treatments-xml/data/F2/76/69/F2766946996E5998A7A7C4E7F3BBD70E.xml

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<document ID-CLB-Dataset="24300" ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian &amp; Fisher, Brian L." docDate="2020" docId="F2766946996E5998A7A7C4E7F3BBD70E" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole masoala Salata &amp; Fisher 2020, sp. nov." docType="treatment" docUuid="170762F5-C1DE-4BF4-A551-B07B1C763053" docUuidSource="ZooBank" docVersion="7" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1732649601836" updateUser="ExternalLinkService">
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<mods:title id="1EDF76118C053303BD52EAF2EDE62C8B">Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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<mods:namePart id="E13A94E9CE8BDB57C2CD28F9424D63F6">Salata, Sebastian</mods:namePart>
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<mods:namePart id="F16C918D4D0CF7D85FC29C84B4A9139F">Fisher, Brian L.</mods:namePart>
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<mods:typeOfResource id="F5E43F5402FDEF24813B33694C1956BB">text</mods:typeOfResource>
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<mods:date id="066A43B5C7D41CE0DB0D39A41B9F53BF">2020</mods:date>
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<mods:identifier id="6348755466C927E6A51A8176951D6921" type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
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<treatment id="F2766946996E5998A7A7C4E7F3BBD70E" ID-GBIF-Taxon="161776992" LSID="urn:lsid:zoobank.org:act:170762F5-C1DE-4BF4-A551-B07B1C763053" httpUri="http://treatment.plazi.org/id/F2766946996E5998A7A7C4E7F3BBD70E" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="52EBD14F3C60BF113300E02A55414FE1" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="32A00F38637B13B54C286B19ECA3570D" pageId="0" pageNumber="1">
<taxonomicName id="42F9293DF7334B5B3AF22A96019C8724" ID-CoL="CMLGG" LSID="http://zoobank.org/170762F5-C1DE-4BF4-A551-B07B1C763053" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole masoala" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="masoala" status="sp. nov.">Pheidole masoala</taxonomicName>
<taxonomicNameLabel id="5246F74438DFB883EB2E36BC506AEF5D" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation id="5E84FFA0B52B128E44570FF9A08DCD42" captionStart="Figure 77" captionStartId="F77" captionText="Figure 77. Pheidole masoala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923165) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0375363)." figureDoi="10.3897/zookeys.905.39592.figure77" httpUri="https://binary.pensoft.net/fig/373124" pageId="0" pageNumber="1">Figs 77A-F</figureCitation>
<figureCitation id="B9434F6686A52C90EFEE0DEED0EC1EDF" captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">, 85I</figureCitation>
<figureCitation id="2710813C04780876EFBD8D4B81E527B5" captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Distribution. Pheidole lamperos sp. nov. (A). P. longipilosa sp. nov. (B). P. longispinosa Forel (C). P. lutea sp. nov. (D). P. madinika sp. nov. (E). P. mahaboensis sp. nov. (F). P. maizina sp. nov. (G). P. makaensis sp. nov. (H). P. makirovana sp. nov. (I). P. manantenensis sp. nov. (J). P. mantadia sp. nov. (K). P. marieannae sp. nov. (L). P. masoala sp. nov. (M). P. mavesatra sp. nov. (N). P. miramila sp. nov. (O). P. moramanaensis sp. nov. (P). P. navoatrensis sp. nov. (R). P. nemoralis Forel (S). P. ocypodea sp. nov. (T). P. parviocula sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure87" httpUri="https://binary.pensoft.net/fig/373134" pageId="0" pageNumber="1">, 87M</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="87E1B359E24BD6261BA502E0B3D6745E" pageId="0" pageNumber="1" type="type material">
<paragraph id="A5A1D7503BC34AD145A439D9895EC251" pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph id="A8E3CBB2CFC1C19F309C0273D16782DA" pageId="0" pageNumber="1">
<emphasis id="15F148917302F1BDCF42BEA7F8A94AE9" bold="true" pageId="0" pageNumber="1">
<emphasis id="6CEB1AEF53F4CAFDBCF1A19E78C05A84" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
.
</emphasis>
Madagascar. •1 major worker; Antsiranana; Masoala National Park;
<geoCoordinate id="4B94534651A328268E3C10B194F9FB3D" degrees="15.32331" direction="south" orientation="latitude" precision="1" value="-15.32331">-15.32331</geoCoordinate>
,
<geoCoordinate id="BFB746301C592204035385010E06AF48" degrees="50.30751" direction="east" orientation="longitude" precision="1" value="50.30751">50.30751</geoCoordinate>
; alt. 60 m; 12 Mar 2014; B.L. Fisher et al. leg.; BLF32969, CASENT0375363 (CASC).
<emphasis id="98936904DC2A165D63EAF63ADB988478" bold="true" pageId="0" pageNumber="1">
<emphasis id="3D8CA33EFF25BF8ADD6E61B07FC87FAB" italics="true" pageId="0" pageNumber="1">Paratype</emphasis>
.
</emphasis>
Madagascar. •1w.; same data as for holotype; CASENT0923165 (CASC).
</paragraph>
</subSubSection>
<subSubSection id="72CF63D9E93EC87667D280061819ABD7" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="5145A580BDC08B9035C29423D1EDC1F9" pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph id="6C9399B83FBB2C3B9D5096DDB1E2F128" pageId="0" pageNumber="1">
Madagascar. -
<emphasis id="552342BE303D6CE66A5650F436C75498" bold="true" italics="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
: •2w., 1s.; 2.0 km S Andrakata; -14.65, 49.71667; alt. 520 m; 2 Dec 1994; B.L. Fisher leg.; CASENT0198553 (CASC). •3w., 5s., 1q., 1m.;
<normalizedToken id="754FB0FAB764BCEE441A95726D40EBBC" originalValue="Forêt">Foret</normalizedToken>
Ambanitaza, 26.1 km 347° Antalaha;
<geoCoordinate id="4FA6481F77D8B612ACB1B697B6BE145D" degrees="14.67933" direction="south" orientation="latitude" precision="1" value="-14.67933">-14.67933</geoCoordinate>
,
<geoCoordinate id="CE508204761BCC8FD63AFFAF9BBE8CDB" degrees="50.18367" direction="east" orientation="longitude" precision="1" value="50.18367">50.18367</geoCoordinate>
; alt. 240 m; 26 Nov 2004; B.L. Fisher leg.; CASENT0054845, CASENT0054898, CASENT0055597, CASENT0109607, CASENT0109608, CASENT0109634, CASENT0109635 (CASC). •1w., 1s.; Masoala National Park;
<geoCoordinate id="003A67470D965F6445927A69A5E7C8E1" degrees="15.3014" direction="south" orientation="latitude" precision="5" value="-15.3014">-15.3014</geoCoordinate>
,
<geoCoordinate id="12AD07FE6B3932B22FE5845D29575201" degrees="50.22776" direction="east" orientation="longitude" precision="1" value="50.22776">50.22776</geoCoordinate>
; alt. 280 m; 7 Mar 2014; B.L. Fisher et al. leg.; CASENT0377564 (CASC). •1w., 1s.; Masoala National Park;
<geoCoordinate id="33F1BEED6A411542B8ACFEC6FF7E54EB" degrees="15.32331" direction="south" orientation="latitude" precision="1" value="-15.32331">-15.32331</geoCoordinate>
,
<geoCoordinate id="19681CB267F941548148994362800F31" degrees="50.30751" direction="east" orientation="longitude" precision="1" value="50.30751">50.30751</geoCoordinate>
; alt. 60 m; 10 Mar 2014; B.L. Fisher et al. leg.; CASENT0375396 (CASC). -
<emphasis id="11A92E1B17A6C3914307C642CB81459B" bold="true" italics="true" pageId="0" pageNumber="1">Toamasina</emphasis>
: •2w., 4s.; Parc National Mananara-Nord, 7.1 km 261° Antanambe;
<geoCoordinate id="48A1F4DD8BCC4B891DB96C8F86803078" degrees="16.455" direction="south" orientation="latitude" precision="55" value="-16.455">-16.455</geoCoordinate>
,
<geoCoordinate id="C8AC9CC5864B4AE5D6D6C9A83B7A4FB3" degrees="49.7875" direction="east" orientation="longitude" precision="5" value="49.7875">49.7875</geoCoordinate>
; alt. 225 m; 15 Nov 2005; B.L. Fisher et al. leg.; CASENT0067475, CASENT0067490, CASENT0069439, CASENT0069448 (CASC). •1w., 1m.;
<normalizedToken id="52CD625B95C40A4607A02F0D1C2CA06B" originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken id="4F60FFEEECBE52A6493F26D5D06F48B0" originalValue="Spéciale">Speciale</normalizedToken>
Ambatovaky, Sandrangato River;
<geoCoordinate id="7946BF4FBC058A1BFEC36935B8C64E42" degrees="16.7755" direction="south" orientation="latitude" precision="5" value="-16.7755">-16.7755</geoCoordinate>
,
<geoCoordinate id="ADAFA419CB601FA1408A16351176240E" degrees="49.26427" direction="east" orientation="longitude" precision="1" value="49.26427">49.26427</geoCoordinate>
; alt. 430 m; 24 Feb 2010; B.L. Fisher et al. leg.; CASENT0161802 (CASC). •1w., 1s.;
<normalizedToken id="9A58581274F337B4E532555595306333" originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken id="FF3140AA98D57C06D6F42E9A5C05D042" originalValue="Spéciale">Speciale</normalizedToken>
Ambatovaky, Sandrangato River;
<geoCoordinate id="D1396BA35EBF3F4BC0168641B71818FA" degrees="16.77274" direction="south" orientation="latitude" precision="1" value="-16.77274">-16.77274</geoCoordinate>
,
<geoCoordinate id="CBE68D6846B0F55F93F850EEDE9BE14A" degrees="49.26551" direction="east" orientation="longitude" precision="1" value="49.26551">49.26551</geoCoordinate>
; alt. 450 m; 20 Feb 2010; B.L. Fisher et al. leg.; CASENT0162680 (CASC). -
<emphasis id="F4885C02A6AB201014A5E118B415E157" bold="true" italics="true" pageId="0" pageNumber="1">Toliara</emphasis>
: •1w., 1s.;
<normalizedToken id="830432B52AF79E7454A52434C80803DA" originalValue="Forêt">Foret</normalizedToken>
Ivohibe 55.0 km N Tolagnaro;
<geoCoordinate id="95A4D81AE9EC405E1EB0DF0E915E6BD0" degrees="24.569" direction="south" orientation="latitude" precision="55" value="-24.569">-24.569</geoCoordinate>
,
<geoCoordinate id="E6FD4F53335F2BFF1162B9EAB2A77203" degrees="47.204" direction="east" orientation="longitude" precision="55" value="47.204">47.204</geoCoordinate>
; alt. 200 m; 3 Dec 2006; B.L. Fisher et al. leg.; CASENT0122620 (CASC).
</paragraph>
</subSubSection>
<subSubSection id="218B549E1A41AF5123663E29878BEE51" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="A6ED17A77C2D32683CC0071B15133205" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="4C983207DA00111673509F9133175BC0" pageId="0" pageNumber="1">
<emphasis id="D7604602732056F8A615C9909CDB07A0" bold="true" pageId="0" pageNumber="1">
<emphasis id="6CECFFEE9BEF82629C2B7CDC0C83DA80" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
.
</emphasis>
Small species: HL: 0.93-1.17 (1.05), HW: 0.89-1.07 (0.97), WL: 0.76-0.92 (0.84); head in full-face view sub-rectangular, anterior and posterior sides of eyes relatively straight; sides of the head with sparse, moderately long, erect pilosity; antennal scrobes present, well delimited, and forming distinct dorsal concavity beneath frontal carina, scrobe surface foveolate, sometimes with additional sparse, thick, and irregular rugae, delimited ventrally and posteriorly by carinulae; inner hypostomal teeth distinct, big, closely spaced, triangular, with rounded apex and wide base; outer hypostomal teeth smaller and thinner than inner hypostomal teeth, with moderately wide base, triangular; gaster finely shagreened; body dark yellow.
<emphasis id="7AE6839901BC71C9BC6EEE9B898CA8BC" bold="true" pageId="0" pageNumber="1">
<emphasis id="CD115DA11D507963A342218251837216" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
Head foveolate, genae with reduced sculpture or partially smooth; promesonotum low, long, flat or slightly convex, with relatively convex posterior declivity; propodeal spines minute, indistinct; mesosoma foveolate, katepisternum smooth.
</paragraph>
</subSubSection>
<subSubSection id="43C97FD01CEEB5374F308E670FF710AF" pageId="0" pageNumber="1" type="description">
<paragraph id="7CDFCAFC08ED39144153E435F8EF9A0E" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="68BFCC126845B526DABBF59DBF5D7AA5" pageId="0" pageNumber="1">
<emphasis id="364F9D2134C083337441A9FB4F5E4564" bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
Measurements (
<emphasis id="38054EAE80A0D59D2A7C2E43AA3B0A40" italics="true" pageId="0" pageNumber="1">N</emphasis>
= 10): HL: 0.93-1.17 (1.05); HW: 0.89-1.07 (0.97); SL: 0.41-0.49 (0.45); EL: 0.13-0.16 (0.14); WL: 0.76-0.92 (0.84); PSL: 0.14-0.18 (0.16); MTL: 0.44-0.5 (0.45); PNW: 0.53-0.62 (0.58); PTW: 0.13-0.18 (0.15); PPW: 0.37-0.48 (0.43); CI: 90.8-95.9 (93.1); SI: 45.4-49.3 (46.7); PSLI: 13.4-16.2 (15.0); PPI: 31.9-37.5 (34.9); PNI: 56.7-63.7 (59.3); MTI: 44.9-49.1 (46.7).
<emphasis id="6514D8A7C9F0EEC71BC6B2054071C7B3" bold="true" pageId="0" pageNumber="1">
<emphasis id="E1B9559E99C8DC77E284B3C4AC4934A4" italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
In full-face view sub-rectangular, anterior and posterior sides of eyes relatively straight (Fig.
<figureCitation id="89FC3644B84130406A283F38BDB9B142" captionStart="Figure 77" captionStartId="F77" captionText="Figure 77. Pheidole masoala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923165) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0375363)." figureDoi="10.3897/zookeys.905.39592.figure77" httpUri="https://binary.pensoft.net/fig/373124" pageId="0" pageNumber="1">77B</figureCitation>
). In lateral view sub-rectangular; ventral and dorsal faces relatively flat; dorsal face finely depressed posteriorly, forming indistinct transverse depression between frons and occipital lobes; inner hypostomal teeth visible. Sides of the head with sparse, moderately long, erect pilosity; whole head with moderately dense, short, suberect to erect pilosity. Head depressed posteriorly, forming shallow transverse depression between frons and occipital lobes. Antennal scrobes distinct and delimited by carinulae; scrobe surface foveolate, sometimes with additional sparse, thick, and irregular rugae, delimited ventrally and posteriorly by carinulae. Occipital lobes shiny, with thick, sparse, irregular rugae, interspaces smooth; frons with dense, thick, and longitudinal rugae, interspaces smooth, and with fine but distinct foveolae on the upper half of frons; malar area with dense, thick, longitudinal to irregular rugulae, interspaces smooth; genae shiny, smooth to finely foveolate. Centre of clypeus smooth and shiny, lateral sides with longitudinal rugae; median notch present, wide and indistinct; median longitudinal carina absent; lateral longitudinal carinae present. Scape, when laid back, slightly exceeding midlength of head; pilosity suberect to erect (Fig.
<figureCitation id="D27D2E41EFE9932CD99C87395E94A6BD" captionStart="Figure 77" captionStartId="F77" captionText="Figure 77. Pheidole masoala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923165) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0375363)." figureDoi="10.3897/zookeys.905.39592.figure77" httpUri="https://binary.pensoft.net/fig/373124" pageId="0" pageNumber="1">77B, D</figureCitation>
). Inner hypostomal teeth distinct, big, closely spaced, triangular, with rounded apex and wide base; outer hypostomal teeth smaller and thinner than inner hypostomal teeth, with moderately wide base, triangular (Fig.
<figureCitation id="7313D6DB15DEE02031FBF469F449FEDE" captionStart="Figure 85" captionStartId="F85" captionText="Figure 85. Hypostomal teeth. Pheidole madinika sp. nov. (A). P. mahaboensis sp. nov. (B). P. maizina sp. nov. (C). P. makaensis sp. nov. (D). P. makirovana sp. nov. (E). P. manantenensis sp. nov. (F). P. mantadia sp. nov. (G). P. marieannae sp. nov. (H). P. masoala sp. nov. (I). P. mavesatra sp. nov. (J). P. miramila sp. nov. (K). P. moramanaensis sp. nov. (L). P. navoatrensis sp. nov. (M). P. nemoralis Forel (N). P. ocypodea sp. nov. (O). P. parviocula sp. nov. (P). P. petax Forel (Q). P. podargea sp. nov. (R). P. praegrandis sp. nov. (S). P. ranohirensis sp. nov. (T). P. rugocephala sp. nov. (U). P. rugofitarata sp. nov. (V). P. scabrata Forel (W). P. typhlos sp. nov. (X). P. vatovavensis sp. nov. (Y). P. voasara sp. nov. (Z). P. vohemarensis sp. nov. (AA). P. zavamanira sp. nov. (AB)." figureDoi="10.3897/zookeys.905.39592.figure85" httpUri="https://binary.pensoft.net/fig/373132" pageId="0" pageNumber="1">85I</figureCitation>
).
<emphasis id="375C670FCDF4A5AA6CEA18DC8ABCA5F1" bold="true" pageId="0" pageNumber="1">
<emphasis id="01876C0E4C888D4659DAF84D41E63A8C" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum moderately long, relatively low, and evenly angular, tubercle-like projections absent; promesonotal groove absent; metanotal groove absent; propodeal spines small, triangular, with acute apex; humeral area laterally weakly produced (Fig.
<figureCitation id="83EF203F29DE2B41360DBBAF2E5B2812" captionStart="Figure 77" captionStartId="F77" captionText="Figure 77. Pheidole masoala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923165) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0375363)." figureDoi="10.3897/zookeys.905.39592.figure77" httpUri="https://binary.pensoft.net/fig/373124" pageId="0" pageNumber="1">77D</figureCitation>
). Surface shiny, with fine foveolae, additional sparse and thin rugae on promesonotal dorsum and sometimes propodeum, sculpture slightly weakening on lateral surfaces of pronotum; katepisternum smooth. Pilosity sparse, moderately long, and erect (Fig.
<figureCitation id="7654DA3B353027C1FF2D361A1ADC6A0C" captionStart="Figure 77" captionStartId="F77" captionText="Figure 77. Pheidole masoala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923165) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0375363)." figureDoi="10.3897/zookeys.905.39592.figure77" httpUri="https://binary.pensoft.net/fig/373124" pageId="0" pageNumber="1">77D, F</figureCitation>
).
<emphasis id="13CC106017CC86A5CCC598625F8B1FB2" bold="true" pageId="0" pageNumber="1">
<emphasis id="7EB498F182EB113D10F34FE75179E837" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Shiny and foveolate; peduncle short, with distinct horizontal lobes on its basal part; node relatively low, triangular, with rounded apex, in rear view node straight; pilosity moderately sparse and erect (Fig.
<figureCitation id="403821E495198A077D1D5460A392B9C5" captionStart="Figure 77" captionStartId="F77" captionText="Figure 77. Pheidole masoala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923165) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0375363)." figureDoi="10.3897/zookeys.905.39592.figure77" httpUri="https://binary.pensoft.net/fig/373124" pageId="0" pageNumber="1">77D, F</figureCitation>
).
<emphasis id="794D56D21E5560046B8917B1702D8626" bold="true" pageId="0" pageNumber="1">
<emphasis id="8318CEA46C4A0C3C1B16E21488B68995" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Shiny, with fine and sparse foveolae, smooth on dorsum; in dorsal view sides with long, acute, and triangular projections; pilosity long, moderately dense, and erect (Fig.
<figureCitation id="AD2FA5F6524A2190BE7EA41E174A9134" captionStart="Figure 77" captionStartId="F77" captionText="Figure 77. Pheidole masoala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923165) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0375363)." figureDoi="10.3897/zookeys.905.39592.figure77" httpUri="https://binary.pensoft.net/fig/373124" pageId="0" pageNumber="1">77D, F</figureCitation>
).
<emphasis id="802D9E5332DC5BF95CDB2F69942BE011" bold="true" pageId="0" pageNumber="1">
<emphasis id="6B86B5B4EB12A9B6C56255F28F699AD5" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Shiny and finely shagreened; pilosity dense, moderately long, and erect (Fig.
<figureCitation id="0D8E6CEBA1FE3E2A6477A773D52ECB6A" captionStart="Figure 77" captionStartId="F77" captionText="Figure 77. Pheidole masoala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923165) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0375363)." figureDoi="10.3897/zookeys.905.39592.figure77" httpUri="https://binary.pensoft.net/fig/373124" pageId="0" pageNumber="1">77D, F</figureCitation>
).
<emphasis id="55205EE9A8E93B5D9CA3633901771F3F" bold="true" pageId="0" pageNumber="1">
<emphasis id="94695E06CB2E6454620BBC54AE427C03" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Unicolourous, dark yellow; lower part of malar area, frons, and gaster brown (Fig.
<figureCitation id="3CAA07AEA6E797D13B63BD1C056267E4" captionStart="Figure 77" captionStartId="F77" captionText="Figure 77. Pheidole masoala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923165) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0375363)." figureDoi="10.3897/zookeys.905.39592.figure77" httpUri="https://binary.pensoft.net/fig/373124" pageId="0" pageNumber="1">77D, F</figureCitation>
).
</paragraph>
<caption id="5EBC2383975941A75B5C657B786946C3" doi="10.3897/zookeys.905.39592.figure77" httpUri="https://binary.pensoft.net/fig/373124" pageId="0" pageNumber="1" start="Figure 77" startId="F77">
<paragraph id="630AA231E9B48810778FEE53A5900BBD" pageId="0" pageNumber="1">
<emphasis id="C91F7333D2059898471205224D68E48E" bold="true" pageId="0" pageNumber="1">Figure 77.</emphasis>
<taxonomicName id="174BEC454010A76AD482706395ED4370" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole masoala" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="masoala">
<emphasis id="E89B54EFF66E803457C1892BCC148791" italics="true" pageId="0" pageNumber="1">Pheidole masoala</emphasis>
</taxonomicName>
sp. nov., full-face view (
<emphasis id="F48209AF17BDB15FFC6CB325AF2705F0" bold="true" pageId="0" pageNumber="1">A</emphasis>
), profile (
<emphasis id="EF4627B358381E8B3D6787B5308AA656" bold="true" pageId="0" pageNumber="1">C</emphasis>
), and dorsal view (
<emphasis id="C46CE42459AB75B54EB1FC2AFAAAAB4A" bold="true" pageId="0" pageNumber="1">E</emphasis>
) of paratype minor worker (CASENT0923165) and full-face view (
<emphasis id="3CB5CD15C4725DEF7F2FC7EEEADEEE66" bold="true" pageId="0" pageNumber="1">B</emphasis>
), profile (
<emphasis id="DCDFFFDDBBCB0D2D6043AB68196FAD3C" bold="true" pageId="0" pageNumber="1">D</emphasis>
), and dorsal view (
<emphasis id="3F7264B07F6DAC9667B4284E29A160CA" bold="true" pageId="0" pageNumber="1">F</emphasis>
) of holotype major worker (CASENT0375363).
</paragraph>
</caption>
<paragraph id="5F4AD3B2DA122D1C7E96F1216743231E" pageId="0" pageNumber="1">
<emphasis id="AE5CCCAC1FB892883FD0E19480D39313" bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
Measurements (
<emphasis id="F4A1A5C638CF888C8D4300EDCB71873A" italics="true" pageId="0" pageNumber="1">N</emphasis>
= 10): HL: 0.44-0.48 (0.46); HW: 0.39-0.43 (0.42); SL: 0.39-0.42 (0.4); EL: 0.09-0.1 (0.1); WL: 0.52-0.56 (0.54); PSL: 0.06-0.08 (0.07); MTL: 0.3-0.34 (0.32); PNW: 0.27-0.29 (0.27); PTW: 0.05-0.07 (0.06); PPW: 0.1-0.11 (0.1); CI: 88.5-95.2 (91.4); SI: 93.9-99.5 (97.3); PSLI: 13.3-16.5 (14.7); PPI: 50.5-64.6 (57.9); PNI: 63.8-68.5 (65.9); MTI: 73.4-80.0 (76.2).
<emphasis id="DB227901F9B64A7C4EB566A7A4262A4E" bold="true" pageId="0" pageNumber="1">
<emphasis id="B787A34AD01983BBA205D8E7B0394CC1" italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Occipital margin straight or indistinctly concave; occipital carina absent (Fig.
<figureCitation id="A4AF5B58A224D40A8045A37E74A6D04B" captionStart="Figure 77" captionStartId="F77" captionText="Figure 77. Pheidole masoala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923165) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0375363)." figureDoi="10.3897/zookeys.905.39592.figure77" httpUri="https://binary.pensoft.net/fig/373124" pageId="0" pageNumber="1">77A</figureCitation>
). Pilosity moderately sparse, long, erect. Head foveolate, genae with reduced sculpture or partially smooth. Clypeus foveolate; median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by one-fifth of its length; pilosity suberect (Fig.
<figureCitation id="3533C1C195A41038DFBD7596FAE2138C" captionStart="Figure 77" captionStartId="F77" captionText="Figure 77. Pheidole masoala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923165) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0375363)." figureDoi="10.3897/zookeys.905.39592.figure77" httpUri="https://binary.pensoft.net/fig/373124" pageId="0" pageNumber="1">77A, C</figureCitation>
).
<emphasis id="14D1CEE8577B2CEEA6FD0B40FF832D20" bold="true" pageId="0" pageNumber="1">
<emphasis id="BE24AEB565FFFE8C597A0780757A96B7" italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
.
</emphasis>
In lateral view, promesonotum low, long, flat or slightly convex, with relatively convex posterior declivity; promesonotal groove absent; metanotal groove indistinct; propodeal spines minute, indistinct, triangular, apex acute (Fig.
<figureCitation id="022E6D41D33E9E59425DAA92FEF8A215" captionStart="Figure 77" captionStartId="F77" captionText="Figure 77. Pheidole masoala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923165) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0375363)." figureDoi="10.3897/zookeys.905.39592.figure77" httpUri="https://binary.pensoft.net/fig/373124" pageId="0" pageNumber="1">77C</figureCitation>
). Sculpture foveolate, katepisternum smooth. Pilosity moderately sparse, moderately short, and erect (Fig.
<figureCitation id="7F8F52D9EE2A52790683FF795B2CB30E" captionStart="Figure 77" captionStartId="F77" captionText="Figure 77. Pheidole masoala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923165) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0375363)." figureDoi="10.3897/zookeys.905.39592.figure77" httpUri="https://binary.pensoft.net/fig/373124" pageId="0" pageNumber="1">77C, E</figureCitation>
).
<emphasis id="4F76F10D442A4B60F2D5B323AE61C2D6" bold="true" pageId="0" pageNumber="1">
<emphasis id="1E9AADA8769D66CE106C06A4F849224B" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
Peduncle very short and thin with ventral face slightly convex; with few short, erect setae (Fig.
<figureCitation id="A0DC8BC872456D75043935630269C9FF" captionStart="Figure 77" captionStartId="F77" captionText="Figure 77. Pheidole masoala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923165) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0375363)." figureDoi="10.3897/zookeys.905.39592.figure77" httpUri="https://binary.pensoft.net/fig/373124" pageId="0" pageNumber="1">77C, E</figureCitation>
).
<emphasis id="CE520E74B0A8EDD4042137278EECC82F" bold="true" pageId="0" pageNumber="1">
<emphasis id="A0D653674DD9EBFDB7B3FD72966685FA" italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
.
</emphasis>
Short, low, and slightly convex; with few short, erect setae (Fig.
<figureCitation id="196DEFE9E7411B40AF032A91F141AE68" captionStart="Figure 77" captionStartId="F77" captionText="Figure 77. Pheidole masoala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923165) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0375363)." figureDoi="10.3897/zookeys.905.39592.figure77" httpUri="https://binary.pensoft.net/fig/373124" pageId="0" pageNumber="1">77C, E</figureCitation>
).
<emphasis id="2D16ACF46A4E3BF3363FC844DFD57B33" bold="true" pageId="0" pageNumber="1">
<emphasis id="D8CBFB01342D84CDB105B2F982780C4E" italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
.
</emphasis>
With sparse, erect pilosity (Fig.
<figureCitation id="04D6D92625E8B4A254A745F278F9389F" captionStart="Figure 77" captionStartId="F77" captionText="Figure 77. Pheidole masoala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923165) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0375363)." figureDoi="10.3897/zookeys.905.39592.figure77" httpUri="https://binary.pensoft.net/fig/373124" pageId="0" pageNumber="1">77C, E</figureCitation>
).
<emphasis id="0FBDF6789A94865A0CC753CD8F54F774" bold="true" pageId="0" pageNumber="1">
<emphasis id="D15E07DC7F6BC67792079E4C939D1B96" italics="true" pageId="0" pageNumber="1">Colour</emphasis>
.
</emphasis>
Unicolourous, yellow (Fig.
<figureCitation id="A710A7A027F436A03EBCCF572F609A0E" captionStart="Figure 77" captionStartId="F77" captionText="Figure 77. Pheidole masoala sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0923165) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0375363)." figureDoi="10.3897/zookeys.905.39592.figure77" httpUri="https://binary.pensoft.net/fig/373124" pageId="0" pageNumber="1">77C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A1CF979FDAA2C0DCF9D06C90E5000001" pageId="0" pageNumber="1" type="etymology">
<paragraph id="3FD9E9BE7C7C3986A4515DF51FCE0C29" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="3335660B34C91432D25367629BE099DF" pageId="0" pageNumber="1">From the type locality.</paragraph>
</subSubSection>
<subSubSection id="970C80CB5BC7FCF73C1B640448EE2CB7" pageId="0" pageNumber="1" type="biology_ecology">
<paragraph id="9DDB563489DF3CDB40CE29F031A8751A" pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph id="32D06CB326FD2D5D0802BB84DF581770" pageId="0" pageNumber="1">The species was collected between 30-520 m in elevation, in rainforest and disturbed rainforest. Nests were located in rotten logs and sticks on the ground.</paragraph>
</subSubSection>
<subSubSection id="40E671F56BD6C4564BFDD34927C6D8B3" pageId="0" pageNumber="1" type="comments">
<paragraph id="5297C203615B6EA69A78E66751AA148A" pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph id="052C4B3F0CA90572E63270F202220A40" pageId="0" pageNumber="1">
<taxonomicName id="505C82553C5DA094E7EB69A40FBCAEBF" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole masoala" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="masoala">
<emphasis id="5B034D0AA01AA6DF97AFBE9F73FA1F3C" italics="true" pageId="0" pageNumber="1">Pheidole masoala</emphasis>
</taxonomicName>
sp. nov. is most similar to
<taxonomicName id="0EF362A085B0F493FDFE2F0816B8FBCE" lsidName="P. madinika" pageId="0" pageNumber="1" rank="species" species="madinika">
<emphasis id="63B99059B0B941AEF7570EE0901B385F" italics="true" pageId="0" pageNumber="1">P. madinika</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="B1AE5E4554859F45C6D4CC1D3E86AF6B" bold="true" pageId="0" pageNumber="1">
<emphasis id="C24B986CB8C12A70910B96FFF47B52E9" italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
.
</emphasis>
<taxonomicName id="74AD3C5D4CBBCFEF48FFD2A529B15F39" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole masoala" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="masoala">
<emphasis id="EB13693463DEE6D5B7438E9484857FB5" italics="true" pageId="0" pageNumber="1">Pheidole masoala</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName id="31ED021B26EB2EDF133077D2556062FB" lsidName="P. madinika" pageId="0" pageNumber="1" rank="species" species="madinika">
<emphasis id="C1E92C4410BBE8F4A68A36D17CDB82BB" italics="true" pageId="0" pageNumber="1">P. madinika</emphasis>
</taxonomicName>
sp. nov. in dark yellow body colouration, presence of shallow transverse depression between frons and occipital lobes, and well developed antennal scrobes which are delimited ventrally and posteriorly by carinulae.
<emphasis id="B2A19A6C2B58FB7D6EA9D28ABC650D41" bold="true" pageId="0" pageNumber="1">
<emphasis id="35E9109296C1C11D5767B29C22B8A0C7" italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
.
</emphasis>
<taxonomicName id="6A37B83C8F6C66E7DF16E924EF471368" authorityName="Salata &amp; Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole masoala" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="masoala">
<emphasis id="F055F46D6A9C5B370231A2450954AF98" italics="true" pageId="0" pageNumber="1">Pheidole masoala</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName id="22BBDBE54E8926D14700F996D4912B2D" lsidName="P. madinika" pageId="0" pageNumber="1" rank="species" species="madinika">
<emphasis id="0BE07E16EAF594DED1ED503AB0C02477" italics="true" pageId="0" pageNumber="1">P. madinika</emphasis>
</taxonomicName>
sp. nov. by relatively convex posterior declivity of promesonotum, and shallow and indistinct metanotal groove.
</paragraph>
</subSubSection>
</treatment>
</document>