285 lines
32 KiB
XML
285 lines
32 KiB
XML
<document id="F207F625FE06D2E624178F4DC3EB71C1" ID-CLB-Dataset="46667" ID-DOI="10.11646/zootaxa.3754.3.2" ID-GBIF-Dataset="fe375541-8083-43f8-9cf8-b8d75af79599" ID-ISSN="1175-5326" ID-Zenodo-Dep="251551" ID-ZooBank="9E645014-5464-4E7C-8D4A-0B3B52A5AA53" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="carolina" checkinTime="1460368973748" checkinUser="plazi" docAuthor="Kantor, Yuri, Lozouet, Pierre, Puillandre, Nicolas & Bouchet, Philippe" docDate="2014" docId="0F4E87BB7B3FFFACFF22F913A630FE6C" docLanguage="en" docName="zt03754p276.pdf" docOrigin="Zootaxa 3754 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hortia macrocephala Kantor, Lozouet, Puillandre & Bouchet, 2014, new species" docType="treatment" docUuid="12ED529C-73B0-4284-89A7-E4832B1F6A7F" docUuidSource="ZooBank" docVersion="11" lastPageNumber="256" masterDocId="F377FFC37B31FFBDFFB5FFADA776FFE4" masterDocTitle="Lost and found: The Eocene family Pyramimitridae (Neogastropoda) discovered in the Recent fauna of the Indo-Pacific" masterLastPageNumber="276" masterPageNumber="239" pageNumber="253" updateTime="1728431205123" updateUser="ExternalLinkService">
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<mods:mods id="8A3F6AF3C4EF318026D49378CACA5113" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B6409C9FD2E03D7F439EFE2E9AF452F9">
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<mods:title id="7E61FDF4140152DB30514DFE632B09FA">Lost and found: The Eocene family Pyramimitridae (Neogastropoda) discovered in the Recent fauna of the Indo-Pacific</mods:title>
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</mods:titleInfo>
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<mods:name id="55FF3DD037FE77158C45A31CC4409C8B" type="personal">
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<mods:roleTerm id="417818498FB00300CE3DD9B8D6844BC3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C55D4CEE9A4E66FD166FF4B826D175D8">Kantor, Yuri</mods:namePart>
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<mods:namePart id="FAAC2A1DD7E7AD4D10EF2FFE40B2AF65">Lozouet, Pierre</mods:namePart>
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<mods:namePart id="0B7926D5A8409DC4D8F75F73329C69B6">Puillandre, Nicolas</mods:namePart>
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</mods:role>
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<mods:namePart id="3B601F493703AED882F4BA5428EF4AD8">Bouchet, Philippe</mods:namePart>
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<mods:typeOfResource id="FD3EED9FA0F719EA077CEFD47ABC195A">text</mods:typeOfResource>
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<mods:title id="E90D7F86FA7B6407AB58AA13EA2F1900">Zootaxa</mods:title>
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<mods:part id="8D88785FE9450AE24CBC4EF3603BD03D">
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<mods:date id="FC0A1774A7606785B6A36CDC28CA9117">2014</mods:date>
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<mods:detail id="545CCC1BF5D88F9FBB31CEBA69FC5F29" type="volume">
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<mods:number id="4BE18D6ECF0C69662C6379EFF797C4AE">3754</mods:number>
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<mods:detail id="82B7475FBD8F83281BAC67BF9F7692CC" type="issue">
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<mods:number id="2DEFA150F089354B8071C0DF58ED637C">3</mods:number>
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<mods:identifier id="614C5A18EC16933718B232534F9A63A2" type="CLB-Dataset">46667</mods:identifier>
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<mods:identifier id="8ACAE9D3A71CE015D7DEC0A3BBCC9225" type="DOI">10.11646/zootaxa.3754.3.2</mods:identifier>
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<treatment id="0F4E87BB7B3FFFACFF22F913A630FE6C" ID-DOI="http://doi.org/10.5281/zenodo.5676780" ID-GBIF-Taxon="119411399" ID-Zenodo-Dep="5676780" LSID="urn:lsid:zoobank.org:act:12ED529C-73B0-4284-89A7-E4832B1F6A7F" httpUri="http://treatment.plazi.org/id/0F4E87BB7B3FFFACFF22F913A630FE6C" lastPageId="17" lastPageNumber="256" pageId="14" pageNumber="253">
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<subSubSection id="CFFD65267B3FFFB3FF22F913A60EF91D" pageId="14" pageNumber="253" type="nomenclature">
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<paragraph id="875836AD7B3FFFB3FF22F913A546F93C" blockId="14.[151,560,1726,1785]" box="[151,560,1726,1752]" pageId="14" pageNumber="253">
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<heading id="DC1081C17B3FFFB3FF22F913A546F93C" bold="true" box="[151,560,1726,1752]" fontSize="11" level="1" pageId="14" pageNumber="253" reason="1">
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<emphasis id="B593EABF7B3FFFB3FF22F913A546F93C" bold="true" box="[151,560,1726,1752]" pageId="14" pageNumber="253">
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<taxonomicName id="40E74D2E7B3FFFB3FF22F913A6EFF93C" box="[151,409,1726,1752]" class="Magnoliopsida" family="Pyramimitridae" genus="Hortia" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="253" phylum="Mollusca" rank="species" species="macrocephala" status="sp. nov.">
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<emphasis id="B593EABF7B3FFFB3FF22F913A6EFF93C" bold="true" box="[151,409,1726,1752]" italics="true" pageId="14" pageNumber="253">Hortia macrocephala</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="AEA057C47B3FFFB3FE15F913A546F93C" box="[416,560,1726,1752]" pageId="14" pageNumber="253" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="875836AD7B3FFFB3FF22F94DA60EF91D" blockId="14.[151,560,1726,1785]" box="[151,376,1760,1785]" pageId="14" pageNumber="253">
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<figureCitation id="1FDC2A287B3FFFB3FF22F94DA677F91D" box="[151,257,1760,1785]" captionStart="FIGURE 2" captionStartId="6.[151,255,1693,1716]" captionTargetBox="[182,1406,179,1669]" captionTargetId="figure@6.[182,1406,157,1669]" captionTargetPageId="6" captionText="FIGURE 2. Overview of fossil and Recent Pyramimitridae. A. Endiatoma quadricincta (Cossmann, 1883), MNHN. F. A 47757, Lower Eocene, France. B – C. Pyramimitra terebraeformis (Conrad, 1848), MNHN. F. J 08748, Middle Eocene, USA. D – E. Pyramimitra quadralirata Dockery, 1977, MNHN. F. A 47750, Middle Eocene, USA. F. Hortia priabonica n. sp., MNHN. F. A 47752, Upper Eocene, France. G. Hortia arriuensis Lozouet, 1999, MNHN. F. A 47751, Upper Oligocene, France. H. Hortia pseudotaranis n. sp., New Caledonia, SL 8.4 mm. I. Hortia macrocephala n. sp., New Caledonia, SL 10.4 mm. J. Hortia paradrillia n. sp., New Caledonia, SL 8.1 mm. K. Hortia marshalli n. sp., New Zealand, SL 10.6 mm. L. Hortia spenceri n. sp., New Zealand, SL 9.0 mm. M. Hortia aotearoa n. sp., New Zealand, SL 9.0 mm. N. Hortia solitaria n. sp., South of Madagascar, SL 6.5 mm. O. Teremitra fallax n. sp., Solomon Islands, SL 11.8 mm. P. Teremitra efatensis (Aubry, 1999), Vanuatu, SL 10.8 mm. Q. Teremitra? hortensis (Vinassa de Regny, 1898), Upper Eocene, France. R. Teremitra sp., Taiwan, SL 16 mm. S. Vaughanites? superstes n. sp., Philippines, SL 19.1 mm." figureDoi="http://doi.org/10.5281/zenodo.251553" httpUri="https://zenodo.org/record/251553/files/figure.png" pageId="14" pageNumber="253">
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Figures
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<date id="F359106D7B3FFFB3FF47F94CA677F91D" box="[242,257,1761,1785]" pageId="14" pageNumber="253">2</date>
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</figureCitation>
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I, 7A–D, 9
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="CFFD65267B3FFFB3FF22F885A433F811" pageId="14" pageNumber="253" type="materials_examined">
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<paragraph id="875836AD7B3FFFB3FF22F885A4EAF8A5" blockId="14.[151,1436,1832,2037]" box="[151,924,1832,1857]" pageId="14" pageNumber="253">
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<emphasis id="B593EABF7B3FFFB3FF22F885A633F8A5" bold="true" box="[151,325,1832,1857]" pageId="14" pageNumber="253">
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<typeStatus id="585C880F7B3FFFB3FF22F885A7A4F8A5" box="[151,210,1832,1857]" pageId="14" pageNumber="253">Type</typeStatus>
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material.
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||
</emphasis>
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<typeStatus id="585C880F7B3FFFB3FEF9F885A6C2F8A5" box="[332,436,1832,1857]" pageId="14" pageNumber="253" type="holotype">Holotype</typeStatus>
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<collectionCode id="E1F6AE687B3FFFB3FE0EF884A567F8A4" LSID="urn:lsid:biocol.org:col:34988" box="[443,529,1833,1856]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="253">MNHN</collectionCode>
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24571, 5
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<typeStatus id="585C880F7B3FFFB3FD35F884A59DF8A5" box="[640,747,1833,1857]" pageId="14" pageNumber="253" type="paratype">paratypes</typeStatus>
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<collectionCode id="E1F6AE687B3FFFB3FD46F884A43EF8A4" LSID="urn:lsid:biocol.org:col:34988" box="[755,840,1833,1856]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="253">MNHN</collectionCode>
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24572.
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</paragraph>
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<paragraph id="875836AD7B3FFFB3FF72F8E1A3D9F880" blockId="14.[151,1436,1832,2037]" box="[199,1199,1868,1893]" pageId="14" pageNumber="253">
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<emphasis id="B593EABF7B3FFFB3FF72F8E1A613F881" bold="true" box="[199,357,1868,1893]" pageId="14" pageNumber="253">
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<typeStatus id="585C880F7B3FFFB3FF72F8E1A674F881" box="[199,258,1868,1893]" pageId="14" pageNumber="253">Type</typeStatus>
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locality.
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||
</emphasis>
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South of
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||
<collectingCountry id="FFF0763D7B3FFFB3FE66F8E0A5F5F881" box="[467,643,1868,1893]" name="New Caledonia" pageId="14" pageNumber="253">New Caledonia</collectingCountry>
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,
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,
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<geoCoordinate id="E2D3506A7B3FFFB3FD40F8E1A417F881" box="[757,865,1868,1893]" direction="east" orientation="longitude" pageId="14" pageNumber="253" precision="925" value="167.25">167º15’E</geoCoordinate>
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,
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<quantity id="401F9B487B3FFFB3FCD8F8E0A4C2F880" box="[877,948,1869,1893]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="14" pageNumber="253" unit="m" value="440.0">440 m</quantity>
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||
[BIOCAL st. DW77].
|
||
</paragraph>
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||
<paragraph id="875836AD7B3FFFB3FF72F8DDA433F811" blockId="14.[151,1436,1832,2037]" pageId="14" pageNumber="253">
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<emphasis id="B593EABF7B3FFFB3FF72F8DDA6C7F86D" bold="true" box="[199,433,1904,1929]" pageId="14" pageNumber="253">Material examined.</emphasis>
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South of
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<collectingCountry id="FFF0763D7B3FFFB3FD96F8DCA5A2F86D" box="[547,724,1904,1929]" name="New Caledonia" pageId="14" pageNumber="253">New Caledonia</collectingCountry>
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||
. BIOCAL, st. DW 77,
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<geoCoordinate id="E2D3506A7B3FFFB3FC69F8DDA341F86D" box="[988,1079,1904,1929]" direction="south" orientation="latitude" pageId="14" pageNumber="253" precision="925" value="-22.25">22º15’S</geoCoordinate>
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,
|
||
<geoCoordinate id="E2D3506A7B3FFFB3FBF1F8DDA3C6F86D" box="[1092,1200,1904,1929]" direction="east" orientation="longitude" pageId="14" pageNumber="253" precision="925" value="167.25">167º15’E</geoCoordinate>
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,
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<quantity id="401F9B487B3FFFB3FB08F8DCA271F86C" box="[1213,1287,1905,1929]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="14" pageNumber="253" unit="m" value="440.0">440 m</quantity>
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, 35 dd, 11 lv (
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<typeStatus id="585C880F7B3FFFB3FF2BF839A674F849" box="[158,258,1940,1965]" pageId="14" pageNumber="253" type="holotype">holotype</typeStatus>
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and
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<typeStatus id="585C880F7B3FFFB3FE8CF838A6DCF849" box="[313,426,1941,1965]" pageId="14" pageNumber="253" type="paratype">paratypes</typeStatus>
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). EXBODI, st. CP 3786,
|
||
<geoCoordinate id="E2D3506A7B3FFFB3FD71F839A457F849" box="[708,801,1940,1965]" direction="south" orientation="latitude" pageId="14" pageNumber="253" precision="925" value="-22.25">22º15’S</geoCoordinate>
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,
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<geoCoordinate id="E2D3506A7B3FFFB3FC98F839A4EEF849" box="[813,920,1940,1965]" direction="east" orientation="longitude" pageId="14" pageNumber="253" precision="925" value="167.21666">167º13’E</geoCoordinate>
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,
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<quantity id="401F9B487B3FFFB3FC11F838A353F848" box="[932,1061,1941,1965]" metricMagnitude="2" metricUnit="m" metricValue="4.25" metricValueMax="4.42" metricValueMin="4.08" pageId="14" pageNumber="253" unit="m" value="425.0" valueMax="442.0" valueMin="408.0">408–442 m</quantity>
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, 4 dd, 2 lv; st. CP 3787,
|
||
<geoCoordinate id="E2D3506A7B3FFFB3FA8EF838A2EEF849" box="[1339,1432,1941,1965]" direction="south" orientation="latitude" pageId="14" pageNumber="253" precision="925" value="-22.216667">22º13’S</geoCoordinate>
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,
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<geoCoordinate id="E2D3506A7B3FFFB3FF22F815A674F835" box="[151,258,1976,2001]" direction="east" orientation="longitude" pageId="14" pageNumber="253" precision="925" value="167.1">167º06’E</geoCoordinate>
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,
|
||
<quantity id="401F9B487B3FFFB3FEBBF814A6F8F834" box="[270,398,1977,2001]" metricMagnitude="2" metricUnit="m" metricValue="2.36" metricValueMax="2.4899999999999998" metricValueMin="2.23" pageId="14" pageNumber="253" unit="m" value="236.0" valueMax="249.0" valueMin="223.0">223–249 m</quantity>
|
||
, 1 dd; st. CP 3811,
|
||
<geoCoordinate id="E2D3506A7B3FFFB3FDD2F814A5B5F835" box="[615,707,1977,2001]" direction="south" orientation="latitude" pageId="14" pageNumber="253" precision="925" value="-21.733334">21º44’S</geoCoordinate>
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,
|
||
<geoCoordinate id="E2D3506A7B3FFFB3FD7AF815A44CF835" box="[719,826,1976,2001]" direction="east" orientation="longitude" pageId="14" pageNumber="253" precision="925" value="166.63333">166º38’E</geoCoordinate>
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||
,
|
||
<quantity id="401F9B487B3FFFB3FCF3F815A4B1F834" box="[838,967,1976,2001]" metricMagnitude="2" metricUnit="m" metricValue="5.275" metricValueMax="6.68" metricValueMin="3.87" pageId="14" pageNumber="253" unit="m" value="527.5" valueMax="668.0" valueMin="387.0">387–668 m</quantity>
|
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, 1 dd. TERRASSE, st. DW3086, Terrasse sud-est, passe de la Sarcelle,
|
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<geoCoordinate id="E2D3506A7B3FFFB3FE69F871A541F811" box="[476,567,2012,2037]" direction="south" orientation="latitude" pageId="14" pageNumber="253" precision="925" value="-21.916666">21°55'S</geoCoordinate>
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,
|
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<geoCoordinate id="E2D3506A7B3FFFB3FDF7F871A5DAF811" box="[578,684,2012,2037]" direction="east" orientation="longitude" pageId="14" pageNumber="253" precision="925" value="167.21666">167°13'E</geoCoordinate>
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,
|
||
<quantity id="401F9B487B3FFFB3FD0DF870A477F810" box="[696,769,2013,2037]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="253" unit="m" value="400.0">400 m</quantity>
|
||
, 4 dd.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="D39866257B3EFFB2FF22F9B8A2E6F988" ID-DOI="http://doi.org/10.5281/zenodo.251558" httpUri="https://zenodo.org/record/251558/files/figure.png" pageId="15" pageNumber="254" targetBox="[188,1398,195,1533]" targetPageId="15">
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<paragraph id="875836AD7B3EFFB2FF22F9B8A2E6F988" blockId="15.[151,1437,1557,1644]" pageId="15" pageNumber="254">
|
||
<emphasis id="B593EABF7B3EFFB2FF22F9B8A66EF9C8" bold="true" box="[151,280,1557,1580]" pageId="15" pageNumber="254">FIGURE 7.</emphasis>
|
||
Radulae of Recent
|
||
<taxonomicName id="40E74D2E7B3EFFB2FE5CF9B8A546F9C8" box="[489,560,1557,1580]" class="Magnoliopsida" family="Pyramimitridae" genus="Hortia" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="254" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B593EABF7B3EFFB2FE5CF9B8A546F9C8" box="[489,560,1557,1580]" italics="true" pageId="15" pageNumber="254">Hortia</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
<emphasis id="B593EABF7B3EFFB2FD25F9B8A5BDF9C8" bold="true" box="[656,715,1557,1580]" pageId="15" pageNumber="254">A–D.</emphasis>
|
||
<taxonomicName id="40E74D2E7B3EFFB2FD64F9B8A4C0F9C8" box="[721,950,1557,1580]" class="Magnoliopsida" family="Pyramimitridae" genus="Hortia" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="254" phylum="Mollusca" rank="species" species="macrocephala" status="sp. nov.">
|
||
<emphasis id="B593EABF7B3EFFB2FD64F9B8A4C0F9C8" box="[721,950,1557,1580]" italics="true" pageId="15" pageNumber="254">Hortia macrocephala</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="AEA057C47B3EFFB2FC09F9B8A480F9C8" box="[956,1014,1557,1580]" pageId="15" pageNumber="254" rank="species">n. sp.</taxonomicNameLabel>
|
||
, holotype.
|
||
<emphasis id="B593EABF7B3EFFB2FBD9F9B8A3F3F9C8" bold="true" box="[1132,1157,1557,1580]" pageId="15" pageNumber="254">A.</emphasis>
|
||
Entire radular membrane.
|
||
<emphasis id="B593EABF7B3EFFB2FF22F998A7D9F9A8" bold="true" box="[151,175,1589,1612]" pageId="15" pageNumber="254">B.</emphasis>
|
||
Enlarged lateral teeth.
|
||
<emphasis id="B593EABF7B3EFFB2FE1EF998A6B2F9A8" bold="true" box="[427,452,1589,1612]" pageId="15" pageNumber="254">C.</emphasis>
|
||
Enlarged anteriormost part of radular membrane.
|
||
<emphasis id="B593EABF7B3EFFB2FC56F998A48BF9A8" bold="true" box="[995,1021,1589,1612]" pageId="15" pageNumber="254">D.</emphasis>
|
||
Enlarged rachidian teeth.
|
||
<emphasis id="B593EABF7B3EFFB2FAACF998A23BF9A8" bold="true" box="[1305,1357,1589,1612]" pageId="15" pageNumber="254">E–F.</emphasis>
|
||
<taxonomicName id="40E74D2E7B3EFFB2FAE0F998A67AF988" class="Magnoliopsida" family="Pyramimitridae" genus="Hortia" kingdom="Animalia" order="Neogastropoda" pageId="15" pageNumber="254" phylum="Mollusca" rank="species" species="paradrillia" status="sp. nov.">
|
||
<emphasis id="B593EABF7B3EFFB2FAE0F998A67AF988" italics="true" pageId="15" pageNumber="254">Hortia paradrillia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="AEA057C47B3EFFB2FEA6F9F8A63BF988" box="[275,333,1621,1644]" pageId="15" pageNumber="254" rank="species">n. sp.</taxonomicNameLabel>
|
||
, holotype.
|
||
<emphasis id="B593EABF7B3EFFB2FE76F9F8A6ADF988" bold="true" box="[451,475,1621,1644]" pageId="15" pageNumber="254">E.</emphasis>
|
||
Dorsal view of central part of radular membrane.
|
||
<emphasis id="B593EABF7B3EFFB2FC41F9F8A373F988" bold="true" box="[1012,1029,1621,1644]" pageId="15" pageNumber="254">F.</emphasis>
|
||
Enlarged rachidian and lateral teeth.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="CFFD65267B3EFFADFF72F938A5F7F87E" lastPageId="16" lastPageNumber="255" pageId="15" pageNumber="254" type="description">
|
||
<paragraph id="875836AD7B3EFFADFF72F938A5E7F97A" blockId="15.[151,1437,1685,2034]" lastBlockId="16.[151,1437,1561,2018]" lastPageId="16" lastPageNumber="255" pageId="15" pageNumber="254">
|
||
<emphasis id="B593EABF7B3EFFB2FF72F938A627F94A" bold="true" box="[199,337,1685,1710]" pageId="15" pageNumber="254">Description</emphasis>
|
||
(
|
||
<typeStatus id="585C880F7B3EFFB2FED5F938A6B0F94A" box="[352,454,1685,1710]" pageId="15" pageNumber="254" type="holotype">holotype</typeStatus>
|
||
) (
|
||
<figureCitation id="1FDC2A287B3EFFB2FE69F938A569F94A" box="[476,543,1685,1710]" captionStart="FIGURE 9" captionStartId="17.[151,255,1701,1724]" captionTargetBox="[188,1398,435,1676]" captionTargetId="figure@17.[188,1398,435,1676]" captionTargetPageId="17" captionText="FIGURE 9. Shells of Hortia macrocephala n. sp. A – D. Holotype, SL 10.4 mm. D. Enlarged protoconch. E. Paratype, SL 11.2 mm. F. Paratype, SL 8.8 mm. G. Paratype, SL 9.5 mm. H – I. SEM photographs of shell (H) and protoconch (I) of the specimen from the station DW 77, SL 8.7 mm. All shells at the same scale." figureDoi="http://doi.org/10.5281/zenodo.251560" httpUri="https://zenodo.org/record/251560/files/figure.png" pageId="15" pageNumber="254">Fig. 9</figureCitation>
|
||
A–D). Shell small, narrowly fusiform, with high, elevated spire (LWL/SL 0.46) and attenuated, medium long canal. Shell of ca 1.5 protoconch whorls and 6.5 slightly convex teleoconch whorls. Protoconch (
|
||
<figureCitation id="1FDC2A287B3EFFB2FE90F970A61EF912" box="[293,360,1757,1782]" captionStart="FIGURE 9" captionStartId="17.[151,255,1701,1724]" captionTargetBox="[188,1398,435,1676]" captionTargetId="figure@17.[188,1398,435,1676]" captionTargetPageId="17" captionText="FIGURE 9. Shells of Hortia macrocephala n. sp. A – D. Holotype, SL 10.4 mm. D. Enlarged protoconch. E. Paratype, SL 11.2 mm. F. Paratype, SL 8.8 mm. G. Paratype, SL 9.5 mm. H – I. SEM photographs of shell (H) and protoconch (I) of the specimen from the station DW 77, SL 8.7 mm. All shells at the same scale." figureDoi="http://doi.org/10.5281/zenodo.251560" httpUri="https://zenodo.org/record/251560/files/figure.png" pageId="15" pageNumber="254">Fig. 9</figureCitation>
|
||
D) paucispiral, semitransparent, globose, smooth, diameter 670 Μm, height 610 Μm, transition to teleoconch marked by two weak opisthocline ribs. Suture impressed. Besides inconspicuous growth lines, axial sculpture of narrow, closely spaced, sinuous and slightly prosocline ribs, 18 on penultimate whorl, 19 on last whorl. Ribs run from suture to suture on spire whorls and become obsolete on shell base at transition to canal. Spiral sculpture of one weak spiral thread abapically of suture and distinct, broadly spaced, narrow cords, rounded in section, two on first whorl, three starting from second whorl and 19 on last whorl, of which 13 on shell base and canal, more closely spaced. Cords on spire whorls and four cords on last whorl form small but distinct rounded beads at intersection with axial ribs. Aperture low, narrow elongate, 0.37 (together with canal) of SL. Outer lip partially broken, concave at transition to long, narrow, and nearly straight canal. Shallow notch at shoulder. Columella short, straight, smooth. Callus narrow, of thin transparent glaze overlying parietal region. Siphonal notch absent. Canal well differentiated from aperture, recurved abaxially to left. Shell color off-white. Periostracum transparent, yellowish, peeling.
|
||
</paragraph>
|
||
<caption id="D39866257B21FFADFF22FA98A7AFFA08" ID-DOI="http://doi.org/10.5281/zenodo.251559" httpUri="https://zenodo.org/record/251559/files/figure.png" pageId="16" pageNumber="255" targetBox="[170,1416,195,1303]" targetPageId="16">
|
||
<paragraph id="875836AD7B21FFADFF22FA98A7AFFA08" blockId="16.[151,1437,1333,1516]" pageId="16" pageNumber="255">
|
||
<emphasis id="B593EABF7B21FFADFF22FA98A66CFAA8" bold="true" box="[151,282,1333,1356]" pageId="16" pageNumber="255">FIGURE 8.</emphasis>
|
||
Anatomy of
|
||
<taxonomicName id="40E74D2E7B21FFADFE1EFA98A519FAA8" box="[427,623,1333,1356]" class="Magnoliopsida" family="Pyramimitridae" genus="Hortia" kingdom="Animalia" order="Neogastropoda" pageId="16" pageNumber="255" phylum="Mollusca" rank="species" species="paradrillia" status="sp. nov.">
|
||
<emphasis id="B593EABF7B21FFADFE1EFA98A519FAA8" box="[427,623,1333,1356]" italics="true" pageId="16" pageNumber="255">Hortia paradrillia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="AEA057C47B21FFADFDC2FA98A5C4FAA8" box="[631,690,1333,1356]" pageId="16" pageNumber="255" rank="species">n. sp.</taxonomicNameLabel>
|
||
<emphasis id="B593EABF7B21FFADFD0CFA98A584FAA8" bold="true" box="[697,754,1333,1356]" pageId="16" pageNumber="255">A–B.</emphasis>
|
||
Anterior part of the body, removed from the shell.
|
||
<emphasis id="B593EABF7B21FFADFA95FA98A24DFAA8" bold="true" box="[1312,1339,1333,1356]" pageId="16" pageNumber="255">C.</emphasis>
|
||
Anterior part of the mantle.
|
||
<emphasis id="B593EABF7B21FFADFEDAFAF8A6DFFA88" bold="true" box="[367,425,1365,1388]" pageId="16" pageNumber="255">D–E.</emphasis>
|
||
Right and left views of the organs of the anterior foregut.
|
||
<emphasis id="B593EABF7B21FFADFBF7FAF8A322FA88" bold="true" box="[1090,1108,1365,1388]" pageId="16" pageNumber="255">F.</emphasis>
|
||
Mid-gut gland, ventral view. Abbreviations: cep.t, cephalic tentacles; col.m, columellar muscle; ct, ctenidium; gmg, granulated part of the mid-gut gland; hg, hypobranchial gland; nep, nephridium; nr, nerve ring; od, odontophore with radula; oe, esophagus; os, osphradium; p, penis; pr, proboscis; prp, propodium; r,
|
||
<taxonomicName id="40E74D2E7B21FFADFD56FA18A447FA28" box="[739,817,1461,1484]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" kingdom="Animalia" pageId="16" pageNumber="255" phylum="Mollusca" rank="species" species="rectum">rectum</taxonomicName>
|
||
; s, siphon; sg, salivary gland; tmg, tubular part of mid-gut gland.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="875836AD7B21FFADFF72F907A677F901" blockId="16.[151,1437,1561,2018]" pageId="16" pageNumber="255">
|
||
Measurements (
|
||
<typeStatus id="585C880F7B21FFADFECFF904A695F926" box="[378,483,1705,1730]" pageId="16" pageNumber="255" type="holotype">holotype</typeStatus>
|
||
): SL
|
||
<quantity id="401F9B487B21FFADFD94F907A5FFF925" box="[545,649,1706,1730]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="16" pageNumber="255" unit="mm" value="10.4">10.4 mm</quantity>
|
||
, LWL
|
||
<quantity id="401F9B487B21FFADFD6CF907A444F925" box="[729,818,1706,1730]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="16" pageNumber="255" unit="mm" value="4.8">4.8 mm</quantity>
|
||
, AL (with canal)
|
||
<quantity id="401F9B487B21FFADFC4BF907A32EF925" box="[1022,1112,1706,1730]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="16" pageNumber="255" unit="mm" value="3.8">3.8 mm</quantity>
|
||
, SW
|
||
<quantity id="401F9B487B21FFADFB22F904A384F925" box="[1175,1266,1705,1730]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="16" pageNumber="255" unit="mm" value="2.6">2.6 mm</quantity>
|
||
. Maximum SL
|
||
<quantity id="401F9B487B21FFADFF22F963A78BF901" box="[151,253,1742,1766]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="16" pageNumber="255" unit="mm" value="11.2">11.2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="875836AD7B21FFADFF72F95CA5F7F87E" blockId="16.[151,1437,1561,2018]" pageId="16" pageNumber="255">
|
||
Radula of
|
||
<typeStatus id="585C880F7B21FFADFE89F95CA6E8F8EE" box="[316,414,1777,1802]" pageId="16" pageNumber="255" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="1FDC2A287B21FFADFE1BF95CA686F8EE" box="[430,496,1777,1802]" captionStart="FIGURE 7" captionStartId="15.[151,255,1557,1580]" captionTargetBox="[188,1398,195,1533]" captionTargetId="figure@15.[188,1398,195,1533]" captionTargetPageId="15" captionText="FIGURE 7. Radulae of Recent Hortia species. A – D. Hortia macrocephala n. sp., holotype. A. Entire radular membrane. B. Enlarged lateral teeth. C. Enlarged anteriormost part of radular membrane. D. Enlarged rachidian teeth. E – F. Hortia paradrillia n. sp., holotype. E. Dorsal view of central part of radular membrane. F. Enlarged rachidian and lateral teeth." figureDoi="http://doi.org/10.5281/zenodo.251558" httpUri="https://zenodo.org/record/251558/files/figure.png" pageId="16" pageNumber="255">Fig. 7</figureCitation>
|
||
A–D) short, about 320 Μm long (8.4% of AL), consisting of about 35 rows of teeth, 4–5 nascent, with strong alary processes embracing anterior part of odontophore. Membrane folded along both sides of rachidian. Rachidian teeth with broad base, strongly arched anteriorly, and large cusp, sometimes with indistinct additional serration. Lateral teeth high, triangular, shallowly scoop-shaped. Inner side adjoining rachidian with one to three small additional denticles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFFD65267B21FFADFF72F808A334F859" box="[199,1090,1957,1982]" pageId="16" pageNumber="255" type="distribution">
|
||
<paragraph id="875836AD7B21FFADFF72F808A334F859" blockId="16.[151,1437,1561,2018]" box="[199,1090,1957,1982]" pageId="16" pageNumber="255">
|
||
<emphasis id="B593EABF7B21FFADFF72F808A616F85A" bold="true" box="[199,352,1957,1982]" pageId="16" pageNumber="255">Distribution.</emphasis>
|
||
South of
|
||
<collectingCountry id="FFF0763D7B21FFADFE7BF80BA508F85A" box="[462,638,1957,1982]" name="New Caledonia" pageId="16" pageNumber="255">New Caledonia</collectingCountry>
|
||
, alive in
|
||
<quantity id="401F9B487B21FFADFD56F80BA412F859" box="[739,868,1958,1982]" metricMagnitude="2" metricUnit="m" metricValue="4.41" metricValueMax="4.42" metricValueMin="4.4" pageId="16" pageNumber="255" unit="m" value="441.0" valueMax="442.0" valueMin="440.0">440–442 m</quantity>
|
||
, shells from
|
||
<specimenCount id="91E1FD247B21FFADFC41F80BA334F859" box="[1012,1090,1958,1982]" pageId="16" pageNumber="255" type="generic">
|
||
<quantity id="401F9B487B21FFADFC41F80BA348F859" box="[1012,1086,1958,1982]" metricMagnitude="2" metricUnit="m" metricValue="2.4899999999999998" pageId="16" pageNumber="255" unit="m" value="249.0">249 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFFD65267B21FFADFF72F864A3E3F806" box="[199,1173,1993,2018]" pageId="16" pageNumber="255" type="etymology">
|
||
<paragraph id="875836AD7B21FFADFF72F864A3E3F806" blockId="16.[151,1437,1561,2018]" box="[199,1173,1993,2018]" pageId="16" pageNumber="255">
|
||
<emphasis id="B593EABF7B21FFADFF72F864A638F806" bold="true" box="[199,334,1993,2018]" pageId="16" pageNumber="255">Etymology.</emphasis>
|
||
The name reflects the large, non-planktotrophic protoconch of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFFD65267B20FFACFF72FF3AA630FE6C" pageId="17" pageNumber="256" type="discussion">
|
||
<paragraph id="875836AD7B20FFACFF72FF3AA4DAFEF9" blockId="17.[151,1436,151,392]" pageId="17" pageNumber="256">
|
||
<emphasis id="B593EABF7B20FFACFF72FF3AA64DFF54" bold="true" box="[199,315,151,176]" pageId="17" pageNumber="256">Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="585C880F7B20FFACFEC1FF35A6A9FF54" box="[372,479,152,176]" pageId="17" pageNumber="256" type="paratype">paratypes</typeStatus>
|
||
and other specimens are very similar to the
|
||
<typeStatus id="585C880F7B20FFACFC7BFF3AA346FF54" box="[974,1072,151,176]" pageId="17" pageNumber="256" type="holotype">holotype</typeStatus>
|
||
in shell outline. Specimens may have a columella with slight swellings that look like weak columellar plaits, but are caused by threads of the spiral sculpture that are only partially overlaid by a very weak callus. In some specimens, the subsutural spiral thread is more pronounced and also forms beads at intersections with axial ribs.
|
||
</paragraph>
|
||
<paragraph id="875836AD7B20FFACFF72FE84A630FE6C" blockId="17.[151,1436,151,392]" pageId="17" pageNumber="256">
|
||
<taxonomicName id="40E74D2E7B20FFACFF72FE84A6CEFEA4" box="[199,440,295,320]" class="Magnoliopsida" family="Pyramimitridae" genus="Hortia" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="256" phylum="Mollusca" rank="species" species="macrocephala">
|
||
<emphasis id="B593EABF7B20FFACFF72FE84A6CEFEA4" box="[199,440,295,320]" italics="true" pageId="17" pageNumber="256">Hortia macrocephala</emphasis>
|
||
</taxonomicName>
|
||
is most similar in shell outline to
|
||
<taxonomicName id="40E74D2E7B20FFACFC83FE84A351FEA4" box="[822,1063,295,320]" class="Magnoliopsida" family="Pyramimitridae" genus="Hortia" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="256" phylum="Mollusca" rank="species" species="pseudotaranis" status="sp. nov.">
|
||
<emphasis id="B593EABF7B20FFACFC83FE84A351FEA4" box="[822,1063,295,320]" italics="true" pageId="17" pageNumber="256">Hortia pseudotaranis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="AEA057C47B20FFACFB98FE85A31DFEDB" box="[1069,1131,296,319]" pageId="17" pageNumber="256" rank="species">n. sp.</taxonomicNameLabel>
|
||
, but differs in having more numerous and more closely spaced axial ribs (19 on last whorl in
|
||
<taxonomicName id="40E74D2E7B20FFACFC7BFEE0A3B0FE81" box="[974,1222,332,357]" class="Magnoliopsida" family="Pyramimitridae" genus="Hortia" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="256" phylum="Mollusca" rank="species" species="macrocephala">
|
||
<emphasis id="B593EABF7B20FFACFC7BFEE0A3B0FE81" box="[974,1222,332,357]" italics="true" pageId="17" pageNumber="256">Hortia macrocephala</emphasis>
|
||
</taxonomicName>
|
||
vs.
|
||
<quantity id="401F9B487B20FFACFAB6FEE1A235FE81" box="[1283,1347,332,357]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="17" pageNumber="256" unit="in" value="13.0">13 in</quantity>
|
||
<taxonomicName id="40E74D2E7B20FFACFAE4FEE0A640FE6C" class="Magnoliopsida" family="Pyramimitridae" genus="Hortia" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="256" phylum="Mollusca" rank="species" species="pseudotaranis">
|
||
<emphasis id="B593EABF7B20FFACFAE4FEE0A640FE6C" italics="true" pageId="17" pageNumber="256">Hortia pseudotaranis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |