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<document id="4F0761EE1A787E13540C48CEB0585AA5" ID-CLB-Dataset="29309" ID-DOI="10.11646/zootaxa.4450.3.6" ID-GBIF-Dataset="576197a6-015b-4293-ac10-d8dcf3360f9f" ID-ISSN="1175-5326" ID-Zenodo-Dep="1444864" ID-ZooBank="EA15F553-F271-48D1-A2A4-9D453AC44353" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538634640006" checkinUser="plazi" docAuthor="Heppner, John B. &amp; Bae, Yang-Seop" docDate="2018" docId="81788781FFD6FFB4FF3D1C12D596542B" docLanguage="en" docName="zootaxa.4450.3.6.pdf" docOrigin="Zootaxa 4450 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anthozela cypriflammella Heppner &amp; Bae 2018, new species" docType="treatment" docVersion="7" lastPageNumber="398" masterDocId="7D41FFF9FFD7FFB0FFAA1E75D4115119" masterDocTitle="A new species of Anthozela from Vietnam, with a list of species in the genus (Lepidoptera: Tortricidae: Olethreutinae: Enarmoniini)" masterLastPageNumber="400" masterPageNumber="394" pageNumber="395" updateTime="1698564360139" updateUser="ExternalLinkService">
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<mods:title id="0CDB972D57D554E56840E905272DA7E3">A new species of Anthozela from Vietnam, with a list of species in the genus (Lepidoptera: Tortricidae: Olethreutinae: Enarmoniini)</mods:title>
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<mods:namePart id="46A66AC03FD6D1490263445640BBF582">Heppner, John B.</mods:namePart>
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<mods:namePart id="A5C8EE66FFD26F5B0056B353E9796254">Bae, Yang-Seop</mods:namePart>
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<treatment id="81788781FFD6FFB4FF3D1C12D596542B" ID-DOI="http://doi.org/10.5281/zenodo.5949519" ID-GBIF-Taxon="148462431" ID-Zenodo-Dep="5949519" LSID="urn:lsid:plazi:treatment:81788781FFD6FFB4FF3D1C12D596542B" httpUri="http://treatment.plazi.org/id/81788781FFD6FFB4FF3D1C12D596542B" lastPageId="4" lastPageNumber="398" pageId="1" pageNumber="395">
<subSubSection id="41CB651CFFD6FFB1FF3D1C12D50053BA" pageId="1" pageNumber="395" type="nomenclature">
<paragraph id="096E3697FFD6FFB1FF3D1C12D728539B" blockId="1.[151,825,615,675]" box="[151,825,615,642]" pageId="1" pageNumber="395">
<heading id="522681FBFFD6FFB1FF3D1C12D728539B" bold="true" box="[151,825,615,642]" fontSize="11" level="1" pageId="1" pageNumber="395" reason="1">
<emphasis id="3BA5EA85FFD6FFB1FF3D1C12D728539B" bold="true" box="[151,825,615,642]" pageId="1" pageNumber="395">
<taxonomicName id="CED14D14FFD6FFB1FF3D1C12D68C539B" authority="Heppner &amp; Bae" authorityName="Heppner &amp; Bae" authorityYear="2018" box="[151,669,615,642]" class="Insecta" family="Tortricidae" genus="Anthozela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="395" phylum="Arthropoda" rank="species" species="cypriflammella" status="sp. nov.">
<emphasis id="3BA5EA85FFD6FFB1FF3D1C12D5DF5398" bold="true" box="[151,462,615,641]" italics="true" pageId="1" pageNumber="395">Anthozela cypriflammella</emphasis>
Heppner &amp; Bae
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,
<taxonomicNameLabel id="209657FEFFD6FFB1FD031C1DD728539B" box="[681,825,616,642]" pageId="1" pageNumber="395" rank="species">new species</taxonomicNameLabel>
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</heading>
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(
<figureCitation id="91EA2A12FFD6FFB1FF351CFFD51953BA" box="[159,264,650,675]" captionStart-0="FIGURES 12" captionStart-1="FIGURES 36" captionStart-2="FIGURES 7" captionStart-3="FIGURES 8" captionStartId-0="2.[151,264,1728,1751]" captionStartId-1="3.[151,264,1878,1901]" captionStartId-2="4.[151,264,1171,1194]" captionStartId-3="5.[151,264,1997,2020]" captionTargetBox-0="[172,1401,357,1700]" captionTargetBox-1="[151,1435,409,1858]" captionTargetBox-2="[166,1426,195,1149]" captionTargetBox-3="[246,1336,556,1970]" captionTargetId-0="figure@2.[161,1421,337,1711]" captionTargetId-1="figure@3.[151,1435,409,1858]" captionTargetId-2="figure@4.[166,1426,195,1158]" captionTargetId-3="figure@5.[246,1336,555,1970]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURES 12. Anthozela cypriflammella n. sp., Vietnam. 1) Male holotype (10.8 mm wingspan); 2) Female paratype (10.8mm wingspan)." captionText-1="FIGURES 36. Anthozela cypriflammella n. sp., close-up of head (lateral and dorsal views). 34) Male; 56) Female." captionText-2="FIGURES 7. Anthozela cypriflammella n. sp.: male genitalia. a) phallus; b) central genital close-up; c) pregenital sternal plate (genitalia slide JBH 3491)." captionText-3="FIGURES 8. Anthozela cypriflammella n. sp.: female genitalia. a) sterigma and ostium; b) signa (genitalia slide JBH 3323)." httpUri-0="https://zenodo.org/record/1444866/files/figure.png" httpUri-1="https://zenodo.org/record/1444868/files/figure.png" httpUri-2="https://zenodo.org/record/1444870/files/figure.png" httpUri-3="https://zenodo.org/record/1444872/files/figure.png" pageId="1" pageNumber="395">Figs. 18</figureCitation>
)
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<subSubSection id="41CB651CFFD6FFB1FF3D1CA7D0BF53F2" box="[151,1198,722,747]" pageId="1" pageNumber="395" type="materials_examined">
<paragraph id="096E3697FFD6FFB1FF3D1CA7D0BF53F2" blockId="1.[151,1437,722,2007]" box="[151,1198,722,747]" pageId="1" pageNumber="395">
<materialsCitation id="B9B93CCAFFD6FFB1FF3D1CA7D0BF53F2" ID-GBIF-Occurrence="1914123839" box="[151,1198,722,747]" country="Vietnam" elevation="155" location="Mac Lake" pageId="1" pageNumber="395" specimenCount="1" stateProvince="Ninh Binh Province" typeStatus="holotype">
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<typeStatus id="D66A8835FFD6FFB1FF3D1CA7D4C353F2" box="[151,210,722,747]" pageId="1" pageNumber="395">Type</typeStatus>
locality:
</emphasis>
Cuc Phuong Natl. Park,
<location id="0C0E604CFFD6FFB1FDFA1CA6D6D053F2" LSID="urn:lsid:plazi:treatment:81788781FFD6FFB4FF3D1C12D596542B:0C0E604CFFD6FFB1FDFA1CA6D6D053F2" box="[592,705,722,747]" country="Vietnam" name="Mac Lake" pageId="1" pageNumber="395" stateProvince="Ninh Binh Province">Mac Lake</location>
(alt.
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<quantity id="CE299B72FFD6FFB1FD511CA7D75453F2" box="[763,837,722,747]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="1" pageNumber="395" unit="m" value="155.0">155 m</quantity>
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),
<collectingRegion id="CB15F875FFD6FFB1FCF31CA7D02D53F2" box="[857,1084,722,747]" country="Vietnam" name="Ninh Binh" pageId="1" pageNumber="395">
Ninh
<collectingRegion id="CB15F875FFD6FFB1FC321CA7D02D53F2" box="[920,1084,722,747]" inferredBySuffix="true" pageId="1" pageNumber="395">Binh Province</collectingRegion>
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,
<collectingCountry id="71C67607FFD6FFB1FBED1CA7D0BB53F2" box="[1095,1194,722,747]" name="Vietnam" pageId="1" pageNumber="395">Vietnam</collectingCountry>
.
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</paragraph>
</subSubSection>
<subSubSection id="41CB651CFFD6FFB1FF6D1C83D0805262" pageId="1" pageNumber="395" type="diagnosis">
<paragraph id="096E3697FFD6FFB1FF6D1C83D0805262" blockId="1.[151,1437,722,2007]" pageId="1" pageNumber="395">
<emphasis id="3BA5EA85FFD6FFB1FF6D1C83D52B5216" bold="true" box="[199,314,758,783]" pageId="1" pageNumber="395">Diagnosis</emphasis>
.
<taxonomicName id="CED14D14FFD6FFB1FEE11C83D67F5216" authorityName="Heppner &amp; Bae" authorityYear="2018" box="[331,622,758,783]" class="Insecta" family="Tortricidae" genus="Anthozela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="395" phylum="Arthropoda" rank="species" species="cypriflammella">
<emphasis id="3BA5EA85FFD6FFB1FEE11C83D67F5216" box="[331,622,758,783]" italics="true" pageId="1" pageNumber="395">Anthozela cypriflammella</emphasis>
</taxonomicName>
is superficially similar to other species of
<taxonomicName id="CED14D14FFD6FFB1FBCB1C83D0C05216" box="[1121,1233,758,783]" class="Insecta" family="Tortricidae" genus="Anthozela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="395" phylum="Arthropoda" rank="genus">
<emphasis id="3BA5EA85FFD6FFB1FBCB1C83D0C05216" box="[1121,1233,758,783]" italics="true" pageId="1" pageNumber="395">Anthozela</emphasis>
</taxonomicName>
, even those form Africa, but can be distinguished by the strong carinate ridge on the sacculus of the male genitalia and the absence of strong posterior spines on the 8th sternal plate. The female genitalia lack an accessory bursa but have a wide colliculum (usually narrower in other congeners) and a subtriangular, funnel-shape ostium.
</paragraph>
</subSubSection>
<subSubSection id="41CB651CFFD6FFB2FF6D1DF3D53B5004" lastPageId="2" lastPageNumber="396" pageId="1" pageNumber="395" type="description">
<paragraph id="096E3697FFD6FFB1FF6D1DF3D56B563A" blockId="1.[151,1437,722,2007]" pageId="1" pageNumber="395">
<emphasis id="3BA5EA85FFD6FFB1FF6D1DF3D5405286" bold="true" box="[199,337,902,927]" pageId="1" pageNumber="395">Description</emphasis>
. Male (
<figureCitation id="91EA2A12FFD6FFB1FE061DF3D5E15286" box="[428,496,902,927]" captionStart="FIGURES 12" captionStartId="2.[151,264,1728,1751]" captionTargetBox="[172,1401,357,1700]" captionTargetId="figure@2.[161,1421,337,1711]" captionTargetPageId="2" captionText="FIGURES 12. Anthozela cypriflammella n. sp., Vietnam. 1) Male holotype (10.8 mm wingspan); 2) Female paratype (10.8mm wingspan)." httpUri="https://zenodo.org/record/1444866/files/figure.png" pageId="1" pageNumber="395">Fig. 1</figureCitation>
). Head (
<figureCitation id="91EA2A12FFD6FFB1FDFD1DF3D6D75286" box="[599,710,902,927]" captionStart-0="FIGURES 12" captionStart-1="FIGURES 36" captionStartId-0="2.[151,264,1728,1751]" captionStartId-1="3.[151,264,1878,1901]" captionTargetBox-0="[172,1401,357,1700]" captionTargetBox-1="[151,1435,409,1858]" captionTargetId-0="figure@2.[161,1421,337,1711]" captionTargetId-1="figure@3.[151,1435,409,1858]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 12. Anthozela cypriflammella n. sp., Vietnam. 1) Male holotype (10.8 mm wingspan); 2) Female paratype (10.8mm wingspan)." captionText-1="FIGURES 36. Anthozela cypriflammella n. sp., close-up of head (lateral and dorsal views). 34) Male; 56) Female." httpUri-0="https://zenodo.org/record/1444866/files/figure.png" httpUri-1="https://zenodo.org/record/1444868/files/figure.png" pageId="1" pageNumber="395">Figs. 23</figureCitation>
): Vertex yellow-tan, mixed with some dark brown; frons same; antenna dark brown with few yellow-tan scales; scape yellow-tan, with dark brown streak dorsally; labial palpus light yellow, with apical segment light brown, mesally same. Thorax: Light yellow mixed over dark brown, more yellow posteriorly; tegulae same; venter tan-white, with light yellow-tan at neck; legs white basally and femurs, with tibia tan and with two angled dark brown rings; midtibia with spine-like scales on dorsal margin; hindtibia lustrous gray; tarsal segments alternating gray and pale tan. Wingspan
<quantity id="CE299B72FFD6FFB1FC1A1A4FD045554B" box="[944,1108,1082,1107]" metricMagnitude="-2" metricUnit="m" metricValue="1.065" metricValueMax="1.08" metricValueMin="1.05" pageId="1" pageNumber="395" unit="mm" value="10.65" valueMax="10.8" valueMin="10.5">10.510.8 mm</quantity>
in males (n = 2),
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in female (n = 1). Forewing with yellow scales scattered over black-brown on basal half of forewing, with indistinct black-brown costal strigulae, and with a cross-wing vertical fascia of silver-gray, straight mesally but very irregular distally, nearly to costal margin but interrupted by thin line of burnt-orange and a fine silver line angled from costal black-brown strigula towards apex; apical half of costa with several black-brown strigulae angled towards tornus, each separated by burnt orange, and on costal margin by light yellow; apical half of forewing otherwise reddishburnt-orange with several irregular lines and spots of silver-gray, subtermen and tornus with similar coloration; fringe black-brown, with a near-apical and near-tornal patch of light orange. Venter gray-brown with pale white on cell and cubitus and dorsal margin, with tan and dark brown strigulae on costal margin as in dorsal surface. Hindwing dark brown-gray with slightly paler area near midwing. Venter similar to dorsum but duller. Abdomen: Brown-gray; venter pale tan-white, with brown anterior line on each sternite; genital tuft brown-gray; pregenital plates as simple posteriorly concave tergal plate (
<figureCitation id="91EA2A12FFD6FFB1FD611BB3D73154C6" box="[715,800,1478,1503]" captionStart="FIGURES 7" captionStartId="4.[151,264,1171,1194]" captionTargetBox="[166,1426,195,1149]" captionTargetId="figure@4.[166,1426,195,1158]" captionTargetPageId="4" captionText="FIGURES 7. Anthozela cypriflammella n. sp.: male genitalia. a) phallus; b) central genital close-up; c) pregenital sternal plate (genitalia slide JBH 3491)." httpUri="https://zenodo.org/record/1444870/files/figure.png" pageId="1" pageNumber="395">Fig. 7b</figureCitation>
), intersegmental membrane connected to male genitalia very long (subequal to 8th segment), with dorsal setal patch on each side. Male genitalia (
<figureCitation id="91EA2A12FFD6FFB1FB031B9FD0FE571A" box="[1193,1263,1514,1539]" captionStart="FIGURES 7" captionStartId="4.[151,264,1171,1194]" captionTargetBox="[166,1426,195,1149]" captionTargetId="figure@4.[166,1426,195,1158]" captionTargetPageId="4" captionText="FIGURES 7. Anthozela cypriflammella n. sp.: male genitalia. a) phallus; b) central genital close-up; c) pregenital sternal plate (genitalia slide JBH 3491)." httpUri="https://zenodo.org/record/1444870/files/figure.png" pageId="1" pageNumber="395">Fig. 7</figureCitation>
) with tegumen elongated as strong reversed V-shape, with strongly convex uncus; socius laterally as curved concave plate with strong row of stout, comb-like spines directed mesally, of uniform length (
<figureCitation id="91EA2A12FFD6FFB1FC401847D02C5752" box="[1002,1085,1586,1611]" captionStart="FIGURES 7" captionStartId="4.[151,264,1171,1194]" captionTargetBox="[166,1426,195,1149]" captionTargetId="figure@4.[166,1426,195,1158]" captionTargetPageId="4" captionText="FIGURES 7. Anthozela cypriflammella n. sp.: male genitalia. a) phallus; b) central genital close-up; c) pregenital sternal plate (genitalia slide JBH 3491)." httpUri="https://zenodo.org/record/1444870/files/figure.png" pageId="1" pageNumber="395">Fig. 7a</figureCitation>
); valva elongated, setose, with slightly bulbous distal end, and saccular margin with a strong dorsally-directed carinate ridge about 1/6 of valval length from base; vinculum simple, V-shaped; juxta a narrow band with lateral flattened quadratic basal plate, elongated dorsally to narrow arms to anellus, latter with a longer emarginated flattened lateral flange-like plate; caulis dorsally with groove-like invaginated apex with wedge-shaped lateral point around phallus; phallus short and slender, slightly bulged medially, vesica with patch of about 30 cornuti as thumbtack-like spines (points directed outwardly);
</paragraph>
<paragraph id="096E3697FFD6FFB2FF6D195BD53B5004" blockId="1.[151,1437,722,2007]" lastBlockId="2.[151,1436,151,285]" lastPageId="2" lastPageNumber="396" pageId="1" pageNumber="395">
Female (
<figureCitation id="91EA2A12FFD6FFB1FE82195BD57F565E" box="[296,366,1838,1863]" captionStart="FIGURES 12" captionStartId="2.[151,264,1728,1751]" captionTargetBox="[172,1401,357,1700]" captionTargetId="figure@2.[161,1421,337,1711]" captionTargetPageId="2" captionText="FIGURES 12. Anthozela cypriflammella n. sp., Vietnam. 1) Male holotype (10.8 mm wingspan); 2) Female paratype (10.8mm wingspan)." httpUri="https://zenodo.org/record/1444866/files/figure.png" pageId="1" pageNumber="395">Fig. 2</figureCitation>
). Head and Thorax: Essentially as in male, but head with somewhat paler yellow-white scaling, particularly on labial palpus; hindwing with midwing pale area somewhat more contrasting than in male. Abdomen: Female genitalia (
<figureCitation id="91EA2A12FFD6FFB1FE761903D60F5696" box="[476,542,1910,1935]" captionStart="FIGURES 8" captionStartId="5.[151,264,1997,2020]" captionTargetBox="[246,1336,556,1970]" captionTargetId="figure@5.[246,1336,555,1970]" captionTargetPageId="5" captionText="FIGURES 8. Anthozela cypriflammella n. sp.: female genitalia. a) sterigma and ostium; b) signa (genitalia slide JBH 3323)." httpUri="https://zenodo.org/record/1444872/files/figure.png" pageId="1" pageNumber="395">Fig. 8</figureCitation>
) elongated, with short ovipositor, papilla anales setose and flattened; sterigma a simple semicircular ring at posterior margin to simple, subtriangular funnel-shaped ostium bursae (
<figureCitation id="91EA2A12FFD6FFB1FB4F19EFD12556AA" box="[1253,1332,1946,1971]" captionStart="FIGURES 8" captionStartId="5.[151,264,1997,2020]" captionTargetBox="[246,1336,556,1970]" captionTargetId="figure@5.[246,1336,555,1970]" captionTargetPageId="5" captionText="FIGURES 8. Anthozela cypriflammella n. sp.: female genitalia. a) sterigma and ostium; b) signa (genitalia slide JBH 3323)." httpUri="https://zenodo.org/record/1444872/files/figure.png" pageId="1" pageNumber="395">Fig. 8a</figureCitation>
) equal in width to ductus bursae; colliculum somewhat indistinct and of similar width as ductus bursae; length of ductus bursae subequal to that of corpus bursae, with sinuous entry into corpus bursa as the ductus and bursae gradually widen to full bursal width; ductus seminalis emergent at posterior end of corpus bursa; corpus bursae large, ovate, with simple scobinations, and two large, elongated, thorn-like signa on lateral wall of bursa (
<figureCitation id="91EA2A12FFD5FFB2FB151EAAD10551E1" box="[1215,1300,223,248]" captionStart="FIGURES 8" captionStartId="5.[151,264,1997,2020]" captionTargetBox="[246,1336,556,1970]" captionTargetId="figure@5.[246,1336,555,1970]" captionTargetPageId="5" captionText="FIGURES 8. Anthozela cypriflammella n. sp.: female genitalia. a) sterigma and ostium; b) signa (genitalia slide JBH 3323)." httpUri="https://zenodo.org/record/1444872/files/figure.png" pageId="2" pageNumber="396">Fig. 8b</figureCitation>
); secondary bursa absent.
</paragraph>
</subSubSection>
<caption id="5DAE661FFFD5FFB2FF3D18B5D57B57EC" httpUri="https://zenodo.org/record/1444866/files/figure.png" pageId="2" pageNumber="396" startId="2.[151,264,1728,1751]" targetBox="[172,1401,357,1700]" targetPageId="2">
<paragraph id="096E3697FFD5FFB2FF3D18B5D57B57EC" blockId="2.[151,1436,1728,1781]" pageId="2" pageNumber="396">
<emphasis id="3BA5EA85FFD5FFB2FF3D18B5D52E57CF" bold="true" box="[151,319,1728,1751]" pageId="2" pageNumber="396">FIGURES 12.</emphasis>
<taxonomicName id="CED14D14FFD5FFB2FEE118B4D64757CE" authority="Heppner &amp; Bae, 2018" authorityName="Heppner &amp; Bae" authorityYear="2018" box="[331,598,1729,1751]" class="Insecta" family="Tortricidae" genus="Anthozela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="396" phylum="Arthropoda" rank="species" species="cypriflammella" status="sp. nov.">
<emphasis id="3BA5EA85FFD5FFB2FEE118B4D64757CE" box="[331,598,1729,1751]" italics="true" pageId="2" pageNumber="396">Anthozela cypriflammella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="209657FEFFD5FFB2FDCB18B4D6B157CF" box="[609,672,1729,1750]" pageId="2" pageNumber="396" rank="species">
<emphasis id="3BA5EA85FFD5FFB2FDCB18B4D6B157CF" bold="true" box="[609,672,1729,1750]" pageId="2" pageNumber="396">n. sp.</emphasis>
</taxonomicNameLabel>
, Vietnam. 1) Male holotype (10.8 mm wingspan); 2) Female paratype (10.8mm wingspan).
</paragraph>
</caption>
<subSubSection id="41CB651CFFD5FFB2FF6D1957D78356BE" pageId="2" pageNumber="396" type="materials_examined">
<paragraph id="096E3697FFD5FFB2FF6D1957D1435646" blockId="2.[151,1436,1826,2031]" pageId="2" pageNumber="396">
<materialsCitation id="B9B93CCAFFD5FFB2FF6D1957D15E5646" ID-GBIF-Occurrence="1914123840" collectingDate="2009-05-04" collectingDateMax="2009-05-07" collectingDateMin="2009-05-04" collectionCode="MGCL, PPRI" collectorName="J. B. Heppner" country="Vietnam" elevation="155" location="Mac Lake" pageId="2" pageNumber="396" specimenCount="1" specimenCount-male="1" stateProvince="Ninh Binh Province" typeStatus="holotype">
<emphasis id="3BA5EA85FFD5FFB2FF6D1957D52C5622" bold="true" box="[199,317,1826,1851]" pageId="2" pageNumber="396">
<typeStatus id="D66A8835FFD5FFB2FF6D1957D5295622" box="[199,312,1826,1851]" pageId="2" pageNumber="396" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="1FD7FD1EFFD5FFB2FEEE1956D54A5623" box="[324,347,1827,1850]" pageId="2" pageNumber="396" type="male"></specimenCount>
,
<collectingCountry id="71C67607FFD5FFB2FEC21957D5DB5622" box="[360,458,1826,1851]" name="Vietnam" pageId="2" pageNumber="396">Vietnam</collectingCountry>
,
<collectingRegion id="CB15F875FFD5FFB2FE7C1957D6AB5622" box="[470,698,1826,1851]" country="Vietnam" name="Ninh Binh" pageId="2" pageNumber="396">
Ninh
<collectingRegion id="CB15F875FFD5FFB2FDBF1957D6AB5622" box="[533,698,1826,1851]" inferredBySuffix="true" pageId="2" pageNumber="396">Binh Province</collectingRegion>
</collectingRegion>
, Cuc Phuong National Park.,
<location id="0C0E604CFFD5FFB2FBA31956D06C5622" LSID="urn:lsid:plazi:treatment:81788781FFD6FFB4FF3D1C12D596542B:0C0E604CFFD5FFB2FBA31956D06C5622" box="[1033,1149,1826,1851]" country="Vietnam" name="Mac Lake" pageId="2" pageNumber="396" stateProvince="Ninh Binh Province">Mac Lake</location>
,
<elevation id="82FCD1A4FFD5FFB2FB231957D0DA5622" box="[1161,1227,1826,1851]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="2" pageNumber="396" unit="m" value="155.0">
<quantity id="CE299B72FFD5FFB2FB231957D0DA5622" box="[1161,1227,1826,1851]" metricMagnitude="2" metricUnit="m" metricValue="1.55" pageId="2" pageNumber="396" unit="m" value="155.0">155m</quantity>
</elevation>
,
<collectingDate id="6D2BE9BFFFD5FFB2FB7D1956D16E5622" box="[1239,1407,1827,1851]" pageId="2" pageNumber="396" value="2009-05-04" valueMax="2009-05-07" valueMin="2009-05-04">
<date id="7D6F1057FFD5FFB2FB7D1956D16E5622" box="[1239,1407,1827,1851]" pageId="2" pageNumber="396" value="2009-05-04" valueMax="2009-05-07" valueMin="2009-05-04">47 May 2009</date>
</collectingDate>
,
<collectorName id="A4245341FFD5FFB2FA201956D5095647" pageId="2" pageNumber="396">J. B. Heppner</collectorName>
(genitalia slide JBH 3491), deposited in
<collectionCode id="6FC0AE52FFD5FFB2FD491932D7255646" box="[739,820,1863,1887]" pageId="2" pageNumber="396">MGCL</collectionCode>
on indefinite loan from
<collectionCode id="6FC0AE52FFD5FFB2FBEF1932D0915647" box="[1093,1152,1863,1886]" country="South Africa" httpUri="http://grbio.org/cool/6j3y-rabv" name="ARC-Plant Protection Research Institute, National Collection of Fungi: Culture Collection" pageId="2" pageNumber="396">PPRI</collectionCode>
(
<collectingRegion id="CB15F875FFD5FFB2FB251933D0C65646" box="[1167,1239,1862,1887]" country="Vietnam" name="Ha Noi" pageId="2" pageNumber="396">Hanoi</collectingRegion>
,
<collectingCountry id="71C67607FFD5FFB2FB491933D1565646" box="[1251,1351,1862,1887]" name="Vietnam" pageId="2" pageNumber="396">Vietnam</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="096E3697FFD5FFB2FF6D191FD78356BE" blockId="2.[151,1436,1826,2031]" pageId="2" pageNumber="396">
<emphasis id="3BA5EA85FFD5FFB2FF6D191FD52E569A" bold="true" box="[199,319,1898,1923]" pageId="2" pageNumber="396">Paratypes</emphasis>
(1 ♂, 1 ♀):
<materialsCitation id="B9B93CCAFFD5FFB2FE63191ED777569A" ID-GBIF-Occurrence="1914123837" box="[457,870,1898,1923]" collectingDate="2009-05-04" collectingDateMax="2009-05-07" collectingDateMin="2009-05-04" collectionCode="MGCL" collectorName="J. B. Heppner" country="Vietnam" elevation="155" location="Mac Lake" pageId="2" pageNumber="396" specimenCount="1" specimenCount-male="1" stateProvince="Ninh Binh Province">
<specimenCount id="1FD7FD1EFFD5FFB2FE63191ED5E4569B" box="[457,501,1899,1923]" pageId="2" pageNumber="396" type="male">1 ♂</specimenCount>
, same data as holotype (
<collectionCode id="6FC0AE52FFD5FFB2FCAC191ED74C569A" box="[774,861,1899,1923]" pageId="2" pageNumber="396">MGCL</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="B9B93CCAFFD5FFB2FCDB191ED79F56BE" ID-GBIF-Occurrence="1914123838" collectingDate="2008-06-21" collectingDateMax="2008-06-26" collectingDateMin="2008-06-21" collectionCode="MGCL" collectorName="J. B. Heppner" country="Vietnam" elevation="240" location="Bac Kan Province" pageId="2" pageNumber="396" specimenCount="1" specimenCount-female="1" stateProvince="Bac Kan Province">
<specimenCount id="1FD7FD1EFFD5FFB2FCDB191ED78A569B" box="[881,923,1899,1923]" pageId="2" pageNumber="396" type="female">1 ♀</specimenCount>
, Ba Be Natl. Park,
<elevation id="82FCD1A4FFD5FFB2FBDF191ED0AF569B" box="[1141,1214,1899,1923]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="396" unit="m" value="240.0">
<quantity id="CE299B72FFD5FFB2FBDF191ED0AF569B" box="[1141,1214,1899,1923]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="396" unit="m" value="240.0">240 m</quantity>
</elevation>
,
<collectingRegion id="CB15F875FFD5FFB2FB60191ED188569A" box="[1226,1433,1898,1923]" country="Vietnam" name="Bac Kan" pageId="2" pageNumber="396">
Bac
<collectingRegion id="CB15F875FFD5FFB2FB56191ED188569A" box="[1276,1433,1898,1923]" inferredBySuffix="true" pageId="2" pageNumber="396">Kan Province</collectingRegion>
</collectingRegion>
,
<collectingDate id="6D2BE9BFFFD5FFB2FF3D19FBD55C56BE" box="[151,333,1934,1959]" pageId="2" pageNumber="396" value="2008-06-21" valueMax="2008-06-26" valueMin="2008-06-21">
<date id="7D6F1057FFD5FFB2FF3D19FBD55C56BE" box="[151,333,1934,1959]" pageId="2" pageNumber="396" value="2008-06-21" valueMax="2008-06-26" valueMin="2008-06-21">2126 Jun 2008</date>
</collectingDate>
,
<collectorName id="A4245341FFD5FFB2FEF219FAD5E056BF" box="[344,497,1935,1958]" pageId="2" pageNumber="396">J. B. Heppner</collectorName>
(genitalia slide JBH 3323) (
<collectionCode id="6FC0AE52FFD5FFB2FC8419FAD79456BE" box="[814,901,1935,1959]" pageId="2" pageNumber="396">MGCL</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="41CB651CFFD5FFB3FF6D19C7D1865059" lastPageId="3" lastPageNumber="397" pageId="2" pageNumber="396" type="biology_ecology">
<paragraph id="096E3697FFD5FFB3FF6D19C7D1865059" blockId="2.[151,1436,1826,2031]" lastBlockId="3.[151,1436,151,357]" lastPageId="3" lastPageNumber="397" pageId="2" pageNumber="396">
<emphasis id="3BA5EA85FFD5FFB2FF6D19C7D50E56D2" bold="true" box="[199,287,1970,1995]" pageId="2" pageNumber="396">Biology</emphasis>
. The biology of the new species is unknown. Larval hosts have been reported for three species of
<taxonomicName id="CED14D14FFD5FFB2FF3D19A3D51656F6" box="[151,263,2006,2031]" class="Insecta" family="Tortricidae" genus="Anthozela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="3BA5EA85FFD5FFB2FF3D19A3D51656F6" box="[151,263,2006,2031]" italics="true" pageId="2" pageNumber="396">Anthozela</emphasis>
</taxonomicName>
:
<taxonomicName id="CED14D14FFD5FFB2FEB319A2D63656F6" authority="Ghesquiere" authorityName="Ghesquiere" box="[281,551,2006,2031]" class="Insecta" family="Tortricidae" genus="Anthozela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="396" phylum="Arthropoda" rank="species" species="anonidii">
<emphasis id="3BA5EA85FFD5FFB2FEB319A2D58856F6" box="[281,409,2006,2031]" italics="true" pageId="2" pageNumber="396">
<collectingRegion id="CB15F875FFD5FFB2FEB319A2D53C56F7" box="[281,301,2007,2030]" country="Nigeria" name="Rivers" pageId="2" pageNumber="396">A</collectingRegion>
. anonidii
</emphasis>
Ghesquière
</taxonomicName>
, which has been recorded from the fruit of
<taxonomicName id="CED14D14FFD5FFB2FB8419A3D18D56F6" box="[1070,1436,2006,2031]" class="Magnoliopsida" family="Annonaceae" genus="Anonidium" higherTaxonomySource="GBIF" kingdom="Plantae" order="Magnoliales" pageId="2" pageNumber="396" phylum="Tracheophyta" rank="species" species="manii">
<emphasis id="3BA5EA85FFD5FFB2FB8419A3D0E556F7" box="[1070,1268,2006,2031]" italics="true" pageId="2" pageNumber="396">Anonidium manii</emphasis>
(Annonaceae)
</taxonomicName>
(
<bibRefCitation id="6D404B66FFD4FFB3FF341EE2D57451A9" box="[158,357,151,176]" pageId="3" pageNumber="397" refId="ref2866" refString="Ghesquiere, J. (1940) Catalogues Raisonnes de la Faune Entomologique du Congo Belge. Lepidopteres, Microlepidopteres. Tervuren: Musee du Congo Belge, 120 pp., 5 pl. [in Annales du Musee du Congo Belge. C. Zoologie, Series 3, 7, 1 - 120]" type="book chapter">Ghesquière 1940</bibRefCitation>
);
<taxonomicName id="CED14D14FFD4FFB3FED11EEDD68F51A9" authority="Meyrick" authorityName="Meyrick" authorityYear="1913" box="[379,670,151,176]" class="Insecta" family="Tortricidae" genus="Anthozela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="397" phylum="Arthropoda" rank="species" species="chrysoxantha">
<emphasis id="3BA5EA85FFD4FFB3FED11EEDD62251A9" box="[379,563,151,176]" italics="true" pageId="3" pageNumber="397">
<collectingRegion id="CB15F875FFD4FFB3FED11EEDD59E51B6" box="[379,399,152,175]" country="Nigeria" name="Rivers" pageId="3" pageNumber="397">A</collectingRegion>
. chrysoxantha
</emphasis>
Meyrick
</taxonomicName>
, which has been recorded from
<taxonomicName id="CED14D14FFD4FFB3FBA51EECD07551A9" authorityName="Linnaeus" authorityYear="1753" box="[1039,1124,153,176]" class="Magnoliopsida" family="Rubiaceae" genus="Pavetta" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="397" phylum="Tracheophyta" rank="genus">
<emphasis id="3BA5EA85FFD4FFB3FBA51EECD07551A9" box="[1039,1124,153,176]" italics="true" pageId="3" pageNumber="397">Pavetta</emphasis>
</taxonomicName>
sp. (Rubiacacae) (
<bibRefCitation id="6D404B66FFD4FFB3FA911EE2D4C251CC" pageId="3" pageNumber="397" refId="ref3125" refString="Meyrick, E. (1936) Exotic Microlepidoptera. Fol. 5. Printed by Taylor and Francis, London, 73 pp. [pp. 24 - 96]" type="book chapter">Meyrick 1936</bibRefCitation>
); and
<taxonomicName id="CED14D14FFD4FFB3FEB21EC9D63351CC" authority="(Meyrick)" authorityName="Meyrick" baseAuthorityName="Meyrick" baseAuthorityYear="1907" box="[280,546,188,213]" class="Insecta" family="Tortricidae" genus="Anthozela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="397" phylum="Arthropoda" rank="species" species="hemidoxa">
<emphasis id="3BA5EA85FFD4FFB3FEB21EC9D5B451CC" box="[280,421,188,213]" italics="true" pageId="3" pageNumber="397">
<collectingRegion id="CB15F875FFD4FFB3FEB21EC9D53D51CA" box="[280,300,188,211]" country="Nigeria" name="Rivers" pageId="3" pageNumber="397">A</collectingRegion>
. hemidoxa
</emphasis>
(Meyrick)
</taxonomicName>
, the pepper shoot borer or top shoot borer, which has been reared on shoots of pepper vine,
<taxonomicName id="CED14D14FFD4FFB3FE801E94D64851E1" box="[298,601,223,248]" class="Magnoliopsida" family="Piperaceae" genus="Piper" higherTaxonomySource="CoL" kingdom="Plantae" order="Piperales" pageId="3" pageNumber="397" phylum="Magnoliopsida" rank="species" species="nigrum">
<emphasis id="3BA5EA85FFD4FFB3FE801E94D5D051E1" box="[298,449,225,248]" italics="true" pageId="3" pageNumber="397">Piper nigrum</emphasis>
(Piperaceae)
</taxonomicName>
, in
<collectingCountry id="71C67607FFD4FFB3FD291EAAD6AD51E1" box="[643,700,223,248]" name="India" pageId="3" pageNumber="397">India</collectingCountry>
(
<bibRefCitation id="6D404B66FFD4FFB3FD661EAAD77D51E1" box="[716,876,223,248]" pageId="3" pageNumber="397" refId="ref2826" refString="Fletcher, T. B. (1921) Life-histories of Indian insects. Microlepidoptera. Memoires of the Department of Agriculture in India, Entomology Series, 6, 1 - 217, 68 pls. [1920]" type="journal article">Fletcher 1921</bibRefCitation>
,
<bibRefCitation id="6D404B66FFD4FFB3FCD31EAAD10F51E1" box="[889,1310,223,248]" pageId="3" pageNumber="397" refId="ref2502" refString="Devasahayam, S. &amp; Abdulla Koya, K. M. (1993) Seasonal incidence of hymenopterous parasitoids of top shoot borer, Cydia hemidoxa Meyrick (Lepidoptera: Tortricidae) on black pepper. Journal of Entomological Research, New Delhi, 17 (3), 205 - 207." type="journal article">Devasahayam &amp; Abdulla Koya 1993</bibRefCitation>
,
<bibRefCitation id="6D404B66FFD4FFB3FA801E95D17251E1" box="[1322,1379,224,248]" pageId="3" pageNumber="397" refId="ref2556" refString="Devasahayam, S. &amp; Abdula Koya, K. M. (1994) Seasonal incidence of Hexamermis sp. (Dor., Merminthidae) parasitizing larvae of top shoot borer Cydia hemidoxa Meyr. (Lep., Tortricidae) on black pepper. Journal of Applied Entomology, 117, 31 - 34. https: // doi. org / 10.1111 / j. 1439 - 0418.1994. tb 00704. x" type="journal article">1994</bibRefCitation>
) and
<collectingCountry id="71C67607FFD4FFB3FF3D1F71D5155004" box="[151,260,260,285]" name="Indonesia" pageId="3" pageNumber="397">Indonesia</collectingCountry>
(
<bibRefCitation id="6D404B66FFD4FFB3FEBF1F71D5C55004" box="[277,468,260,285]" pageId="3" pageNumber="397" refId="ref2951" refString="Kalshoven, L. G. E. (1950) De Plagen van de Cultuurgewassen in Indonesie. Fol. 1. W. Van Hoeve, The Hague, 512 pp. [English translation, 1980: Pests of Crops in Indonesia, Jakarta, 701 pp.]" type="book">Kalshoven 1950</bibRefCitation>
).
<bibRefCitation id="6D404B66FFD4FFB3FE401F71D6955004" box="[490,644,260,285]" pageId="3" pageNumber="397" refId="ref2923" refString="Horak, M. (2006) Olethreutine Moths of Australia (Lepidoptera: Tortricidae). Monographs on Australia Lepidoptera, 10, 1 - 522." type="journal article">Horak (2006)</bibRefCitation>
notes that larvae of
<taxonomicName id="CED14D14FFD4FFB3FCDE1F71D0125004" baseAuthorityName="Meyrick" baseAuthorityYear="1907" box="[884,1027,260,285]" class="Insecta" family="Tortricidae" genus="Anthozela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="397" phylum="Arthropoda" rank="species" species="hemidoxa">
<emphasis id="3BA5EA85FFD4FFB3FCDE1F71D0125004" box="[884,1027,260,285]" italics="true" pageId="3" pageNumber="397">
<collectingRegion id="CB15F875FFD4FFB3FCDE1F71D7995002" box="[884,904,260,283]" country="Nigeria" name="Rivers" pageId="3" pageNumber="397">A</collectingRegion>
. hemidoxa
</emphasis>
</taxonomicName>
have been found on
<taxonomicName id="CED14D14FFD4FFB3FB501F70D5335059" authorityName="Ohwi (Choisy" authorityYear="1934" class="Magnoliopsida" family="Piperaceae" genus="Piper" higherTaxonomySource="CoL" kingdom="Plantae" order="Piperales" pageId="3" pageNumber="397" phylum="Magnoliopsida" rank="species" species="kadsura">
<emphasis id="3BA5EA85FFD4FFB3FB501F70D18D5004" box="[1274,1436,260,285]" italics="true" pageId="3" pageNumber="397">Piper kadsura</emphasis>
(Piperaceae)
</taxonomicName>
in the Ryukyu Islands, but comments that there may be more than one species of
<taxonomicName id="CED14D14FFD4FFB3FB1C1F52D1375059" box="[1206,1318,295,320]" class="Insecta" family="Tortricidae" genus="Anthozela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="397" phylum="Arthropoda" rank="genus">
<emphasis id="3BA5EA85FFD4FFB3FB1C1F52D1375059" box="[1206,1318,295,320]" italics="true" pageId="3" pageNumber="397">Anthozela</emphasis>
</taxonomicName>
involved.
</paragraph>
</subSubSection>
<subSubSection id="41CB651CFFD4FFB3FF6D1F39D7C6507C" box="[199,983,332,357]" pageId="3" pageNumber="397" type="distribution">
<paragraph id="096E3697FFD4FFB3FF6D1F39D7C6507C" blockId="3.[151,1436,151,357]" box="[199,983,332,357]" pageId="3" pageNumber="397">
<emphasis id="3BA5EA85FFD4FFB3FF6D1F39D548507C" bold="true" box="[199,345,332,357]" pageId="3" pageNumber="397">Distribution</emphasis>
. The new species is known only from northern
<collectingCountry id="71C67607FFD4FFB3FCDA1F39D7C2507C" box="[880,979,332,357]" name="Vietnam" pageId="3" pageNumber="397">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="5DAE661FFFD4FFB3FF3D1923D1595674" box="[151,1352,1878,1901]" httpUri="https://zenodo.org/record/1444868/files/figure.png" pageId="3" pageNumber="397" startId="3.[151,264,1878,1901]" targetBox="[151,1435,409,1858]" targetPageId="3">
<paragraph id="096E3697FFD4FFB3FF3D1923D1595674" blockId="3.[151,1352,1878,1901]" box="[151,1352,1878,1901]" pageId="3" pageNumber="397">
<emphasis id="3BA5EA85FFD4FFB3FF3D1923D52B5675" bold="true" box="[151,314,1878,1901]" pageId="3" pageNumber="397">FIGURES 36.</emphasis>
<taxonomicName id="CED14D14FFD4FFB3FEEA1922D6545674" authority="Heppner &amp; Bae, 2018" authorityName="Heppner &amp; Bae" authorityYear="2018" box="[320,581,1879,1901]" class="Insecta" family="Tortricidae" genus="Anthozela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="397" phylum="Arthropoda" rank="species" species="cypriflammella" status="sp. nov.">
<emphasis id="3BA5EA85FFD4FFB3FEEA1922D6545674" box="[320,581,1879,1901]" italics="true" pageId="3" pageNumber="397">Anthozela cypriflammella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="209657FEFFD4FFB3FDE11922D6955675" box="[587,644,1879,1900]" pageId="3" pageNumber="397" rank="species">
<emphasis id="3BA5EA85FFD4FFB3FDE11922D6955675" bold="true" box="[587,644,1879,1900]" pageId="3" pageNumber="397">n. sp.</emphasis>
</taxonomicNameLabel>
, close-up of head (lateral and dorsal views). 34) Male; 56) Female.
</paragraph>
</caption>
<caption id="5DAE661FFFD3FFB4FF3D1AE6D5B955D0" httpUri="https://zenodo.org/record/1444870/files/figure.png" pageId="4" pageNumber="398" startId="4.[151,264,1171,1194]" targetBox="[166,1426,195,1149]" targetPageId="4">
<paragraph id="096E3697FFD3FFB4FF3D1AE6D5B955D0" blockId="4.[151,1435,1171,1225]" pageId="4" pageNumber="398">
<emphasis id="3BA5EA85FFD3FFB4FF3D1AE6D53055B0" bold="true" box="[151,289,1171,1194]" pageId="4" pageNumber="398">FIGURES 7.</emphasis>
<taxonomicName id="CED14D14FFD3FFB4FE8D1AE1D63D55B3" authority="Heppner &amp; Bae, 2018" authorityName="Heppner &amp; Bae" authorityYear="2018" box="[295,556,1172,1194]" class="Insecta" family="Tortricidae" genus="Anthozela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="398" phylum="Arthropoda" rank="species" species="cypriflammella" status="sp. nov.">
<emphasis id="3BA5EA85FFD3FFB4FE8D1AE1D63D55B3" box="[295,556,1172,1194]" italics="true" pageId="4" pageNumber="398">Anthozela cypriflammella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="209657FEFFD3FFB4FD991AE1D67D55B0" box="[563,620,1172,1193]" pageId="4" pageNumber="398" rank="species">
<emphasis id="3BA5EA85FFD3FFB4FD991AE1D67D55B0" bold="true" box="[563,620,1172,1193]" pageId="4" pageNumber="398">n. sp.</emphasis>
</taxonomicNameLabel>
: male genitalia. a) phallus; b) central genital close-up; c) pregenital sternal plate (genitalia slide JBH 3491).
</paragraph>
</caption>
<subSubSection id="41CB651CFFD3FFB4FF6D1A80D596542B" pageId="4" pageNumber="398" type="etymology">
<paragraph id="096E3697FFD3FFB4FF6D1A80D596542B" blockId="4.[151,1436,1269,1330]" pageId="4" pageNumber="398">
<emphasis id="3BA5EA85FFD3FFB4FF6D1A80D5575417" bold="true" box="[199,326,1269,1294]" pageId="4" pageNumber="398">Etymology</emphasis>
. The specific epithet refers to the red and yellow forewing coloration, latinized for &quot;beautiful little flame-colored&quot; moth.
</paragraph>
</subSubSection>
</treatment>
</document>