308 lines
38 KiB
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308 lines
38 KiB
XML
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<mods:title id="D9B3A70C4D742934CCC7D359CAAFCDF9">On the taxonomy of Lygosoma bampfyldei Bartlett, 1895 (Squamata: Scincidae) with descriptions of new species from Borneo and Peninsular Malaysia and the resurrection of Lygosoma schneideri Werner, 1900</mods:title>
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<treatment id="AA2B87D3D914FF858682F936FB0FFF04" ID-DOI="http://doi.org/10.5281/zenodo.5988922" ID-GBIF-Taxon="146019955" ID-Zenodo-Dep="5988922" LSID="urn:lsid:plazi:treatment:AA2B87D3D914FF858682F936FB0FFF04" httpUri="http://treatment.plazi.org/id/AA2B87D3D914FF858682F936FB0FFF04" lastPageId="10" lastPageNumber="537" pageId="8" pageNumber="536">
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<heading id="797581A9D914FF878682F936FDEDF972" bold="true" box="[151,580,1670,1697]" fontSize="11" level="1" pageId="8" pageNumber="536" reason="1">
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<taxonomicName id="E5824D46D914FF878682F936FDEDF972" authority="Werner, 1900" authorityName="Werner" authorityYear="1900" box="[151,580,1670,1697]" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="536" phylum="Chordata" rank="species" species="schneideri">
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<emphasis id="10F6EAD7D914FF878682F936FDEDF972" bold="true" box="[151,580,1670,1697]" pageId="8" pageNumber="536">
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<emphasis id="10F6EAD7D914FF878682F936FE3FF973" bold="true" box="[151,406,1670,1696]" italics="true" pageId="8" pageNumber="536">Lygosoma schneideri</emphasis>
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<bibRefCitation id="46134B34D914FF878788F937FDEDF972" box="[413,580,1671,1697]" pageId="8" pageNumber="536" refId="ref9532" refString="Werner, F. (1900) Reptilien und batrachier aus Sumatra, gesammelt von Herrn Gustav Schneider jr. im Jaher 1897 - 98. Zoologische Jahbucher System, 7, 487 - 502." type="journal article">Werner, 1900</bibRefCitation>
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<paragraph id="223D36C5D914FF878682F91AFE09F911" blockId="8.[151,580,1670,1766]" box="[151,416,1705,1730]" pageId="8" pageNumber="536">Sumatran Supple Skink</paragraph>
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<paragraph id="223D36C5D914FF878682F97DFEBFF935" blockId="8.[151,580,1670,1766]" box="[151,278,1741,1766]" pageId="8" pageNumber="536">
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<figureCitation id="BAB92A40D914FF878682F97DFF4AF935" box="[151,227,1741,1766]" captionStart="FIGURE 4" captionStartId="8.[151,250,1451,1473]" captionTargetBox="[176,1396,723,1417]" captionTargetId="figure@8.[151,1436,700,1430]" captionTargetPageId="8" captionText="FIGURE 4. Comparisons of body stature of the types of A, Lygosoma scheideri (BM 4743) from Djpura, Indragiri, Sumatra, Indonesia; B, syntype of Lygosoma bampfyldei (BMNH 1946.8.10.84) from the Rejan River, Sarawak, East Malaysia; C, L. kinabatanganensis sp. nov. (FMNH 76226) from the Kinabatangan District, Deramakot camp (Deramakot Sabah Forestry Department), Sabah, East Malaysia; and D, L. peninsulare sp. nov. (LSUHC 13857) from 13.5 km east of Jeli, Kelantan, Peninsular Malaysia." httpUri="https://zenodo.org/record/1298212/files/figure.png" pageId="8" pageNumber="536">Figs. 4</figureCitation>
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&
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<figureCitation id="BAB92A40D914FF878712F97DFEBFF935" box="[263,278,1741,1766]" captionStart="FIGURE 5" captionStartId="9.[151,250,1095,1117]" captionTargetBox="[151,1411,210,1047]" captionTargetId="figure@9.[151,1436,195,1073]" captionTargetPageId="9" captionText="FIGURE 5. A. Right lateral view of the heads of A, the syntype of Lygosoma bampfyldei (BMNH 1946.8.10.84) from the Rejan River, Sarawak, East Malaysia; B, the holotype of L. kinabatanganensis sp. nov. (FMNH 76226) from the Kinabatangan District, Deramakot camp (Deramakot Sabah Forestry Department), Sabah, East Malaysia; C, L. peninsulare sp. nov. (LSUHC 13857) from 13.5 km east of Jeli, Kelantan, Peninsular Malaysia; and D, L. scheideri (BM 4743) from Djpura, Indragiri, Sumatra, Indonesia. Photos by LLG." httpUri="https://zenodo.org/record/1298214/files/figure.png" pageId="8" pageNumber="536">5</figureCitation>
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<taxonomicName id="E5824D46D914FF878682F8A6FDEBF8F8" authority="Boulenger, 1912: 93" authorityName="Boulenger" authorityPageNumber="93" authorityYear="1912" box="[151,578,1813,1835]" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="536" phylum="Chordata" rank="species" species="bampfyldii">
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<emphasis id="10F6EAD7D914FF878682F8A6FED9F8F8" box="[151,368,1813,1835]" italics="true" pageId="8" pageNumber="536">Lygosoma bampfyldii</emphasis>
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:93
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:36
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<emphasis id="10F6EAD7D914FF878682F885FEDEF899" box="[151,375,1844,1866]" italics="true" pageId="8" pageNumber="536">Lygosoma bampfyldei</emphasis>
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de Rooji, 1915:263
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;
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:915
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;
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Welch
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<emphasis id="10F6EAD7D914FF878547F885FC24F899" box="[850,909,1844,1866]" italics="true" pageId="8" pageNumber="536">et al.,</emphasis>
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1990
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:83
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</treatmentCitation>
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;
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<bibRefCitation id="46134B34D914FF8785E2F885FB7CF899" box="[1015,1237,1844,1866]" pageId="8" pageNumber="536" refId="ref7791" refString="Das, I. & Norsham, Y. (2007) Status of knowledge of the Malaysian herpetofauna. In: Chua, L. (Ed.), Status of Biological Diversity inMalaysia and Threat Assessment of Plant Species in Malaysia. Forest Research Institute Malaysia, Kepong, pp. 31 - 81. https: // doi. org / 10.21475 / poj. 10.04.17. pne 32" type="book chapter">Das & Yaakob, 2007</bibRefCitation>
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:78
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;
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:238
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;
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:613
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.
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<taxonomicName id="E5824D46D914FF878682F8C2FE5FF854" authority="Smith 1937: 228" authorityName="Smith" authorityPageNumber="228" authorityYear="1937" box="[151,502,1905,1927]" class="Reptilia" family="Scincidae" genus="Riopa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="536" phylum="Chordata" rank="species" species="bampfyldei">
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<emphasis id="10F6EAD7D914FF878682F8C2FEE3F854" box="[151,330,1905,1927]" italics="true" pageId="8" pageNumber="536">Riopa bampfyldei</emphasis>
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:228
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;
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<treatmentCitation id="A32310D4D914FF878415F8C1FCFFF854" author="" box="[512,854,1905,1927]" page="275" pageId="8" pageNumber="536">
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<bibRefCitation id="46134B34D914FF878415F8C1FC83F854" box="[512,810,1905,1927]" pageId="8" pageNumber="536" refId="ref9112" refString="Manthey, U. & W. Grossman (1997) Amphibien and Reptilien S ¸ dostasiens." type="book">Manthey & Grossmann 1997</bibRefCitation>
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:275
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;
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<treatmentCitation id="A32310D4D914FF878574F8C1FBF2F854" author="" box="[865,1115,1905,1927]" page="27" pageId="8" pageNumber="536">
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<bibRefCitation id="46134B34D914FF878574F8C1FB95F854" box="[865,1084,1905,1927]" pageId="8" pageNumber="536" refId="ref7710" refString="Chan-ard, T., Grossman, W., Gumprecht, A. & Schultz, K. - D. (1999) Amphibians and Reptiles of Peninsular Malaysia and Thailand. Bushmaster Publications, Wuerselen, 240 pp." type="book">
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Chan-ard
|
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<emphasis id="10F6EAD7D914FF8785D0F8C2FC56F854" box="[965,1023,1905,1927]" italics="true" pageId="8" pageNumber="536">et al.,</emphasis>
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1999
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:27
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="223D36C5D914FF878682F820FDE9F875" blockId="8.[151,1436,1813,1989]" box="[151,576,1936,1958]" pageId="8" pageNumber="536">
|
||
<treatmentCitationGroup id="029211EBD914FF878682F820FDE9F875" box="[151,576,1936,1958]" pageId="8" pageNumber="536">
|
||
<taxonomicName id="E5824D46D914FF878682F820FDE9F875" authority="Mittleman, 1952: 22" authorityName="Mittleman" authorityPageNumber="22" authorityYear="1952" box="[151,576,1936,1958]" class="Reptilia" family="Scincidae" genus="Mochlus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="536" phylum="Chordata" rank="species" species="bampfyledei">
|
||
<emphasis id="10F6EAD7D914FF878682F820FED9F875" box="[151,368,1936,1958]" italics="true" pageId="8" pageNumber="536">Mochlus bampfyledei</emphasis>
|
||
<treatmentCitation id="A32310D4D914FF878763F820FDE9F875" author="" box="[374,576,1936,1958]" page="22" pageId="8" pageNumber="536">
|
||
<bibRefCitation id="46134B34D914FF878763F820FD88F875" box="[374,545,1936,1958]" pageId="8" pageNumber="536" refId="ref9141" refString="Mittleman, M. B. (1952) A generic synopsis of the lizards of the subfamily Lygosominae. Smithsonian Miscellaneous Collections, 117, 1 - 35." type="journal article">Mittleman, 1952</bibRefCitation>
|
||
:22
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="223D36C5D914FF878682F800FDD3F816" blockId="8.[151,1436,1813,1989]" box="[151,634,1967,1989]" pageId="8" pageNumber="536">
|
||
<treatmentCitationGroup id="029211EBD914FF878682F800FDD3F816" box="[151,634,1967,1989]" pageId="8" pageNumber="536">
|
||
<taxonomicName id="E5824D46D914FF878682F800FDD3F816" authority="Denzer & Manthey, 1991: 317" authorityName="Denzer & Manthey" authorityPageNumber="317" authorityYear="1991" box="[151,634,1967,1989]" class="Reptilia" family="Scincidae" genus="Riopa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="536" phylum="Chordata" rank="species" species="bampfyldii">
|
||
<emphasis id="10F6EAD7D914FF878682F800FEEFF816" box="[151,326,1967,1989]" italics="true" pageId="8" pageNumber="536">Riopa bampfyldii</emphasis>
|
||
<treatmentCitation id="A32310D4D914FF878759F800FDD3F816" author="" box="[332,634,1967,1989]" page="317" pageId="8" pageNumber="536">
|
||
<bibRefCitation id="46134B34D914FF878759F800FDE7F816" box="[332,590,1967,1989]" pageId="8" pageNumber="536" refId="ref8050" refString="Denzer, W. & Manthey, U. (1991) A nominal checklist of the lizards inhabiting Peninsular Malaysia and Singapore. Raffles Bulletin of Zoology, 39, 309 - 322." type="journal article">Denzer & Manthey, 1991</bibRefCitation>
|
||
:317
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="76FD664DD915FF868682FBF7FDAEFB0B" httpUri="https://zenodo.org/record/1298214/files/figure.png" pageId="9" pageNumber="537" startId="9.[151,250,1095,1117]" targetBox="[151,1411,210,1047]" targetPageId="9">
|
||
<paragraph id="223D36C5D915FF868682FBF7FDAEFB0B" blockId="9.[151,1436,1095,1240]" pageId="9" pageNumber="537">
|
||
<emphasis id="10F6EAD7D915FF868682FBF7FEBCFB8F" bold="true" box="[151,277,1095,1117]" pageId="9" pageNumber="537">FIGURE 5.</emphasis>
|
||
A. Right lateral view of the heads of A, the syntype of
|
||
<taxonomicName id="E5824D46D915FF868573FBF8FBEEFB8E" authorityName="de Rooji" authorityYear="1915" box="[870,1095,1095,1117]" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="537" phylum="Chordata" rank="species" species="bampfyldei">
|
||
<emphasis id="10F6EAD7D915FF868573FBF8FBEEFB8E" box="[870,1095,1095,1117]" italics="true" pageId="9" pageNumber="537">Lygosoma bampfyldei</emphasis>
|
||
</taxonomicName>
|
||
(BMNH 1946.8.10.84) from the Rejan River, Sarawak, East Malaysia; B, the holotype of
|
||
<taxonomicName id="E5824D46D915FF8684CCFBD7FC1BFBAF" authority="Grismer & Quah & Duzulkafly & Yambun, 2018" authorityName="Grismer & Quah & Duzulkafly & Yambun" authorityYear="2018" box="[729,946,1126,1148]" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="537" phylum="Chordata" rank="species" species="kinabatanganensis" status="sp. nov.">
|
||
<emphasis id="10F6EAD7D915FF8684CCFBD7FC1BFBAF" box="[729,946,1126,1148]" italics="true" pageId="9" pageNumber="537">L. kinabatanganensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0BC557ACD915FF8685ADFBD7FBAEFBAF" box="[952,1031,1127,1148]" pageId="9" pageNumber="537" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(FMNH 76226) from the Kinabatangan District, Deramakot camp (Deramakot Sabah Forestry Department), Sabah, East Malaysia; C,
|
||
<taxonomicName id="E5824D46D915FF868241FB36FB4CFB49" authority="Grismer & Quah & Duzulkafly & Yambun, 2018" authorityName="Grismer & Quah & Duzulkafly & Yambun" authorityYear="2018" box="[1108,1253,1156,1179]" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="537" phylum="Chordata" rank="species" species="peninsulare" status="sp. nov.">
|
||
<emphasis id="10F6EAD7D915FF868241FB36FB4CFB49" box="[1108,1253,1156,1179]" italics="true" pageId="9" pageNumber="537">L. peninsulare</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0BC557ACD915FF8682F9FB35FA92FB49" box="[1260,1339,1157,1178]" pageId="9" pageNumber="537" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(LSUHC 13857) from 13.5 km east of Jeli, Kelantan, Peninsular Malaysia; and D,
|
||
<taxonomicName id="E5824D46D915FF8685B2FB14FB8AFB6A" box="[935,1059,1187,1209]" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="537" phylum="Chordata" rank="species" species="scheideri">
|
||
<emphasis id="10F6EAD7D915FF8685B2FB14FB8AFB6A" box="[935,1059,1187,1209]" italics="true" pageId="9" pageNumber="537">L. scheideri</emphasis>
|
||
</taxonomicName>
|
||
(BM 4743) from Djpura, Indragiri, Sumatra, Indonesia. Photos by LLG.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="6A98654ED915FF868682FAB5FC0BFA92" pageId="9" pageNumber="537" type="materials_examined">
|
||
<paragraph id="223D36C5D915FF868682FAB5FC0BFA92" blockId="9.[151,1437,1284,2029]" pageId="9" pageNumber="537">
|
||
<materialsCitation id="92EA3C98D915FF868682FAB5FC0BFA92" ID-GBIF-Occurrence="1851456851" collectingDate="1900-01-01" collectorName="G. Schneider" country="Indonesia" location="Sumatra" pageId="9" pageNumber="537" specimenCount="1" stateProvince="Sumatra" typeStatus="holotype">
|
||
<emphasis id="10F6EAD7D915FF868682FAB5FEADFACE" bold="true" box="[151,260,1285,1309]" pageId="9" pageNumber="537">
|
||
<typeStatus id="FD398867D915FF868682FAB5FEADFACE" box="[151,260,1285,1309]" pageId="9" pageNumber="537" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
. Adult male (BM 4743) from “Indragiri, Djapura”,
|
||
<collectingRegion id="E046F827D915FF868593FAB5FC4FFACE" box="[902,998,1285,1309]" country="Indonesia" name="Sumatera Utara" pageId="9" pageNumber="537">Sumatra</collectingRegion>
|
||
,
|
||
<collectingCountry id="5A957655D915FF8685ECFAB4FBCFFACE" box="[1017,1126,1284,1309]" name="Indonesia" pageId="9" pageNumber="537">Indonesia</collectingCountry>
|
||
and accessioned into the Naturhistorischen Museum Basel by
|
||
<collectorName id="8F775313D915FF868423FA99FD6CFA92" box="[566,709,1320,1345]" pageId="9" pageNumber="537">G. Schneider</collectorName>
|
||
on
|
||
<date id="563C1005D915FF8684FAFA99FC37FA92" box="[751,926,1321,1345]" pageId="9" pageNumber="537" value="1900-01-01">
|
||
<collectingDate id="4678E9EDD915FF8684FAFA99FC37FA92" box="[751,926,1321,1345]" pageId="9" pageNumber="537" value="1900-01-01">1 January 1900</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6A98654ED915FF8686D2FAFCFF05F8C6" pageId="9" pageNumber="537" type="diagnosis">
|
||
<paragraph id="223D36C5D915FF8686D2FAFCFF05F8C6" blockId="9.[151,1437,1284,2029]" pageId="9" pageNumber="537">
|
||
<emphasis id="10F6EAD7D915FF8686D2FAFCFE93FAB6" bold="true" box="[199,314,1356,1381]" pageId="9" pageNumber="537">Diagnosis</emphasis>
|
||
.
|
||
<taxonomicName id="E5824D46D915FF86875DFAFEFD9FFAB6" authorityName="Werner" authorityYear="1900" box="[328,566,1356,1381]" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="537" phylum="Chordata" rank="species" species="schneideri">
|
||
<emphasis id="10F6EAD7D915FF86875DFAFEFD9FFAB6" box="[328,566,1356,1381]" italics="true" pageId="9" pageNumber="537">Lygosoma schneideri</emphasis>
|
||
</taxonomicName>
|
||
can be differentiated from all other
|
||
<taxonomicName id="E5824D46D915FF8685D2FAFEFB90FAB6" authorityName="Bampfyldii Boulenger" authorityYear="1900" box="[967,1081,1358,1381]" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="537" phylum="Chordata" rank="genus">
|
||
<emphasis id="10F6EAD7D915FF8685D2FAFEFB90FAB6" box="[967,1081,1358,1381]" italics="true" pageId="9" pageNumber="537">Lygosoma</emphasis>
|
||
</taxonomicName>
|
||
by having the combination of a relatively large (SVL =
|
||
<quantity id="E57A9B20D915FF8687A7FAC1FDBCFA5A" box="[434,533,1393,1417]" metricMagnitude="-1" metricUnit="m" metricValue="1.29" pageId="9" pageNumber="537" unit="mm" value="129.0">129 mm</quantity>
|
||
) robust body (PEC/SVL = 0.15); seven supralabials and infralabials; midline contact of the supranasals; prefrontals not in contact; frontoparietal contacting three supraoculars; large postinterparietal present; eight superciliaries; two postsuboculars, the first being small; one or two primary and three secondary and tertiary temporals; eight nuchal scales; a deep postnasal groove extending from the nasal scale to below the anterior portion of the eye and lying below the anterior loreals and lower preocular and above the second and third supralabials; scaly lower eyelid, no window; 45 midbody scale rows; 95 paravertebral scale rows; 98 ventral scale rows; 34 caudal scale rows at the tenth subcaudal; eight large precloacal scales; smooth to weakly keeled subdigital finger lamellae, 10 lamellae on third finger; keeled subdigital toe lamellae, 16 lamellae on fourth toe; low, round, small, palmar scales numbering seven across the base of the palm; head pattern consisting of a dark, continuous frontal and occipital band separated by a yellowish band; dark occipital band not confluent with lighter color of dorsum and tail. These characters are scored across all species in the
|
||
<taxonomicName id="E5824D46D915FF868240F96AFB58F922" authorityName="de Rooji" authorityYear="1915" box="[1109,1265,1752,1777]" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="537" phylum="Chordata" rank="species" species="bampfyldei">
|
||
<emphasis id="10F6EAD7D915FF868240F96AFB58F922" box="[1109,1265,1752,1777]" italics="true" pageId="9" pageNumber="537">L. bampfyldei</emphasis>
|
||
</taxonomicName>
|
||
group in
|
||
<tableCitation id="6F00037ED915FF86834BF968FF01F8C6" captionStart="TABLE 2" captionStartId="16.[147,226,490,511]" captionText="TABLE 2. Stanđarđ đescriptive anđ điagnostic (shađeđ) characters among đifferent combinations of species of the Lygosoma bampfyldei group. All measurements are in millimeters. / = đata unobtainable. Abbreviations are listeđ in the Materials anđ methođs." pageId="9" pageNumber="537">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6A98654ED915FF8686D2F890FD24F852" pageId="9" pageNumber="537" type="distribution">
|
||
<paragraph id="223D36C5D915FF8686D2F890FD24F852" blockId="9.[151,1437,1284,2029]" pageId="9" pageNumber="537">
|
||
<emphasis id="10F6EAD7D915FF8686D2F890FEC9F8EA" bold="true" box="[199,352,1824,1849]" pageId="9" pageNumber="537">Distribution.</emphasis>
|
||
<taxonomicName id="E5824D46D915FF868772F892FDFFF8EA" authorityName="Werner" authorityYear="1900" box="[359,598,1824,1849]" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="537" phylum="Chordata" rank="species" species="schneideri">
|
||
<emphasis id="10F6EAD7D915FF868772F892FDFFF8EA" box="[359,598,1824,1849]" italics="true" pageId="9" pageNumber="537">Lygosoma schneideri</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<typeStatus id="FD398867D915FF86857EF891FC32F8EA" box="[875,923,1825,1849]" pageId="9" pageNumber="537">type</typeStatus>
|
||
locality.
|
||
<bibRefCitation id="46134B34D915FF868216F891FB28F8EA" box="[1027,1153,1825,1849]" pageId="9" pageNumber="537" refId="ref7752" refString="Das, I. (2010) A Field Guide to the Reptiles of South-east Asia. Bloomsbury, London, 325 pp. https: // doi. org / 10.1108 / RR- 08 - 2016 - 0210" type="book">Das (2010)</bibRefCitation>
|
||
reports this species from “north-western
|
||
<collectingRegion id="E046F827D915FF86875DF8F5FE0CF88E" box="[328,421,1861,1885]" country="Indonesia" name="Sumatera Utara" pageId="9" pageNumber="537">Sumatra</collectingRegion>
|
||
(Indrajiri,
|
||
<collectingRegion id="E046F827D915FF868436F8F4FD60F88E" box="[547,713,1860,1885]" country="Indonesia" name="Riau" pageId="9" pageNumber="537">Riau Province</collectingRegion>
|
||
)” however the
|
||
<typeStatus id="FD398867D915FF86856DF8F5FC01F88E" box="[888,936,1861,1885]" pageId="9" pageNumber="537">type</typeStatus>
|
||
locality listed by
|
||
<bibRefCitation id="46134B34D915FF86826DF8F5FAB7F88E" box="[1144,1310,1861,1885]" pageId="9" pageNumber="537" refId="ref9568" refString="Werner, F. (1910) Uber neue oder seltene Reptilien des Naturhistorischen Museums in Hamburg. ii. Eidechsen des Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 27 (1909), 1 (Supplement), 1910, 1 - 46." type="journal article">Werner (1910)</bibRefCitation>
|
||
“Indragiri, Djapura” is in southeastern
|
||
<collectingRegion id="E046F827D915FF8687DBF8D9FD82F852" box="[462,555,1897,1921]" country="Indonesia" name="Sumatera Utara" pageId="9" pageNumber="537">Sumatra</collectingRegion>
|
||
(
|
||
<figureCitation id="BAB92A40D915FF86842EF8D8FDD5F852" box="[571,636,1896,1921]" captionStart="FIGURE 1" captionStartId="1.[151,250,1219,1241]" captionTargetBox="[151,1436,483,1197]" captionTargetId="figure@1.[151,1436,483,1197]" captionTargetPageId="1" captionText="FIGURE 1. Distribution of the species of the Lygosoma bampfyldei group in Sundaland. Stars designate the location of the type localities for L. bampfyldei along the Rajang River, Sarawak, East Malaysia; L. peninsulare sp. nov. from 13.5 km east of Jeli, Kelantan, Peninsular Malaysia; L. kinabatanganensis sp. nov. from the Kinabatangan District, Deramakot camp (Deramakot Sabah Forestry Department), Sabah, East Malaysia; and L. schneideri from Indragiri, Djapura, Sumatra, Indonesia. Colors are coded to match those of the Figure 3." httpUri="https://zenodo.org/record/1298206/files/figure.png" pageId="9" pageNumber="537">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6A98654ED915FF8586D2F83DFB0FFF04" lastPageId="10" lastPageNumber="538" pageId="9" pageNumber="537" type="discussion">
|
||
<paragraph id="223D36C5D915FF8586D2F83DFB0FFF04" blockId="9.[151,1437,1284,2029]" lastBlockId="10.[151,1436,151,215]" lastPageId="10" lastPageNumber="538" pageId="9" pageNumber="537">
|
||
<emphasis id="10F6EAD7D915FF8686D2F83DFE92F876" bold="true" box="[199,315,1933,1957]" pageId="9" pageNumber="537">Remarks.</emphasis>
|
||
<bibRefCitation id="46134B34D915FF868759F83CFDB4F876" box="[332,541,1932,1957]" pageId="9" pageNumber="537" refId="ref7660" refString="Boulenger, G. A. (1912) A Fertebrate Fauna of the Malay Peninsula from the Isthmus of Kra to Singapore Including Adjacent Islands. Taylor and Francis, London, 314 pp. https: // doi. org / 10.5962 / bhl. title. 10813" type="book">Boulenger (1912)</bibRefCitation>
|
||
placed
|
||
<taxonomicName id="E5824D46D915FF86849CF83EFC2BF876" authorityName="Werner" authorityYear="1900" box="[649,898,1932,1957]" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="537" phylum="Chordata" rank="species" species="schneideri">
|
||
<emphasis id="10F6EAD7D915FF86849CF83EFC2BF876" box="[649,898,1932,1957]" italics="true" pageId="9" pageNumber="537">Lygosoma schneideri</emphasis>
|
||
</taxonomicName>
|
||
in the synonymy of
|
||
<taxonomicName id="E5824D46D915FF86828BF83EFAEAF876" authorityName="de Rooji" authorityYear="1915" box="[1182,1347,1932,1957]" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="537" phylum="Chordata" rank="species" species="bampfyldei">
|
||
<emphasis id="10F6EAD7D915FF86828BF83EFAEAF876" box="[1182,1347,1932,1957]" italics="true" pageId="9" pageNumber="537">L. bampfyldei</emphasis>
|
||
</taxonomicName>
|
||
in his redescription of the latter. Our analyses indicate that
|
||
<taxonomicName id="E5824D46D915FF8684FAF802FC2AF81A" authorityName="Werner" authorityYear="1900" box="[751,899,1968,1993]" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="537" phylum="Chordata" rank="species" species="schneideri">
|
||
<emphasis id="10F6EAD7D915FF8684FAF802FC2AF81A" box="[751,899,1968,1993]" italics="true" pageId="9" pageNumber="537">L. schneideri</emphasis>
|
||
</taxonomicName>
|
||
is considerably different from
|
||
<taxonomicName id="E5824D46D915FF8682F6F802FAD6F81A" authorityName="de Rooji" authorityYear="1915" box="[1251,1407,1968,1993]" class="Reptilia" family="Scincidae" genus="Lygosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="537" phylum="Chordata" rank="species" species="bampfyldei">
|
||
<emphasis id="10F6EAD7D915FF8682F6F802FAD6F81A" box="[1251,1407,1968,1993]" italics="true" pageId="9" pageNumber="537">L. bampfyldei</emphasis>
|
||
</taxonomicName>
|
||
in having two vs. three suboculars, a postparietal scale (
|
||
<figureCitation id="BAB92A40D915FF86851CF864FCF8F83E" box="[777,849,2004,2029]" captionStart="FIGURE 2" captionStartId="2.[151,250,886,908]" captionTargetBox="[200,1376,213,823]" captionTargetId="figure@2.[151,1436,193,865]" captionTargetPageId="2" captionText="FIGURE 2. A) Right lateral view of the head of the holotype of Lygosoma kinabatanganensis sp. nov. (FMNH 76226) from the Kinabatangan District, Deramakot camp (Deramakot Sabah Forestry Department), Sabah, East Malaysia and B) dorsal view of head of the holotype of L. schneideri from Djpura, Indragiri, Sumatra, Indonesia illustrating the positional relationship and size of some of the head scales. Terminology is adapted from Taylor (1935). AL = anterior loreal, Cs = chinshield; F = frontal, Fn = frontonasal, Fp = frontoparietal, IL = infralabial, Ip = interparietal, M = mental, N = nasal, Nu = nuchal, P = parietal, Pf = prefrontal, PI = postinterparietal, PL = posterior loreal, Pm = postmental, Pos = postsubocular, Pro = preocular, Prs = presubocular, PT = primary temporal, Psl = postsupralabial, Pto = postocular, R = rostral, Sl = supralabial, Sn = supranasal, So = supraocular, ST = secondary temporal, and TT = tertiary temporal. * = superciliary. Illustration by LLG." httpUri="https://zenodo.org/record/1298208/files/figure.png" pageId="9" pageNumber="537">Fig. 2</figureCitation>
|
||
), 45 vs. 36–40 midbody scale rows, 95 vs. 82–85 paravertebral scale rows, 34 vs. 28–30 scale rows around the tail at the 10th subcaudal, and a significantly (
|
||
<emphasis id="10F6EAD7D916FF858356FF2CFAE2FF60" box="[1347,1355,156,179]" italics="true" pageId="10" pageNumber="538">t</emphasis>
|
||
= -6,
|
||
<emphasis id="10F6EAD7D916FF85839BFF2CFA35FF60" box="[1422,1436,156,179]" italics="true" pageId="10" pageNumber="538">p</emphasis>
|
||
= 0.004) more robust (PEC/SVL = 0.15) vs. a slender (PEC/SVL = 0.12–0.13) body (
|
||
<figureCitation id="BAB92A40D916FF858241FF0EFB30FF04" box="[1108,1177,190,215]" captionStart="FIGURE 4" captionStartId="8.[151,250,1451,1473]" captionTargetBox="[176,1396,723,1417]" captionTargetId="figure@8.[151,1436,700,1430]" captionTargetPageId="8" captionText="FIGURE 4. Comparisons of body stature of the types of A, Lygosoma scheideri (BM 4743) from Djpura, Indragiri, Sumatra, Indonesia; B, syntype of Lygosoma bampfyldei (BMNH 1946.8.10.84) from the Rejan River, Sarawak, East Malaysia; C, L. kinabatanganensis sp. nov. (FMNH 76226) from the Kinabatangan District, Deramakot camp (Deramakot Sabah Forestry Department), Sabah, East Malaysia; and D, L. peninsulare sp. nov. (LSUHC 13857) from 13.5 km east of Jeli, Kelantan, Peninsular Malaysia." httpUri="https://zenodo.org/record/1298212/files/figure.png" pageId="10" pageNumber="538">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |