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<document id="60E523AA879658CF7C027D0FB0CB628C" ID-DOI="10.11646/zootaxa.3904.3.1" ID-GBIF-Dataset="5319c775-933f-499c-8c2a-6eb810fdd42b" ID-ISSN="1175-5326" ID-Zenodo-Dep="253414" ID-ZooBank="FC00E6FA-7442-4F9C-84F4-543D93311FE5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460887662619" checkinUser="plazi" docAuthor="Krolow, Tiago Kütter, Henriques, Augusto Loureiro, Gorayeb, Inocêncio De Sousa, Limeira-De-Oliveira, Francisco &amp; Buestán, Jaime" docDate="2015" docId="F96D878BAA32BD01FBC4F987F8B2FEC5" docLanguage="en" docName="zt03904p333.pdf" docOrigin="Zootaxa 3904 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pityocera (Pseudelaphella) pernaquila Gorayeb &amp; Krolow, sp. nov." docType="treatment" docVersion="8" lastPageNumber="324" masterDocId="0554FFF3AA26BD16FB53FFA8FC5AFFA0" masterDocTitle="Taxonomic revision of the Neotropical genus Pityocera Giglio-Tos, 1896 (Diptera: Tabanidae: Scionini)" masterLastPageNumber="333" masterPageNumber="301" pageNumber="321" updateTime="1698359274381" updateUser="plazi">
<mods:mods id="916979D94E5BDD5AC97BF268C5CC7063" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="65CCF82F31F50A3B708858E3D35A26E6">
<mods:title id="257A8E95F8AE95CD830B76C83F09D57E">Taxonomic revision of the Neotropical genus Pityocera Giglio-Tos, 1896 (Diptera: Tabanidae: Scionini)</mods:title>
</mods:titleInfo>
<mods:name id="7DE99C9866D90973B0260813956579E5" type="personal">
<mods:role id="11531B265AE72D826A194B5F153A5AC9">
<mods:roleTerm id="3819834A96E76911977EAED55C85E24A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5C98B7FE06C8D436B94CD46771029D06">Krolow, Tiago Kütter</mods:namePart>
</mods:name>
<mods:name id="D65E0D9E31D5496F8031FD00F1727480" type="personal">
<mods:role id="0EFDA59E0D4DD939C5208A60F81B7616">
<mods:roleTerm id="07F6B9799506A117BA25AFCD1A10B9DE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0E7B1C1366802D8EF1F099DAFA43470C">Henriques, Augusto Loureiro</mods:namePart>
</mods:name>
<mods:name id="86F62CEF19D93CD48D8E4DE11A7A25C9" type="personal">
<mods:role id="3F9FB4FB3B83EE34795041FEEF99289B">
<mods:roleTerm id="A688A8E198B1286B22D73699C1F359BA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="94EE990607D247857BBD2ADD8D7162F9">Gorayeb, Inocêncio De Sousa</mods:namePart>
</mods:name>
<mods:name id="1EB4ABAEF5A8F0F06D5B5D9BE8DDB92A" type="personal">
<mods:role id="FA585A01C6B042CD1B9E2C82F69BCE13">
<mods:roleTerm id="A2C2CCD38C0171C875F04BAA84920E62">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="49C3A267A1D24EEF9E9C3FCCCC1EFB35">Limeira-De-Oliveira, Francisco</mods:namePart>
</mods:name>
<mods:name id="0F2A1AA58D441D4AA2473DBFB63F412B" type="personal">
<mods:role id="B5AB236DD6349541D24C9202236A490A">
<mods:roleTerm id="3C2082FCA915F205D6257B6779AA606A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="56CF6A9AC42E6B83312FF438A6F74308">Buestán, Jaime</mods:namePart>
</mods:name>
<mods:typeOfResource id="C1AF0FF392BA8DAA8B683D5FD8F17F96">text</mods:typeOfResource>
<mods:relatedItem id="C9FA87E45FB273F785021F6C6661ED3B" type="host">
<mods:titleInfo id="3A66019B39590E9A3DE4BD7FCA392919">
<mods:title id="0ADEAD4775C23CA5036A86731AEEDD7A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D19F0DC1C7E9B517C81648DC1C596694">
<mods:date id="F8FF30EC7C1D6EBC21EAAE9EE59342F6">2015</mods:date>
<mods:detail id="2A544EAFC1D3751E8DA76D3B176AA007" type="volume">
<mods:number id="33B59ACBE01ADF1E9E33482ACB623D3A">3904</mods:number>
</mods:detail>
<mods:detail id="EB8EA9F3D14EE9DC90C79ABCB65FD2D8" type="issue">
<mods:number id="3A44A6D809BE110877C87F7475762380">3</mods:number>
</mods:detail>
<mods:extent id="7F62B82B1C695BCFCF56D2D19E7CE1C5" unit="page">
<mods:start id="FD0464AE8800FBE6D7F43FEEF94B9D7B">301</mods:start>
<mods:end id="D18676D835CDCA8684E73D7839D5FEB6">333</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="4CA0DDBF1B422542251661DBAE8EAE75">journal article</mods:classification>
<mods:identifier id="1A7B4D2AFE302B0C2E98BDB399AF6F78" type="DOI">10.11646/zootaxa.3904.3.1</mods:identifier>
<mods:identifier id="D467EB0E93035FE2BE3DFBCD576213DA" type="GBIF-Dataset">5319c775-933f-499c-8c2a-6eb810fdd42b</mods:identifier>
<mods:identifier id="04432CDF0B7D4BEB6DB3DE5005E32E50" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8CB12367B3C0C0FE1C1D50E620169829" type="Zenodo-Dep">253414</mods:identifier>
<mods:identifier id="B5B6D81B6AB20FDF2634AFE65E2C2781" type="ZooBank">FC00E6FA-7442-4F9C-84F4-543D93311FE5</mods:identifier>
</mods:mods>
<treatment id="F96D878BAA32BD01FBC4F987F8B2FEC5" ID-DOI="http://doi.org/10.5281/zenodo.6100347" ID-GBIF-Taxon="119586787" ID-Zenodo-Dep="6100347" LSID="urn:lsid:plazi:treatment:F96D878BAA32BD01FBC4F987F8B2FEC5" httpUri="http://treatment.plazi.org/id/F96D878BAA32BD01FBC4F987F8B2FEC5" lastPageId="23" lastPageNumber="324" pageId="20" pageNumber="321">
<subSubSection id="39DE6516AA32BD02FBC4F987FD16F9CB" pageId="20" pageNumber="321" type="nomenclature">
<paragraph id="717B369DAA32BD02FBC4F987FF9FF9EA" blockId="20.[151,965,1583,1643]" box="[151,965,1583,1610]" pageId="20" pageNumber="321">
<heading id="2A3381F1AA32BD02FBC4F987FF9FF9EA" bold="true" box="[151,965,1583,1610]" fontSize="11" level="1" pageId="20" pageNumber="321" reason="1">
<emphasis id="43B0EA8FAA32BD02FBC4F987FF9FF9EA" bold="true" box="[151,965,1583,1610]" pageId="20" pageNumber="321">
<taxonomicName id="B6C44D1EAA32BD02FBC4F987FF3AF9EA" ID-CoL="9HTWD" authority="Gorayeb &amp; Krolow" authorityName="Gorayeb &amp; Krolow" box="[151,864,1583,1610]" class="Insecta" family="Tabanidae" genus="Pityocera" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="pernaquila" status="sp. nov." subGenus="Pseudelaphella">
<emphasis id="43B0EA8FAA32BD02FBC4F987FD53F9E9" bold="true" box="[151,265,1583,1609]" italics="true" pageId="20" pageNumber="321">Pityocera</emphasis>
(
<emphasis id="43B0EA8FAA32BD02FA49F987FD88F9E9" bold="true" box="[282,466,1583,1609]" italics="true" pageId="20" pageNumber="321">Pseudelaphella</emphasis>
)
<emphasis id="43B0EA8FAA32BD02FAB0F987FE3DF9E9" bold="true" box="[483,615,1583,1609]" italics="true" pageId="20" pageNumber="321">pernaquila</emphasis>
Gorayeb &amp; Krolow
</taxonomicName>
<taxonomicNameLabel id="588357F4AA32BD02F834F998FF9FF9EA" box="[871,965,1584,1610]" pageId="20" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="717B369DAA32BD02FBC4F9FAFD16F9CB" blockId="20.[151,965,1583,1643]" box="[151,332,1618,1643]" pageId="20" pageNumber="321">(figures 12A–F)</paragraph>
</subSubSection>
<subSubSection id="39DE6516AA32BD02FBC4F933F857F910" box="[151,1037,1690,1712]" pageId="20" pageNumber="321" type="reference_group">
<paragraph id="717B369DAA32BD02FBC4F933F857F910" blockId="20.[151,1037,1690,1712]" box="[151,1037,1690,1712]" pageId="20" pageNumber="321">
<treatmentCitationGroup id="51D411B3AA32BD02FBC4F933F857F910" box="[151,1037,1690,1712]" pageId="20" pageNumber="321">
<taxonomicName id="B6C44D1EAA32BD02FBC4F933FF77F910" ID-CoL="8DDH5" authority="Henriques &amp; Gorayeb, 1993: 5" authorityName="Henriques &amp; Gorayeb" authorityPageNumber="5" authorityYear="1993" box="[151,813,1690,1712]" class="Insecta" family="Tabanidae" genus="Pityocera" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="nana" subGenus="Pseudelaphella">
<emphasis id="43B0EA8FAA32BD02FBC4F933FCA0F910" box="[151,250,1691,1712]" italics="true" pageId="20" pageNumber="321">Pityocera</emphasis>
(
<emphasis id="43B0EA8FAA32BD02FA5BF932FDFEF910" box="[264,420,1690,1712]" italics="true" pageId="20" pageNumber="321">Pseudelaphella</emphasis>
)
<emphasis id="43B0EA8FAA32BD02FAE0F933FDBFF910" box="[435,485,1691,1712]" italics="true" pageId="20" pageNumber="321">nana</emphasis>
,
<treatmentCitation id="F065108CAA32BD02FAA1F932FF77F910" author="Henriques" box="[498,813,1690,1712]" page="5" pageId="20" pageNumber="321" year="1993">
<bibRefCitation id="15554B6CAA32BD02FAA1F932FF4FF910" author="Henriques" box="[498,789,1690,1712]" pageId="20" pageNumber="321" refString="Henriques, A. L. &amp; Gorayeb, I. S. (1993) A colecao de Tabanidae (Insecta: Diptera) do Museu Paraense Emilio Goeldi (MPEG), Belem, Para, Brasil. Goeldiana Zoologia, 20, 1 - 23." type="journal article" year="1993">Henriques &amp; Gorayeb, 1993</bibRefCitation>
: 5
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="F065108CAA32BD02F86BF932F853F910" author="Henriques" box="[824,1033,1690,1712]" page="62" pageId="20" pageNumber="321" year="1997">
<bibRefCitation id="15554B6CAA32BD02F86BF932FFB9F910" author="Henriques" box="[824,995,1690,1712]" pageId="20" pageNumber="321" refString="Henriques, A. L. (1997) A colecao de Tabanidae (Insecta: Diptera) do Instituto Nacional de Pesquisas da Amazonia (INPA), Manaus, Amazonas, Brasil. Boletim do Museu Paraense Emilio Goeldi, serie Zoologia, 11 (1), 57 - 99. [1995]" type="journal article" year="1997">Henriques, 1997</bibRefCitation>
: 62
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="39DE6516AA32BD02FBC4F975FE02F955" box="[151,600,1757,1782]" pageId="20" pageNumber="321" type="materials_examined">
<paragraph id="717B369DAA32BD02FBC4F975FE02F955" blockId="20.[151,1437,1757,2033]" box="[151,600,1757,1782]" pageId="20" pageNumber="321">
<emphasis id="43B0EA8FAA32BD02FBC4F975FD6FF956" bold="true" box="[151,309,1757,1782]" pageId="20" pageNumber="321">
<typeStatus id="AE7F883FAA32BD02FBC4F975FC88F956" box="[151,210,1757,1782]" pageId="20" pageNumber="321">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="09D3760DAA32BD02FA6FF975FDD9F956" box="[316,387,1757,1782]" name="Brazil" pageId="20" pageNumber="321">Brazil</collectingCountry>
, Pará, Serra Norte.
</paragraph>
</subSubSection>
<subSubSection id="39DE6516AA32BD02FB94F8A9FCBCF8C2" pageId="20" pageNumber="321" type="diagnosis">
<paragraph id="717B369DAA32BD02FB94F8A9FCBCF8C2" blockId="20.[151,1437,1757,2033]" pageId="20" pageNumber="321">
<emphasis id="43B0EA8FAA32BD02FB94F8A9FD1BF8BA" bold="true" box="[199,321,1793,1818]" pageId="20" pageNumber="321">Diagnosis.</emphasis>
size varying from
<quantity id="B63C9B78AA32BD02F97FF8A9FEE6F8BA" box="[556,700,1793,1818]" metricMagnitude="-2" metricUnit="m" metricValue="1.0399999999999998" metricValueMax="1.16" metricValueMin="0.9199999999999999" pageId="20" pageNumber="321" unit="mm" value="10.399999999999999" valueMax="11.6" valueMin="9.2">9.2–11.6mm</quantity>
; frons parallel sides; face, gena and parafacial pruinose, only clypeus smooth and shiny; beard white; antennal flagellum orange; first flagellomere as wide as long; legs dark brown.
</paragraph>
</subSubSection>
<subSubSection id="39DE6516AA32BD03FB94F8C5FDD9FB34" lastPageId="21" lastPageNumber="322" pageId="20" pageNumber="321" type="description">
<paragraph id="717B369DAA32BD02FB94F8C5F914F825" blockId="20.[151,1437,1757,2033]" box="[199,1358,1901,1926]" pageId="20" pageNumber="321">
<emphasis id="43B0EA8FAA32BD02FB94F8C5FD96F826" bold="true" box="[199,460,1901,1926]" pageId="20" pageNumber="321">
Description,
<typeStatus id="AE7F883FAA32BD02FA0CF8C5FD96F826" box="[351,460,1901,1926]" pageId="20" pageNumber="321" type="holotype">Holotype</typeStatus>
</emphasis>
♀: body length
<quantity id="B63C9B78AA32BD02F9D0F8C5FEBDF825" box="[643,743,1901,1926]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="20" pageNumber="321" unit="mm" value="10.5">10.5 mm</quantity>
(fig. 12A); wing length
<quantity id="B63C9B78AA32BD02F8ABF8C6F805F825" box="[1016,1119,1902,1926]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="20" pageNumber="321" unit="mm" value="10.1">10.1 mm</quantity>
; wing width
<quantity id="B63C9B78AA32BD02FFA3F8C6F913F825" box="[1264,1353,1902,1926]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="20" pageNumber="321" unit="mm" value="3.2">3.2 mm</quantity>
.
</paragraph>
<paragraph id="717B369DAA32BD02FB94F839F9C7F851" blockId="20.[151,1437,1757,2033]" pageId="20" pageNumber="321">
<emphasis id="43B0EA8FAA32BD02FB94F839FD5DF80A" bold="true" box="[199,263,1937,1962]" pageId="20" pageNumber="321">Head</emphasis>
(fig. 12B): eyes without color pattern when dry, densely pilose. Occiput with white hairs. Frons parallel, slightly convergent at base, D.I. 1.1, F.I. 2.8; with yellowish brown pruinosity and brown pilosity. Subcallus glabrous, with pruinosity identical to frons. Face with pruinosity same as frons, and sparse light-brown hair.
</paragraph>
<paragraph id="717B369DAA33BD03FBC4FF3FFF86FE25" blockId="21.[151,1437,151,2024]" pageId="21" pageNumber="322">Parafacial and gena with greyish pruinosity and white beard. Clypeus smooth and shiny. Antenna (fig. 12C), scape and pedicel with yellowish brown pruinosity and black hairs; flagellum dark orange, without pruinosity; first flagellomere as high as long, with tuft of dorsal hairs; flagellomeres 1–3 fused, 4–7 free; apical flagellomere with black hairs. Palpus with first segment tubular and a little smaller than second; second segment with enlargement occupying more than half of the length and hairs only on external margins, central area dark orange and bare; distal half narrowed and completely pilose; proboscis slender and very long, over twice the length of the head; theca membranous; labella slender (without expansion) and partially sclerotized;</paragraph>
<paragraph id="717B369DAA33BD03FB94FE38FDD3FD20" blockId="21.[151,1437,151,2024]" pageId="21" pageNumber="322">
<emphasis id="43B0EA8FAA33BD03FB94FE38FD73FE09" bold="true" box="[199,297,400,425]" pageId="21" pageNumber="322">Thorax:</emphasis>
scutum dark brown, with black hair and three very distinct longitudinal bands of greyish white pruinosity; post-pronotal lobe, notopleura, post-alar callus and scutellum apex with greyish-white pruinosity. Notopleura with tuft of black hair (dorso-lateral view). Pleura with dark integument covered by greyish white pruinosity and predominantly white hair. Legs slender; coxae with pruinosity and pilosity similar to pleura; trochanters, tibiae, femora and tarsi dark brown with predominantly black pilosity, with some sparse brown hairs; tibiae with black hair in dorsal view. Wing fumose; r5 cell closed with very long petiole; appendix of R4 very short; venation dark-brown.
</paragraph>
<paragraph id="717B369DAA33BD03FB94FD23FEA3FD4B" blockId="21.[151,1437,151,2024]" pageId="21" pageNumber="322">
<emphasis id="43B0EA8FAA33BD03FB94FD23FD1FFD04" bold="true" box="[199,325,651,676]" pageId="21" pageNumber="322">Abdomen:</emphasis>
dark brown, tergites with posterior transverse bands bright; tergites 1–2 bright (yellowish) with dark spot in the middle; tergites 3–7 dark brown with light posterior transverse bands; pilosity predominantly black, with white hairs starting from the fourth tergite.
</paragraph>
<paragraph id="717B369DAA33BD03FB94FD5DFD52FB8A" blockId="21.[151,1437,151,2024]" pageId="21" pageNumber="322">
Terminalia (
<typeStatus id="AE7F883FAA33BD03FA00FD5EFDE0FCAE" box="[339,442,758,782]" pageId="21" pageNumber="322" type="paratype">paratype</typeStatus>
). Tergites IX and X, cercus and hypoproct as in figure (fig. 12D). Tergite IX fused, narrowed in center. Tergite X divided into two pieces with a membranous space between the contact area of the internal lateral margins and the tergite IX. Cercus with margins rounded and apex slightly acuminate. Hypoproct rounded, reaching the first third of the cercus length. Hypogynium + hypogynial valve (fig. 12E). Hypogynium subtriangular with lateral margins straight from middle to apex and slightly rounded from middle to basis; apex with a v-shaped slit reaching the hypogynial valve. Genital fork (fig. 12F) with arms slender, but with distal expansions sclerotized; anterior margin straight with slight median indentation; lateral extremities rounded; spermathecal ducts long and little sclerotized; spermatheca (fig. 12F) heavily pigmented, elongated, with apex narrowed.
</paragraph>
<paragraph id="717B369DAA33BD03FB94FB9DFDDDFBD1" blockId="21.[151,1437,151,2024]" pageId="21" pageNumber="322">
<emphasis id="43B0EA8FAA33BD03FB94FB9DFD92FBEE" bold="true" box="[199,456,1077,1102]" pageId="21" pageNumber="322">
<typeStatus id="AE7F883FAA33BD03FB94FB9DFD65FBEE" box="[199,319,1077,1102]" pageId="21" pageNumber="322" type="paratype">Paratypes</typeStatus>
variations.
</emphasis>
Body length
<quantity id="B63C9B78AA33BD03F93AFB9DFF5EFBED" box="[617,772,1077,1102]" metricMagnitude="-2" metricUnit="m" metricValue="1.0399999999999998" metricValueMax="1.16" metricValueMin="0.9199999999999999" pageId="21" pageNumber="322" unit="mm" value="10.399999999999999" valueMax="11.6" valueMin="9.2">9.2–11.6 mm</quantity>
; wing length
<quantity id="B63C9B78AA33BD03F8F6FB9DF81AFBED" box="[933,1088,1077,1102]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" metricValueMax="1.06" metricValueMin="0.96" pageId="21" pageNumber="322" unit="mm" value="10.1" valueMax="10.6" valueMin="9.6">9.6–10.6 mm</quantity>
; wing width 3.0–
<quantity id="B63C9B78AA33BD03FE5EFB9DF933FBED" box="[1293,1385,1077,1101]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="21" pageNumber="322" unit="mm" value="3.2">3.2 mm</quantity>
; F.I. 2.7–2.9; D.I. 1.0–1.1.
</paragraph>
<paragraph id="717B369DAA33BD03FB94FBD3FDD9FB34" blockId="21.[151,1437,151,2024]" box="[199,387,1147,1172]" pageId="21" pageNumber="322">
<emphasis id="43B0EA8FAA33BD03FB94FBD3FD57FB34" bold="true" box="[199,269,1147,1172]" pageId="21" pageNumber="322">Male:</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="39DE6516AA33BD03FB94FB37FE23FB18" box="[199,633,1183,1208]" pageId="21" pageNumber="322" type="distribution">
<paragraph id="717B369DAA33BD03FB94FB37FE23FB18" blockId="21.[151,1437,151,2024]" box="[199,633,1183,1208]" pageId="21" pageNumber="322">
<emphasis id="43B0EA8FAA33BD03FB94FB37FD3AFB18" bold="true" box="[199,352,1183,1208]" pageId="21" pageNumber="322">Distribution.</emphasis>
<collectingCountry id="09D3760DAA33BD03FA34FB37FDF1FB18" box="[359,427,1183,1208]" name="Brazil" pageId="21" pageNumber="322">Brazil</collectingCountry>
(Pará, Rondônia).
</paragraph>
</subSubSection>
<subSubSection id="39DE6516AA33BD03FB94FB6AFEC8F8B3" pageId="21" pageNumber="322" type="materials_examined">
<paragraph id="717B369DAA33BD03FB94FB6AF8ACF96C" blockId="21.[151,1437,151,2024]" pageId="21" pageNumber="322">
<emphasis id="43B0EA8FAA33BD03FB94FB6AFDA2FB7B" bold="true" box="[199,504,1218,1243]" pageId="21" pageNumber="322">
<typeStatus id="AE7F883FAA33BD03FB94FB6AFD58FB7B" box="[199,258,1218,1243]" pageId="21" pageNumber="322">Type</typeStatus>
material.
<typeStatus id="AE7F883FAA33BD03FAD8FB6AFDA2FB7B" box="[395,504,1218,1243]" pageId="21" pageNumber="322" type="holotype">Holotype</typeStatus>
</emphasis>
♀. “
<collectingCountry id="09D3760DAA33BD03F968FB6BFEFFFB7B" box="[571,677,1219,1243]" name="Brazil" pageId="21" pageNumber="322">BRAZIL</collectingCountry>
, Pará\Serra Norte\3 alfa\
<date id="057A105DAA33BD03F884FB6AF811FB7B" box="[983,1099,1218,1243]" pageId="21" pageNumber="322" value="1985-06-26">26.vi.1985</date>
” “
<collectingCountry id="09D3760DAA33BD03FF3CFB6AF8ECFB7B" box="[1135,1206,1218,1243]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
, Pará\M F Torres” (
<collectionCode id="17D5AE58AA33BD03FBCDFB4FFCB4FB5F" box="[158,238,1255,1279]" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="21" pageNumber="322">MPEG</collectionCode>
<collectionCode id="17D5AE58AA33BD03FBA9FB4FFD7DFB5E" box="[250,295,1255,1278]" pageId="21" pageNumber="322">DIP</collectionCode>
12001513).
<emphasis id="43B0EA8FAA33BD03FAECFB4EFE04FB5F" bold="true" box="[447,606,1254,1279]" pageId="21" pageNumber="322">
<typeStatus id="AE7F883FAA33BD03FAECFB4EFE6DFB5F" box="[447,567,1254,1279]" pageId="21" pageNumber="322" type="paratype">Paratypes</typeStatus>
18
</emphasis>
♀. “Pará, Tucuruí\ 08.7[vii].1978” “
<collectingCountry id="09D3760DAA33BD03FF56FB4EF810FB5F" box="[1029,1098,1254,1279]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
Pará\W.
<collectingCountry id="09D3760DAA33BD03FFE9FB4FF953FB5F" box="[1210,1289,1255,1279]" name="France" pageId="21" pageNumber="322">França</collectingCountry>
” “
<taxonomicName id="B6C44D1EAA33BD03FE7BFB4EF9CEFB5F" box="[1320,1428,1254,1279]" class="Insecta" family="Tabanidae" genus="Pityocera" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="genus">Pityocera</taxonomicName>
\ (Pseudelaphella)\
<taxonomicName id="B6C44D1EAA33BD03FA30FAA2FD8BFA82" authority="Wlk" authorityName="Wlk" box="[355,465,1289,1314]" class="Insecta" family="Tabanidae" genus="Pityocera" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="nana">nana Wlk</taxonomicName>
\ det. G.B. Fairchild, 1981” (♀ MPEG-DIP 12006434); “
<collectingCountry id="09D3760DAA33BD03FF33FAA1F8FDFA82" box="[1120,1191,1289,1314]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
, Pará\ Serra Norte\ 3 Alfa\
<date id="057A105DAA33BD03FB8BFA85FD0BFAE6" box="[216,337,1325,1350]" pageId="21" pageNumber="322" value="1985-06-26">26.vi.1985</date>
” “
<collectingCountry id="09D3760DAA33BD03FA3DFA85FDEFFAE6" box="[366,437,1325,1350]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
, Pará\M F Torres” (3♀ MPEG-DIP 12001511, 12001514, 12001515); “
<collectingCountry id="09D3760DAA33BD03FF88FA85F97AFAE6" box="[1243,1312,1325,1350]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
Pará\Serra Norte\Serraria\
<date id="057A105DAA33BD03FA1FFAF8FDE2FAC9" box="[332,440,1360,1385]" pageId="21" pageNumber="322" value="1985-06-15">15.vi.1985</date>
” “
<collectingCountry id="09D3760DAA33BD03FA85FAF8FE47FAC9" box="[470,541,1360,1385]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
, Pará\M. F. Torres” (♀
<collectionCode id="17D5AE58AA33BD03F879FAF9FFDAFAC9" box="[810,896,1361,1385]" pageId="21" pageNumber="322">CZMA</collectionCode>
); “
<collectingCountry id="09D3760DAA33BD03F8CDFAF8FFB9FAC9" box="[926,995,1360,1385]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
Pará\Serra Norte\Serraria\
<date id="057A105DAA33BD03FE4BFAF8F9CBFAC9" box="[1304,1425,1360,1385]" pageId="21" pageNumber="322" value="1985-06-22">22.vi.1985</date>
” “
<collectingCountry id="09D3760DAA33BD03FBF2FADCFCB2FA2D" box="[161,232,1396,1421]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
, Pará\M. Zanuto” (♀ MPEG-DIP 12001518); “
<collectingCountry id="09D3760DAA33BD03F863FADCFF2FFA2D" box="[816,885,1396,1421]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
Pará\Serra Norte\Serraria\
<date id="057A105DAA33BD03FFEEFADCF976FA2D" box="[1213,1324,1396,1421]" pageId="21" pageNumber="322" value="1985-06-27">27.vi.1985</date>
” “
<collectingCountry id="09D3760DAA33BD03FE03FADCF9CDFA2D" box="[1360,1431,1396,1421]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
, Pará\W.
<collectingCountry id="09D3760DAA33BD03FBAAFA30FD12FA10" box="[249,328,1432,1456]" name="France" pageId="21" pageNumber="322">França</collectingCountry>
” (2♀ MPEG-DIP 12001522, 12001525); “
<collectingCountry id="09D3760DAA33BD03F86BFA3FFF27FA10" box="[824,893,1431,1456]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
Pará\Serra Norte\Serraria\
<date id="057A105DAA33BD03FFEBFA3FF90DFA10" box="[1208,1367,1431,1456]" pageId="21" pageNumber="322" value="1985-06-20" valueMax="1985-06-24" valueMin="1985-06-20">20–24.vi.1985</date>
\Arm. Malaise” “
<collectingCountry id="09D3760DAA33BD03FA7AFA13FD2AFA74" box="[297,368,1467,1492]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
, Pará\F. Ramos” (2♀ MPEG-DIP 12001520, 12001526); “
<collectingCountry id="09D3760DAA33BD03FF93FA13F95FFA74" box="[1216,1285,1467,1492]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
Pará\Serra Norte\Serraria\
<date id="057A105DAA33BD03FA19FA76FDE3FA57" box="[330,441,1502,1527]" pageId="21" pageNumber="322" value="1985-06-29">29.vi.1985</date>
” “
<collectingCountry id="09D3760DAA33BD03F957FA76FE11FA57" box="[516,587,1502,1527]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
, Pará\H. Andrade” (♀
<collectionCode id="17D5AE58AA33BD03FF5AFA77F805FA57" box="[1033,1119,1503,1527]" pageId="21" pageNumber="322">CZMA</collectionCode>
); “
<collectingCountry id="09D3760DAA33BD03FFFFFA76F8ABFA57" box="[1196,1265,1502,1527]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
Pará\Serra Norte\Serraria\
<date id="057A105DAA33BD03FA19F9AAFDA8F9BB" box="[330,498,1538,1563]" pageId="21" pageNumber="322" value="1985-08-01" valueMax="1985-08-03" valueMin="1985-08-01">01–03.viii.1985</date>
” “
<collectingCountry id="09D3760DAA33BD03F94EF9AAFE3EF9BB" box="[541,612,1538,1563]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
, Pará\M. Zanuto” (♀ MPEG-DIP 12001510); “
<collectingCountry id="09D3760DAA33BD03FF9EF9AAF948F9BB" box="[1229,1298,1538,1563]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
Pará\Serra Norte\Pojuca\
<date id="057A105DAA33BD03FA6FF98DFD8FF99E" box="[316,469,1573,1598]" pageId="21" pageNumber="322" value="1985-06-20" valueMax="1985-06-24" valueMin="1985-06-20">20–24.vi.1985</date>
\Arm. Malaise” “
<collectingCountry id="09D3760DAA33BD03F9F2F98DFEB2F99E" box="[673,744,1573,1598]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
, Pará\ F. Ramos” (2♀ MPEG-DIP 12001516, 12001519); “
<collectingCountry id="09D3760DAA33BD03FBF2F9E0FCB2F9C1" box="[161,232,1608,1633]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
, Pará\Serra Norte\Fofoca\
<date id="057A105DAA33BD03F94DF9E0FEC2F9C1" box="[542,664,1608,1633]" pageId="21" pageNumber="322" value="1985-06-22">22.vi.1985</date>
” “
<collectingCountry id="09D3760DAA33BD03F9EBF9E0FEA5F9C1" box="[696,767,1608,1633]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
, Pará\N. Bittencourt” (♀ MPEG-DIP 12001517); “
<collectingCountry id="09D3760DAA33BD03FE03F9E0F9CDF9C1" box="[1360,1431,1608,1633]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
, Pará\Serra Norte\Salobo\
<date id="057A105DAA33BD03FAEEF9C4FE67F925" box="[445,573,1644,1669]" pageId="21" pageNumber="322" value="1984-08-05">05.viii.1984</date>
” “
<collectingCountry id="09D3760DAA33BD03F90DF9C4FEFFF925" box="[606,677,1644,1669]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
, Pará\M. Zanuto” (♀ MPEG-DIP 12001524); “
<collectingCountry id="09D3760DAA33BD03FF83F9C4F94DF925" box="[1232,1303,1644,1669]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
, Pará\Serra Norte\N.I.C Rupestre\
<date id="057A105DAA33BD03FACAF927FE51F908" box="[409,523,1679,1704]" pageId="21" pageNumber="322" value="1985-06-20">20.vi.1985</date>
” “
<collectingCountry id="09D3760DAA33BD03F97BF927FE35F908" box="[552,623,1679,1704]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
, Pará\M. Torres” (♀ MPEG-DIP 12001512); “
<collectingCountry id="09D3760DAA33BD03FFD0F927F890F908" box="[1155,1226,1679,1704]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
, Pará\Serra Norte, N1\Arm. Suspensa
<quantity id="B63C9B78AA33BD03FA22F91BFDE0F96C" box="[369,442,1715,1740]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="6.0" metricValueMin="1.0" pageId="21" pageNumber="322" unit="m" value="3.5" valueMax="6.0" valueMin="1.0">1–6 m</quantity>
\
<date id="057A105DAA33BD03FA97F91BFE31F96C" box="[452,619,1715,1740]" pageId="21" pageNumber="322" value="1985-07-12" valueMax="1985-07-15" valueMin="1985-07-12">12–15.vii.1985</date>
” “
<collectingCountry id="09D3760DAA33BD03F9DBF91BFE95F96C" box="[648,719,1715,1740]" name="Brazil" pageId="21" pageNumber="322">Brasil</collectingCountry>
, Pará\R. D. Thomas” (♀ MPEG-DIP 12001523).
</paragraph>
<paragraph id="717B369DAA33BD03FB94F97EFEC8F8B3" blockId="21.[151,1437,151,2024]" pageId="21" pageNumber="322">
<emphasis id="43B0EA8FAA33BD03FB94F97EFE66F94F" bold="true" box="[199,572,1750,1775]" pageId="21" pageNumber="322">Additional material examined:</emphasis>
<collectingCountry id="09D3760DAA33BD03F916F97FFEF7F94F" box="[581,685,1751,1775]" name="Brazil" pageId="21" pageNumber="322">BRAZIL</collectingCountry>
,
<emphasis id="43B0EA8FAA33BD03F9E8F97EFF72F94F" box="[699,808,1750,1775]" italics="true" pageId="21" pageNumber="322">Rondônia</emphasis>
, Ouro Preto do Oeste, Rio Boa Vista,
<date id="057A105DAA33BD03FFB8F97EF9C2F94F" box="[1259,1432,1750,1775]" pageId="21" pageNumber="322" value="1995-07-06" valueMax="1995-07-12" valueMin="1995-07-06">06–12.vii.1995</date>
, J.A. Rafael &amp; J. Vidal (5♀
<collectionCode id="17D5AE58AA33BD03FA94F953FE53F8B3" box="[455,521,1787,1811]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="21" pageNumber="322">INPA</collectionCode>
, ♀
<collectionCode id="17D5AE58AA33BD03F963F953FEDCF8B3" box="[560,646,1787,1811]" pageId="21" pageNumber="322">CZMA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="39DE6516AA33BD03FB94F8B5F9C7F896" box="[199,1437,1821,1846]" pageId="21" pageNumber="322" type="etymology">
<paragraph id="717B369DAA33BD03FB94F8B5F9C7F896" blockId="21.[151,1437,151,2024]" box="[199,1437,1821,1846]" pageId="21" pageNumber="322">
<emphasis id="43B0EA8FAA33BD03FB94F8B5FD14F896" bold="true" box="[199,334,1821,1846]" pageId="21" pageNumber="322">Etymology.</emphasis>
the name derives from Latin and is named for its dark legs;
<emphasis id="43B0EA8FAA33BD03F8BCF8B7F86AF896" box="[1007,1072,1823,1846]" italics="true" pageId="21" pageNumber="322">perna</emphasis>
= legs +
<emphasis id="43B0EA8FAA33BD03FFCBF8B5F8B1F896" box="[1176,1259,1821,1846]" italics="true" pageId="21" pageNumber="322">aquilus</emphasis>
= dark colored.
</paragraph>
</subSubSection>
<subSubSection id="39DE6516AA33BD01FB94F8E9F8B2FEC5" lastPageId="23" lastPageNumber="324" pageId="21" pageNumber="322" type="discussion">
<paragraph id="717B369DAA33BD03FB94F8E9FCBAF848" blockId="21.[151,1437,151,2024]" pageId="21" pageNumber="322">
<emphasis id="43B0EA8FAA33BD03FB94F8E9FD16F8FA" bold="true" box="[199,332,1857,1882]" pageId="21" pageNumber="322">Discussion.</emphasis>
here described as a new species, but previously treated as the female of
<taxonomicName id="B6C44D1EAA33BD03FFDDF8EAF8B9F8F9" box="[1166,1251,1858,1881]" class="Insecta" family="Tabanidae" genus="Pityocera" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="nana">
<emphasis id="43B0EA8FAA33BD03FFDDF8EAF8B9F8F9" box="[1166,1251,1858,1881]" italics="true" pageId="21" pageNumber="322">P. nana</emphasis>
</taxonomicName>
in
<bibRefCitation id="15554B6CAA33BD03FE59F8E9FD13F8DD" author="Henriques" pageId="21" pageNumber="322" refString="Henriques, A. L. &amp; Gorayeb, I. S. (1993) A colecao de Tabanidae (Insecta: Diptera) do Museu Paraense Emilio Goeldi (MPEG), Belem, Para, Brasil. Goeldiana Zoologia, 20, 1 - 23." type="journal article" year="1993">Henriques &amp; Gorayeb (1993)</bibRefCitation>
and
<bibRefCitation id="15554B6CAA33BD03FA2CF8CCFE1DF8DD" author="Henriques" box="[383,583,1892,1917]" pageId="21" pageNumber="322" refString="Henriques, A. L. (1997) A colecao de Tabanidae (Insecta: Diptera) do Instituto Nacional de Pesquisas da Amazonia (INPA), Manaus, Amazonas, Brasil. Boletim do Museu Paraense Emilio Goeldi, serie Zoologia, 11 (1), 57 - 99. [1995]" type="journal article" year="1997">Henriques (1997)</bibRefCitation>
. This mistake is possibly related to the fact that a specimen from the MPEG was identified as such by Fairchild in 1981. Such occurrence raises the possibility that
<taxonomicName id="B6C44D1EAA33BD03FF36F821F8E2F800" box="[1125,1208,1929,1952]" class="Insecta" family="Tabanidae" genus="Pityocera" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="nana">
<emphasis id="43B0EA8FAA33BD03FF36F821F8E2F800" box="[1125,1208,1929,1952]" italics="true" pageId="21" pageNumber="322">P. nana</emphasis>
</taxonomicName>
may also have been misidentified in other works (
<bibRefCitation id="15554B6CAA33BD03FABFF803FEA1F864" author="Fairchild" box="[492,763,1963,1988]" pageId="21" pageNumber="322" refString="Fairchild, G. B. &amp; Leon, L. A. (1986) Lista provisional de Tabanidae (Diptera) del Ecuador. Publicaciones del Museo Ecuatoriano de Ciencias Naturales, 5, 97 - 122." type="journal article" year="1986">Fairchild &amp; León 1986</bibRefCitation>
;
<bibRefCitation id="15554B6CAA33BD03F85AF803FFAAF864" author="Chainey" box="[777,1008,1963,1988]" pageId="21" pageNumber="322" refString="Chainey, J. E., Hall, M. J. R., Amayo, J. L. &amp; Bettella, P. (1994) A preliminary checklist and key to the genera and subgenera of Tabanidae (Diptera) of Bolivia with particular reference to Santa Cruz Department. Memorias do Instituto Oswaldo Cruz, 89, 321 - 345. http: // dx. doi. org / 10.1590 / s 0074 - 02761994000300007" type="journal article" year="1994">
Chainey
<emphasis id="43B0EA8FAA33BD03F823F805FFFEF864" box="[880,932,1963,1988]" italics="true" pageId="21" pageNumber="322">et al</emphasis>
. 1994
</bibRefCitation>
;
<bibRefCitation id="15554B6CAA33BD03F8ADF804F8BBF864" author="Buestan" box="[1022,1249,1963,1988]" pageId="21" pageNumber="322" refString="Buestan J. A., Navarrete, R. A. &amp; Mejia, M. T. (2007) Lista actualizada de tabanos (Diptera: Tabanidae) del Ecuador. Revista Ecuatoriana de Higiene y Medicina Tropical, 44 (1), 23 - 78." type="journal article" year="2007">
Buestán
<emphasis id="43B0EA8FAA33BD03FF31F805F8C7F864" box="[1122,1181,1963,1988]" italics="true" pageId="21" pageNumber="322">et al.</emphasis>
2007
</bibRefCitation>
;
<bibRefCitation id="15554B6CAA33BD03FFBCF803FC8EF848" author="Cardenas" pageId="21" pageNumber="322" refString="Cardenas, R. E., Buestan, J. A. &amp; Dangles, O. (2009) Diversity and distribution models of horse flies (Diptera: Tabanidae) from Ecuador. Annales de la Societe Entomologique de France (Nouvelle Serie), 45 (4), 511 - 528. http: // dx. doi. org / 10.1080 / 00379271.2009.10697633" type="journal article" year="2009">
Cárdenas
<emphasis id="43B0EA8FAA33BD03FE32F805F9C6F864" box="[1377,1436,1963,1988]" italics="true" pageId="21" pageNumber="322">et al.</emphasis>
2009
</bibRefCitation>
).
</paragraph>
<caption id="25BB6615AA30BD00FBC4F80AFEFDF856" httpUri="https://zenodo.org/record/253426/files/figure.png" pageId="22" pageNumber="323" targetBox="[192,1398,208,1916]" targetPageId="22">
<paragraph id="717B369DAA30BD00FBC4F80AFEFDF856" blockId="22.[151,1436,1954,2038]" pageId="22" pageNumber="323">
<emphasis id="43B0EA8FAA30BD00FBC4F80AFD79F818" bold="true" box="[151,291,1954,1976]" pageId="22" pageNumber="323">FIGURE 12.</emphasis>
<taxonomicName id="B6C44D1EAA30BD00FA7EF80BFEE5F818" box="[301,703,1954,1976]" class="Insecta" family="Tabanidae" genus="Pityocera" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="pernaquila" status="sp. nov." subGenus="Pseudelaphella">
<emphasis id="43B0EA8FAA30BD00FA7EF80BFDCAF818" box="[301,400,1955,1976]" italics="true" pageId="22" pageNumber="323">Pityocera</emphasis>
(
<emphasis id="43B0EA8FAA30BD00FAF1F80AFE64F818" box="[418,574,1954,1976]" italics="true" pageId="22" pageNumber="323">Pseudelaphella</emphasis>
)
<emphasis id="43B0EA8FAA30BD00F903F80AFEE5F818" box="[592,703,1954,1976]" italics="true" pageId="22" pageNumber="323">pernaquila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="588357F4AA30BD00F99AF80AFF47F818" box="[713,797,1954,1976]" pageId="22" pageNumber="323" rank="species">
<emphasis id="43B0EA8FAA30BD00F99AF80AFF47F818" bold="true" box="[713,797,1954,1976]" pageId="22" pageNumber="323">sp. nov.</emphasis>
</taxonomicNameLabel>
♀ (A–B Holotype, C–F Paratype). A—habitus dorsal view; B—head front-lateral view; C—antenna lateral view; D—tergites 9 + 10, cerci and hypoproct dorsal view; E—hypogynium + hypogynial valve; F—genital fork and spermatheca.
</paragraph>
</caption>
<paragraph id="717B369DAA31BD01FB94FF3FF8B2FEC5" blockId="23.[151,1436,151,357]" pageId="23" pageNumber="324">
This species can be differentiated from the others by its habitus characteristically darker than other
<emphasis id="43B0EA8FAA31BD01FBC4FF14FD19FF75" box="[151,323,188,213]" italics="true" pageId="23" pageNumber="324">Pseudelaphella</emphasis>
, associated with dark brown legs with predominantly black pilosity. It shows substantial color variation, in that the abdomens of specimens collected in Pará have the two first tergites yellow with a black spot in the center, while specimens from Rondônia have all tergites dark brown. Legs and antenna are also darker in the specimens from Rondônia. Therefore, specimens from Rondônia were left out of the
<typeStatus id="AE7F883FAA31BD01FF20FE80F8F9FEE0" box="[1139,1187,296,320]" pageId="23" pageNumber="324">type</typeStatus>
series, given that the addition of new material and/or males can define the unification or separation of these two groups.
</paragraph>
</subSubSection>
</treatment>
</document>
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