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<mods:title id="6DBB2A34F4BA0E18F44E21330A344BB4">Taxonomy of Morellia Robineau-Desvoidy (Diptera: Muscidae): revision of the subgenera Morellia s. str. and Parapyrellia Townsend</mods:title>
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<mods:namePart id="74AB33E9A75C9846A89E2C4A3BE95E46">Pamplona, Denise</mods:namePart>
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<mods:namePart id="66948BC1576F63283E1BEC43A218A86F">Nihei, Silvio S.</mods:namePart>
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<mods:namePart id="EB8D4A7BC5F655CABCB8EA615262018F">Couri, Márcia S.</mods:namePart>
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<paragraph id="D744368C4951FF8BA2951B71FB87FDE0" blockId="92.[151,1090,592,652]" box="[151,1090,592,618]" pageId="92" pageNumber="93">
<heading id="8C0C81E04951FF8BA2951B71FB87FDE0" bold="true" box="[151,1090,592,618]" fontSize="11" level="1" pageId="92" pageNumber="93" reason="1">
<emphasis id="E58FEA9E4951FF8BA2951B71FB87FDE0" bold="true" box="[151,1090,592,618]" pageId="92" pageNumber="93">
1.
<taxonomicName id="10FB4D0F4951FF8BA2B61B71FC8DFDE0" ID-CoL="44CYQ" authority="Pamplona &amp; Couri, 1995" authorityName="Pamplona &amp; Couri" authorityYear="1995" box="[180,840,592,618]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="dendropanacis" subGenus="Parapyrellia">
<emphasis id="E58FEA9E4951FF8BA2B61B71FED9FDE0" bold="true" box="[180,284,592,618]" italics="true" pageId="92" pageNumber="93">Morellia</emphasis>
(
<emphasis id="E58FEA9E4951FF8BA32F1B71FEF9FDE0" bold="true" box="[301,316,592,618]" italics="true" pageId="92" pageNumber="93">P</emphasis>
.)
<emphasis id="E58FEA9E4951FF8BA3571B71FDC3FDE0" bold="true" box="[341,518,592,618]" italics="true" pageId="92" pageNumber="93">dendropanacis</emphasis>
<bibRefCitation id="B36A4B7D4951FF8BA00C1B71FC8DFDE0" author="Pamplona" box="[526,840,592,618]" pageId="92" pageNumber="93" refString="Pamplona, D. &amp; Couri, M. S. (1995) Morellia dendropanacis, a new species, and other species with spotted wings: characterization and comparison (Diptera, Muscidae, Muscinae). Proceedings of the Biological Society of Washington, 108, 451 - 457." type="journal article" year="1995">Pamplona &amp; Couri, 1995</bibRefCitation>
</taxonomicName>
, subgen. comb. nov.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9FE165074951FF8BA2951B52FD8DFD06" box="[151,584,627,652]" pageId="92" pageNumber="93" type="description">
<paragraph id="D744368C4951FF8BA2951B52FD8DFD06" blockId="92.[151,1090,592,652]" box="[151,584,627,652]" pageId="92" pageNumber="93">
(figs
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)
</paragraph>
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<paragraph id="D744368C4951FF8BA2951B9BFE10FD65" blockId="92.[151,1437,698,751]" pageId="92" pageNumber="93">
<materialsCitation id="67933CD14951FF8BA2951B9BFE10FD65" ID-GBIF-Occurrence="1306560188" collectionCode="MNRJ" country="Brazil" location="Campinas" pageId="92" pageNumber="93" specimenCount="2" stateProvince="Sao Paulo" typeStatus="holotype">
<treatmentCitationGroup id="F7EB11A24951FF8BA2951B9BFE10FD65" pageId="92" pageNumber="93">
<taxonomicName id="10FB4D0F4951FF8BA2951B9BFD00FD5B" ID-CoL="44CYQ" authority="Pamplona &amp; Couri, 1995: 451" authorityName="Pamplona &amp; Couri" authorityPageNumber="451" authorityYear="1995" box="[151,709,698,721]" class="Insecta" family="Muscidae" genus="Morellia" kingdom="Animalia" order="Diptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="dendropanacis">
<emphasis id="E58FEA9E4951FF8BA2951B9BFE49FD5A" bold="true" box="[151,396,698,720]" italics="true" pageId="92" pageNumber="93">Morellia dendropanacis</emphasis>
<treatmentCitation id="565A109D4951FF8BA3961B9AFD00FD5B" author="Pamplona" box="[404,709,699,721]" page="451" pageId="92" pageNumber="93" year="1995">
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: 451
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</taxonomicName>
.
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locality:
<collectingCountry id="AFEC761C4951FF8BA1621B9AFC78FD5A" box="[864,957,699,720]" name="Brazil" pageId="92" pageNumber="93">BRAZIL</collectingCountry>
,
<collectingRegion id="153FF86E4951FF8BA1C51B9AFBEBFD5B" box="[967,1070,699,721]" country="Brazil" name="Sao Paulo" pageId="92" pageNumber="93">São Paulo</collectingRegion>
,
<location id="D22460574951FF8BA63B1B9AFB5AFD5B" LSID="urn:lsid:plazi:treatment:5F52879A4951FF8AA2951B71FCC9FBD1:D22460574951FF8BA63B1B9AFB5AFD5B" box="[1081,1183,699,721]" country="Brazil" name="Campinas" pageId="92" pageNumber="93" stateProvince="Sao Paulo">Campinas</location>
, Fazenda Santa Genebra.
<typeStatus id="0840882E4951FF8BA2C51BF8FEE6FD65" box="[199,291,729,751]" pageId="92" pageNumber="93" type="holotype">Holotype</typeStatus>
Ƌ,
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(seen).
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</paragraph>
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<subSubSection id="9FE165074951FF8AA2951A3DFC12FC65" lastPageId="93" lastPageNumber="94" pageId="92" pageNumber="93" type="description">
<paragraph id="D744368C4951FF8BA2951A3DFC2EFB87" blockId="92.[151,1437,796,2012]" pageId="92" pageNumber="93">
<emphasis id="E58FEA9E4951FF8BA2951A3DFE81FCBF" bold="true" box="[151,324,796,821]" pageId="92" pageNumber="93">Redescription.</emphasis>
Colour. Black, with blue reflections. Frontal vitta and fronto-orbital plate shining black with grey pruinosity on upper half, face and parafacial black with grey pruinosity, mouth margin yellowish-brown, gena brown; antenna yellowish-brown, basal half of arista yellow; palpus yellow; mesonotum metallic black with a slight blue reflection and with one silver-dusted central vitta in front of the acrostichals; legs brown; haltere white; calypters faintly brownish, with brownish edge, upper calypter on outer half white and opaque; wing with dark markings on humeral crossvein, on the adjacent area of basal radial cell, on subcostal apex connected to a spot on rm, on apex of r1 cell and on cross-vein dm-cu. Abdomen without pruinosity.
</paragraph>
<paragraph id="D744368C4951FF8BA2C51D3AFCA4FBB9" blockId="92.[151,1437,796,2012]" box="[199,865,1051,1076]" pageId="92" pageNumber="93">
<emphasis id="E58FEA9E4951FF8BA2C51D3AFECEFBBE" bold="true" box="[199,267,1051,1076]" pageId="92" pageNumber="93">Male.</emphasis>
Body length: 4.55.0 mm; wing length: 4.55.0 mm.
</paragraph>
<paragraph id="D744368C4951FF8BA2C51D1EFD97FA81" blockId="92.[151,1437,796,2012]" pageId="92" pageNumber="93">Head. Eyes bare, separated at level of anterior ocellus by 0.160.19 of head width; frons divergent towards lunule; ocellar triangle with a pair of setae and a few setulae; 1113 weak pairs of frontal setae; postpedicel 1.82.2 times the length of pedicel; pedicel with 2 setae and a few setulae; arista plumose, with 1114 cilia, bare apicad, and with no secondary plumes; vibrissa shorter than arista; 1 supravibrissal seta, less than half the length of vibrissa, and few setulae that extend to level of middle of postpedicel; 46 subvibrissal setae followed by 810 genal setulae. Palpus expanded at apex.</paragraph>
<paragraph id="D744368C4951FF8AA2C51C36FE81FE01" blockId="92.[151,1437,796,2012]" lastBlockId="93.[151,1437,151,1115]" lastPageId="93" lastPageNumber="94" pageId="92" pageNumber="93">Thorax. Acrostichals 0+1; dorsocentrals 0+2 (weak); intra-alars 1+1; presutural 1; prealar 1, longer than first supra-alar; supra-alars 3; postpronotals 3; notopleuron with 2 long, subequal setae and covered with groundsetulae; postalars 2; no intrapostalar; scutellum with 1 basal, 1 latero-discal and 1 apical pairs of setae, and lateroventrally with many setae, the basalmost ones downcurved; postalar wall and suprasquamal ridge bare; proepisternal depression with short setulae; proepimeron covered with white ground-setulae and several white fine setulae; prosternum broad and bare; anepisternum with a posterior row of about 12 setae, without upcurved setae on upper-anterior corner, and covered with long, white setulae; katepisternals 1+2 and katepisternum with long white setulae; anepimeron extensively setulose with setae short on anterior half and long on posterior half; katepimeron bare with some pile; anatergite bare; katatergite setulose. Legs: fore femur with a posterodorsal row of setae on apical two-thirds, a complete posteroventral row and a posterior row of setae on basal two-thirds; fore tibia with 57 long posteroventral setae and many setulae on apical half, 1 dorsal and 1 posterodorsal preapical setae, and anterior surface with a ctenidium of golden setulae; fore tarsus with some long setae on basal segment; mid femur with a tubercle at apex of anterodorsal to dorsal surfaces bearing about 3 agglutinated short spines that continue as a row on posterior to posteroventral surfaces, an excavation at apex of ventral surface, 4 oblique anterior subapical setae, and no ventral setae; mid tibia with an anterodorsal comb-like ctenidium of very short spinose setulae on basal half and fine setae on apical half, 3 strong setae on posterior surface, and 3 anteroventral, 1 anterodorsal, 1 dorsal, 2 posteroventral and 1 ventral apical setae; hind femur with an anteroventral row of setae, an anterodorsal row of setae on basal two-thirds, about 5 weak posteroventral setae on apical fourth, and 2 dorsal preapical setae; hind tibia with 67 long and weak anteroventral setae on apical third, anterodorsal surface with 2 setae on basal third, 1 on middle third and 1 on apical third, posterior surface with 1 weak seta on middle third, a dorsal row of short but strong setae, posteroventral surface with 78 long setulae on apical third that continue with many setulae as far as the anteroventral setae, with 1 dorsal and 1 anterodorsal preapical setae, 1 anteroventral and 2 weak posteroventral apical setae, and a posterior brush-like ctenidium; hind tarsus with strong setae ventrally. Wing with a uniform covering of microtrichia; dm-cu straight; stem-vein bare before, and with 5 short setulae after, humeral cross-vein on dorsal surface, and ventrally with about 3 short setulae before, and bare after, humeral crossvein; R2+3 bare on dorsal and ventral surfaces; R4+5 setulose from base to beyond r-m on dorsal surface and bare on ventral surface.</paragraph>
<paragraph id="D744368C4950FF8AA2C518B7FC37FE59" blockId="93.[151,1437,151,1115]" pageId="93" pageNumber="94">Abdomen. With a lateral tuft of setulae on syntergite 1+2, tergites 35 with normal setulae; sternite 1 setulose; posterior margin of sternite 5 deeply concave and with setae; basal half bare.</paragraph>
<paragraph id="D744368C4950FF8AA2C518FFFDDAFDE9" blockId="93.[151,1437,151,1115]" pageId="93" pageNumber="94">
Terminalia (figs
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<bibRefCitation id="B36A4B7D4950FF8AA3DB18FFFD32FE7D" author="Pamplona" box="[473,759,478,503]" pageId="93" pageNumber="94" refString="Pamplona, D. &amp; Couri, M. S. (1995) Morellia dendropanacis, a new species, and other species with spotted wings: characterization and comparison (Diptera, Muscidae, Muscinae). Proceedings of the Biological Society of Washington, 108, 451 - 457." type="journal article" year="1995">Pamplona &amp; Couri 1995</bibRefCitation>
). Outer middle third of cercal plate slightly convex; cercal plate curved anteriorly; surstylus with about 4 setulae on apical half; aedeagal apodeme narrowed apically; aedeagus with spicules on anterior membrane of distiphallus; gonopod with a long and strong seta on basal third; paramere with some setae apically.
</paragraph>
<paragraph id="D744368C4950FF8AA2C51B4FFCBEFD0D" blockId="93.[151,1437,151,1115]" box="[199,891,622,647]" pageId="93" pageNumber="94">
<emphasis id="E58FEA9E4950FF8AA2C51B4FFEE0FD0D" bold="true" box="[199,293,622,647]" pageId="93" pageNumber="94">Female.</emphasis>
Body length. 4.55.0 mm; wing length: 4.55.0 mm.
</paragraph>
<paragraph id="D744368C4950FF8AA2C51BB3FC04FCB1" blockId="93.[151,1437,151,1115]" pageId="93" pageNumber="94">Similar to male, differing as follows: lower half of face reddish, frontal plate with white pruinosity beside reddish area, and gena brownish-yellow; eyes separated at level of anterior ocellus by 0.320.38 of head width; fronto-orbital plate with 45 proclinate setae and 2 reclinate setulae; fore tibia without posteroventral setae; mid femur without a tubercle and with only 2 posterodorsal to posterior preapical setae; hind tibia with no posteroventral setae and with only 2 anteroventral setae on middle third.</paragraph>
<paragraph id="D744368C4950FF8AA2C51A67FC12FC65" blockId="93.[151,1437,151,1115]" pageId="93" pageNumber="94">
Ovipositor (figs
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<bibRefCitation id="B36A4B7D4950FF8AA3D81A67FD34FCD5" author="Pamplona" box="[474,753,838,863]" pageId="93" pageNumber="94" refString="Pamplona, D. &amp; Couri, M. S. (1995) Morellia dendropanacis, a new species, and other species with spotted wings: characterization and comparison (Diptera, Muscidae, Muscinae). Proceedings of the Biological Society of Washington, 108, 451 - 457." type="journal article" year="1995">Pamplona &amp; Couri 1995</bibRefCitation>
). Tergite 6 shaped as 2 golf-club-like rods, posteriorly with 6 setulae; tergite7 as 2 rods, anteriorly enlarged, posteriorly with 2 plates with spines; tergite 8 as 2 rods, posteriorly with 2 plates with strong spines; sternite 6 with 6 setulae; sternite 7 as 3 plates with setae; sternite 8 as 2 plates with setae; enlarged spicules on intersegmental membrane between segments 7 and 8; epiproct with many setae; hypoproct longer than broad with many setae; cercus cylindrical and long.
</paragraph>
</subSubSection>
<subSubSection id="9FE165074950FF8AA2C51ADBFD4FFBBD" pageId="93" pageNumber="94" type="materials_examined">
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<materialsCitation id="67933CD14950FF8AA2C51ADBFD43FBBD" ID-GBIF-Occurrence="1306560176" collectingDate="1992-07-03" collectionCode="MNRJ" collectorName="E. Pombal" country="Brazil" county="Material" location="Fazenda Santa Genebra" municipality="Campinas" pageId="93" pageNumber="94" specimenCount="4" specimenCount-female="2" stateProvince="Sao Paulo" typeStatus="holotype">
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<collectingCounty id="3E254E004950FF8AA2C51ADBFEEAFB99" box="[199,303,1018,1043]" pageId="93" pageNumber="94">Material</collectingCounty>
examined.
</emphasis>
<collectingCountry id="AFEC761C4950FF8AA3B51ADAFDE4FB99" box="[439,545,1019,1043]" name="Brazil" pageId="93" pageNumber="94">BRAZIL</collectingCountry>
:
<collectingRegion id="153FF86E4950FF8AA0291ADAFD5AFB99" box="[555,671,1018,1043]" country="Brazil" name="Sao Paulo" pageId="93" pageNumber="94">São Paulo</collectingRegion>
,
<collectingMunicipality id="3720ACF64950FF8AA0AB1ADBFCD9FB99" box="[681,796,1018,1043]" pageId="93" pageNumber="94">Campinas</collectingMunicipality>
,
<location id="D22460574950FF8AA1251ADBFBE8FB99" LSID="urn:lsid:plazi:treatment:5F52879A4951FF8AA2951B71FCC9FBD1:D22460574950FF8AA1251ADBFBE8FB99" box="[807,1069,1018,1043]" country="Brazil" county="Material" municipality="Campinas" name="Fazenda Santa Genebra" pageId="93" pageNumber="94" stateProvince="Sao Paulo">Fazenda Santa Genebra</location>
,
<date id="A345104C4950FF8AA63A1ADBFB69FB99" box="[1080,1196,1018,1043]" pageId="93" pageNumber="94" value="1992-07-03">
<collectingDate id="B301E9A44950FF8AA63A1ADBFB69FB99" box="[1080,1196,1018,1043]" pageId="93" pageNumber="94" value="1992-07-03">3.vii.1992</collectingDate>
</date>
,
<collectorName id="7A0E535A4950FF8AA6B51ADAFAEBFB99" box="[1207,1326,1018,1043]" pageId="93" pageNumber="94">E. Pombal</collectorName>
,
<typeStatus id="0840882E4950FF8AA7381ADBFA59FB99" box="[1338,1436,1018,1043]" pageId="93" pageNumber="94" type="holotype">holotype</typeStatus>
Ƌ, 1 Ƌ
<typeStatus id="0840882E4950FF8AA2E91D01FE8EFBB2" box="[235,331,1056,1080]" pageId="93" pageNumber="94" type="paratype">paratype</typeStatus>
and
<specimenCount id="C1FDFD054950FF8AA3801D3EFE69FBBD" box="[386,428,1055,1079]" pageId="93" pageNumber="94" type="female">2 ♀</specimenCount>
<typeStatus id="0840882E4950FF8AA3B11D01FDDBFBB2" box="[435,542,1056,1080]" pageId="93" pageNumber="94" type="paratype">paratypes</typeStatus>
(
<collectionCode id="B1EAAE494950FF8AA02F1D3EFDBBFBBD" box="[557,638,1055,1079]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="93" pageNumber="94">MNRJ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9FE165074950FF8AA2C51D63FCC9FBD1" box="[199,780,1090,1115]" pageId="93" pageNumber="94" type="distribution">
<paragraph id="D744368C4950FF8AA2C51D63FCC9FBD1" blockId="93.[151,1437,151,1115]" box="[199,780,1090,1115]" pageId="93" pageNumber="94">
<emphasis id="E58FEA9E4950FF8AA2C51D63FEA5FBD1" bold="true" box="[199,352,1090,1115]" pageId="93" pageNumber="94">Distribution.</emphasis>
NEOTROPICAL:
<collectingCountry id="AFEC761C4950FF8AA03A1D63FDB9FBD1" box="[568,636,1090,1115]" name="Brazil" pageId="93" pageNumber="94">Brazil</collectingCountry>
(
<collectingRegion id="153FF86E4950FF8AA0891D62FCC5FBD1" box="[651,768,1090,1115]" country="Brazil" name="Sao Paulo" pageId="93" pageNumber="94">São Paulo</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>