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<document ID-DOI="http://doi.org/10.5281/zenodo.5621361" ID-GBIF-Dataset="362dc26a-0302-4b89-98c2-a84515dc5b40" ID-GBIF-Taxon="126094925" ID-Zenodo-Dep="5621361" checkinTime="1483706356665" checkinUser="plazi" docAuthor="A. Ionescu &amp; P. - A. Eyer" docDate="2016" docId="5F2F5A00FF8AFFE7FE0B44A96F79C013" docLanguage="en" docName="Ionescu_Eyer_2016_IJE_CataglyphisIsrael.pdf" docOrigin="Israel Journal Of Entomology 46" docStyle="DocumentStyle{}" docTitle="Cataglyphis israelensis Ionescu &amp; Eyer, 2016, n. sp." docType="treatment" docVersion="6" lastPageId="11" lastPageNumber="119" masterDocId="A3162278FF8FFFECFFB1417B6E26C415" masterDocTitle="Notes on Cataglyphis Foerster, 1850 of the bicolor species-group in Israel, with description of a new species (Hymenoptera: Formicidae)" masterLastPageNumber="131" masterPageNumber="109" pageId="5" pageNumber="114" updateTime="1635548064805" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Notes on Cataglyphis Foerster, 1850 of the bicolor species-group in Israel, with description of a new species (Hymenoptera: Formicidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>A. Ionescu</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>P. - A. Eyer</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Israel Journal Of Entomology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>46</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>109</mods:start>
<mods:end>131</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.221456</mods:identifier>
<mods:identifier type="GBIF-Dataset">362dc26a-0302-4b89-98c2-a84515dc5b40</mods:identifier>
<mods:identifier type="Zenodo-Dep">221456</mods:identifier>
<mods:identifier type="ZooBank">0FE0E11B-3411-4204-A63D-DB23DDE5A379</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5621361" ID-GBIF-Taxon="126094925" ID-Zenodo-Dep="5621361" LSID="urn:lsid:plazi:treatment:5F2F5A00FF8AFFE7FE0B44A96F79C013" httpUri="http://treatment.plazi.org/id/5F2F5A00FF8AFFE7FE0B44A96F79C013" lastPageId="11" lastPageNumber="119" pageId="5" pageNumber="114">
<subSubSection box="[442,775,1490,1517]" pageId="5" pageNumber="114" type="nomenclature">
<paragraph blockId="5.[151,1066,1459,1597]" box="[442,775,1490,1517]" pageId="5" pageNumber="114">
<heading box="[442,775,1490,1517]" centered="true" fontSize="11" level="3" pageId="5" pageNumber="114" reason="1">
<taxonomicName box="[442,707,1490,1517]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="114" phylum="Arthropoda" rank="species" species="israelensis" status="n. sp.">
<emphasis bold="true" box="[442,707,1490,1517]" italics="true" pageId="5" pageNumber="114">Cataglyphis israelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[715,775,1491,1517]" pageId="5" pageNumber="114" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[485,732,1530,1557]" pageId="5" pageNumber="114" type="description">
<paragraph blockId="5.[151,1066,1459,1597]" box="[485,732,1530,1557]" pageId="5" pageNumber="114">
<heading box="[485,732,1530,1557]" centered="true" fontSize="11" level="3" pageId="5" pageNumber="114" reason="1">
(
<figureCitation box="[494,565,1531,1557]" captionStart="Figs 2 6" captionStartId="4.[144,183,1406,1428]" captionTargetBox="[155,1031,406,1383]" captionTargetId="figure@4.[147,1054,406,1386]" captionTargetPageId="4" captionText="Figs 2 6. Measurements for ant descriptions: (2) head of large worker in full-face view; (3) Frst (F 1) and second (F 2) funiculus segments of C. isis worker; (4) C. israelensis large worker, mesosoma outline in lateral view, pilosity omitted; (5) petiole node outline of large worker in lateral view, pilosity omitted; (6) male subgenital plate outline in ventral view. Abbreviations: iOD inter-Ocular Distance, ML Mesosoma Length, OcClD Ocellus-Clypeal Distance, PrH Propodeum Height, PrL Propodeum Length, PNA Petiole Node Angle, PNH Petiole Node Height, PNL Petiole Node Length, lSAL lateral Subgenital Plate Appendage Length, mSAL median Subgenital Plate Appendage Length." pageId="5" pageNumber="114">Figs 4</figureCitation>
,
<figureCitation box="[577,637,1531,1557]" captionStart-0="Figs 7, 8" captionStart-1="Figs 9 11" captionStart-2="Figs 12, 13" captionStartId-0="6.[168,207,1084,1106]" captionStartId-1="7.[151,190,594,616]" captionStartId-2="8.[144,183,1128,1150]" captionTargetBox-1="[161,1053,234,574]" captionTargetBox-2="[185,1017,235,1107]" captionTargetId-1="figure@7.[161,1053,234,574]" captionTargetId-2="figure@8.[185,1017,235,723]" captionText-0="Figs 7, 8. C. israelensis holotype worker in lateral (7) and dorsal (8) views. Scale bars, 1.5 mm." captionText-1="Figs 9 11. C. israelensis worker, head in full-face view: (9) holotype, (10) large paratype, (11) small paratype. Scale bars, 1.5 mm." captionText-2="Figs 12, 13. C. israelensis, propodeum and petiole node in lateral view: (12) small paratype worker," pageId="5" pageNumber="114">713</figureCitation>
, 2325)
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,1066,1570,1597]" pageId="5" pageNumber="114" type="nomenclature">
<paragraph blockId="5.[151,1066,1459,1597]" box="[151,1066,1570,1597]" pageId="5" pageNumber="114">
<emphasis bold="true" box="[151,226,1570,1597]" pageId="5" pageNumber="114">LSID:</emphasis>
urn:lsid:zoobank.org:act:53502C30-9AA0-4A01-AAC9-C630CFEE6D19.
</paragraph>
</subSubSection>
<caption box="[168,1035,1084,1106]" pageId="6" pageNumber="115">
<paragraph blockId="6.[168,1035,1084,1106]" box="[168,1035,1084,1106]" pageId="6" pageNumber="115">
Figs 7, 8.
<taxonomicName box="[258,381,1084,1106]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="115" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[258,381,1084,1106]" italics="true" pageId="6" pageNumber="115">C. israelensis</emphasis>
</taxonomicName>
holotype worker in lateral (7) and dorsal (8) views. Scale bars, 1.5 mm.
</paragraph>
</caption>
<subSubSection box="[144,983,1175,1201]" pageId="6" pageNumber="115" type="etymology">
<paragraph blockId="6.[144,1059,1175,1597]" box="[144,983,1175,1201]" pageId="6" pageNumber="115">
<emphasis bold="true" box="[144,282,1175,1201]" pageId="6" pageNumber="115">Etymology:</emphasis>
This species is named after Israel, its known distribution area.
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="118" pageId="6" pageNumber="115" type="description">
<paragraph blockId="6.[144,1059,1175,1597]" pageId="6" pageNumber="115">
<emphasis bold="true" box="[144,400,1214,1241]" pageId="6" pageNumber="115">Description: Worker:</emphasis>
<emphasis box="[406,467,1215,1241]" italics="true" pageId="6" pageNumber="115">Head</emphasis>
. Head of large worker subquadrate with emarginated occipital margin, and slightly convex lateral margins converging apically (
<figureCitation box="[962,1044,1247,1274]" captionStart="Figs 9 11" captionStartId="7.[151,190,594,616]" captionTargetBox="[161,1053,234,574]" captionTargetId="figure@7.[161,1053,234,574]" captionText="Figs 9 11. C. israelensis worker, head in full-face view: (9) holotype, (10) large paratype, (11) small paratype. Scale bars, 1.5 mm." pageId="6" pageNumber="115">Fig. 10</figureCitation>
); head of minor worker (
<figureCitation box="[403,482,1280,1306]" captionStart="Figs 9 11" captionStartId="7.[151,190,594,616]" captionTargetBox="[161,1053,234,574]" captionTargetId="figure@7.[161,1053,234,574]" captionText="Figs 9 11. C. israelensis worker, head in full-face view: (9) holotype, (10) large paratype, (11) small paratype. Scale bars, 1.5 mm." pageId="6" pageNumber="115">Fig. 11</figureCitation>
) distinctly elongate, with rounded occipital margin posterior to eye and parallel lateral margins, HW/HL=0.860.99; head width shows accelerating growth rate with head length; head width with ocellus-clypeal distance, and interocular distance with head length show isometrical growth; scape long (SL/HW=0.981.37), shows decelerating growth rate with head width; scape length variable among localities, shortest in ants from northern Coastal Plain, longest in ants from Judean Desert (
<tableCitation box="[546,633,1473,1499]" captionStart="Table 2" captionStartId="18.[144,197,236,257]" captionTargetBox="[144,1058,337,1237]" captionTargetPageId="18" captionText="Table 2: Least square means ± 95 % conFdence interval at covariate mean, and sample size for four variables per sampling localities grouped by taxon and geographic region; see Materials and Methods for full names of variables and Fig. 1 for numbers and names of geographic regions." httpUri="http://table.plazi.org/id/83F9BB9EFF9DFFFEFF2141976DAAC524" pageId="6" pageNumber="115" tableUuid="83F9BB9EFF9DFFFEFF2141976DAAC524">Table 2</tableCitation>
); F1L/F2L=1.642.02.
</paragraph>
<paragraph blockId="6.[144,1059,1175,1597]" lastBlockId="7.[151,1066,679,1603]" lastPageId="7" lastPageNumber="116" pageId="6" pageNumber="115">
<emphasis box="[170,290,1506,1531]" italics="true" pageId="6" pageNumber="115">Mesosoma</emphasis>
. Mesonotum in lateral view distinctly raised over pronotum (
<figureCitation captionStart="Figs 2 6" captionStartId="4.[144,183,1406,1428]" captionTargetBox="[155,1031,406,1383]" captionTargetId="figure@4.[147,1054,406,1386]" captionTargetPageId="4" captionText="Figs 2 6. Measurements for ant descriptions: (2) head of large worker in full-face view; (3) Frst (F 1) and second (F 2) funiculus segments of C. isis worker; (4) C. israelensis large worker, mesosoma outline in lateral view, pilosity omitted; (5) petiole node outline of large worker in lateral view, pilosity omitted; (6) male subgenital plate outline in ventral view. Abbreviations: iOD inter-Ocular Distance, ML Mesosoma Length, OcClD Ocellus-Clypeal Distance, PrH Propodeum Height, PrL Propodeum Length, PNA Petiole Node Angle, PNH Petiole Node Height, PNL Petiole Node Length, lSAL lateral Subgenital Plate Appendage Length, mSAL median Subgenital Plate Appendage Length." pageId="6" pageNumber="115">Figs 4</figureCitation>
,
<figureCitation box="[174,189,1538,1564]" captionStart="Figs 7, 8" captionStartId="6.[168,207,1084,1106]" captionText="Figs 7, 8. C. israelensis holotype worker in lateral (7) and dorsal (8) views. Scale bars, 1.5 mm." pageId="6" pageNumber="115">7</figureCitation>
) in 96 of 108 specimens; outline of propodeum dorsum smoothly arched in lateral view (
<figureCitation box="[293,368,1570,1596]" captionStart="Figs 2 6" captionStartId="4.[144,183,1406,1428]" captionTargetBox="[155,1031,406,1383]" captionTargetId="figure@4.[147,1054,406,1386]" captionTargetPageId="4" captionText="Figs 2 6. Measurements for ant descriptions: (2) head of large worker in full-face view; (3) Frst (F 1) and second (F 2) funiculus segments of C. isis worker; (4) C. israelensis large worker, mesosoma outline in lateral view, pilosity omitted; (5) petiole node outline of large worker in lateral view, pilosity omitted; (6) male subgenital plate outline in ventral view. Abbreviations: iOD inter-Ocular Distance, ML Mesosoma Length, OcClD Ocellus-Clypeal Distance, PrH Propodeum Height, PrL Propodeum Length, PNA Petiole Node Angle, PNH Petiole Node Height, PNL Petiole Node Length, lSAL lateral Subgenital Plate Appendage Length, mSAL median Subgenital Plate Appendage Length." pageId="6" pageNumber="115">Figs 4</figureCitation>
,
<figureCitation box="[382,398,1571,1597]" captionStart="Figs 7, 8" captionStartId="6.[168,207,1084,1106]" captionText="Figs 7, 8. C. israelensis holotype worker in lateral (7) and dorsal (8) views. Scale bars, 1.5 mm." pageId="6" pageNumber="115">7</figureCitation>
,
<figureCitation box="[411,485,1570,1596]" captionStart="Figs 12, 13" captionStartId="8.[144,183,1128,1150]" captionTargetBox="[185,1017,235,1107]" captionTargetId="figure@8.[185,1017,235,723]" captionText="Figs 12, 13. C. israelensis, propodeum and petiole node in lateral view: (12) small paratype worker," pageId="6" pageNumber="115">12, 13</figureCitation>
); propodeum height variable, highest in ants from northern Coastal Plain (
<figureCitation box="[429,516,679,705]" captionStart="Figs 12, 13" captionStartId="8.[144,183,1128,1150]" captionTargetBox="[185,1017,235,1107]" captionTargetId="figure@8.[185,1017,235,723]" captionText="Figs 12, 13. C. israelensis, propodeum and petiole node in lateral view: (12) small paratype worker," pageId="7" pageNumber="116">Fig. 13</figureCitation>
), lowest in ants from Judean Desert (
<tableCitation box="[960,1051,679,705]" captionStart="Table 2" captionStartId="18.[144,197,236,257]" captionTargetBox="[144,1058,337,1237]" captionTargetPageId="18" captionText="Table 2: Least square means ± 95 % conFdence interval at covariate mean, and sample size for four variables per sampling localities grouped by taxon and geographic region; see Materials and Methods for full names of variables and Fig. 1 for numbers and names of geographic regions." httpUri="http://table.plazi.org/id/83F9BB9EFF9DFFFEFF2141976DAAC524" pageId="7" pageNumber="116" tableUuid="83F9BB9EFF9DFFFEFF2141976DAAC524">Table 2</tableCitation>
); petiole node of largest workers high, dorsally slightly rounded (
<figureCitation box="[856,928,711,738]" captionStart="Figs 7, 8" captionStartId="6.[168,207,1084,1106]" captionText="Figs 7, 8. C. israelensis holotype worker in lateral (7) and dorsal (8) views. Scale bars, 1.5 mm." pageId="7" pageNumber="116">Figs 7</figureCitation>
,
<figureCitation box="[939,969,711,737]" captionStart="Figs 12, 13" captionStartId="8.[144,183,1128,1150]" captionTargetBox="[185,1017,235,1107]" captionTargetId="figure@8.[185,1017,235,723]" captionText="Figs 12, 13. C. israelensis, propodeum and petiole node in lateral view: (12) small paratype worker," pageId="7" pageNumber="116">13</figureCitation>
), petiole node of medium-small workers lower, more elongate (
<figureCitation box="[758,841,743,769]" captionStart="Figs 12, 13" captionStartId="8.[144,183,1128,1150]" captionTargetBox="[185,1017,235,1107]" captionTargetId="figure@8.[185,1017,235,723]" captionText="Figs 12, 13. C. israelensis, propodeum and petiole node in lateral view: (12) small paratype worker," pageId="7" pageNumber="116">Fig. 12</figureCitation>
); petiole node angle variable, largest in ants from northern Coastal Plain (
<tableCitation box="[757,846,775,801]" captionStart="Table 2" captionStartId="18.[144,197,236,257]" captionTargetBox="[144,1058,337,1237]" captionTargetPageId="18" captionText="Table 2: Least square means ± 95 % conFdence interval at covariate mean, and sample size for four variables per sampling localities grouped by taxon and geographic region; see Materials and Methods for full names of variables and Fig. 1 for numbers and names of geographic regions." httpUri="http://table.plazi.org/id/83F9BB9EFF9DFFFEFF2141976DAAC524" pageId="7" pageNumber="116" tableUuid="83F9BB9EFF9DFFFEFF2141976DAAC524">Table 2</tableCitation>
;
<figureCitation box="[860,943,775,801]" captionStart="Figs 12, 13" captionStartId="8.[144,183,1128,1150]" captionTargetBox="[185,1017,235,1107]" captionTargetId="figure@8.[185,1017,235,723]" captionText="Figs 12, 13. C. israelensis, propodeum and petiole node in lateral view: (12) small paratype worker," pageId="7" pageNumber="116">Fig. 13</figureCitation>
); hindtibia 1.21.4 times longer than midfemur.
</paragraph>
<caption pageId="7" pageNumber="116" targetBox="[161,1053,234,574]" targetPageId="7">
<paragraph blockId="7.[151,1065,594,640]" pageId="7" pageNumber="116">
Figs 911.
<taxonomicName box="[253,377,594,616]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="116" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[253,377,594,616]" italics="true" pageId="7" pageNumber="116">C. israelensis</emphasis>
</taxonomicName>
worker, head in full-face view: (9) holotype, (10) large paratype, (11) small paratype. Scale bars, 1.5 mm.
</paragraph>
</caption>
<paragraph blockId="7.[151,1066,679,1603]" pageId="7" pageNumber="116">
<emphasis box="[178,284,839,865]" italics="true" pageId="7" pageNumber="116">Sculpture</emphasis>
. Entire body with piligerous pits; head, mesosoma and gaster dorsally and laterally reticulate/shagreened, matte; seashore populations with more accentuated sculpture, but part of specimens with laterally shiny gaster; mandible, leg and ventral surface of head and gaster Fnely striate/carinulate, shiny.
</paragraph>
<paragraph blockId="7.[151,1066,679,1603]" pageId="7" pageNumber="116">
<emphasis box="[178,264,967,993]" italics="true" pageId="7" pageNumber="116">Pilosity</emphasis>
. Pilosity white to yellowish brown: clypeus, frons, vertex, mesosoma, petiole node dorsum and gaster segments with sparse, erect, hairs; dorsum of promesonotum with 114 hairs, number not correlated with size; propodeum dorsum, from metanotal groove to propodeal spiracle, with 14 hairs (in 37/109 specimens); ventral surface of head, mandible, coxa and petiole node with erect to decumbent setae, on latter occasionally lacking; hind tibia with two rows of decumbent bristles on ventral surface; body covered with very short, appressed, white pubescence, more abundant on head and propodeum; convex face of scape with dense, appressed to subappressed pubescence on distal ¼–½ of length.
</paragraph>
<paragraph blockId="7.[151,1066,679,1603]" pageId="7" pageNumber="116">
<emphasis box="[178,243,1255,1281]" italics="true" pageId="7" pageNumber="116">Color</emphasis>
. Highly variable: head, mesosoma and petiole bright red to ochraceousorange, leg brownish to black, gaster black (
<figureCitation box="[642,713,1287,1314]" captionStart="Figs 7, 8" captionStartId="6.[168,207,1084,1106]" captionText="Figs 7, 8. C. israelensis holotype worker in lateral (7) and dorsal (8) views. Scale bars, 1.5 mm." pageId="7" pageNumber="116">Figs 7</figureCitation>
,
<figureCitation box="[724,753,1287,1313]" captionStart="Figs 12, 13" captionStartId="8.[144,183,1128,1150]" captionTargetBox="[185,1017,235,1107]" captionTargetId="figure@8.[185,1017,235,723]" captionText="Figs 12, 13. C. israelensis, propodeum and petiole node in lateral view: (12) small paratype worker," pageId="7" pageNumber="116">12</figureCitation>
) in populations from Lower Galilee, Yizreel Valley, Sea of Galilee area and Upper Jordan Valley; head, mesosoma and petiole dark red to dark reddish brown, leg concolorous, gaster black in populations from Upper Galilee, Lower Hermon, Golan Heights and from Carmel Ridge to Judean Hills; and uniformly black in populations from northern Coastal Plain (Atlit, Hof haBonim) and Judean Desert (Allon, Maale Adummim); in bicolored nest series medium workers (MW=1.01.5) always lighter colored than larger specimens, smallest workers black.
</paragraph>
<paragraph blockId="7.[151,1066,679,1603]" lastBlockId="8.[144,1059,1224,1603]" lastPageId="8" lastPageNumber="117" pageId="7" pageNumber="116">
<emphasis box="[178,340,1545,1570]" italics="true" pageId="7" pageNumber="116">Measurements</emphasis>
: TL=6.413.3, HL=1.483.20, HW=1.373.07, EL=0.490.86, iOD=1.052.30, OcClD=0.801.43, SL=1.733.26, F1L=0.430.85, F2L=0.24 0.49, ML=2.594.45, MW=0.942.01, PrL=0.891.54, PrH=0.190.45, PNL= 0.490.84, PNW=0.290.63, PNH=0.280.53, PNA=6584°, mFmL=1.953.91, hTbL=2.544.94 (n=110).
</paragraph>
<caption pageId="8" pageNumber="117" targetBox="[185,1017,235,1107]" targetPageId="8">
<paragraph blockId="8.[144,1048,1128,1174]" pageId="8" pageNumber="117">
Figs 12, 13.
<taxonomicName box="[256,380,1128,1150]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="117" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[256,380,1128,1150]" italics="true" pageId="8" pageNumber="117">C. israelensis</emphasis>
</taxonomicName>
, propodeum and petiole node in lateral view: (12) small paratype worker, (13) worker from Hof haBonim. Scale bars, 1.5 mm.
</paragraph>
</caption>
<paragraph blockId="8.[144,1059,1224,1603]" lastBlockId="9.[151,1067,236,1603]" lastPageId="9" lastPageNumber="118" pageId="8" pageNumber="117">
<emphasis bold="true" box="[144,409,1320,1347]" pageId="8" pageNumber="117">Differential diagnosis:</emphasis>
<taxonomicName box="[421,576,1321,1347]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="117" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[421,576,1321,1347]" italics="true" pageId="8" pageNumber="117">C. israelensis</emphasis>
</taxonomicName>
is a large species of the
<taxonomicName box="[878,958,1321,1347]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="117" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis box="[878,958,1321,1347]" italics="true" pageId="8" pageNumber="117">bicolor</emphasis>
</taxonomicName>
speciesgroup with habitus and color pattern similar to
<emphasis box="[715,778,1353,1379]" italics="true" pageId="8" pageNumber="117">drusa</emphasis>
and
<emphasis box="[858,938,1353,1378]" italics="true" pageId="8" pageNumber="117">assyria</emphasis>
(AntWeb CASENT0912219 and CASENT0912218). It has symmetrically and smoothly arched propodeum outline in lateral view (
<figureCitation box="[633,707,1417,1443]" captionStart="Figs 7, 8" captionStartId="6.[168,207,1084,1106]" captionText="Figs 7, 8. C. israelensis holotype worker in lateral (7) and dorsal (8) views. Scale bars, 1.5 mm." pageId="8" pageNumber="117">Figs 7</figureCitation>
,
<figureCitation box="[722,751,1417,1443]" captionStart="Figs 12, 13" captionStartId="8.[144,183,1128,1150]" captionTargetBox="[185,1017,235,1107]" captionTargetId="figure@8.[185,1017,235,723]" captionText="Figs 12, 13. C. israelensis, propodeum and petiole node in lateral view: (12) small paratype worker," pageId="8" pageNumber="117">13</figureCitation>
), similar to
<emphasis box="[903,966,1417,1443]" italics="true" pageId="8" pageNumber="117">drusa</emphasis>
and
<taxonomicName class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="117" phylum="Arthropoda" rank="species" species="nodus">
<emphasis italics="true" pageId="8" pageNumber="117">
C.
<taxonomicName box="[144,210,1449,1475]" pageId="8" pageNumber="117" rank="species" species="nodus">nodus</taxonomicName>
</emphasis>
</taxonomicName>
(AntWeb CASENT0911115 and CASENT0911116), differing from higher and steeply rounded outline in
<taxonomicName box="[493,585,1481,1507]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="117" phylum="Arthropoda" rank="species" species="niger">
<emphasis box="[493,585,1481,1507]" italics="true" pageId="8" pageNumber="117">C. niger</emphasis>
</taxonomicName>
(Fig. 16),
<taxonomicName box="[707,869,1481,1507]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="117" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[707,726,1481,1507]" italics="true" pageId="8" pageNumber="117">C</emphasis>
. cf.
<emphasis box="[777,869,1481,1506]" italics="true" pageId="8" pageNumber="117">savignyi</emphasis>
</taxonomicName>
(Fig. 17) and
<taxonomicName class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="117" phylum="Arthropoda" rank="species" species="holgerseni">
<emphasis italics="true" pageId="8" pageNumber="117">C. holgerseni</emphasis>
</taxonomicName>
(Fig. 14), but higher and less Fattened than in
<taxonomicName box="[781,850,1513,1539]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="117" phylum="Arthropoda" rank="species" species="isis">
<emphasis box="[781,850,1513,1539]" italics="true" pageId="8" pageNumber="117">C. isis</emphasis>
</taxonomicName>
(Fig. 15;
<tableCitation box="[959,1041,1513,1539]" captionStart="Table 1" captionStartId="16.[144,197,236,257]" captionTargetBox="[144,1058,310,1593]" captionTargetPageId="16" captionText="Table 1: Least square means ± 95 % conFdence interval at covariate mean, and sample size for 17 va-" httpUri="http://table.plazi.org/id/83F9BB9EFF9FFFFCFF2141976A04C514" pageId="8" pageNumber="117" tableUuid="83F9BB9EFF9FFFFCFF2141976A04C514">Table 1</tableCitation>
); populations from Atlit and Hof haBonim have high propodeum (
<figureCitation box="[878,959,1545,1571]" captionStart="Figs 12, 13" captionStartId="8.[144,183,1128,1150]" captionTargetBox="[185,1017,235,1107]" captionTargetId="figure@8.[185,1017,235,723]" captionText="Figs 12, 13. C. israelensis, propodeum and petiole node in lateral view: (12) small paratype worker," pageId="8" pageNumber="117">Fig. 13</figureCitation>
), similar to neighboring
<taxonomicName box="[314,474,1577,1603]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="117" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[314,333,1577,1603]" italics="true" pageId="8" pageNumber="117">C</emphasis>
. cf.
<emphasis box="[382,474,1577,1602]" italics="true" pageId="8" pageNumber="117">savignyi</emphasis>
</taxonomicName>
populations from northern and central Coastal Plain (
<tableCitation box="[160,246,236,262]" captionStart="Table 2" captionStartId="18.[144,197,236,257]" captionTargetBox="[144,1058,337,1237]" captionTargetPageId="18" captionText="Table 2: Least square means ± 95 % conFdence interval at covariate mean, and sample size for four variables per sampling localities grouped by taxon and geographic region; see Materials and Methods for full names of variables and Fig. 1 for numbers and names of geographic regions." httpUri="http://table.plazi.org/id/83F9BB9EFF9DFFFEFF2141976DAAC524" pageId="9" pageNumber="118" tableUuid="83F9BB9EFF9DFFFEFF2141976DAAC524">Table 2</tableCitation>
).
<taxonomicName box="[268,420,236,263]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[268,420,236,263]" italics="true" pageId="9" pageNumber="118">C. israelensis</emphasis>
</taxonomicName>
has midfemur length similar to
<taxonomicName box="[785,877,237,263]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="niger">
<emphasis box="[785,877,237,263]" italics="true" pageId="9" pageNumber="118">C. niger</emphasis>
</taxonomicName>
(
<tableCitation box="[895,976,236,262]" captionStart="Table 1" captionStartId="16.[144,197,236,257]" captionTargetBox="[144,1058,310,1593]" captionTargetPageId="16" captionText="Table 1: Least square means ± 95 % conFdence interval at covariate mean, and sample size for 17 va-" httpUri="http://table.plazi.org/id/83F9BB9EFF9FFFFCFF2141976A04C514" pageId="9" pageNumber="118" tableUuid="83F9BB9EFF9FFFFCFF2141976A04C514">Table 1</tableCitation>
) and to
<taxonomicName box="[151,318,268,295]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[151,170,269,295]" italics="true" pageId="9" pageNumber="118">C</emphasis>
. cf.
<emphasis box="[226,318,269,294]" italics="true" pageId="9" pageNumber="118">savignyi</emphasis>
</taxonomicName>
from northern and central Coastal Plain (
<tableCitation box="[803,892,268,294]" captionStart="Table 2" captionStartId="18.[144,197,236,257]" captionTargetBox="[144,1058,337,1237]" captionTargetPageId="18" captionText="Table 2: Least square means ± 95 % conFdence interval at covariate mean, and sample size for four variables per sampling localities grouped by taxon and geographic region; see Materials and Methods for full names of variables and Fig. 1 for numbers and names of geographic regions." httpUri="http://table.plazi.org/id/83F9BB9EFF9DFFFEFF2141976DAAC524" pageId="9" pageNumber="118" tableUuid="83F9BB9EFF9DFFFEFF2141976DAAC524">Table 2</tableCitation>
), but distinctly shorter than in
<taxonomicName box="[322,484,300,327]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[322,341,301,327]" italics="true" pageId="9" pageNumber="118">C</emphasis>
. cf.
<emphasis box="[392,484,301,326]" italics="true" pageId="9" pageNumber="118">savignyi</emphasis>
</taxonomicName>
from Negev and Arava Valley (
<tableCitation box="[854,942,300,326]" captionStart="Table 2" captionStartId="18.[144,197,236,257]" captionTargetBox="[144,1058,337,1237]" captionTargetPageId="18" captionText="Table 2: Least square means ± 95 % conFdence interval at covariate mean, and sample size for four variables per sampling localities grouped by taxon and geographic region; see Materials and Methods for full names of variables and Fig. 1 for numbers and names of geographic regions." httpUri="http://table.plazi.org/id/83F9BB9EFF9DFFFEFF2141976DAAC524" pageId="9" pageNumber="118" tableUuid="83F9BB9EFF9DFFFEFF2141976DAAC524">Table 2</tableCitation>
), and from
<taxonomicName box="[151,304,332,359]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="holgerseni">
<emphasis box="[151,304,332,359]" italics="true" pageId="9" pageNumber="118">C. holgerseni</emphasis>
</taxonomicName>
and
<taxonomicName box="[365,438,333,359]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="isis">
<emphasis box="[365,438,333,359]" italics="true" pageId="9" pageNumber="118">C. isis</emphasis>
</taxonomicName>
(
<tableCitation box="[457,542,332,358]" captionStart="Table 1" captionStartId="16.[144,197,236,257]" captionTargetBox="[144,1058,310,1593]" captionTargetPageId="16" captionText="Table 1: Least square means ± 95 % conFdence interval at covariate mean, and sample size for 17 va-" httpUri="http://table.plazi.org/id/83F9BB9EFF9FFFFCFF2141976A04C514" pageId="9" pageNumber="118" tableUuid="83F9BB9EFF9FFFFCFF2141976A04C514">Table 1</tableCitation>
).
<taxonomicName box="[569,723,332,359]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[569,723,332,359]" italics="true" pageId="9" pageNumber="118">C. israelensis</emphasis>
</taxonomicName>
has petiole node angle larger than the other species from Israel with Fattened anterior surface of petiole (
<tableCitation captionStart="Table 1" captionStartId="16.[144,197,236,257]" captionTargetBox="[144,1058,310,1593]" captionTargetPageId="16" captionText="Table 1: Least square means ± 95 % conFdence interval at covariate mean, and sample size for 17 va-" httpUri="http://table.plazi.org/id/83F9BB9EFF9FFFFCFF2141976A04C514" pageId="9" pageNumber="118" tableUuid="83F9BB9EFF9FFFFCFF2141976A04C514">Table 1</tableCitation>
); this speciFc difference especially marked between populations from northern and central Coastal Plain (cf.
<figureCitation box="[487,571,428,454]" captionStart="Figs 12, 13" captionStartId="8.[144,183,1128,1150]" captionTargetBox="[185,1017,235,1107]" captionTargetId="figure@8.[185,1017,235,723]" captionText="Figs 12, 13. C. israelensis, propodeum and petiole node in lateral view: (12) small paratype worker," pageId="9" pageNumber="118">Figs 13</figureCitation>
vs. 16;
<tableCitation box="[663,749,428,454]" captionStart="Table 2" captionStartId="18.[144,197,236,257]" captionTargetBox="[144,1058,337,1237]" captionTargetPageId="18" captionText="Table 2: Least square means ± 95 % conFdence interval at covariate mean, and sample size for four variables per sampling localities grouped by taxon and geographic region; see Materials and Methods for full names of variables and Fig. 1 for numbers and names of geographic regions." httpUri="http://table.plazi.org/id/83F9BB9EFF9DFFFEFF2141976DAAC524" pageId="9" pageNumber="118" tableUuid="83F9BB9EFF9DFFFEFF2141976DAAC524">Table 2</tableCitation>
).
<taxonomicName box="[771,923,428,455]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[771,923,428,455]" italics="true" pageId="9" pageNumber="118">C. israelensis</emphasis>
</taxonomicName>
differs from
<taxonomicName box="[151,250,460,487]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="nodus">
<emphasis box="[151,250,460,487]" italics="true" pageId="9" pageNumber="118">C. nodus</emphasis>
</taxonomicName>
by raised mesonotum over pronotum in 89% (n=108) vs. 12% (n=48) of workers, respectively, character state intermediate in
<taxonomicName box="[735,824,493,519]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="niger">
<emphasis box="[735,824,493,519]" italics="true" pageId="9" pageNumber="118">C. niger</emphasis>
</taxonomicName>
and
<taxonomicName box="[877,1036,492,519]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[877,896,493,519]" italics="true" pageId="9" pageNumber="118">C</emphasis>
. cf.
<emphasis box="[944,1036,493,518]" italics="true" pageId="9" pageNumber="118">savignyi</emphasis>
</taxonomicName>
. It has scarce pilosity on propodeum dorsum, similar to
<taxonomicName box="[729,818,525,551]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="niger">
<emphasis box="[729,818,525,551]" italics="true" pageId="9" pageNumber="118">C. niger</emphasis>
</taxonomicName>
and
<taxonomicName box="[871,1028,524,551]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[871,890,525,551]" italics="true" pageId="9" pageNumber="118">C</emphasis>
. cf.
<emphasis box="[936,1028,525,550]" italics="true" pageId="9" pageNumber="118">savignyi</emphasis>
</taxonomicName>
vs. abundant pilosity in most examined
<taxonomicName box="[544,640,556,583]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="nodus">
<emphasis box="[544,640,556,583]" italics="true" pageId="9" pageNumber="118">C. nodus</emphasis>
</taxonomicName>
(119 setae), and all
<taxonomicName box="[869,937,557,583]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="isis">
<emphasis box="[869,937,557,583]" italics="true" pageId="9" pageNumber="118">C. isis</emphasis>
</taxonomicName>
. Pilosity on propodeum dorsum and petiole node is less abundant than in the type of
<emphasis box="[946,1009,588,614]" italics="true" pageId="9" pageNumber="118">drusa</emphasis>
, but never completely absent as in the type of
<emphasis box="[610,690,621,646]" italics="true" pageId="9" pageNumber="118">assyria</emphasis>
. Pubescence of ventral surface of hindtibia Fne and appressed, similar to other Israeli species vs. coarse in
<taxonomicName box="[944,1065,653,679]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="nigripes">
<emphasis box="[944,1065,653,679]" italics="true" pageId="9" pageNumber="118">C. nigripes</emphasis>
</taxonomicName>
Arnoldi. In bicolored worker series, small to medium specimens (MW=1.01.5) have coxae and petiole concolorous with mesosoma pleura (
<figureCitation box="[817,887,716,743]" captionStart="Figs 7, 8" captionStartId="6.[168,207,1084,1106]" captionText="Figs 7, 8. C. israelensis holotype worker in lateral (7) and dorsal (8) views. Scale bars, 1.5 mm." pageId="9" pageNumber="118">Figs 7</figureCitation>
,
<figureCitation box="[899,929,716,742]" captionStart="Figs 12, 13" captionStartId="8.[144,183,1128,1150]" captionTargetBox="[185,1017,235,1107]" captionTargetId="figure@8.[185,1017,235,723]" captionText="Figs 12, 13. C. israelensis, propodeum and petiole node in lateral view: (12) small paratype worker," pageId="9" pageNumber="118">12</figureCitation>
), or to some extent darker, as opposed to black in
<taxonomicName box="[574,737,748,775]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[574,593,749,775]" italics="true" pageId="9" pageNumber="118">C</emphasis>
. cf.
<emphasis box="[645,737,749,774]" italics="true" pageId="9" pageNumber="118">savignyi</emphasis>
</taxonomicName>
(Fig. 17). Light colored specimens of
<taxonomicName box="[263,412,780,807]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[263,412,780,807]" italics="true" pageId="9" pageNumber="118">C. israelensis</emphasis>
</taxonomicName>
differ from similarly colored
<taxonomicName authority="Fabricius" authorityName="Fabricius" box="[734,965,780,807]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="viatica">
<emphasis box="[734,753,781,807]" italics="true" pageId="9" pageNumber="118">C</emphasis>
.
<emphasis box="[764,839,781,806]" italics="true" pageId="9" pageNumber="118">viatica</emphasis>
(Fabricius)
</taxonomicName>
by matte gaster, seldom laterally shiny, vs. completely shiny gaster in
<taxonomicName box="[820,895,813,838]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="viatica">
<emphasis box="[820,895,813,838]" italics="true" pageId="9" pageNumber="118">viatica</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="118" type="materials_examined">
<paragraph blockId="9.[151,1067,236,1603]" pageId="9" pageNumber="118">
<emphasis bold="true" box="[151,347,852,874]" pageId="9" pageNumber="118">Material examined:</emphasis>
<taxonomicName box="[358,487,852,874]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[358,487,852,874]" italics="true" pageId="9" pageNumber="118">C. israelensis</emphasis>
</taxonomicName>
:
<materialsCitation ID-GBIF-Occurrence="1423744320" collectingDate="2016-05-06" collectorName="P. - A. Eyer" country="Israel" latitude="32.653946" location="Daverat" longLatPrecision="1" longitude="35.377693" pageId="9" pageNumber="118" specimenCount="1" specimenCount-queen="1" typeStatus="holotype">
<typeStatus box="[504,588,852,874]" pageId="9" pageNumber="118">Holotype</typeStatus>
<specimenCount box="[599,616,851,875]" pageId="9" pageNumber="118" type="queen"></specimenCount>
,
<emphasis bold="true" box="[633,695,853,874]" pageId="9" pageNumber="118">
<collectingCountry box="[633,692,853,874]" name="Israel" pageId="9" pageNumber="118">Israel</collectingCountry>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:5F2F5A00FF8AFFE7FE0B44A96F79C013:D259BDCDFF86FFE5FD73422E6D2CC77F" box="[706,778,853,874]" country="Israel" latitude="32.653946" longLatPrecision="1" longitude="35.377693" name="Daverat" pageId="9" pageNumber="118">Daverat</location>
[
<geoCoordinate box="[795,920,852,874]" direction="north" orientation="latitude" pageId="9" pageNumber="118" precision="1" value="32.653946">32°39'14.2&quot;N</geoCoordinate>
<geoCoordinate box="[931,1056,852,874]" direction="east" orientation="longitude" pageId="9" pageNumber="118" precision="1" value="35.377693">35°22'39.7&quot;E</geoCoordinate>
],
<collectingDate box="[151,231,876,898]" pageId="9" pageNumber="118" value="2016-05-06">
<date box="[151,231,876,898]" pageId="9" pageNumber="118" value="2016-05-06">6.v.2016</date>
</collectingDate>
,
<collectorName box="[240,335,876,898]" pageId="9" pageNumber="118">P.-A. Eyer</collectorName>
</materialsCitation>
. Paratypes: same collection data as holotype (6☿); Montfort, 6.v.2016, P.-A. Eyer (7☿); Dishon, 17.v.1973, J. Kugler (3☿); Nahal Senir, 26.iv.1984, J. Kugler (2☿); Merom Golan, 15.vi.1971, J. Kugler (1Ƌ); Tel Dan, 17.vi.1971, H. Bytinski-Salz (1☿, 1♀, 2Ƌ); Panyas (Banyas), 12.v.1971, H. Bytinski-Salz (2☿); Agmon haHula #1, 2012, R. Zeltzer (3☿), #3, 2012, R. Zeltzer (4☿); Hula, 17.vii.2010, C. Drees (2☿); Ammiad #2, 2012, R. Zeltzer (1☿), #3, 2012, R. Zeltzer (1☿), #4, 2012, R. Zeltzer (1☿), #5, 2012, R. Zeltzer (5☿); Teverya [Tiberias], 12.v.19xx, H. Bytinski- Salz (1☿, 1♀, 2Ƌ); Nahal Kanaf, 28.xi.1981, W. Ferguson (2☿); Tel Qazir, iv.1955, J. Wahrman (3Ƌ); Hefzi Bah, 8.ix.2005, M. Vonshak (1☿); Atlit AT888, 12.iv.2016, T. Reiner-Brodetzki (1☿), AT889, 12.iv.2016, T. Reiner-Brodetzki (3☿), AT900, 12.iv.2016, T. Reiner-Brodetzki (3☿), AT901, 12.iv.2016, T. Reiner-Brodetzki (3☿), AT902, 12.iv.2016, T. Reiner-Brodetzki (3☿), AT910, 12.iv.2016, T. Reiner-Brodetzki (3☿), AT912, 12.iv.2016, T. Reiner-Brodetzki (8☿, 5♀), AT916, 12.iv.2016, T. Reiner-Brodetzki (6☿, 12Ƌ); Hof haBonim HB884, 12.iv.2016, T. Reiner-Brodetzki (8☿), HB888, 12.iv.2016, T. Reiner-Brodetzki (4☿), HB890, 12.iv.2016, T. Reiner-Brodetzki (3☿), HB894, 12.iv.2016, T. Reiner-Brodetzki (2☿); Qedumim, 15.vi.2016, A.L.L. Friedman (1☿); Yerushalayim [Jerusalem], 12.vi.1941, H. Bytinski-Salz (3☿, 1♀, 1Ƌ); Allon, 20.iii.2011, R. Zeltzer (6☿), #3, 20.iii.2011, R. Zeltzer (1☿), #4, 20.iii.2011, R. Zeltzer (7☿), #5, 20.iii.2011, R. Zeltzer (8☿); Maale Adummim, 20.iii.1979, J. Kugler (5☿).
</paragraph>
<paragraph blockId="9.[151,1067,236,1603]" pageId="9" pageNumber="118">
<taxonomicName box="[178,261,1287,1309]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="nodus">
<emphasis box="[178,193,1288,1309]" italics="true" pageId="9" pageNumber="118">C</emphasis>
.
<emphasis box="[207,261,1287,1309]" italics="true" pageId="9" pageNumber="118">nodus</emphasis>
</taxonomicName>
:
<emphasis bold="true" box="[276,347,1287,1309]" pageId="9" pageNumber="118">Serbia:</emphasis>
Belgrade, 13.vii.1928, F.S. Bodenheimer (2☿); 4.ix.1928, F.S. Bodenheimer (2☿).
<emphasis bold="true" box="[209,334,1312,1333]" pageId="9" pageNumber="118">Montenegro:</emphasis>
Budua, 21.iii.1923 (1☿).
<emphasis bold="true" box="[581,655,1311,1333]" pageId="9" pageNumber="118">Greece:</emphasis>
Saloniki, 12.v.1972, Poldi (1☿); Marathon, 18.iv.1962, H. Bytinski-Salz (9☿); Nauplion, 13.vii.1928, H. Bytinski-Salz (3☿); Samos, 22.iv.1962, H. Bytinski-Salz (1☿).
<emphasis bold="true" box="[359,436,1360,1381]" pageId="9" pageNumber="118">Turkey:</emphasis>
Edirne [Adrianopel], C. Menozzi (1☿); Istanbul, Saudiye, 4.viii.1929, F.S. Bodenheimer (4☿), 11.iv.1972, H. Bytinski-Salz (7☿); Ankara, 28.iv.1939, F.S. Bodenheimer (6☿); 12.v.1939, F.S. Bodenheimer (3☿); 18.v.1972, H. Bytinski-Salz (3☿); Sardes, 1973, A. Shor (2☿); Sivrice, 15.viii.1939, F.S. Bodenheimer (3☿); Plage Sigirak, 25.v.1939, F.S. Bodenheimer (1☿); Diyarbakır, 17.v.1972, H. Bytinski-Salz (6☿); Slovia, 2.vii.1921, (1☿).
<emphasis bold="true" box="[812,862,1456,1477]" pageId="9" pageNumber="118">Iran:</emphasis>
Fars, Akabad-Kamin, 27.vii.1969, C. Felton (1☿).
</paragraph>
<paragraph blockId="9.[151,1067,236,1603]" box="[178,1015,1504,1528]" pageId="9" pageNumber="118">
<taxonomicName authority="Fabricius" authorityName="Fabricius" box="[178,374,1506,1528]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="viatica">
<emphasis box="[178,193,1506,1527]" italics="true" pageId="9" pageNumber="118">C</emphasis>
.
<emphasis box="[204,266,1507,1528]" italics="true" pageId="9" pageNumber="118">viatica</emphasis>
(Fabricius)
</taxonomicName>
:
<emphasis bold="true" box="[383,476,1506,1527]" pageId="9" pageNumber="118">Morroco:</emphasis>
Oued Zate, 14.iv.1966 (1☿).
<emphasis bold="true" box="[744,823,1506,1528]" pageId="9" pageNumber="118">Algeria:</emphasis>
Sidi bel Abbes (1☿).
</paragraph>
</subSubSection>
<subSubSection lastPageId="10" lastPageNumber="119" pageId="9" pageNumber="118" type="discussion">
<paragraph blockId="9.[151,1067,236,1603]" lastBlockId="10.[144,1059,236,1601]" lastPageId="10" lastPageNumber="119" pageId="9" pageNumber="118">
<emphasis bold="true" box="[151,227,1544,1570]" pageId="9" pageNumber="118">Notes:</emphasis>
Ants from Israel were compared by Kugler (pers. comm., 2005) with types of
<emphasis box="[194,258,1576,1602]" italics="true" pageId="9" pageNumber="118">drusa</emphasis>
and
<emphasis box="[334,415,1577,1602]" italics="true" pageId="9" pageNumber="118">assyria</emphasis>
in the Santschi collection, and with types of
<taxonomicName authority="Fore" authorityName="Fore" class="Insecta" family="Formicidae" genus="Myrmecocystus" kingdom="Animalia" lastPageId="10" lastPageNumber="119" order="Hymenoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="variety" species="viaticus" variety="orientalis">
<emphasis italics="true" lastPageId="10" lastPageNumber="119" pageId="9" pageNumber="118">Myrmecocystus viaticus</emphasis>
var.
<emphasis box="[363,470,236,262]" italics="true" pageId="10" pageNumber="119">orientalis</emphasis>
Fore
</taxonomicName>
(junior synonym of
<taxonomicName box="[761,828,236,262]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nodus">
<emphasis box="[761,828,236,262]" italics="true" pageId="10" pageNumber="119">nodus</emphasis>
</taxonomicName>
) in the Forel collection (there is is a specimen from Jerusalem among Forels syntypes of
<taxonomicName box="[935,1042,268,294]" class="Insecta" family="Formicidae" genus="Myrmecocystus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="variety" species="viaticus" variety="orientalis">
<emphasis box="[935,1042,268,294]" italics="true" pageId="10" pageNumber="119">orientalis</emphasis>
</taxonomicName>
). Specimens from northern Israel have the shape of the petiole, pilosity and color intermediate between the examined types, in accordance with Santschi (1929, 1934). Consequently,
<emphasis box="[389,415,364,390]" italics="true" pageId="10" pageNumber="119">C.</emphasis>
sp. IL0 2 (part) and
<emphasis box="[639,664,364,390]" italics="true" pageId="10" pageNumber="119">C.</emphasis>
sp. IL04 (part) were considered by Kugler as closely related to
<emphasis box="[465,529,395,421]" italics="true" pageId="10" pageNumber="119">drusa</emphasis>
, and published as
<emphasis box="[744,770,396,422]" italics="true" pageId="10" pageNumber="119">C.</emphasis>
n. sp. near
<taxonomicName authority="Kugler 1988" authorityName="Kugler" authorityYear="1988" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nodus">
<emphasis box="[898,965,395,421]" italics="true" pageId="10" pageNumber="119">nodus</emphasis>
(Kugler 1988)
</taxonomicName>
.
<taxonomicName box="[235,345,427,454]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="druzus">
<emphasis box="[235,345,427,454]" italics="true" pageId="10" pageNumber="119">C. druzus</emphasis>
</taxonomicName>
together with
<taxonomicName authority="Fabricius" authorityName="Fabricius" box="[523,772,427,454]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis box="[523,640,427,454]" italics="true" pageId="10" pageNumber="119">C. bicolor</emphasis>
(Fabricius)
</taxonomicName>
were reported as widely distributed in Lebanon by Tohmé (1969) and Tohmé &amp; Tohmé (2014), whereas Knaden
<emphasis box="[241,293,491,517]" italics="true" pageId="10" pageNumber="119">et al</emphasis>
. (2012) reported
<taxonomicName box="[498,615,491,518]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis box="[498,517,491,517]" italics="true" pageId="10" pageNumber="119">C</emphasis>
.
<emphasis box="[534,615,491,517]" italics="true" pageId="10" pageNumber="119">bicolor</emphasis>
</taxonomicName>
and
<taxonomicName box="[676,788,491,518]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="viatica">
<emphasis box="[676,695,491,517]" italics="true" pageId="10" pageNumber="119">C</emphasis>
.
<emphasis box="[712,788,492,517]" italics="true" pageId="10" pageNumber="119">viatica</emphasis>
</taxonomicName>
from the Beqaa Valley (Lebanon). However, the relationship of these species to
<emphasis box="[786,812,523,549]" italics="true" pageId="10" pageNumber="119">C.</emphasis>
sp. IL0 2 is unknown, and according to D. Agosti (pers. comm., 1994) all reports of
<emphasis box="[849,913,555,581]" italics="true" pageId="10" pageNumber="119">drusa</emphasis>
or
<taxonomicName box="[958,1058,555,581]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nodus">
<emphasis box="[958,1058,555,581]" italics="true" pageId="10" pageNumber="119">C. nodus</emphasis>
</taxonomicName>
from Israel are misidentiFcations. Furthermore, samples from the Beqaa Valley that were collected less than 5km from the type locality of
<emphasis box="[800,864,618,644]" italics="true" pageId="10" pageNumber="119">drusa</emphasis>
by Knaden
<emphasis box="[1004,1051,618,644]" italics="true" pageId="10" pageNumber="119">et al</emphasis>
. (2012) cluster with
<taxonomicName box="[367,468,650,677]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nodus">
<emphasis box="[367,386,651,677]" italics="true" pageId="10" pageNumber="119">C</emphasis>
.
<emphasis box="[401,468,650,676]" italics="true" pageId="10" pageNumber="119">nodus</emphasis>
</taxonomicName>
, at a distance from the Israeli species (Eyer unpubl. data).
</paragraph>
<paragraph blockId="10.[144,1059,236,1601]" pageId="10" pageNumber="119">
Males and gyne of
<taxonomicName box="[383,483,714,741]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nodus">
<emphasis box="[383,483,714,741]" italics="true" pageId="10" pageNumber="119">C. nodus</emphasis>
</taxonomicName>
were not available for this study; however, the species diagnosis based on the male and gyne morphology proposed by Agosti (1994) indicates for males SL/HW&lt;1.70 in
<taxonomicName box="[551,650,778,804]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nodus">
<emphasis box="[551,650,778,804]" italics="true" pageId="10" pageNumber="119">C. nodus</emphasis>
</taxonomicName>
vs. SL/HW&gt;1.70 in
<taxonomicName box="[884,1010,778,804]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[884,1010,778,804]" italics="true" pageId="10" pageNumber="119">C. savignyi</emphasis>
</taxonomicName>
and for gyne HW/HL&lt;0.95 in
<taxonomicName box="[429,527,810,836]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nodus">
<emphasis box="[429,527,810,836]" italics="true" pageId="10" pageNumber="119">C. nodus</emphasis>
</taxonomicName>
vs. HW/HL&gt;0.95 in
<taxonomicName box="[759,883,810,836]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[759,883,810,836]" italics="true" pageId="10" pageNumber="119">C. savignyi</emphasis>
</taxonomicName>
; in both species gyne with the red head and mesosoma. The present study found for males SL/ HW=1.571.74 (n=21) in
<taxonomicName box="[435,587,873,900]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[435,587,873,900]" italics="true" pageId="10" pageNumber="119">C. israelensis</emphasis>
</taxonomicName>
, similar to
<taxonomicName box="[712,873,873,900]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[712,731,874,900]" italics="true" pageId="10" pageNumber="119">C</emphasis>
. cf.
<emphasis box="[780,873,874,899]" italics="true" pageId="10" pageNumber="119">savignyi</emphasis>
</taxonomicName>
from Israel (SL/ HW=1.621.77, n=14) and
<taxonomicName box="[462,556,906,932]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="niger">
<emphasis box="[462,556,906,932]" italics="true" pageId="10" pageNumber="119">C. niger</emphasis>
</taxonomicName>
(SL/HW=1.611.73, n=10);
<taxonomicName box="[891,1058,905,932]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[891,910,906,932]" italics="true" pageId="10" pageNumber="119">C</emphasis>
. cf.
<emphasis box="[965,1058,906,931]" italics="true" pageId="10" pageNumber="119">savignyi</emphasis>
</taxonomicName>
from Sinai have SL/HW=1.711.85 (n=4). For gyne, in contrast to Agosti (1994), HW/HL=0.960.98 (n=8) in
<taxonomicName box="[470,621,969,995]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[470,621,969,995]" italics="true" pageId="10" pageNumber="119">C. israelensis</emphasis>
</taxonomicName>
and HW/HL=0.910.99 (n=5) in
<taxonomicName class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[998,1017,969,995]" italics="true" pageId="10" pageNumber="119">C</emphasis>
. cf.
<emphasis box="[144,237,1002,1027]" italics="true" pageId="10" pageNumber="119">savignyi</emphasis>
</taxonomicName>
, in both species color variable, similar to associated workers. According to Agosti (1990) the apical appendages of the subgenital plate are of similar length in
<taxonomicName box="[171,270,1065,1091]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nodus">
<emphasis box="[171,270,1065,1091]" italics="true" pageId="10" pageNumber="119">C. nodus</emphasis>
</taxonomicName>
(Agosti 1990, Fg. 23), whereas in
<taxonomicName box="[653,804,1065,1091]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[653,804,1065,1091]" italics="true" pageId="10" pageNumber="119">C. israelensis</emphasis>
</taxonomicName>
the median appendage of the subgenital plate is shorter than the lateral appendages (mSAL/lSAL=0.52 0.71, n=7) (Figs 23, 24), similar to
<taxonomicName box="[531,622,1129,1155]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="niger">
<emphasis box="[531,622,1129,1155]" italics="true" pageId="10" pageNumber="119">C. niger</emphasis>
</taxonomicName>
(0.550.73, n=8) and to
<taxonomicName box="[899,1058,1128,1155]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[899,918,1129,1155]" italics="true" pageId="10" pageNumber="119">C</emphasis>
. cf.
<emphasis box="[965,1058,1129,1154]" italics="true" pageId="10" pageNumber="119">savignyi</emphasis>
</taxonomicName>
(0.590.63, n=7) (Figs 26, 27, 29, 30). Likewise, in
<taxonomicName box="[715,866,1160,1187]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[715,866,1160,1187]" italics="true" pageId="10" pageNumber="119">C. israelensis</emphasis>
</taxonomicName>
the stipites in the lateral view (Fig. 25) are similar to
<taxonomicName box="[536,695,1192,1219]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[536,555,1192,1218]" italics="true" pageId="10" pageNumber="119">C</emphasis>
. cf.
<emphasis box="[602,695,1193,1218]" italics="true" pageId="10" pageNumber="119">savignyi</emphasis>
</taxonomicName>
(Fig. 31), intermediate between, but closer to
<taxonomicName box="[288,380,1224,1250]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="niger">
<emphasis box="[288,380,1224,1250]" italics="true" pageId="10" pageNumber="119">C. niger</emphasis>
</taxonomicName>
(Fig. 28) than to
<taxonomicName box="[575,675,1224,1250]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nodus">
<emphasis box="[575,675,1224,1250]" italics="true" pageId="10" pageNumber="119">C. nodus</emphasis>
</taxonomicName>
as illustrated in Agosti (1990, Fgs 45, 46). Therefore, contrary to the proposed afFnity between
<emphasis box="[845,864,1256,1282]" italics="true" pageId="10" pageNumber="119">C</emphasis>
. sp. IL0 2 and
<taxonomicName authority="in Vonshak and Ionescu-Hirsch (2010)" authorityName="in Vonshak and Ionescu-Hirsch" authorityYear="2010" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nodus">
<emphasis box="[1033,1052,1256,1282]" italics="true" pageId="10" pageNumber="119">C</emphasis>
.
<emphasis box="[144,211,1288,1314]" italics="true" pageId="10" pageNumber="119">nodus</emphasis>
in Vonshak and Ionescu-Hirsch (2010)
</taxonomicName>
,
<taxonomicName box="[661,813,1288,1314]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[661,813,1288,1314]" italics="true" pageId="10" pageNumber="119">C. israelensis</emphasis>
</taxonomicName>
cannot be included in the
<taxonomicName box="[186,253,1319,1345]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nodus">
<emphasis box="[186,253,1319,1345]" italics="true" pageId="10" pageNumber="119">nodus</emphasis>
</taxonomicName>
species-complex as diagnosed by Agosti (1990). It is possible that the diagnosis of the
<taxonomicName box="[331,398,1351,1377]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="nodus">
<emphasis box="[331,398,1351,1377]" italics="true" pageId="10" pageNumber="119">nodus</emphasis>
</taxonomicName>
species-complex is too restrictive, thus not accounting for the variation in this widely distributed group (see Emery 1906, Fgs 2830; Knaden
<emphasis box="[191,240,1415,1441]" italics="true" pageId="10" pageNumber="119">et al</emphasis>
. 2012, Fg. 4).
</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="120" pageId="10" pageNumber="119" type="distribution">
<paragraph blockId="10.[144,1059,236,1601]" lastBlockId="11.[151,1067,236,1030]" lastPageId="11" lastPageNumber="120" pageId="10" pageNumber="119">
<taxonomicName box="[170,435,1447,1473]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[170,435,1447,1473]" italics="true" pageId="10" pageNumber="119">Cataglyphis israelensis</emphasis>
</taxonomicName>
is distributed uniformly in northern and north-eastern Israel, parapatric with
<taxonomicName box="[400,565,1479,1505]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[400,419,1479,1505]" italics="true" pageId="10" pageNumber="119">C</emphasis>
. cf.
<emphasis box="[472,565,1480,1505]" italics="true" pageId="10" pageNumber="119">savignyi</emphasis>
</taxonomicName>
, and over most of its range the ants are bicolored, thus identiFable straightforwardly by their color pattern and the shape of the propodeum. However, in areas of the overlap between species containing black morphs, as in the northern Coastal Plain, Jerusalem and the Judean Desert, identiFcation of individual ants is problematic. The Frst discriminant analysis on residuals of measurements on all specimens in four groups—
<taxonomicName box="[860,978,268,294]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="holgerseni">
<emphasis box="[860,978,268,294]" italics="true" pageId="11" pageNumber="120">holgerseni</emphasis>
</taxonomicName>
,
<taxonomicName box="[1012,1049,269,294]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="isis">
<emphasis box="[1012,1049,269,294]" italics="true" pageId="11" pageNumber="120">isis</emphasis>
</taxonomicName>
,
<taxonomicName box="[161,281,300,326]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[161,281,300,326]" italics="true" pageId="11" pageNumber="120">israelensis</emphasis>
</taxonomicName>
and
<taxonomicName box="[359,418,301,326]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="niger">
<emphasis box="[359,418,301,326]" italics="true" pageId="11" pageNumber="120">niger</emphasis>
</taxonomicName>
complex—revealed 100% correct classiFcation of the unproblematic
<taxonomicName box="[324,478,332,359]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="holgerseni">
<emphasis box="[324,478,332,359]" italics="true" pageId="11" pageNumber="120">C. holgerseni</emphasis>
</taxonomicName>
and
<taxonomicName box="[540,614,333,359]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="isis">
<emphasis box="[540,614,333,359]" italics="true" pageId="11" pageNumber="120">C. isis</emphasis>
</taxonomicName>
, 81% correct classiFcation of
<taxonomicName class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis italics="true" pageId="11" pageNumber="120">C. israelensis</emphasis>
</taxonomicName>
and 94% correct classiFcation of ants in the
<taxonomicName box="[752,811,365,390]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="niger">
<emphasis box="[752,811,365,390]" italics="true" pageId="11" pageNumber="120">niger</emphasis>
</taxonomicName>
species-complex. The second analysis was limited to three groups comprising 73
<taxonomicName box="[832,987,396,423]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[832,987,396,423]" italics="true" pageId="11" pageNumber="120">C. israelensis</emphasis>
</taxonomicName>
(19 of them bicolored), 39
<taxonomicName box="[388,482,429,455]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="niger">
<emphasis box="[388,482,429,455]" italics="true" pageId="11" pageNumber="120">C. niger</emphasis>
</taxonomicName>
and 102
<taxonomicName box="[595,763,428,455]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[595,614,429,455]" italics="true" pageId="11" pageNumber="120">C</emphasis>
. cf.
<emphasis box="[670,763,429,454]" italics="true" pageId="11" pageNumber="120">savignyi</emphasis>
</taxonomicName>
specimens from localities sampled for molecular analyses. The discriminant analysis (Wilks Lambda: 0.366 approx. F (16,408)=16.675, p&lt;0.001) resulted in 86 % correct classiFcation of ants belonging to
<taxonomicName box="[300,453,524,551]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[300,453,524,551]" italics="true" pageId="11" pageNumber="120">C. israelensis</emphasis>
</taxonomicName>
, 84% correct classiFcation of
<taxonomicName box="[798,961,524,551]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[798,817,525,551]" italics="true" pageId="11" pageNumber="120">C</emphasis>
. cf.
<emphasis box="[868,961,525,550]" italics="true" pageId="11" pageNumber="120">savignyi</emphasis>
</taxonomicName>
and only 45 % correct classiFcation of
<taxonomicName box="[483,575,557,583]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="niger">
<emphasis box="[483,502,557,583]" italics="true" pageId="11" pageNumber="120">C</emphasis>
.
<emphasis box="[516,575,557,582]" italics="true" pageId="11" pageNumber="120">niger</emphasis>
</taxonomicName>
. The scrutiny of the results showed that accounting for color increased the correct classiFcation rate of
<taxonomicName box="[846,1000,588,615]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[846,1000,588,615]" italics="true" pageId="11" pageNumber="120">C. israelensis</emphasis>
</taxonomicName>
: only four were misidentiFed as
<taxonomicName box="[450,612,620,647]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[450,469,621,647]" italics="true" pageId="11" pageNumber="120">C</emphasis>
. cf.
<emphasis box="[519,612,621,646]" italics="true" pageId="11" pageNumber="120">savignyi</emphasis>
</taxonomicName>
and one as
<taxonomicName box="[745,837,621,647]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="niger">
<emphasis box="[745,764,621,647]" italics="true" pageId="11" pageNumber="120">C</emphasis>
.
<emphasis box="[778,837,621,646]" italics="true" pageId="11" pageNumber="120">niger</emphasis>
</taxonomicName>
; and eight
<taxonomicName class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[963,982,621,647]" italics="true" pageId="11" pageNumber="120">C</emphasis>
. cf.
<emphasis italics="true" pageId="11" pageNumber="120">savignyi</emphasis>
</taxonomicName>
and two
<taxonomicName box="[325,417,653,679]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="niger">
<emphasis box="[325,344,653,679]" italics="true" pageId="11" pageNumber="120">C</emphasis>
.
<emphasis box="[358,417,653,678]" italics="true" pageId="11" pageNumber="120">niger</emphasis>
</taxonomicName>
were misidentiFed as
<taxonomicName box="[672,825,652,679]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[672,825,652,679]" italics="true" pageId="11" pageNumber="120">C. israelensis</emphasis>
</taxonomicName>
. A direct comparison of 54 black
<taxonomicName box="[292,448,684,711]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[292,448,684,711]" italics="true" pageId="11" pageNumber="120">C. israelensis</emphasis>
</taxonomicName>
and 49
<taxonomicName box="[545,712,684,711]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[545,564,685,711]" italics="true" pageId="11" pageNumber="120">C</emphasis>
. cf.
<emphasis box="[619,712,685,710]" italics="true" pageId="11" pageNumber="120">savignyi</emphasis>
</taxonomicName>
collected on the Coastal Plain was carried out by recalculating the average regression lines between the two groups for PrH, PNL, PNH and PNA. For
<taxonomicName box="[631,784,748,775]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[631,784,748,775]" italics="true" pageId="11" pageNumber="120">C. israelensis</emphasis>
</taxonomicName>
, a low propodeum, short petiole node, high petiole node and large petiole node angle relative to average, and raised mesonotum over pronotum were considered characteristic. Forty-nine
<taxonomicName box="[151,306,844,871]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[151,306,844,871]" italics="true" pageId="11" pageNumber="120">C. israelensis</emphasis>
</taxonomicName>
and 37
<taxonomicName box="[403,570,844,871]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="savignyi">
<emphasis box="[403,422,845,871]" italics="true" pageId="11" pageNumber="120">C</emphasis>
. cf.
<emphasis box="[477,570,845,870]" italics="true" pageId="11" pageNumber="120">savignyi</emphasis>
</taxonomicName>
displayed the characteristic condition in at least three variables, an 8% rate of misclassiFcation vs. 50 % in a random choice. Consequently, appropriate sample size and number of samples may support a reasonably high correct identiFcation rate (
<emphasis box="[611,673,941,966]" italics="true" pageId="11" pageNumber="120">sensu</emphasis>
Seifert 2014) of
<taxonomicName box="[874,1029,940,967]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="israelensis">
<emphasis box="[874,1029,940,967]" italics="true" pageId="11" pageNumber="120">C. israelensis</emphasis>
</taxonomicName>
by morphology, despite a high intraspeciFc variability within nest series and among local populations.
</paragraph>
</subSubSection>
</treatment>
</document>