257 lines
31 KiB
XML
257 lines
31 KiB
XML
<document ID-CLB-Dataset="9289" ID-DOI="http://dx.doi.org/10.3897/zookeys.1004.57526" ID-GBIF-Dataset="4e633bc1-16fc-4e83-b8a0-cb9289897616" ID-PMC="PMC7758311" ID-Pensoft-Pub="1313-2970-1004-27" ID-Pensoft-UUID="BEECC18115E95A798CA5250990A350E3" ID-PubMed="33384565" ID-ZooBank="320559CF19B5423CB7FB72555290241A" ModsDocID="1313-2970-1004-27" checkinTime="1608157838683" checkinUser="pensoft" docAuthor="Maddison, Wayne P., Beattie, Imara, Marathe, Kiran, Ng, Paul Y. C., Kanesharatnam, Nilani, Benjamin, Suresh P. & Kunte, Krushnamegh" docDate="2020" docId="F09C8524C1A1506C9730F94F97B92D4D" docLanguage="en" docName="ZooKeys 1004: 27-97" docOrigin="ZooKeys 1004" docPubDate="2020-12-16" docSource="http://dx.doi.org/10.3897/zookeys.1004.57526" docTitle="Maripanthus reinholdae Maddison, sp. nov." docType="treatment" docVersion="6" id="BEECC18115E95A798CA5250990A350E3" lastPageNumber="27" masterDocId="BEECC18115E95A798CA5250990A350E3" masterDocTitle="A phylogenetic and taxonomic review of baviine jumping spiders (Araneae, Salticidae, Baviini)" masterLastPageNumber="97" masterPageNumber="27" pageNumber="27" updateTime="1732936594770" updateUser="ExternalLinkService">
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<mods:titleInfo id="1C896F10C4CAA3EB2F6B44C4C4AB849E">
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<mods:title id="7AA8C3F2E25BA87577031C08113E5511">A phylogenetic and taxonomic review of baviine jumping spiders (Araneae, Salticidae, Baviini)</mods:title>
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<mods:namePart id="75FAB630A5CA9C5DCD4A04BE6EEF5457">Maddison, Wayne P.</mods:namePart>
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<mods:affiliation id="F4E83EB50FF837C9D94A39ED05997A79">Departments of Zoology and Botany and Beaty Biodiversity Museum, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada</mods:affiliation>
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<mods:nameIdentifier id="F579D7ED74280BF1D72117467C368320" type="email">wmaddisn@mail.ubc.ca</mods:nameIdentifier>
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<mods:namePart id="086F522AF0337E776792747851CF8BCB">Beattie, Imara</mods:namePart>
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<mods:affiliation id="4F0FEB3952102CD6C977FEC1E7CC5541">Department of Zoology, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada</mods:affiliation>
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<mods:namePart id="F5A8E96BE175CEA071CA04E924C6C7B8">Marathe, Kiran</mods:namePart>
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<mods:affiliation id="60D0D55EE099B550CCCD5979DB79CF15">Department of Zoology, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada & National Centre for Biological Sciences, Tata Institute of Fundamental Research, GKVK Campus, Bellary Road, Bengaluru 560065, India</mods:affiliation>
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<mods:namePart id="5AD29C100A5EFD18710D6F68D28D5D2F">Ng, Paul Y. C.</mods:namePart>
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<mods:affiliation id="17E94218672A767A24288E7A7DA6007D">205 River Valley Road, # 16 - 53, Singapore 238274, Republic of Singapore</mods:affiliation>
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<mods:namePart id="21EEB7DB82B6A1648DD903BF80592D00">Kanesharatnam, Nilani</mods:namePart>
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<mods:affiliation id="4840D56B7C570608B04E42F0DC064229">National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka & Department of Zoology, Faculty of Science, Eastern University, Vantharumoolai, Sri Lanka</mods:affiliation>
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<mods:namePart id="0996AF3E3680DF4223B5F7496FF287E0">Benjamin, Suresh P.</mods:namePart>
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<mods:affiliation id="735D28FF88DDD2B19A5CA518A943D608">National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka</mods:affiliation>
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<mods:namePart id="7B79F68E04BB74992400EF640388F1D6">Kunte, Krushnamegh</mods:namePart>
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<mods:affiliation id="B01F4C066951E76BA6F581B72228B2E6">National Centre for Biological Sciences, Tata Institute of Fundamental Research, GKVK Campus, Bellary Road, Bengaluru 560065, India</mods:affiliation>
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<mods:title id="9E5027C815D0807C9FB1D0CF331F3CF4">ZooKeys</mods:title>
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<mods:date id="848B3AFAD08C2DE9150CB03373D3B49F">2020</mods:date>
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<mods:number id="F1CAD44F04AE71C5E7485EA657931F0F">2020-12-16</mods:number>
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<mods:number id="EBAAD873B3E3CD4479C3B0A67CC6B50C">1004</mods:number>
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<mods:classification id="743BF46B8B81E9E400C503C8CB604739">journal article</mods:classification>
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<mods:identifier id="336716C16221C4EC6A137ABFF866E6E5" type="DOI">http://dx.doi.org/10.3897/zookeys.1004.57526</mods:identifier>
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<treatment id="F09C8524C1A1506C9730F94F97B92D4D" ID-GBIF-Taxon="176088102" LSID="urn:lsid:zoobank.org:act:5B8BB349-2D63-42E2-AD0A-4426B4BAD0D9" httpUri="http://treatment.plazi.org/id/F09C8524C1A1506C9730F94F97B92D4D" lastPageNumber="27" pageId="0" pageNumber="27" scope_family="Salticidae" scope_order="Araneae" scope_tribe="Baviini">
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<subSubSection id="B9669601C766D07ABDA539B91F7F7350" pageId="0" pageNumber="27" type="nomenclature">
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<paragraph id="8F51C38F0A5954612A7BE5C760969D07" pageId="0" pageNumber="27">
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<taxonomicName id="C9610ED8829D3ED82376D4FA7DFEB944" LSID="http://zoobank.org/5B8BB349-2D63-42E2-AD0A-4426B4BAD0D9" authority="Maddison, sp. nov." authorityName="Maddison, sp. nov." class="Magnoliopsida" family="Pittosporaceae" genus="Maripanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Maripanthus reinholdae" order="Apiales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="species" species="reinholdae">Maripanthus reinholdae Maddison, sp. nov.</taxonomicName>
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<figureCitation id="A086F96D57EF3E911F600C91F93C14FB" captionStart="Figures 4–35" captionStartId="F2" captionText="Figures 4 - 35. Variation in traits among baviines 4, 5 oblique views of female prosoma 4 Bavia nessagyna (specimen IDWM. 20004; note convex front face of chelicerae, and typical thorax) 5 Indopadilla redunca (holotype IDWM. 20011; note concave front face of chelicerae, and thoracic bulges) 6 - 12 left male endite (9 also with chelicera) 6 Bavia nessagyna (holotype IDWM. 20005) 7 Indopadilla redunca (specimen SWK 12 - M 0009) 8 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 9 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 10 Padillothorax badut (specimen SWK 12 - 4688) 11 Maripanthus jubatus (specimen AS 19.4373) 12 Piranthus planolancis (specimen AS 19.5970) 13 - 20 carapace (all females except 15) 13 Bavia nessagyna (specimen IDWM. 20004) 14 Indopadilla redunca (holotype IDWM. 20011) 15 Stagetillus opaciceps male (specimen JK. 13.02.26.0017) 16 Stagetillus cf. opaciceps female (specimen MRB 079). 17 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 18 Padillothorax mulu (from Mulu Nat. Pk.) 19 Maripanthus reinholdae (specimen SWK 12 - 1934) 20 Piranthus planolancis (specimen AS 19.5940) 21 - 28 Metatarsus of first leg, retrolateral view. All female except 24 male. All are of left legs except 23, 25, 28 which are of right leg, digitally flipped 21 Bavia nessagyna (specimen IDWM. 20004) 22 Indopadilla kahariana (specimen SWK 12 - 1876) 23 Stagetillus cf. opaciceps (specimen MRB 079) 24 Padillothorax semiostrinus (specimen SWK 12 - EP 0105) 25 Padillothorax mulu (specimen SWK 12 - EP 0105) 26 Maripanthus reinholdae (specimen SWK 12 - 1934) 27 Maripanthus draconis (from Gunung Belemut, Johor) 28 Piranthus planolancis (specimen AS 19.5940) 29 - 35 prolateral surface of male first leg femur. 29 Bavia cf. capistrata (specimen from Singapore) 30 Indopadilla kahariana (specimen from Lambir Hills Nat. Pk.) 31 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 32 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 33 Padillothorax badut (specimen SWK 12 - 4688) 34 Maripanthus draconis (specimen from Johor, Gunung Lambak) 35 Piranthus bakau (holotype). Scale bars: 0.1 mm (6 - 12, 21 - 35), 1.0 mm (4, 5, 13 - 20)." figureDoi="10.3897/zookeys.1004.57526.figures4-35" httpUri="https://binary.pensoft.net/fig/487533" pageId="0" pageNumber="27">Figs 19</figureCitation>
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<figureCitation id="0A342E90D4AE4664F3A79FCA2329E0F6" captionStart="Figures 4–35" captionStartId="F2" captionText="Figures 4 - 35. Variation in traits among baviines 4, 5 oblique views of female prosoma 4 Bavia nessagyna (specimen IDWM. 20004; note convex front face of chelicerae, and typical thorax) 5 Indopadilla redunca (holotype IDWM. 20011; note concave front face of chelicerae, and thoracic bulges) 6 - 12 left male endite (9 also with chelicera) 6 Bavia nessagyna (holotype IDWM. 20005) 7 Indopadilla redunca (specimen SWK 12 - M 0009) 8 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 9 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 10 Padillothorax badut (specimen SWK 12 - 4688) 11 Maripanthus jubatus (specimen AS 19.4373) 12 Piranthus planolancis (specimen AS 19.5970) 13 - 20 carapace (all females except 15) 13 Bavia nessagyna (specimen IDWM. 20004) 14 Indopadilla redunca (holotype IDWM. 20011) 15 Stagetillus opaciceps male (specimen JK. 13.02.26.0017) 16 Stagetillus cf. opaciceps female (specimen MRB 079). 17 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 18 Padillothorax mulu (from Mulu Nat. Pk.) 19 Maripanthus reinholdae (specimen SWK 12 - 1934) 20 Piranthus planolancis (specimen AS 19.5940) 21 - 28 Metatarsus of first leg, retrolateral view. All female except 24 male. All are of left legs except 23, 25, 28 which are of right leg, digitally flipped 21 Bavia nessagyna (specimen IDWM. 20004) 22 Indopadilla kahariana (specimen SWK 12 - 1876) 23 Stagetillus cf. opaciceps (specimen MRB 079) 24 Padillothorax semiostrinus (specimen SWK 12 - EP 0105) 25 Padillothorax mulu (specimen SWK 12 - EP 0105) 26 Maripanthus reinholdae (specimen SWK 12 - 1934) 27 Maripanthus draconis (from Gunung Belemut, Johor) 28 Piranthus planolancis (specimen AS 19.5940) 29 - 35 prolateral surface of male first leg femur. 29 Bavia cf. capistrata (specimen from Singapore) 30 Indopadilla kahariana (specimen from Lambir Hills Nat. Pk.) 31 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 32 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 33 Padillothorax badut (specimen SWK 12 - 4688) 34 Maripanthus draconis (specimen from Johor, Gunung Lambak) 35 Piranthus bakau (holotype). Scale bars: 0.1 mm (6 - 12, 21 - 35), 1.0 mm (4, 5, 13 - 20)." figureDoi="10.3897/zookeys.1004.57526.figures4-35" httpUri="https://binary.pensoft.net/fig/487533" pageId="0" pageNumber="27">, 26</figureCitation>
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<figureCitation id="9BEDCB2989737257F0B65A508D1A75F2" captionStart="Figures 215–224" captionStartId="F23" captionText="Figures 215 - 224. Maripanthus reinholdae sp. nov. and M. cf. menghaiensis 215 - 222 Maripanthus reinholdae 215 male left palp, prolateral view (holotype JK. 11.12.24.0006) 216 same, ventral view 217 same, retrolateral view 218 epigyne, ventral (specimen SWK 12 - 1934) 219 vulva, dorsal 220, 221 female SWK 12 - 1934 222 holotype male 223, 224 male M. cf. menghaiensis (specimen IDWM. 20013). Scale bars on genitalia 0.1 mm; on body 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures215-224" httpUri="https://binary.pensoft.net/fig/487554" pageId="0" pageNumber="27">, 215-222</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="7308FA0918FC57D34021B904A93A5C58" pageId="0" pageNumber="27" type="type material">
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<paragraph id="C7479F2D4872BB43E4FCD630A8DC8143" pageId="0" pageNumber="27">Type material.</paragraph>
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<paragraph id="66F1495C375B051901256543F4FD501D" pageId="0" pageNumber="27">
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<emphasis id="3E8BCB7E592F1C9F9C30AE212C248A6F" bold="true" italics="true" pageId="0" pageNumber="27">Holotype</emphasis>
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male (specimen JK 11.12.24.0006), in LKCNHM, from Brunei: Ulu Temburong National Park, Canopy Walk Trail,
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<geoCoordinate id="6A5185F1AA72756C0E719B87FFC8EBFB" degrees="4.5522" direction="north" orientation="latitude" precision="5" value="4.5522">4.5522°N</geoCoordinate>
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,
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<geoCoordinate id="FF1EA2F589E478285982B517A5AB20A3" degrees="115.1578" direction="east" orientation="longitude" precision="5" value="115.1578">115.1578°E</geoCoordinate>
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, J. K. H. Koh 24 Dec. 2011.
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<emphasis id="2D7F409D1E63A42B3EB2AD0385B81D83" bold="true" italics="true" pageId="0" pageNumber="27">Paratype</emphasis>
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female (specimen SWK12-1934, in UBCZ), from Malaysia: Sarawak: Mulu Nat. Pk., Botanical Trail,
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<geoCoordinate id="76E3457EA3F409C545C443144D37043E" degrees="4.0406" direction="north" orientation="latitude" precision="5" value="4.0406">4.0406°N</geoCoordinate>
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,
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<geoCoordinate id="EB1DCF027B2D78346C1A23B6CE768870" degrees="114.8170" direction="east" orientation="longitude" precision="5" value="114.817">114.8170°E</geoCoordinate>
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to
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<geoCoordinate id="E842D6F6CA9F45DA9053183E68E285AB" degrees="4.0404" direction="north" orientation="latitude" precision="5" value="4.0404">4.0404°N</geoCoordinate>
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,
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<geoCoordinate id="717CA51A8D8B830C2D963970D591BB05" degrees="114.8176" direction="east" orientation="longitude" precision="5" value="114.8176">114.8176°E</geoCoordinate>
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, 50 m el. 16 March 2012 Maddison/Piascik/Ang WPM#12-044.
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<emphasis id="665D26B07AD81FABEE5D0FC530386262" bold="true" italics="true" pageId="0" pageNumber="27">Paratype</emphasis>
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female (specimen SWK12-1991, in UBCZ), also from Mulu Nat. Pk., Kenyalang Trail,
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<geoCoordinate id="416D667945F62EBCE09F325F64CC6753" degrees="4.0229" direction="north" orientation="latitude" precision="5" value="4.0229">4.0229°N</geoCoordinate>
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,
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<geoCoordinate id="2D1338A9D11BA757850A5AAC3300CB4B" degrees="114.8128" direction="east" orientation="longitude" precision="5" value="114.8128">114.8128°E</geoCoordinate>
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to
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<geoCoordinate id="503F5BF944DE8C2E81341285317F6F6F" degrees="4.0228" direction="north" orientation="latitude" precision="5" value="4.0228">4.0228°N</geoCoordinate>
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,
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<geoCoordinate id="752B7A90C5CC6EE7EA6FF409377DBC18" degrees="114.8134" direction="east" orientation="longitude" precision="5" value="114.8134">114.8134°E</geoCoordinate>
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, 55 m el. 17 March 2012 Maddison/Piascik/Ang WPM#12-047.
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</paragraph>
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</subSubSection>
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<subSubSection id="EE0522397F3503C7C83D081A971FC528" pageId="0" pageNumber="27" type="etymology">
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<paragraph id="BCE8DA696C779E30B9785F797AC3711A" pageId="0" pageNumber="27">Etymology.</paragraph>
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<paragraph id="88C72DAFEB2EC4303C031E939DB81822" pageId="0" pageNumber="27">
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Named in honour of Christa Deeleman-Reinhold, whose extensive work on southeast Asian spiders has greatly increased our knowledge of the
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<normalizedToken id="650B713069FE31FF9A0CCE5C323EC547" originalValue="area’s">area's</normalizedToken>
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fauna. She has discovered and described over 350 new species, including 54 new salticids.
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<emphasis id="184996FFB35E3925B456A77FE21B12B2" italics="true" pageId="0" pageNumber="27">Other names</emphasis>
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: In
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<bibRefCitation id="654ED3DD0B8F6A194EF5B9C46221C15F" author="Maddison, WP" journalOrPublisher="Journal of Arachnology" pageId="0" pageNumber="27" publicationUrl="http://salticidae.org/salticidImages/" refId="B23" refString="Maddison, WP, 2015b. Salticid images. http://salticidae.org/salticidImages/" title="Salticid images." url="http://salticidae.org/salticidImages/" year="2015 b">Maddison (2015b)</bibRefCitation>
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and
|
||
<normalizedToken id="B2D412BB0DCF24068A75F483F475914A" originalValue="WPM’s">WPM's</normalizedToken>
|
||
field or lab notebooks the informal code for this species was
|
||
<normalizedToken id="F67AAA42F7CE5726AED73FA086E665CB" originalValue="“BVBGB”">"BVBGB"</normalizedToken>
|
||
. This species is shown in
|
||
<bibRefCitation id="A64D88A88AA759111A9512F6CA48DFF4" author="Koh, JKH" journalOrPublisher="Sabah Forestry Department, city" pageId="0" pageNumber="27" refId="B18" refString="Koh, JKH, Bay, N, 2019. Borneo Spiders: A photographic field guide. Sabah Forestry Department, city" title="Borneo Spiders: A photographic field guide." year="2019">Koh and Bay 2019</bibRefCitation>
|
||
as "
|
||
<taxonomicName id="5EA83A76BD6BF8AC2E1BFAD1A998430A" authorityName="Simon" authorityYear="1877" class="Arachnida" family="Pittosporaceae" genus="Bavia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bavia" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="330F4E4A66045A5E6DC3101DDA955B8C" italics="true" pageId="0" pageNumber="27">Bavia</emphasis>
|
||
</taxonomicName>
|
||
" sp. C Two-lined long-bellied jumping spider; the upper photograph on p. 208 is of the holotype.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DAF461E36801866155F9E842167B5637" pageId="0" pageNumber="27" type="diagnosis">
|
||
<paragraph id="57C7154EF1213A21F2D930E1FE1DDA3F" pageId="0" pageNumber="27">Diagnosis.</paragraph>
|
||
<paragraph id="85EFCB57A5D02150EC41B1CBCF638CA9" pageId="0" pageNumber="27">
|
||
Very similar to
|
||
<taxonomicName id="4F98B3C3A8AF5B857B0B42A7C06E7FC7" lsidName="M. menghaiensis" pageId="0" pageNumber="27" rank="species" species="menghaiensis">
|
||
<emphasis id="8D3CE367B0CA519ABC414536439C7861" italics="true" pageId="0" pageNumber="27">M. menghaiensis</emphasis>
|
||
</taxonomicName>
|
||
, and like it smaller and more
|
||
<taxonomicName id="BC3BE51A2C024E47DA8C4B8BF9FDF7A6" authorityName="Caleb & Sankaran" authorityYear="2019" class="Arachnida" family="Salticidae" genus="Indopadilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Indopadilla" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A6D2FE22C9A6A6AA53679CA458C53110" italics="true" pageId="0" pageNumber="27">Indopadilla</emphasis>
|
||
</taxonomicName>
|
||
-like in body form than
|
||
<taxonomicName id="D1B05717C9D59B4A008BAA6A410D39D0" lsidName="M. draconis" pageId="0" pageNumber="27" rank="species" species="draconis">
|
||
<emphasis id="470F3C100BD96A4BFC7AD8D546A3C849" italics="true" pageId="0" pageNumber="27">M. draconis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C53D35D1EF674A7E09A8CD88A85C4576" lsidName="M. jubatus" pageId="0" pageNumber="27" rank="species" species="jubatus">
|
||
<emphasis id="809E1EC6DEABA5713235578AB1225DE8" italics="true" pageId="0" pageNumber="27">M. jubatus</emphasis>
|
||
</taxonomicName>
|
||
(shorter first legs, more elongate abdomen). Differs from
|
||
<taxonomicName id="EC8887394434BD709751134B94990C71" lsidName="M. menghaiensis" pageId="0" pageNumber="27" rank="species" species="menghaiensis">
|
||
<emphasis id="69C10C94A512DA6433267A656128CC60" italics="true" pageId="0" pageNumber="27">M. menghaiensis</emphasis>
|
||
</taxonomicName>
|
||
in the longer epigynal openings, and in details of the
|
||
<normalizedToken id="91801DBABF4F7C1CA5390BD9BA260F2C" originalValue="palp’s">palp's</normalizedToken>
|
||
bulb. In view from the retrolateral, the embolus is first directed to the distal then quickly turns dorsally (
|
||
<taxonomicName id="C5AB864E910B63BB682F71AA19FDBFB4" lsidName="M. menghaiensis" pageId="0" pageNumber="27" rank="species" species="menghaiensis">
|
||
<emphasis id="7031B50A3BF626D68010A44391321C57" italics="true" pageId="0" pageNumber="27">M. menghaiensis</emphasis>
|
||
</taxonomicName>
|
||
, embolus begins toward the dorsal). When the embolus comes out from behind the tegulum it is directed slightly proximally (slightly distally in
|
||
<taxonomicName id="9A3049E0D1E978B66D89369CA06538A2" lsidName="M. menghaiensis" pageId="0" pageNumber="27" rank="species" species="menghaiensis">
|
||
<emphasis id="6FA4C7A5FB4541AAD8586C3409783E30" italics="true" pageId="0" pageNumber="27">M. menghaiensis</emphasis>
|
||
</taxonomicName>
|
||
). The embolus is thinner near the tip than in
|
||
<taxonomicName id="106DB748AB29D5BAD0C22C7239A365F2" lsidName="M. menghaiensis" pageId="0" pageNumber="27" rank="species" species="menghaiensis">
|
||
<emphasis id="40EF7145BEE8B5FB5948389785F9ABB1" italics="true" pageId="0" pageNumber="27">M. menghaiensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="82251EA750B26B2966526F2F8D9026B0" pageId="0" pageNumber="27" type="description">
|
||
<paragraph id="15EB5A255FB3B170E1DFF33788D4759E" pageId="0" pageNumber="27">Description.</paragraph>
|
||
<paragraph id="C87A492F6C8A7504E5F21E3673DC068C" pageId="0" pageNumber="27">
|
||
<emphasis id="0AA6D846A4F96280644584A66F0DCB04" bold="true" pageId="0" pageNumber="27">Male</emphasis>
|
||
(based on holotype, specimen JK.11.12.24.0006; living holotype shown on p. 208 of
|
||
<bibRefCitation id="17BE03E0B353F9E2DA4FBA433A0714BC" author="Koh, JKH" journalOrPublisher="Sabah Forestry Department, city" pageId="0" pageNumber="27" refId="B18" refString="Koh, JKH, Bay, N, 2019. Borneo Spiders: A photographic field guide. Sabah Forestry Department, city" title="Borneo Spiders: A photographic field guide." year="2019">Koh and Bay 2019</bibRefCitation>
|
||
). Carapace length 2.9; abdomen length 4.0.
|
||
<emphasis id="A2C4A112C834CC196F030F01BAAD3BCD" bold="true" italics="true" pageId="0" pageNumber="27">Carapace</emphasis>
|
||
(Fig.
|
||
<figureCitation id="4B391A7B8E391A0AE7C9A69979F29677" captionStart="Figures 215–224" captionStartId="F23" captionText="Figures 215 - 224. Maripanthus reinholdae sp. nov. and M. cf. menghaiensis 215 - 222 Maripanthus reinholdae 215 male left palp, prolateral view (holotype JK. 11.12.24.0006) 216 same, ventral view 217 same, retrolateral view 218 epigyne, ventral (specimen SWK 12 - 1934) 219 vulva, dorsal 220, 221 female SWK 12 - 1934 222 holotype male 223, 224 male M. cf. menghaiensis (specimen IDWM. 20013). Scale bars on genitalia 0.1 mm; on body 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures215-224" httpUri="https://binary.pensoft.net/fig/487554" pageId="0" pageNumber="27">222</figureCitation>
|
||
) black except for brown around fovea. Cream scales form a band crossing the thorax behind the PLEs, a small streak on the midline of the thoracic slope, and a narrow band along the lateral margins.
|
||
<emphasis id="E6B7E8D8CE4A4E2D01C4E2B27125F7C7" bold="true" italics="true" pageId="0" pageNumber="27">Clypeus</emphasis>
|
||
dark but centrally with a cluster of erect white scales that overhang the chelicerae.
|
||
<emphasis id="C2243251DB52FF83714849EA10E4D619" bold="true" italics="true" pageId="0" pageNumber="27">Chelicerae</emphasis>
|
||
simple, vertical, dark. Plurident.
|
||
<emphasis id="47B2BAC1FADA1670DE23BF130EC683D2" bold="true" italics="true" pageId="0" pageNumber="27">Palp</emphasis>
|
||
pale except for dark femur. Embolus long, arising retrolaterally before curling under the bulb prolaterally then proceeding distally (Figs
|
||
<figureCitation id="7264FFF0997C8A197A6C6FF0C8CD8DB6" captionStart="Figures 215–224" captionStartId="F23" captionText="Figures 215 - 224. Maripanthus reinholdae sp. nov. and M. cf. menghaiensis 215 - 222 Maripanthus reinholdae 215 male left palp, prolateral view (holotype JK. 11.12.24.0006) 216 same, ventral view 217 same, retrolateral view 218 epigyne, ventral (specimen SWK 12 - 1934) 219 vulva, dorsal 220, 221 female SWK 12 - 1934 222 holotype male 223, 224 male M. cf. menghaiensis (specimen IDWM. 20013). Scale bars on genitalia 0.1 mm; on body 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures215-224" httpUri="https://binary.pensoft.net/fig/487554" pageId="0" pageNumber="27">215-217</figureCitation>
|
||
). A short spur diverges from the embolus before it turns distally (Figs
|
||
<figureCitation id="E69958F2DB393447375FE0A3532E54A4" captionStart="Figures 215–224" captionStartId="F23" captionText="Figures 215 - 224. Maripanthus reinholdae sp. nov. and M. cf. menghaiensis 215 - 222 Maripanthus reinholdae 215 male left palp, prolateral view (holotype JK. 11.12.24.0006) 216 same, ventral view 217 same, retrolateral view 218 epigyne, ventral (specimen SWK 12 - 1934) 219 vulva, dorsal 220, 221 female SWK 12 - 1934 222 holotype male 223, 224 male M. cf. menghaiensis (specimen IDWM. 20013). Scale bars on genitalia 0.1 mm; on body 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures215-224" httpUri="https://binary.pensoft.net/fig/487554" pageId="0" pageNumber="27">215</figureCitation>
|
||
,
|
||
<figureCitation id="B31A6294F1F77F81320DB7CA7FE7D193" captionStart="Figures 215–224" captionStartId="F23" captionText="Figures 215 - 224. Maripanthus reinholdae sp. nov. and M. cf. menghaiensis 215 - 222 Maripanthus reinholdae 215 male left palp, prolateral view (holotype JK. 11.12.24.0006) 216 same, ventral view 217 same, retrolateral view 218 epigyne, ventral (specimen SWK 12 - 1934) 219 vulva, dorsal 220, 221 female SWK 12 - 1934 222 holotype male 223, 224 male M. cf. menghaiensis (specimen IDWM. 20013). Scale bars on genitalia 0.1 mm; on body 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures215-224" httpUri="https://binary.pensoft.net/fig/487554" pageId="0" pageNumber="27">216</figureCitation>
|
||
). RTA vertical except for a bend and curl distally. Endite with small sharp corner, similar but smaller to that in the larger species of
|
||
<taxonomicName id="461275C414F2A49F3F265AB3F3FD926D" authorityName="Maddison" authorityYear="2020" class="Magnoliopsida" family="Pittosporaceae" genus="Maripanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Maripanthus" order="Apiales" pageId="0" pageNumber="27" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="8834F480D9A9B65219A6DFBCA9F6F8A8" italics="true" pageId="0" pageNumber="27">Maripanthus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="2647608A9B3ABDA9D69260254EBA2E21" bold="true" italics="true" pageId="0" pageNumber="27">Legs</emphasis>
|
||
pale honey coloured except for first, which is black in all segments except the tarsus. Third and fourth legs darker near more distal joints.
|
||
<emphasis id="DFB8FE82AB8A598E3D8B09EF9619B975" bold="true" italics="true" pageId="0" pageNumber="27">Abdomen</emphasis>
|
||
thin, dark, with a pale mid-dorsal band, just lateral to which are indistinct longitudinal streaks of white scales.
|
||
</paragraph>
|
||
<paragraph id="C199AC953CD3E2B1D978A4761FBA07A1" pageId="0" pageNumber="27">
|
||
<emphasis id="4DA08F206D3F20A5553D8B1CE2F3EEB3" bold="true" pageId="0" pageNumber="27">Female</emphasis>
|
||
(based on paratype, specimen SWK12-1934). Carapace length 3.7; abdomen length 5.0.
|
||
<emphasis id="315BEEBABE0C80C40ADF2D50BDF6E248" bold="true" italics="true" pageId="0" pageNumber="27">Carapace</emphasis>
|
||
as in male, but slightly paler in integument.
|
||
<emphasis id="1FA35BDB8C7E80EDE4770D2B9FBB76EA" bold="true" italics="true" pageId="0" pageNumber="27">Chelicerae</emphasis>
|
||
with three promarginal and five retromarginal teeth.
|
||
<emphasis id="E0207CC7E1A1390D0F60620D1C2B3EC7" bold="true" italics="true" pageId="0" pageNumber="27">Legs</emphasis>
|
||
as in male but with first legs only slightly darker than the others.
|
||
<emphasis id="E5478B3121709B456D8096E7E2FDE634" bold="true" italics="true" pageId="0" pageNumber="27">Abdomen</emphasis>
|
||
brown with central pale chevroned band, and thin white streaks as in male.
|
||
<emphasis id="20EF3DD3B247EEAE4B03457F4AA5F86A" bold="true" italics="true" pageId="0" pageNumber="27">Epigyne</emphasis>
|
||
(Fig.
|
||
<figureCitation id="0AD90E93C74A623F2FFC9FA3B53D54C1" captionStart="Figures 215–224" captionStartId="F23" captionText="Figures 215 - 224. Maripanthus reinholdae sp. nov. and M. cf. menghaiensis 215 - 222 Maripanthus reinholdae 215 male left palp, prolateral view (holotype JK. 11.12.24.0006) 216 same, ventral view 217 same, retrolateral view 218 epigyne, ventral (specimen SWK 12 - 1934) 219 vulva, dorsal 220, 221 female SWK 12 - 1934 222 holotype male 223, 224 male M. cf. menghaiensis (specimen IDWM. 20013). Scale bars on genitalia 0.1 mm; on body 1.0 mm." figureDoi="10.3897/zookeys.1004.57526.figures215-224" httpUri="https://binary.pensoft.net/fig/487554" pageId="0" pageNumber="27">218</figureCitation>
|
||
) similar in conformation to
|
||
<taxonomicName id="8E12A7CCC6436EF0A334DCC04536B266" lsidName="M. draconis" pageId="0" pageNumber="27" rank="species" species="draconis">
|
||
<emphasis id="0C39435A6DDFA1820F7B73DAB01E05C1" italics="true" pageId="0" pageNumber="27">M. draconis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="75D9185D429088A839A0D140932968D8" lsidName="M. jubatus" pageId="0" pageNumber="27" rank="species" species="jubatus">
|
||
<emphasis id="0275BCE846CB8B7E142DF61C8384C4DA" italics="true" pageId="0" pageNumber="27">M. jubatus</emphasis>
|
||
</taxonomicName>
|
||
, but with atria smaller and copulatory ducts not so compacted.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="71F4381EB0FC45513238D2046A33076D" pageId="0" pageNumber="27" type="materials_examined">
|
||
<paragraph id="223CE0ED9055B9C24BEC0261A1D42CF0" pageId="0" pageNumber="27">Additional material examined.</paragraph>
|
||
<paragraph id="4563D569CCEC2CFAD16A2D81E4A43A2D" pageId="0" pageNumber="27">
|
||
One female (specimen JK.12.02.04.0010, in LKCNHM) from Brunei: Belait, Trail To Wasai Teraja Secondary Forest,
|
||
<geoCoordinate id="FB2DF7AB013C5BC9E18D86E9903B4CE3" degrees="4.2911" direction="north" orientation="latitude" precision="5" value="4.2911">4.2911°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="5A8CA782FB9E8EE3C5FF245C83636C47" degrees="114.4231" direction="east" orientation="longitude" precision="5" value="114.4231">114.4231°E</geoCoordinate>
|
||
, J. K. H. Koh 4 February 2012.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |