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<mods:title id="95BAB6285A6423F3EDC7F0D1FC6338EA">Lost and found: The Eocene family Pyramimitridae (Neogastropoda) discovered in the Recent fauna of the Indo-Pacific</mods:title>
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<paragraph id="875836AD7B28FFA4FF22FC5CA69AFBEF" blockId="25.[151,492,1009,1035]" box="[151,492,1009,1035]" pageId="25" pageNumber="264">
<heading id="DC1081C17B28FFA4FF22FC5CA69AFBEF" bold="true" box="[151,492,1009,1035]" fontSize="11" level="1" pageId="25" pageNumber="264" reason="1">
<emphasis id="B593EABF7B28FFA4FF22FC5CA69AFBEF" bold="true" box="[151,492,1009,1035]" pageId="25" pageNumber="264">
Genus
<taxonomicName id="40E74D2E7B28FFA4FF58FC5CA615FBEF" ID-CoL="7PJ7J" box="[237,355,1009,1035]" class="Gastropoda" family="Pyramimitridae" genus="Teremitra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="264" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B593EABF7B28FFA4FF58FC5CA615FBEF" bold="true" box="[237,355,1009,1035]" italics="true" pageId="25" pageNumber="264">Teremitra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="AEA057C47B28FFA4FEDEFC5CA69AFBEF" box="[363,492,1009,1035]" pageId="25" pageNumber="264" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="CFFD65267B28FFA4FF22FB95A514FBAA" box="[151,610,1080,1102]" pageId="25" pageNumber="264" type="materials_examined">
<paragraph id="875836AD7B28FFA4FF22FB95A514FBAA" blockId="25.[151,610,1080,1102]" box="[151,610,1080,1102]" pageId="25" pageNumber="264">
<emphasis id="B593EABF7B28FFA4FF22FB95A654FBAA" bold="true" box="[151,290,1080,1102]" pageId="25" pageNumber="264">
<typeStatus id="585C880F7B28FFA4FF22FB95A7BAFBAA" box="[151,204,1080,1102]" pageId="25" pageNumber="264">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="40E74D2E7B28FFA4FE9CFB95A528FBAA" authority="Aubry, 1999" authorityName="Aubry" authorityYear="1999" box="[297,606,1080,1102]" class="Gastropoda" family="Terebridae" genus="Terebra" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="264" phylum="Mollusca" rank="species" species="efatensis">
<emphasis id="B593EABF7B28FFA4FE9CFB95A6A1FBAA" box="[297,471,1080,1102]" italics="true" pageId="25" pageNumber="264">Terebra efatensis</emphasis>
<bibRefCitation id="E3764B5C7B28FFA4FE68FB95A528FBAA" author="Aubry" box="[477,606,1080,1102]" pageId="25" pageNumber="264" refString="Aubry, U. (1999) New Terebras and Ancient Verses. L'Informatore Piceno, Ancona, 47 pp." type="book" year="1999">Aubry, 1999</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CFFD65267B28FFA4FF22FBD7A21CFAFA" pageId="25" pageNumber="264" type="diagnosis">
<paragraph id="875836AD7B28FFA4FF22FBD7A21CFAFA" blockId="25.[151,1436,1146,1310]" pageId="25" pageNumber="264">
<emphasis id="B593EABF7B28FFA4FF22FBD7A667FB77" bold="true" box="[151,273,1146,1171]" pageId="25" pageNumber="264">Diagnosis.</emphasis>
Shell small, terebriform, narrow elongate, with very high spire and short siphonal canal. Protoconch paucispiral or multispiral up to 3+ whorls. Spiral sculpture of strong spiral cords forming beads at intersection with narrow, closely spaced, axial ribs. Last whorl 0.300.45 of SL. Columella with two weakly pronounced oblique folds. Operculum absent. Radula: rachidian teeth with broad base shallowly arched anteriorly and large cusp, sometimes with indistinct additional serration; high triangular lateral teeth with additional cusps on inner side.
</paragraph>
</subSubSection>
<subSubSection id="CFFD65267B28FFA4FF22FAE4A314FA62" pageId="25" pageNumber="264" type="etymology">
<paragraph id="875836AD7B28FFA4FF22FAE4A5F5FA85" blockId="25.[151,1436,1353,1738]" box="[151,643,1353,1378]" pageId="25" pageNumber="264">
<emphasis id="B593EABF7B28FFA4FF22FAE4A668FA86" bold="true" box="[151,286,1353,1378]" pageId="25" pageNumber="264">Etymology.</emphasis>
Apocope of
<taxonomicName id="40E74D2E7B28FFA4FE1AFAE4A572FA86" box="[431,516,1353,1378]" class="Gastropoda" family="Terebridae" genus="Terebra" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="264" phylum="Mollusca" rank="genus">
<emphasis id="B593EABF7B28FFA4FE1AFAE4A572FA86" box="[431,516,1353,1378]" italics="true" pageId="25" pageNumber="264">Terebra</emphasis>
</taxonomicName>
and
<taxonomicName id="40E74D2E7B28FFA4FD89FAE7A50AFA85" box="[572,636,1354,1377]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="264" phylum="Mollusca" rank="genus">
<emphasis id="B593EABF7B28FFA4FD89FAE7A50AFA85" box="[572,636,1354,1377]" italics="true" pageId="25" pageNumber="264">Mitra</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="875836AD7B28FFA4FF72FAC0A314FA62" blockId="25.[151,1436,1353,1738]" box="[199,1122,1389,1414]" pageId="25" pageNumber="264">
<emphasis id="B593EABF7B28FFA4FF72FAC0A610FA62" bold="true" box="[199,358,1389,1414]" pageId="25" pageNumber="264">Locality/Age.</emphasis>
Upper Eocene of Europe; Recent, West Pacific from
<collectingCountry id="FFF0763D7B28FFA4FC0BFAC0A366FA62" box="[958,1040,1389,1414]" name="Taiwan" pageId="25" pageNumber="264">Taiwan</collectingCountry>
to
<collectingCountry id="FFF0763D7B28FFA4FB86FAC0A32BFA62" box="[1075,1117,1389,1414]" name="Fiji" pageId="25" pageNumber="264">Fiji</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="CFFD65267B28FFA4FF72FA3CA2E0F92D" pageId="25" pageNumber="264" type="discussion">
<paragraph id="875836AD7B28FFA4FF72FA3CA312F942" blockId="25.[151,1436,1353,1738]" pageId="25" pageNumber="264">
<emphasis id="B593EABF7B28FFA4FF72FA3CA64DFA4E" bold="true" box="[199,315,1425,1450]" pageId="25" pageNumber="264">Remarks.</emphasis>
The combination of terebriform shell with a short siphonal canal, presence of columellar plaits and spiral sculpture with strong beads is unique in
<taxonomicName id="40E74D2E7B28FFA4FD2BFA18A426FA2A" box="[670,848,1461,1486]" class="Gastropoda" family="Pyramimitridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="264" phylum="Mollusca" rank="family">Pyramimitridae</taxonomicName>
. The one species already named has previously been included in the
<taxonomicName id="40E74D2E7B28FFA4FEF2FA74A6B7FA16" box="[327,449,1497,1522]" class="Gastropoda" family="Terebridae" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="264" phylum="Mollusca" rank="family">Terebridae</taxonomicName>
, as
<taxonomicName id="40E74D2E7B28FFA4FE59FA74A535FA16" box="[492,579,1497,1522]" class="Gastropoda" family="Terebridae" genus="Terebra" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="264" phylum="Mollusca" rank="genus">
<emphasis id="B593EABF7B28FFA4FE59FA74A535FA16" box="[492,579,1497,1522]" italics="true" pageId="25" pageNumber="264">Terebra</emphasis>
</taxonomicName>
or
<taxonomicName id="40E74D2E7B28FFA4FDDFFA74A58CFA16" box="[618,762,1497,1522]" class="Gastropoda" family="Terebridae" genus="Pristiterebra" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="264" phylum="Mollusca" rank="genus">
<emphasis id="B593EABF7B28FFA4FDDFFA74A58CFA16" box="[618,762,1497,1522]" italics="true" pageId="25" pageNumber="264">Pristiterebra</emphasis>
</taxonomicName>
(
<bibRefCitation id="E3764B5C7B28FFA4FCBCFA74A4E3FA16" author="Aubry" box="[777,917,1497,1522]" pageId="25" pageNumber="264" refString="Aubry, U. (1999) New Terebras and Ancient Verses. L'Informatore Piceno, Ancona, 47 pp." type="book" year="1999">Aubry 1999</bibRefCitation>
;
<bibRefCitation id="E3764B5C7B28FFA4FC17FA77A359FA16" author="Terryn" box="[930,1071,1498,1522]" pageId="25" pageNumber="264" refString="Terryn, Y. (2007) Terebridae. A Collectors Guide. Conchbooks &amp; Natural Arts, 57 pp, 65 plates." type="book" year="2007">Terryn 2007</bibRefCitation>
;
<bibRefCitation id="E3764B5C7B28FFA4FB8EFA77A231FA16" author="Terryn" box="[1083,1351,1497,1522]" pageId="25" pageNumber="264" refString="Terryn, Y. &amp; Holford, M. (2008) The Terebridae of Vanuatu with a revision of the genus Granuliterebra Oyama, 1961. Visaya, Supplement 3, 1 - 118." type="journal article" year="2008">Terryn &amp; Holford 2008</bibRefCitation>
), based on their superficial resemblance in shell shape. The Eocene species of
<emphasis id="B593EABF7B28FFA4FC01FA53A335F9F1" box="[948,1091,1534,1557]" italics="true" pageId="25" pageNumber="264">Pyramimitra</emphasis>
(see
<bibRefCitation id="E3764B5C7B28FFA4FBCAFA50A267F9F2" author="Palmer" box="[1151,1297,1533,1558]" pageId="25" pageNumber="264" refString="Palmer, K. V. W. (1937) The Claibornian Scaphopoda, Gastropoda and Dibranchiate Cephalopoda of the southern United States. Bulletins of American Paleontology, 7, No. 32, Part I, Text, 1 - 548, Part II Plates, plates 1 - 90." type="journal article" year="1937">Palmer 1937</bibRefCitation>
, figs 1215;
<bibRefCitation id="E3764B5C7B28FFA4FF22F98CA64AF9DE" author="Dockery" box="[151,316,1569,1594]" pageId="25" pageNumber="264" refString="Dockery, D. (1977) Mollusca of the Moddys Branch Formation, Mississippi. Mississippi Geological Survey Bulletin, 120, 1 - 212, 28 pls." type="journal article" year="1977">Dockery 1977</bibRefCitation>
; herein
<figureCitation id="1FDC2A287B28FFA4FE2EF98CA69CF9DE" box="[411,490,1569,1594]" captionStart="FIGURE 2" captionStartId="6.[151,255,1693,1716]" captionTargetBox="[182,1406,179,1669]" captionTargetId="figure@6.[182,1406,157,1669]" captionTargetPageId="6" captionText="FIGURE 2. Overview of fossil and Recent Pyramimitridae. A. Endiatoma quadricincta (Cossmann, 1883), MNHN. F. A 47757, Lower Eocene, France. B C. Pyramimitra terebraeformis (Conrad, 1848), MNHN. F. J 08748, Middle Eocene, USA. D E. Pyramimitra quadralirata Dockery, 1977, MNHN. F. A 47750, Middle Eocene, USA. F. Hortia priabonica n. sp., MNHN. F. A 47752, Upper Eocene, France. G. Hortia arriuensis Lozouet, 1999, MNHN. F. A 47751, Upper Oligocene, France. H. Hortia pseudotaranis n. sp., New Caledonia, SL 8.4 mm. I. Hortia macrocephala n. sp., New Caledonia, SL 10.4 mm. J. Hortia paradrillia n. sp., New Caledonia, SL 8.1 mm. K. Hortia marshalli n. sp., New Zealand, SL 10.6 mm. L. Hortia spenceri n. sp., New Zealand, SL 9.0 mm. M. Hortia aotearoa n. sp., New Zealand, SL 9.0 mm. N. Hortia solitaria n. sp., South of Madagascar, SL 6.5 mm. O. Teremitra fallax n. sp., Solomon Islands, SL 11.8 mm. P. Teremitra efatensis (Aubry, 1999), Vanuatu, SL 10.8 mm. Q. Teremitra? hortensis (Vinassa de Regny, 1898), Upper Eocene, France. R. Teremitra sp., Taiwan, SL 16 mm. S. Vaughanites? superstes n. sp., Philippines, SL 19.1 mm." httpUri="https://zenodo.org/record/251553/files/figure.png" pageId="25" pageNumber="264">Figs. 2</figureCitation>
BE, 3AB) have an outer lip that is dentate inside, more pronounced columellar folds, a sculpture of sharp, narrow, spiral cords without beads and obtuse axial varices. The whole surface in
<emphasis id="B593EABF7B28FFA4FF22F9C7A650F965" box="[151,294,1642,1665]" italics="true" pageId="25" pageNumber="264">Pyramimitra</emphasis>
is marked with sharp axial striae. From species of
<taxonomicName id="40E74D2E7B28FFA4FCC2F9C4A376F966" box="[887,1024,1641,1666]" class="Gastropoda" family="Turridae" genus="Vaughanites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="264" phylum="Mollusca" rank="genus">
<emphasis id="B593EABF7B28FFA4FCC2F9C4A376F966" box="[887,1024,1641,1666]" italics="true" pageId="25" pageNumber="264">Vaughanites</emphasis>
</taxonomicName>
, that also possess columellar plaits,
<taxonomicName id="40E74D2E7B28FFA4FF22F923A675F941" box="[151,259,1678,1701]" class="Gastropoda" family="Pyramimitridae" genus="Teremitra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="264" phylum="Mollusca" rank="genus">
<emphasis id="B593EABF7B28FFA4FF22F923A675F941" box="[151,259,1678,1701]" italics="true" pageId="25" pageNumber="264">Teremitra</emphasis>
</taxonomicName>
differs in its coarser sculpture and shorter siphonal canal deflected abaxially.
</paragraph>
<paragraph id="875836AD7B28FFA4FF72F91CA2E0F92D" blockId="25.[151,1436,1353,1738]" box="[199,1430,1713,1738]" pageId="25" pageNumber="264">
<taxonomicName id="40E74D2E7B28FFA4FF72F91CA68AF92E" authority="Vinassa" authorityName="Vinassa" box="[199,508,1713,1738]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="264" phylum="Mollusca" rank="species" species="hortense">
<emphasis id="B593EABF7B28FFA4FF72F91CA6E9F92E" box="[199,415,1713,1738]" italics="true" pageId="25" pageNumber="264">Cerithium hortense</emphasis>
Vinassa
</taxonomicName>
de
<bibRefCitation id="E3764B5C7B28FFA4FD96F91FA5C4F92E" author="Regny" box="[547,690,1714,1738]" pageId="25" pageNumber="264" refString="Vinassa de Regny, P. E. (1898 [&quot; 1897 &quot;]) Synopsis dei molluschi terziari delle Alpi venete (Continuazione e fine della parte prima): IV. Zovencedo (Tufo glauconitico). - V. Monte Pulii, Caldiero, Bolca ecc. - Parte seconda: Strati oligocenici. - VI. Via degli Orti, Valle Orcagna, Priabona ecc. Palaeontographia Italica, 3, 145 - 200, pls 19 - 20 (6 - 7)." type="journal article" year="1898">Regny, 1898</bibRefCitation>
, from the Upper Eocene of Europe, is here transfered to
<taxonomicName id="40E74D2E7B28FFA4FA97F91FA2F8F92D" box="[1314,1422,1714,1737]" class="Gastropoda" family="Pyramimitridae" genus="Teremitra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="25" pageNumber="264" phylum="Mollusca" rank="genus">
<emphasis id="B593EABF7B28FFA4FA97F91FA2F8F92D" box="[1314,1422,1714,1737]" italics="true" pageId="25" pageNumber="264">Teremitra</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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