber="105">Text-fig. 4a–d</figureCitation>
|
||
(S266392)]
|
||
</paragraph>
|
||
<paragraph id="92225C29F339FFA5FCC53711FA09EB3C" blockId="2.[881,1482,1633,1657]" box="[881,1482,1633,1657]" pageId="2" pageNumber="105">R e p o s i t o r y. Palaeobotanical Collections,</paragraph>
|
||
<paragraph id="92225C29F339FFA5FCFF37F4FB40EBFB" blockId="2.[843,1482,1668,1726]" pageId="2" pageNumber="105">
|
||
Department of Palaeobiology, Swedish Museum of Natural History,
|
||
<collectingRegion id="505992CBF339FFA5FC1237D6FBDFEBFB" box="[934,1052,1702,1726]" country="Sweden" name="Stockholm" pageId="2" pageNumber="105">Stockholm</collectingRegion>
|
||
,
|
||
<collectingCountry id="EA8A1CB9F339FFA5FB9337D6FBBCEBFB" box="[1063,1151,1702,1726]" name="Sweden" pageId="2" pageNumber="105">Sweden</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="92225C29F339FFA5FCC537A9FBF9EA56" blockId="2.[843,1482,1752,1811]" pageId="2" pageNumber="105">E t y m o l o g y. The specific epithet refers to the incurved leaf margins.</paragraph>
|
||
<paragraph id="92225C29F339FFA5FCC5365DFA09EA00" blockId="2.[881,1482,1836,1861]" box="[881,1482,1836,1861]" pageId="2" pageNumber="105">
|
||
<materialsCitation id="22F55674F339FFA5FCC5365DFA09EA00" box="[881,1482,1836,1861]" collectionCode="T" latitude="39.058334" pageId="2" pageNumber="105" specimenCount="1">
|
||
<collectionCode id="F48CC4ECF339FFA5FCC5365DFC42EA01" box="[881,897,1837,1860]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="105">T</collectionCode>
|
||
y p e l o c a l i t y. Catefica (
|
||
<geoCoordinate id="F7A93AEEF339FFA5FB01365DFA8EEA01" box="[1205,1357,1836,1861]" degrees="39" direction="north" minutes="03" orientation="latitude" pageId="2" pageNumber="105" precision="15" seconds="30" value="39.058334">39° 03ʹ 30ʺ N</geoCoordinate>
|
||
, 09°14ʹ 30ʺ
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="92225C29F339FFA5FCFF3620FB11EACF" blockId="2.[843,1481,1872,1930]" pageId="2" pageNumber="105">
|
||
W), between the villages of Catefica and Mugideira, about
|
||
<quantity id="5565F1CCF339FFA5FA083620FCAEEACF" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="105" unit="km" value="4.0">4 km</quantity>
|
||
south of Torres Vedras,
|
||
<collectingCountry id="EA8A1CB9F339FFA5FBC43602FB0DEACF" box="[1136,1230,1906,1930]" name="Portugal" pageId="2" pageNumber="105">Portugal</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="92225C29F339FFA5FCC536D5FA09EAF9" blockId="2.[843,1482,1956,2015]" box="[881,1482,1956,1981]" pageId="2" pageNumber="105">T y p e s t r a t u m a n d a g e. Almargem Formation,</paragraph>
|
||
<paragraph id="92225C29F339FFA5FCFF36B7FB38EA9A" blockId="2.[843,1482,1956,2015]" box="[843,1275,1991,2015]" pageId="2" pageNumber="105">Early Cretaceous (Aptian–early Albian).</paragraph>
|
||
<paragraph id="92225C29F339FFA5FCC53689FA0AE554" blockId="2.[843,1482,2041,2100]" box="[881,1481,2041,2066]" pageId="2" pageNumber="105">
|
||
S p e c i f i c d i a g n o s i s.A species of
|
||
<taxonomicName id="559D27AAF339FFA5FAA83689FA0AE554" box="[1308,1481,2041,2065]" class="Bryopsida" family="Polytrichaceae" genus="Polytrichastrum" kingdom="Plantae" order="Polytrichales" pageId="2" pageNumber="105" phylum="Bryophyta" rank="genus">
|
||
<emphasis id="A0E9803BF339FFA5FAA83689FA0AE554" box="[1308,1481,2041,2065]" italics="true" pageId="2" pageNumber="105">Polytrichastrum</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="92225C29F339FFA4FCFF396CFE40EB85" blockId="2.[843,1482,2041,2100]" lastBlockId="3.[106,387,1704,1728]" lastPageId="3" lastPageNumber="106" pageId="2" pageNumber="105">
|
||
G.L.
|
||
<smallCapsWord id="94C4CAF5F339FFA5FCC8396CFC5FE574" baselines="2094,2093" box="[892,924,2076,2100]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Sm" pageId="2" pageNumber="105">SM</smallCapsWord>
|
||
. with distinctly incurved margins of the leaf limb and upper cells of the lamellae that are thin-walled or only slightly thickened distally.
|
||
</paragraph>
|
||
<caption id="C6E20CA1F338FFA4FFC93451FC85EB7B" ID-DOI="http://doi.org/10.5281/zenodo.10480780" ID-Zenodo-Dep="10480780" httpUri="https://zenodo.org/record/10480780/files/figure.png" pageId="3" pageNumber="106" startId="3.[125,206,1313,1334]" targetBox="[125,1439,151,1297]" targetPageId="3" targetType="figure">
|
||
<paragraph id="92225C29F338FFA4FFC93451FC85EB7B" blockId="3.[125,1440,1313,1598]" pageId="3" pageNumber="106">
|
||
<emphasis id="A0E9803BF338FFA4FFC93451FE31E816" bold="true" pageId="3" pageNumber="106">Text-fig. 1. Scanning electron microscope (SEM, a–d) and synchrotron radiation X-ray tomographic microscopy (SRXTM, e–g) images of shoots and leaves of</emphasis>
|
||
<taxonomicName id="559D27AAF338FFA4FE4F344EFBBEE817" authority="HEDENAS, BOMFLEUR et E. M. FRIIS" authorityName="HEDENAS, BOMFLEUR et E. M. FRIIS" authorityYear="2023" box="[507,1149,1342,1363]" class="Bryopsida" family="Ditrichaceae" genus="Chlorosphagnum" kingdom="Plantae" order="Dicranales" pageId="3" pageNumber="106" phylum="Bryophyta" rank="species" species="cateficense">
|
||
<emphasis id="A0E9803BF338FFA4FE4F344EFBBEE817" bold="true" box="[507,1149,1342,1363]" pageId="3" pageNumber="106">
|
||
<emphasis id="A0E9803BF338FFA4FE4F344EFCDCE816" bold="true" box="[507,799,1342,1363]" italics="true" pageId="3" pageNumber="106">Chlorosphagnum cateficense</emphasis>
|
||
<smallCapsWord id="94C4CAF5F338FFA4FC9C344EFCBCE817" baselines="1358,1358" box="[808,895,1342,1363]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Hedenas" pageId="3" pageNumber="106">HEDENÄS</smallCapsWord>
|
||
,
|
||
<smallCapsWord id="94C4CAF5F338FFA4FC3A344EFC37E817" baselines="1358,1358" box="[910,1012,1342,1363]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Bomfleur" pageId="3" pageNumber="106">BOMFLEUR</smallCapsWord>
|
||
et E.M.
|
||
<smallCapsWord id="94C4CAF5F338FFA4FBFF344EFBBEE817" baselines="1358,1358" box="[1099,1149,1342,1363]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Friis" pageId="3" pageNumber="106">FRIIS</smallCapsWord>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
from the Early Cretaceous Catefica mesofossil flora, Almargem Formation, Portugal. a, b: Shoot apices with leaves showing rows of collapsed cell walls between the transverse fibrils of the hyaline cells. c: Detail of leaf surface showing cell pattern with many hyaline cells collapsed. d: Transverse sections of leaves showing differentiation into hyaline (arrows) and enclosed chlorophyllose cells (arrow heads) of similar size. e: Transverse section (orthoslice xy0550) of shoot apex with several leaves showing differentiation into hyaline (arrows) and chlorophyllose cells (arrow heads) of similar size. f: Longitudinal section (orthoslice yz0258) of shoot apex with leaves. g: Longitudinal section and partial surface view (orthoslice xz0430) of leaf showing chlorophyllose cells and also hyaline cells, some with transverse fibrils. Specimens, Catefica 49-S156535 (a, e; holotype), Catefica 49-S156534 (b, c, f, g), Catefica 49- S107796 (d). Scale bars = 500 µm (a, b, f), 100 µm (c, d), 200 µm (e, g).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="92225C29F338FFA4FF2437ACFD2AEAE0" blockId="3.[144,744,1755,1780]" lastBlockId="3.[106,745,1933,1957]" pageId="3" pageNumber="106">
|
||
D i s t i n g u i s h i n g f e a t u r e s.
|
||
<taxonomicName id="559D27AAF338FFA4FD8F37ABFF0CEA53" authorityName="HEDENAS, BOMFLEUR et E. M. FRIIS" authorityYear="2023" class="Bryopsida" family="Polytrichaceae" genus="Polytrichastrum" kingdom="Plantae" order="Polytrichales" pageId="3" pageNumber="106" phylum="Bryophyta" rank="species" species="incurvum">
|
||
<emphasis id="A0E9803BF338FFA4FD8F37ABFF0CEA53" italics="true" pageId="3" pageNumber="106">Polytrichastrum incurvum</emphasis>
|
||
</taxonomicName>
|
||
differs from all extant species of
|
||
<taxonomicName id="559D27AAF338FFA4FD8F378FFD2BEA52" box="[571,744,1791,1815]" class="Bryopsida" family="Polytrichaceae" genus="Polytrichastrum" kingdom="Plantae" order="Polytrichales" pageId="3" pageNumber="106" phylum="Bryophyta" rank="genus">
|
||
<emphasis id="A0E9803BF338FFA4FD8F378FFD2BEA52" box="[571,744,1791,1815]" italics="true" pageId="3" pageNumber="106">Polytrichastrum</emphasis>
|
||
</taxonomicName>
|
||
in having thin-walled or only slightly distally thickened upper cells of the lamella. Among fossil taxa,
|
||
<taxonomicName id="559D27AAF338FFA4FDAF3637FD5FEA1B" authorityName="HEDENAS, BOMFLEUR et E. M. FRIIS" authorityYear="2023" box="[539,668,1863,1886]" class="Bryopsida" family="Polytrichaceae" genus="Polytrichastrum" kingdom="Plantae" order="Polytrichales" pageId="3" pageNumber="106" phylum="Bryophyta" rank="species" species="incurvum">
|
||
<emphasis id="A0E9803BF338FFA4FDAF3637FD5FEA1B" box="[539,668,1863,1886]" italics="true" pageId="3" pageNumber="106">P. incurvum</emphasis>
|
||
</taxonomicName>
|
||
differs from the gametophytes associated with
|
||
<taxonomicName id="559D27AAF338FFA4FDFE361AFD21EAE1" authority="KONOPKA, HEREND., G. L. MERR. et P. R. CRANE" authorityName="KONOPKA, HEREND., G. L. MERR. et P. R. CRANE" class="Bryopsida" family="Polytrichaceae" genus="Eopolytrichum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Polytrichales" pageId="3" pageNumber="106" phylum="Bryophyta" rank="species" species="antiquum">
|
||
<emphasis id="A0E9803BF338FFA4FDFE361AFF0CEAE0" italics="true" pageId="3" pageNumber="106">Eopolytrichum antiquum</emphasis>
|
||
<smallCapsWord id="94C4CAF5F338FFA4FF5436FEFE82EAE1" baselines="1951,1952" box="[224,321,1934,1957]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Konopka" pageId="3" pageNumber="106">KONOPKA</smallCapsWord>
|
||
,
|
||
<smallCapsWord id="94C4CAF5F338FFA4FEED36FEFE69EAE6" baselines="1951,1951" box="[345,426,1934,1957]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Herend" pageId="3" pageNumber="106">HEREND</smallCapsWord>
|
||
., G.L.
|
||
<smallCapsWord id="94C4CAF5F338FFA4FE4336FDFDF5EAE6" baselines="1951,1951" box="[503,566,1933,1957]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Merr" pageId="3" pageNumber="106">MERR</smallCapsWord>
|
||
. et P.R.
|
||
<smallCapsWord id="94C4CAF5F338FFA4FD2936FDFD21EAE1" baselines="1951,1952" box="[669,738,1933,1957]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Crane" pageId="3" pageNumber="106">CRANE</smallCapsWord>
|
||
</taxonomicName>
|
||
,
|
||
</paragraph>
|
||
<paragraph id="92225C29F338FFA4FFDE36C1FD2AEA8C" blockId="3.[106,745,1969,1993]" box="[106,745,1969,1993]" pageId="3" pageNumber="106">which is of Late Cretaceous (Santonian-Campanian) age</paragraph>
|
||
<paragraph id="92225C29F338FFA4FFDE36A4FD2AE555" blockId="3.[106,744,2004,2029]" lastBlockId="3.[106,745,2040,2064]" pageId="3" pageNumber="106">
|
||
(Konopka et al. 1997), in having widely incurved marginal laminae in the leaf limb.
|
||
<taxonomicName id="559D27AAF338FFA4FECE3689FD21E54A" authority="BIPPUS" authorityName="BIPPUS" box="[378,738,2040,2064]" class="Polytrichopsida" family="Polytrichaceae" genus="Meantoinea" higherTaxonomySource="CoL,GBIF" kingdom="Plantae" order="Polytrichales" pageId="3" pageNumber="106" phylum="Bryophyta" rank="species" species="alophosioides">
|
||
<emphasis id="A0E9803BF338FFA4FECE3689FD56E555" box="[378,661,2040,2064]" italics="true" pageId="3" pageNumber="106">Meantoinea alophosioides</emphasis>
|
||
<smallCapsWord id="94C4CAF5F338FFA4FD293689FD21E54A" baselines="2058,2059" box="[669,738,2041,2064]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Bippus" pageId="3" pageNumber="106">BIPPUS</smallCapsWord>
|
||
</taxonomicName>
|
||
,
|
||
</paragraph>
|
||
<paragraph id="92225C29F338FFA4FFDE396CFD2AE571" blockId="3.[106,745,2076,2100]" box="[106,745,2076,2100]" pageId="3" pageNumber="106">
|
||
<smallCapsWord id="94C4CAF5F338FFA4FFDE396CFF07E577" baselines="2094,2094" box="[106,196,2076,2100]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Stockey" pageId="3" pageNumber="106">STOCKEY</smallCapsWord>
|
||
, G.W.
|
||
<smallCapsWord id="94C4CAF5F338FFA4FEBB396CFEBFE577" baselines="2094,2094" box="[271,380,2076,2100]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Rothwell" pageId="3" pageNumber="106">ROTHWELL</smallCapsWord>
|
||
et
|
||
<smallCapsWord id="94C4CAF5F338FFA4FE1E396CFDCBE577" baselines="2093,2094" box="[426,520,2076,2099]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Tomescu" pageId="3" pageNumber="106">TOMESCU</smallCapsWord>
|
||
, which is of Early
|
||
</paragraph>
|
||
<paragraph id="92225C29F338FFA4FC8037F5FAE2EA15" blockId="3.[820,1459,1669,1693]" lastBlockId="3.[820,1313,1848,1872]" pageId="3" pageNumber="106">
|
||
Cretaceous (Valanginian) age (Bippus et al. 2017), differs in having lower leaf lamellae that are 2–3 cells high (rather than 3–4(–5) cells high in
|
||
<taxonomicName id="559D27AAF338FFA4FBD537BDFB26EBA1" authorityName="HEDENAS, BOMFLEUR et E. M. FRIIS" authorityYear="2023" box="[1121,1253,1741,1764]" class="Bryopsida" family="Polytrichaceae" genus="Polytrichastrum" kingdom="Plantae" order="Polytrichales" pageId="3" pageNumber="106" phylum="Bryophyta" rank="species" species="incurvum">
|
||
<emphasis id="A0E9803BF338FFA4FBD537BDFB26EBA1" box="[1121,1253,1741,1764]" italics="true" pageId="3" pageNumber="106">P. incurvum</emphasis>
|
||
</taxonomicName>
|
||
), also in having a distinctly mammillose adaxial lamina surface, in having finely toothed leaf margins, and in lacking the strongly curved lamina margins typical of
|
||
<taxonomicName id="559D27AAF338FFA4FB2F3649FAD9EA15" authorityName="HEDENAS, BOMFLEUR et E. M. FRIIS" authorityYear="2023" box="[1179,1306,1849,1872]" class="Bryopsida" family="Polytrichaceae" genus="Polytrichastrum" kingdom="Plantae" order="Polytrichales" pageId="3" pageNumber="106" phylum="Bryophyta" rank="species" species="incurvum">
|
||
<emphasis id="A0E9803BF338FFA4FB2F3649FAD9EA15" box="[1179,1306,1849,1872]" italics="true" pageId="3" pageNumber="106">P. incurvum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="92225C29F338FFA4FCEE3619FA6DEAC5" blockId="3.[858,1454,1896,1921]" box="[858,1454,1896,1921]" pageId="3" pageNumber="106">D e s c r i p t i o n a n d C o m m e n t s o n t h e</paragraph>
|
||
<paragraph id="92225C29F338FFA4FC8036FDFA70EAE1" blockId="3.[820,1459,1932,1956]" box="[820,1459,1932,1956]" pageId="3" pageNumber="106">M a t e r i a l. Three shoot fragments (S174251, S266391,</paragraph>
|
||
<paragraph id="92225C29F338FFA4FC8036C0FA77EA8D" blockId="3.[820,1460,1968,1992]" box="[820,1460,1968,1992]" pageId="3" pageNumber="106">
|
||
S266392) are known of which the largest is ca.
|
||
<quantity id="5565F1CCF338FFA4FA9336C0FAB4EA8D" box="[1319,1399,1968,1992]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="106" unit="mm" value="2.5">2.5 mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
<paragraph id="92225C29F338FFA3FC8036A4FD3CE846" blockId="3.[820,1459,2004,2028]" lastBlockId="4.[129,767,1259,1283]" lastPageId="4" lastPageNumber="107" pageId="3" pageNumber="106">
|
||
The stem is compressed, and details are not visible except that the cortex consists of 1(–2) layers of incrassate cells which are much smaller than the cells toward the center of the stem. The leaves are erect on the stem with their distal portions incurved (
|
||
<figureCitation id="0AA640ACF33FFFA3FE3B352FFDD1E93D" box="[399,530,1119,1144]" captionStart="Text-fig" captionStartId="5.[125,206,1270,1291]" captionTargetBox="[125,1439,151,1254]" captionTargetId="figure-267@5.[125,1439,151,1254]" captionTargetPageId="5" captionText="Text-fig. 3. Scanning electron microscope (SEM, a–e) and synchrotron radiation X-ray tomographic microscopy (SRXTM, f–g) images of a shoot and leaves of Polytrichastrum incurvum HEDENÄS, BOMFLEUR et E.M.FRIIS from the Early Cretaceous Catefica mesofossil flora, Almargem Formation, Portugal. a: Shoot apex showing several incurved leaves. b: Leaf showing ventral longitudinal lamellae (arrow) and incurved leaf margins. c: Detail of longitudinal lamellae on the ventral leaf surface (arrow) and the incurved leaf margin. d: Transverse fracture through the upper portion of leaf showing lamellae on the ventral surface (arrows). e: Detail of upper surface of three longitudinal lamellae showing outlines of individual cells. f: Transverse section (orthoslice xy0655) of shoot and several leaves at different levels. Note the lamellae on the ventral surface over the costa in sections in the distal part of the leaf (arrow) and the mostly unistratose leaf lamina with smooth cells (arrowheads) and lacking lamellae in the proximal part of the leaf. g: Longitudinal section (orthoslice yz1435) showing shoot with several leaves. Specimen, Catefica 49-S174251 (a–g; holotype). Scale bars = 1 mm (a), 200 µm (b, f, g), 100 µm (c), 50 µm (d), 10 µm (e)." figureDoi="http://doi.org/10.5281/zenodo.10480784" httpUri="https://zenodo.org/record/10480784/files/figure.png" pageId="4" pageNumber="107">Text-figs 3a</figureCitation>
|
||
,
|
||
<figureCitation id="0AA640ACF33FFFA3FDAB3510FD94E932" box="[543,599,1119,1144]" captionStart="Text-fig" captionStartId="6.[148,229,816,837]" captionTargetBox="[148,1462,151,801]" captionTargetId="figure-561@6.[148,1462,151,801]" captionTargetPageId="6" captionText="Text-fig. 4. Synchrotron radiation X-ray tomographic microscopy (SRXTM) images (volume renderings, a, b; orthoslices, c, d) of a shoot and leaves of Polytrichastrum incurvum HEDENÄS, BOMFLEUR et E.M.FRIIS from the Early Cretaceous Catefica mesofossil flora, Almargem Formation, Portugal. a, b: Shoot apex showing incurved leaves with distinctly incurved leaf margins. Note a section through the proximal part of a leaf (b) showing lamellae (arrow) over the costa and the unistratose leaf lamina (arrow head). c: Transverse section (orthoslice xy1440) of a shoot showing sections through the distal portions of six leaves showing the distinct lamellae (arrows). d: Transverse section (orthoslice xy2160) of a shoot showing sections through the proximal portions of several leaves showing the unistratose leaf lamina (arrow) with smooth cells. Specimen, Catefica 49-S266392 (a–d). Scale bars = 1 mm (a), 500 µm (b), 200 µm (c, d)." figureDoi="http://doi.org/10.5281/zenodo.10480786" httpUri="https://zenodo.org/record/10480786/files/figure.png" pageId="4" pageNumber="107">4a, b</figureCitation>
|
||
). From a broad sheathing base, each leaf narrows to a limb that gradually narrows further in its upper portion to a cucullate, obtuse apex (
|
||
<figureCitation id="0AA640ACF33FFFA3FF7535B8FE87E9A5" box="[193,324,1224,1248]" captionStart="Text-fig" captionStartId="5.[125,206,1270,1291]" captionTargetBox="[125,1439,151,1254]" captionTargetId="figure-267@5.[125,1439,151,1254]" captionTargetPageId="5" captionText="Text-fig. 3. Scanning electron microscope (SEM, a–e) and synchrotron radiation X-ray tomographic microscopy (SRXTM, f–g) images of a shoot and leaves of Polytrichastrum incurvum HEDENÄS, BOMFLEUR et E.M.FRIIS from the Early Cretaceous Catefica mesofossil flora, Almargem Formation, Portugal. a: Shoot apex showing several incurved leaves. b: Leaf showing ventral longitudinal lamellae (arrow) and incurved leaf margins. c: Detail of longitudinal lamellae on the ventral leaf surface (arrow) and the incurved leaf margin. d: Transverse fracture through the upper portion of leaf showing lamellae on the ventral surface (arrows). e: Detail of upper surface of three longitudinal lamellae showing outlines of individual cells. f: Transverse section (orthoslice xy0655) of shoot and several leaves at different levels. Note the lamellae on the ventral surface over the costa in sections in the distal part of the leaf (arrow) and the mostly unistratose leaf lamina with smooth cells (arrowheads) and lacking lamellae in the proximal part of the leaf. g: Longitudinal section (orthoslice yz1435) showing shoot with several leaves. Specimen, Catefica 49-S174251 (a–g; holotype). Scale bars = 1 mm (a), 200 µm (b, f, g), 100 µm (c), 50 µm (d), 10 µm (e)." figureDoi="http://doi.org/10.5281/zenodo.10480784" httpUri="https://zenodo.org/record/10480784/files/figure.png" pageId="4" pageNumber="107">Text-figs 3a</figureCitation>
|
||
,
|
||
<figureCitation id="0AA640ACF33FFFA3FEFB35B8FE45E9A5" box="[335,390,1224,1248]" captionStart="Text-fig" captionStartId="6.[148,229,816,837]" captionTargetBox="[148,1462,151,801]" captionTargetId="figure-561@6.[148,1462,151,801]" captionTargetPageId="6" captionText="Text-fig. 4. Synchrotron radiation X-ray tomographic microscopy (SRXTM) images (volume renderings, a, b; orthoslices, c, d) of a shoot and leaves of Polytrichastrum incurvum HEDENÄS, BOMFLEUR et E.M.FRIIS from the Early Cretaceous Catefica mesofossil flora, Almargem Formation, Portugal. a, b: Shoot apex showing incurved leaves with distinctly incurved leaf margins. Note a section through the proximal part of a leaf (b) showing lamellae (arrow) over the costa and the unistratose leaf lamina (arrow head). c: Transverse section (orthoslice xy1440) of a shoot showing sections through the distal portions of six leaves showing the distinct lamellae (arrows). d: Transverse section (orthoslice xy2160) of a shoot showing sections through the proximal portions of several leaves showing the unistratose leaf lamina (arrow) with smooth cells. Specimen, Catefica 49-S266392 (a–d). Scale bars = 1 mm (a), 500 µm (b), 200 µm (c, d)." figureDoi="http://doi.org/10.5281/zenodo.10480786" httpUri="https://zenodo.org/record/10480786/files/figure.png" pageId="4" pageNumber="107">4a, b</figureCitation>
|
||
). The costa is broad and somewhat narrower in the sheathing portion of the leaf. In the limb,
|
||
</paragraph>
|
||
<caption id="C6E20CA1F33FFFA3FF203240FBC9EEB0" ID-DOI="http://doi.org/10.5281/zenodo.10480782" ID-Zenodo-Dep="10480782" httpUri="https://zenodo.org/record/10480782/files/figure.png" pageId="4" pageNumber="107" startId="4.[148,229,816,837]" targetBox="[148,1462,151,801]" targetPageId="4" targetType="figure">
|
||
<paragraph id="92225C29F33FFFA3FF203240FBC9EEB0" blockId="4.[148,1462,816,1013]" pageId="4" pageNumber="107">
|
||
<emphasis id="A0E9803BF33FFFA3FF203240FEA6EE26" bold="true" pageId="4" pageNumber="107">Text-fig. 2. Synchrotron radiation X-ray tomographic microscopy (SRXTM) images (volume renderings, a, b; orthoslices, c, d) of a shoot and leaves of</emphasis>
|
||
<taxonomicName id="559D27AAF33FFFA3FEDE323EFC18EE27" authority="HEDENAS, BOMFLEUR et E. M. FRIIS" authorityName="HEDENAS, BOMFLEUR et E. M. FRIIS" authorityYear="2023" box="[362,987,846,867]" class="Bryopsida" family="Ditrichaceae" genus="Chlorosphagnum" kingdom="Plantae" order="Dicranales" pageId="4" pageNumber="107" phylum="Bryophyta" rank="species" species="cateficense">
|
||
<emphasis id="A0E9803BF33FFFA3FEDE323EFC18EE27" bold="true" box="[362,987,846,867]" pageId="4" pageNumber="107">
|
||
<emphasis id="A0E9803BF33FFFA3FEDE323EFD49EE26" bold="true" box="[362,650,846,867]" italics="true" pageId="4" pageNumber="107">Chlorosphagnum cateficense</emphasis>
|
||
<smallCapsWord id="94C4CAF5F33FFFA3FD24323EFD24EE27" baselines="862,862" box="[656,743,846,867]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Hedenas" pageId="4" pageNumber="107">HEDENÄS</smallCapsWord>
|
||
,
|
||
<smallCapsWord id="94C4CAF5F33FFFA3FD47323EFC9AEE27" baselines="862,862" box="[755,857,846,867]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Bomfleur" pageId="4" pageNumber="107">BOMFLEUR</smallCapsWord>
|
||
et E.M.
|
||
<smallCapsWord id="94C4CAF5F33FFFA3FC1C323EFC18EE27" baselines="862,862" box="[936,987,846,867]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Friis" pageId="4" pageNumber="107">FRIIS</smallCapsWord>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
from the Early Cretaceous Catefica mesofossil flora, Almargem Formation, Portugal. a, b: Both sides of a shoot apex showing several leaves with chlorophyllose cells (arrows) and hyaline cells, some with transverse fibrils (arrow heads). c: Transverse section (orthoslice xy0950) of shoot apex showing differentiation into hyaline (arrows) and chlorophyllose cells (arrow heads) of similar size. d: Longitudinal section (orthoslice yz0682) of shoot apex showing several leaves with chlorophyllose (arrows) and hyaline cells, some with transverse fibrils (arrow heads). Specimen, Catefica 49-S266380 (a–d). Scale bars = 500 µm (a, b), 100 µm (c, d).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="92225C29F33FFFA3FF35347EFCC3E8F4" blockId="4.[129,768,1294,1457]" pageId="4" pageNumber="107">the costa has an abaxial epidermis of broader cells with thick outer walls, one layer of guide cells ca. 8–13 µm in diameter, and 3–6 layers of abaxial stereids and 0–2 layers of adaxial stereids ca. 2–4 µm in diameter. On the adaxial leaf surface the costa is covered with straight leaf lamellae.</paragraph>
|
||
<paragraph id="92225C29F33FFFA3FF3534CCFCC3E8B2" blockId="4.[129,767,1468,1492]" lastBlockId="4.[129,768,1503,1527]" pageId="4" pageNumber="107">
|
||
Near the leaf apex there are 3–4 lamellae, farther down there are between 11–17(–20) lamellae (
|
||
<figureCitation id="0AA640ACF33FFFA3FD9D34AFFD0FE8B2" box="[553,716,1503,1527]" captionStart="Text-fig" captionStartId="5.[125,206,1270,1291]" captionTargetBox="[125,1439,151,1254]" captionTargetId="figure-267@5.[125,1439,151,1254]" captionTargetPageId="5" captionText="Text-fig. 3. Scanning electron microscope (SEM, a–e) and synchrotron radiation X-ray tomographic microscopy (SRXTM, f–g) images of a shoot and leaves of Polytrichastrum incurvum HEDENÄS, BOMFLEUR et E.M.FRIIS from the Early Cretaceous Catefica mesofossil flora, Almargem Formation, Portugal. a: Shoot apex showing several incurved leaves. b: Leaf showing ventral longitudinal lamellae (arrow) and incurved leaf margins. c: Detail of longitudinal lamellae on the ventral leaf surface (arrow) and the incurved leaf margin. d: Transverse fracture through the upper portion of leaf showing lamellae on the ventral surface (arrows). e: Detail of upper surface of three longitudinal lamellae showing outlines of individual cells. f: Transverse section (orthoslice xy0655) of shoot and several leaves at different levels. Note the lamellae on the ventral surface over the costa in sections in the distal part of the leaf (arrow) and the mostly unistratose leaf lamina with smooth cells (arrowheads) and lacking lamellae in the proximal part of the leaf. g: Longitudinal section (orthoslice yz1435) showing shoot with several leaves. Specimen, Catefica 49-S174251 (a–g; holotype). Scale bars = 1 mm (a), 200 µm (b, f, g), 100 µm (c), 50 µm (d), 10 µm (e)." figureDoi="http://doi.org/10.5281/zenodo.10480784" httpUri="https://zenodo.org/record/10480784/files/figure.png" pageId="4" pageNumber="107">Text-figs. 3b–e</figureCitation>
|
||
,
|
||
<figureCitation id="0AA640ACF33FFFA3FD6334AFFD30E8B2" box="[727,755,1503,1527]" captionStart="Text-fig" captionStartId="6.[148,229,816,837]" captionTargetBox="[148,1462,151,801]" captionTargetId="figure-561@6.[148,1462,151,801]" captionTargetPageId="6" captionText="Text-fig. 4. Synchrotron radiation X-ray tomographic microscopy (SRXTM) images (volume renderings, a, b; orthoslices, c, d) of a shoot and leaves of Polytrichastrum incurvum HEDENÄS, BOMFLEUR et E.M.FRIIS from the Early Cretaceous Catefica mesofossil flora, Almargem Formation, Portugal. a, b: Shoot apex showing incurved leaves with distinctly incurved leaf margins. Note a section through the proximal part of a leaf (b) showing lamellae (arrow) over the costa and the unistratose leaf lamina (arrow head). c: Transverse section (orthoslice xy1440) of a shoot showing sections through the distal portions of six leaves showing the distinct lamellae (arrows). d: Transverse section (orthoslice xy2160) of a shoot showing sections through the proximal portions of several leaves showing the unistratose leaf lamina (arrow) with smooth cells. Specimen, Catefica 49-S266392 (a–d). Scale bars = 1 mm (a), 500 µm (b), 200 µm (c, d)." figureDoi="http://doi.org/10.5281/zenodo.10480786" httpUri="https://zenodo.org/record/10480786/files/figure.png" pageId="4" pageNumber="107">4c</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="92225C29F33FFFA3FF353772FD3CEBC7" blockId="4.[129,768,1538,1596]" lastBlockId="4.[129,767,1642,1666]" pageId="4" pageNumber="107">The lamellae are mostly 3–4 cells high, occasionally up to 5(–6) cells high, with the upper cells thin-walled or slightly thickened. The leaf lamina is unistratose, but towards the costa can be occasionally or entirely bistratose. In the limb,</paragraph>
|
||
<paragraph id="92225C29F33FFFA3FF3537FDFD3CEBE0" blockId="4.[129,767,1677,1701]" box="[129,767,1677,1701]" pageId="4" pageNumber="107">the leaf lamina is broadly incurved over the lateral lamellae</paragraph>
|
||
<paragraph id="92225C29F33FFFA3FF3537C0FE98EA16" blockId="4.[129,767,1712,1736]" lastBlockId="4.[129,347,1851,1875]" pageId="4" pageNumber="107">
|
||
(
|
||
<figureCitation id="0AA640ACF33FFFA3FF3337C0FEEFEB82" box="[135,300,1712,1736]" captionStart="Text-fig" captionStartId="5.[125,206,1270,1291]" captionTargetBox="[125,1439,151,1254]" captionTargetId="figure-267@5.[125,1439,151,1254]" captionTargetPageId="5" captionText="Text-fig. 3. Scanning electron microscope (SEM, a–e) and synchrotron radiation X-ray tomographic microscopy (SRXTM, f–g) images of a shoot and leaves of Polytrichastrum incurvum HEDENÄS, BOMFLEUR et E.M.FRIIS from the Early Cretaceous Catefica mesofossil flora, Almargem Formation, Portugal. a: Shoot apex showing several incurved leaves. b: Leaf showing ventral longitudinal lamellae (arrow) and incurved leaf margins. c: Detail of longitudinal lamellae on the ventral leaf surface (arrow) and the incurved leaf margin. d: Transverse fracture through the upper portion of leaf showing lamellae on the ventral surface (arrows). e: Detail of upper surface of three longitudinal lamellae showing outlines of individual cells. f: Transverse section (orthoslice xy0655) of shoot and several leaves at different levels. Note the lamellae on the ventral surface over the costa in sections in the distal part of the leaf (arrow) and the mostly unistratose leaf lamina with smooth cells (arrowheads) and lacking lamellae in the proximal part of the leaf. g: Longitudinal section (orthoslice yz1435) showing shoot with several leaves. Specimen, Catefica 49-S174251 (a–g; holotype). Scale bars = 1 mm (a), 200 µm (b, f, g), 100 µm (c), 50 µm (d), 10 µm (e)." figureDoi="http://doi.org/10.5281/zenodo.10480784" httpUri="https://zenodo.org/record/10480784/files/figure.png" pageId="4" pageNumber="107">Text-figs 3b, c</figureCitation>
|
||
,
|
||
<figureCitation id="0AA640ACF33FFFA3FE8F37C0FE9AEB8D" box="[315,345,1712,1736]" captionStart="Text-fig" captionStartId="6.[148,229,816,837]" captionTargetBox="[148,1462,151,801]" captionTargetId="figure-561@6.[148,1462,151,801]" captionTargetPageId="6" captionText="Text-fig. 4. Synchrotron radiation X-ray tomographic microscopy (SRXTM) images (volume renderings, a, b; orthoslices, c, d) of a shoot and leaves of Polytrichastrum incurvum HEDENÄS, BOMFLEUR et E.M.FRIIS from the Early Cretaceous Catefica mesofossil flora, Almargem Formation, Portugal. a, b: Shoot apex showing incurved leaves with distinctly incurved leaf margins. Note a section through the proximal part of a leaf (b) showing lamellae (arrow) over the costa and the unistratose leaf lamina (arrow head). c: Transverse section (orthoslice xy1440) of a shoot showing sections through the distal portions of six leaves showing the distinct lamellae (arrows). d: Transverse section (orthoslice xy2160) of a shoot showing sections through the proximal portions of several leaves showing the unistratose leaf lamina (arrow) with smooth cells. Specimen, Catefica 49-S266392 (a–d). Scale bars = 1 mm (a), 500 µm (b), 200 µm (c, d)." figureDoi="http://doi.org/10.5281/zenodo.10480786" httpUri="https://zenodo.org/record/10480786/files/figure.png" pageId="4" pageNumber="107">4b</figureCitation>
|
||
). The leaf margin is entire or slightly uneven, but not dentate or denticulate. The lamina cells of the sheathing portion of the leaf are linear, whereas they are irregularly rectangular or quadrate in the transition to the leaf limb and above.
|
||
</paragraph>
|
||
<paragraph id="92225C29F33FFFA3FF12361DFCC3EAA8" blockId="4.[129,767,1901,1995]" lastBlockId="4.[129,768,2006,2030]" pageId="4" pageNumber="107">
|
||
S y s t e m a t i c R e l a t i o n s h i p s. Extant
|
||
<taxonomicName id="559D27AAF33FFFA3FDC0361DFD3CEAC0" box="[628,767,1901,1925]" class="Bryopsida" kingdom="Plantae" order="Polytrichales" pageId="4" pageNumber="107" phylum="Bryophyta" rank="order">Polytrichales</taxonomicName>
|
||
comprise about 19 extant genera, most of which have photosynthetic lamellae on the upper leaf surface. In gametophyte features,
|
||
<taxonomicName id="559D27AAF33FFFA3FE2336A6FD87EAAB" box="[407,580,2006,2030]" class="Bryopsida" family="Polytrichaceae" genus="Polytrichastrum" kingdom="Plantae" order="Polytrichales" pageId="4" pageNumber="107" phylum="Bryophyta" rank="genus">
|
||
<emphasis id="A0E9803BF33FFFA3FE2336A6FD87EAAB" box="[407,580,2006,2030]" italics="true" pageId="4" pageNumber="107">Polytrichastrum</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
</paragraph>
|
||
<paragraph id="92225C29F33FFFA3FF353689FA09E93D" blockId="4.[129,768,2041,2065]" lastBlockId="4.[843,1482,1120,1144]" pageId="4" pageNumber="107">
|
||
<taxonomicName id="559D27AAF33FFFA3FF353689FE9FE555" authority="HEDW." authorityName="HEDW." box="[129,348,2041,2065]" class="Bryopsida" family="Polytrichaceae" genus="Polytrichum" kingdom="Plantae" order="Polytrichales" pageId="4" pageNumber="107" phylum="Bryophyta" rank="genus">
|
||
<emphasis id="A0E9803BF33FFFA3FF353689FEC6E554" box="[129,261,2041,2065]" italics="true" pageId="4" pageNumber="107">Polytrichum</emphasis>
|
||
<smallCapsWord id="94C4CAF5F33FFFA3FEAC3689FE95E54A" baselines="2058,2059" box="[280,342,2041,2064]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Hedw" pageId="4" pageNumber="107">HEDW</smallCapsWord>
|
||
.
|
||
</taxonomicName>
|
||
, from which it differs mainly in sporophytic characters (Bell and Hyvönen 2010). However, our species is referred to
|
||
<taxonomicName id="559D27AAF33FFFA3FBEB354DFACFE910" box="[1119,1292,1085,1109]" class="Bryopsida" family="Polytrichaceae" genus="Polytrichastrum" kingdom="Plantae" order="Polytrichales" pageId="4" pageNumber="107" phylum="Bryophyta" rank="genus">
|
||
<emphasis id="A0E9803BF33FFFA3FBEB354DFACFE910" box="[1119,1292,1085,1109]" italics="true" pageId="4" pageNumber="107">Polytrichastrum</emphasis>
|
||
</taxonomicName>
|
||
due to its unique similarity to
|
||
<taxonomicName id="559D27AAF33FFFA3FC663510FA09E93D" authority="(FLORKE ex BRID.)" baseAuthorityName="BRID." box="[978,1482,1120,1144]" class="Bryopsida" family="Polytrichaceae" genus="Polytrichastrum" kingdom="Plantae" order="Polytrichales" pageId="4" pageNumber="107" phylum="Bryophyta" rank="species" species="sexangulare">
|
||
<emphasis id="A0E9803BF33FFFA3FC663510FAC5E93D" box="[978,1286,1120,1144]" italics="true" pageId="4" pageNumber="107">Polytrichastrum sexangulare</emphasis>
|
||
(
|
||
<smallCapsWord id="94C4CAF5F33FFFA3FAA23511FAA0E93D" baselines="1138,1139" box="[1302,1379,1121,1144]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Florke" pageId="4" pageNumber="107">FLÖRKE</smallCapsWord>
|
||
ex
|
||
<smallCapsWord id="94C4CAF5F33FFFA3FA3C3511FA79E933" baselines="1138,1138" box="[1416,1466,1121,1144]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Brid" pageId="4" pageNumber="107">BRID</smallCapsWord>
|
||
.)
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="92225C29F33FFFA3FCFF35F4FA0EE9DE" blockId="4.[843,1485,1156,1180]" box="[843,1485,1156,1180]" pageId="4" pageNumber="107">
|
||
G.L.
|
||
<smallCapsWord id="94C4CAF5F33FFFA3FCC835F4FC5FE9DF" baselines="1174,1174" box="[892,924,1156,1180]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Sm" pageId="4" pageNumber="107">SM</smallCapsWord>
|
||
. and some phenotypes of what has been called ’
|
||
<emphasis id="A0E9803BF33FFFA3FA0235F4FA0EE9DE" box="[1462,1485,1156,1179]" italics="true" pageId="4" pageNumber="107">P.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="92225C29F33FFFA3FCFF35D8FB6CE86F" blockId="4.[843,1482,1191,1216]" lastBlockId="4.[843,1199,1298,1322]" pageId="4" pageNumber="107">
|
||
<emphasis id="A0E9803BF33FFFA3FCFF35D8FC0CE9FA" box="[843,975,1192,1215]" italics="true" pageId="4" pageNumber="107">norwegicum</emphasis>
|
||
(
|
||
<smallCapsWord id="94C4CAF5F33FFFA3FC6B35D8FBDEE9F8" baselines="1209,1209" box="[991,1053,1192,1215]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Hedw" pageId="4" pageNumber="107">HEDW</smallCapsWord>
|
||
.)
|
||
<smallCapsWord id="94C4CAF5F33FFFA3FB8035D7FB6CE9FA" baselines="1209,1210,1209" box="[1076,1199,1191,1215]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Schljakov'" pageId="4" pageNumber="107">SCHLJAKOV’</smallCapsWord>
|
||
(nomenclature unclear) in regard to the incurved leaf margins. This feature is not found in
|
||
<taxonomicName id="559D27AAF33FFFA3FCD2359EFC29E843" box="[870,1002,1262,1286]" class="Bryopsida" family="Polytrichaceae" genus="Polytrichum" kingdom="Plantae" order="Polytrichales" pageId="4" pageNumber="107" phylum="Bryophyta" rank="genus">
|
||
<emphasis id="A0E9803BF33FFFA3FCD2359EFC29E843" box="[870,1002,1262,1286]" italics="true" pageId="4" pageNumber="107">Polytrichum</emphasis>
|
||
</taxonomicName>
|
||
and therefore we do not hesitate to place our fossil species in
|
||
<taxonomicName id="559D27AAF33FFFA3FC4E3462FB64E86F" box="[1018,1191,1298,1322]" class="Bryopsida" family="Polytrichaceae" genus="Polytrichastrum" kingdom="Plantae" order="Polytrichales" pageId="4" pageNumber="107" phylum="Bryophyta" rank="genus">
|
||
<emphasis id="A0E9803BF33FFFA3FC4E3462FB64E86F" box="[1018,1191,1298,1322]" italics="true" pageId="4" pageNumber="107">Polytrichastrum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="92225C29F33FFFA3FCC53446FA0DE8FD" blockId="4.[881,1482,1334,1358]" lastBlockId="4.[843,1486,1440,1464]" pageId="4" pageNumber="107">
|
||
<taxonomicName id="559D27AAF33FFFA3FCC53446FB45E808" authorityName="HEDENAS, BOMFLEUR et E. M. FRIIS" authorityYear="2023" box="[881,1158,1334,1358]" class="Bryopsida" family="Polytrichaceae" genus="Polytrichastrum" kingdom="Plantae" order="Polytrichales" pageId="4" pageNumber="107" phylum="Bryophyta" rank="species" species="incurvum">
|
||
<emphasis id="A0E9803BF33FFFA3FCC53446FB45E808" box="[881,1158,1334,1358]" italics="true" pageId="4" pageNumber="107">Polytrichastrum incurvum</emphasis>
|
||
</taxonomicName>
|
||
has leaves smaller than many extant species of
|
||
<taxonomicName id="559D27AAF33FFFA3FBB13429FB62E834" box="[1029,1185,1369,1393]" class="Bryopsida" family="Polytrichaceae" kingdom="Plantae" order="Polytrichales" pageId="4" pageNumber="107" phylum="Bryophyta" rank="family">Polytrichaceae</taxonomicName>
|
||
, which may have shoots up to several decimetres tall and well-developed leaves of more than a centimetre long. However, the size of the leaves of
|
||
<taxonomicName id="559D27AAF33FFFA3FA0234D1FA0DE8FD" box="[1462,1486,1441,1464]" pageId="4" pageNumber="107">
|
||
<emphasis id="A0E9803BF33FFFA3FA0234D1FA0DE8FD" box="[1462,1486,1441,1464]" italics="true" pageId="4" pageNumber="107">P.</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="92225C29F33FFFA3FCFF34B5FA09EB2F" blockId="4.[843,1482,1476,1501]" lastBlockId="4.[843,1482,1618,1642]" pageId="4" pageNumber="107">
|
||
<emphasis id="A0E9803BF33FFFA3FCFF34B5FC6DE899" box="[843,942,1477,1500]" italics="true" pageId="4" pageNumber="107">incurvum</emphasis>
|
||
are well within the range seen among young leaves of the extant species. Assignments to extant genera have also been adopted in systematic treatments of other fossil bryophytes (e.g., Konopka et al. 1998, Bippus et al. 2021) For example, both
|
||
<taxonomicName id="559D27AAF33FFFA3FC533722FA09EB2F" authority="KONOPKA, HEREND." authorityName="KONOPKA, HEREND." box="[999,1482,1618,1642]" class="Bryopsida" family="Dicranaceae" genus="Campylopodium" kingdom="Plantae" order="Dicranales" pageId="4" pageNumber="107" phylum="Bryophyta" rank="species" species="allonense">
|
||
<emphasis id="A0E9803BF33FFFA3FC533722FB3CEB2F" box="[999,1279,1618,1642]" italics="true" pageId="4" pageNumber="107">Campylopodium allonense</emphasis>
|
||
<smallCapsWord id="94C4CAF5F33FFFA3FAB23723FAA6EB2C" baselines="1636,1637" box="[1286,1381,1619,1642]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Konopka" pageId="4" pageNumber="107">KONOPKA</smallCapsWord>
|
||
,
|
||
<smallCapsWord id="94C4CAF5F33FFFA3FAC73723FA00EB2D" baselines="1636,1636" box="[1395,1475,1619,1642]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Herend" pageId="4" pageNumber="107">HEREND</smallCapsWord>
|
||
.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="92225C29F33FFFA3FCFF3706FA09EBCB" blockId="4.[843,1482,1654,1678]" box="[843,1482,1654,1678]" pageId="4" pageNumber="107">
|
||
et P.R.
|
||
<smallCapsWord id="94C4CAF5F33FFFA3FC3B3706FC17EBC8" baselines="1672,1673" box="[911,980,1654,1678]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Crane" pageId="4" pageNumber="107">CRANE</smallCapsWord>
|
||
(Konopka et al. 1998) and
|
||
<taxonomicName id="559D27AAF33FFFA3FB493706FA09EBCB" authorityName="Bomfleur & Hedenäs & Friis & Crane & Raunsgaard & Pedersen & Mendes & Kvaček" authorityYear="2023" box="[1277,1482,1654,1678]" class="Bryopsida" family="Dicranaceae" genus="Cynodontium" kingdom="Plantae" order="Dicranales" pageId="4" pageNumber="107" phylum="Bryophyta" rank="species" species="luthii">
|
||
<emphasis id="A0E9803BF33FFFA3FB493706FA09EBCB" box="[1277,1482,1654,1678]" italics="true" pageId="4" pageNumber="107">Cynodontium luthii</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="92225C29F33FFFA3FCFF37EAFB59EBBD" blockId="4.[843,1482,1690,1714]" lastBlockId="4.[843,1178,1761,1785]" pageId="4" pageNumber="107">
|
||
Bippus, G.W.
|
||
<smallCapsWord id="94C4CAF5F33FFFA3FC6337EAFB80EBF5" baselines="1708,1708" box="[983,1091,1690,1714]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Rothwell" pageId="4" pageNumber="107">ROTHWELL</smallCapsWord>
|
||
et
|
||
<smallCapsWord id="94C4CAF5F33FFFA3FBD637EAFB79EBF5" baselines="1708,1708" box="[1122,1210,1690,1714]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Stockey" pageId="4" pageNumber="107">STOCKEY</smallCapsWord>
|
||
(Bippus et al. 2021) were accepted as members of extant genera in the recent overview by Ignatov and Maslova (2021).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |