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<document id="DA88273D23761084411A1BB597258F3C" ENCODING="UTF-8" ID-GBIF-Dataset="d42acefb-3c32-4b96-8400-e1575aa17bc6" ModsDocID="z01578p041" checkinTime="1247322647604" checkinUser="thomas" docAuthor="Paul H. Skelton" docDate="2007" docId="E993771A711B8425B0E1BA124707B4C7" docLanguage="en" docName="2007_Skelton_gg1.xml" docOrigin="Zootaxa 1578" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:91D85038-3B5D-4493-AE8C-6D2ECA205CAA" docTitle="Amphilius dimonikensis Skelton, 2007, sp. nov." docType="treatment" docVersion="11" lastPageNumber="45" masterDocId="69594900A1F947C1E217DCD0031C8796" masterDocTitle="New species of the amphiliid catfish genera Amphilius, Doumea and Phractura and the taxonomy of Paramphilius from West Central Africa (Siluriformes, Amphiliidae)." masterLastPageNumber="68" masterPageNumber="41" pageNumber="44" updateTime="1701314704579" updateUser="plazi">
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<mods:title id="70643D9BFF1B157FCDC2F8298C13DFD3">New species of the amphiliid catfish genera Amphilius, Doumea and Phractura and the taxonomy of Paramphilius from West Central Africa (Siluriformes, Amphiliidae).</mods:title>
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<mods:namePart id="49E4CE2C5043F1E6C056C5730CA3F6AF">Paul H. Skelton</mods:namePart>
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<mods:date id="A5DDBBE3A017B545BA7CC98DDC510C67">2007</mods:date>
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<treatment id="E993771A711B8425B0E1BA124707B4C7" ID-DOI="http://doi.org/10.5281/zenodo.6236486" ID-GBIF-Taxon="100121957" ID-Zenodo-Dep="6236486" LSID="urn:lsid:plazi:treatment:E993771A711B8425B0E1BA124707B4C7" httpUri="http://treatment.plazi.org/id/E993771A711B8425B0E1BA124707B4C7" lastPageNumber="45" pageNumber="44">
<subSubSection id="195106449DF24CF0C6D524FDA5E34860" type="nomenclature">
<paragraph id="2E9439A0A2F637AF4FC0B58CFF12BCF1" pageNumber="44">
<taxonomicName id="B38E289DB8AD7EB340ECF70CB9C0497B" ID-CoL="66FQY" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" order="Siluriformes" pageNumber="44" phylum="Chordata" rank="species" species="dimonikensis">Amphilius dimonikensis</taxonomicName>
<taxonomicNameLabel id="B7AEB65E1D55255F964C949E0D619A52" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C46AB62DADC23B369BC2E7B97CA6D4DF" type="description">
<paragraph id="D0355B4EABA94B88470B687D38684672" pageNumber="44">(Fig. 2)</paragraph>
</subSubSection>
<subSubSection id="24AF7114F66846A9376E1CDC4F82AB26" type="reference_group">
<paragraph id="6D08185B3B3871C630166399D0A439DC" pageNumber="44">
Synonymy.
<taxonomicName id="6196074EE1E02A462222B816431CE24B" LSID-ZBK="urn:lsid:zoobank.org:act:BAC3CEE9-9BBC-4F7F-81FF-9C3D636F655B" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" lsidName-ZBK="Amphilius pulcher Pellegrin 1929:640" order="Siluriformes" pageNumber="44" phylum="Chordata" rank="species" species="pulcher">Amphilius pulcher</taxonomicName>
, non Pellegrin; Mamonekene &amp; Teugels 1993, 44-45.
</paragraph>
</subSubSection>
<subSubSection id="C59E887BBACB89BF1B694BFA1B886A1B" type="nomenclature">
<paragraph id="AC54B64DB3A33788AEF493AC11885622" pageNumber="44">
Type Material.
<materialsCitation id="061B3E47186C8E2A5609A61D51D43DED" ID-GBIF-Occurrence="923906130" country="Republic Of Congo">
<typeStatus id="4F850309C68B9A3C513CE7A388E2E60C">Holotype</typeStatus>
,
<collectionCode id="25E62EAE5DDCA6001561A1E1D2190B5D" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale">MRAC</collectionCode>
92-125-P-324, SL 55.6 mm, Mpoulou River, Kouilou River system, at confluence with Kibouba above the Divenda rapids,
<collectingCountry id="FFAC3252B9FC2E7BE4B21F20758F73F1">Republic of Congo</collectingCountry>
,
<geoCoordinate id="E12AAE150A15F2D4B5F40959C44C55C7" direction="south" orientation="latitude" value="-4.0333333">04° 02' S</geoCoordinate>
,
<geoCoordinate id="64C66C699FD67BAF4A0809FD2144104A" direction="east" orientation="longitude" value="12.35">12° 21 ' E</geoCoordinate>
, V Mamonekene,
<date id="6132FB87A9BC70000E121A678C326DB1" day="09" month="07" value="1992-07-09" year="1992">9 July 1992</date>
</materialsCitation>
;
<materialsCitation id="DAC1DA1F55B2FC4A17391A02CB3A5165" ID-GBIF-Occurrence="923906124" country="Republic Of Congo">
5
<typeStatus id="2CCD6C3A1905D157C4F57F26906CF84C">paratypes</typeStatus>
: 1,
<collectionCode id="982F2D531AA77F68D6EE37AD38133011" collectionName="SAIAB">SAIAB</collectionCode>
79509 (ex MRAC 92-125-P-325), SL 43.5 mm male, same data as holotype
</materialsCitation>
;
<materialsCitation id="BF7A3E5CD0B8E51F09ED77EAB9D9C023" ID-GBIF-Occurrence="923906123" country="Republic Of Congo">
2,
<collectionCode id="61AE0350403A62FF38262562CA294B42" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale">MRAC</collectionCode>
92-125-P-326-327, SL 26.6 mm, 34.2 mm, not sexed, Mpoulou River, tributary of Kouilou River, at confluence of Kiada,
<collectingCountry id="98126B6F0A40D7822AA160CD18CC324B">Republic of Congo</collectingCountry>
,
<geoCoordinate id="F6DB94083FD9B98ADC501631506BEB96" direction="south" orientation="latitude" value="-4.0333333">04°02' S</geoCoordinate>
, \2°2Y E, V.Mamonekene,
<date id="2F42A621D0DA1E49A9C768B1B54EA7D0" day="09" month="08" value="1992-08-09" year="1992">9 August 1992</date>
</materialsCitation>
;
<materialsCitation id="1AA166359CC7AC12D7BD04C5BE4D74E9" ID-GBIF-Occurrence="923906122" country="Republic Of Congo">
2,
<collectionCode id="FF9C5E1BCB0D169102209F160B9824E8" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale">MRAC</collectionCode>
92-125-P-328-329, SL, 21 mm, Kiadi River, tributary of Mpoulou, Kouilou River system, at the confluence,
<collectingCountry id="BE679623E78E6986C0C8351ED84CDDBA">Republic of Congo</collectingCountry>
,
<geoCoordinate id="D117126E8088A6E80EB1E46F43E176F1" direction="south" orientation="latitude" value="-4.0333333">04°02' S</geoCoordinate>
,
<geoCoordinate id="4D10896B42CC7D8140D26DB4106D0A7F" direction="east" orientation="longitude" value="12.35">12°21' E</geoCoordinate>
, V. Mamonekene,
<date id="5282089CD1431CFC9984B86AB2B35D57" day="09" month="08" value="1992-08-09" year="1992">9 August 1992</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3A0DAFC51755F4D5AC29229B7B60CCAC" type="diagnosis">
<paragraph id="56A3C3951E8BB06B39AB7463148BDDD5" lastPageNumber="45" pageNumber="44">
Diagnosis. A combination of characters separate this species from other
<taxonomicName id="0614CE8CD59E2EE9A8F07FC1D445DEFF" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" order="Siluriformes" pageNumber="44" phylum="Chordata" rank="genus">Amphilius</taxonomicName>
. It is one of several
<taxonomicName id="5C4C4A519CFA94A097883059B908C659" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" order="Siluriformes" pageNumber="44" phylum="Chordata" rank="genus">Amphilius</taxonomicName>
characterized by 6+7 principal caudal rays. This character separates it from several species with 8+9 rays found in East and southern Africa (Skelton 1984) and from
<taxonomicName id="4128C97CC8F05E0B33172457ADFEB371" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" order="Siluriformes" pageNumber="44" phylum="Chordata" rank="species" species="longirostris">A. longirostris</taxonomicName>
(with 7+8 principal caudal rays) in the WCA. The position of the eyes is slightly closer the hind margin of the head than the tip of the snout, similar to
<taxonomicName id="79AFDDF3260793CAB50C4ADDEF6611EC" LSID-ZBK="urn:lsid:zoobank.org:act:BAC3CEE9-9BBC-4F7F-81FF-9C3D636F655B" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" lsidName-ZBK="Amphilius pulcher Pellegrin 1929:640" order="Siluriformes" pageNumber="44" phylum="Chordata" rank="species" species="pulcher">A. pulcher</taxonomicName>
but a feature that distinguishes it from other species in the region with 6+7 caudal rays. In the WCA
<taxonomicName id="AB7AA6B166B9473E6AFF404892EF0341" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" order="Siluriformes" pageNumber="44" phylum="Chordata" rank="species" species="dimonikensis">A. dimonikensis</taxonomicName>
is most similar to
<taxonomicName id="4F5381CABFA8A4B6E0CB854B1901FC90" LSID-ZBK="urn:lsid:zoobank.org:act:BAC3CEE9-9BBC-4F7F-81FF-9C3D636F655B" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" lsidName-ZBK="Amphilius pulcher Pellegrin 1929:640" order="Siluriformes" pageNumber="44" phylum="Chordata" rank="species" species="pulcher">A. pulcher</taxonomicName>
, from which it differs most markedly in colour pattern (a regular pattern of brown bands and lighter saddles versus an irregular pattern of striking dark brown or black patches for the latter), as well as in the size of the adipose fin (adipose length = 21% SL vs. 34 % in
<taxonomicName id="C8059FD33740282C015BEDD92C02C58D" LSID-ZBK="urn:lsid:zoobank.org:act:BAC3CEE9-9BBC-4F7F-81FF-9C3D636F655B" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" lsidName-ZBK="Amphilius pulcher Pellegrin 1929:640" order="Siluriformes" pageNumber="44" phylum="Chordata" rank="species" species="pulcher">A. pulcher</taxonomicName>
).
<taxonomicName id="1267759B6DDAC198432AE4E7F28B457F" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" order="Siluriformes" pageNumber="44" phylum="Chordata" rank="species" species="dimonikensis">A. dimonikensis</taxonomicName>
has fewer vertebrae (31-32 total vertebrae vs. 33-34) particularly fewer caudal vertebrae (17-18 vs.19-20) than
<taxonomicName id="C3934930E345F5DF357A62029990CB4D" LSID-ZBK="urn:lsid:zoobank.org:act:BAC3CEE9-9BBC-4F7F-81FF-9C3D636F655B" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" lsidName-ZBK="Amphilius pulcher Pellegrin 1929:640" order="Siluriformes" pageNumber="44" phylum="Chordata" rank="species" species="pulcher">A. pulcher</taxonomicName>
. It differs in pigment pattern from the species described below as
<taxonomicName id="4A555741FECD49440336ADB4FB7EBDAB" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" order="Siluriformes" pageNumber="44" phylum="Chordata" rank="species" species="caudosignatus">Amphilius caudosignatus</taxonomicName>
,
<taxonomicName id="A8CE4DD15EB18080C09451C7894CE04C" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" order="Siluriformes" pageNumber="44" phylum="Chordata" rank="species" species="korupi">Amphilius korupi</taxonomicName>
,
<taxonomicName id="1B47193F340E85649862D251DA819B36" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" order="Siluriformes" pageNumber="44" phylum="Chordata" rank="species" species="mamonekenensis">Amphilius mamonekenensis</taxonomicName>
as well as from
<taxonomicName id="6141409DB03EFC0E2E0E0C97892DF120" LSID-ZBK="urn:lsid:zoobank.org:act:47363F73-A16A-4B41-B358-C4D7C8A73480" class="Actinopterygii" family="Amphiliidae" genus="Amphilius" kingdom="Animalia" lsidName-ZBK="Amphilius nigricaudatus Pellegrin 1909:67" order="Siluriformes" pageNumber="44" phylum="Chordata" rank="species" species="nigricaudatus">Amphilius nigricaudatus</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="9DB04E62C8AED5B9C95A58CF3ED511EA" type="description">
<paragraph id="036C026BD6216E1820CACBA01757B627" pageNumber="45">Description. Measurements and fin ray counts given in Table 1. Body stout, depressed anteriorly, width tapering to a compressed caudal peduncle. Head length 3.5-3.9 times in SL. Head depressed and rounded anteriorly; branchiostegal membrane deeply cleft mid-ventrally; mouth broad, subterminal, lips thick and fleshy; premaxillary teeth caniniform in a medially broad and laterally tapered band and mandibular teeth caniniform in a narrow band; 3 pairs of filamentous barbels, maxillary barbels subequal to head length extending from the upper lips; mandibular barbels on ventral surface of mandible, inner positioned latero-medially behind anterior edge of jaw, outer mandibular pair postero-lateral to inner pair; snout moderately long, half of head length, nares in mid-snout, narrowly separated; eyes about midway or slightly nearer hind margin of head, moderate sized, closely spaced (interorbit 1.2 to 1.9 times orbit diameter), confluent with head (orbital margins attached). Caudal peduncle short (5.9-7.1 times in SL), slightly longer than deep (length 1.2-1.5 times depth). Fins well developed, dorsal base entirely in advance of pelvic fins, origin less than one head length behind head, rectangular with leading simple ray equal to length of leading edge; pectoral fins not reaching pelvic fins, rounded with leading ray curved, padded and pectinate, innermost (posterior) rays align against lateral body wall; pelvic fins broad, with padded pectinate leading ray, not reaching base of anal; anal fin short, not reaching base of caudal; caudal fin with simple epidermal fold over base of rays, hind edge emarginate, ventral lobe slightly longer than dorsal lobe; adipose fin well developed, rounded, base extends from before origin of anal to base of caudal procurrent ridge. Skin smooth, lateral line complete along midline to base of caudal fin. Anus positioned about one third of distance from base of pelvic fins to origin of anal fin. Size reached 55.6 mm SL.</paragraph>
<paragraph id="C2DA4A53D470004ED7C7348FBEBDB0B8" pageNumber="45">Coloration. Head and body dark brown with 4-5 light (creamish) saddle bands on dorsal surface, and a light (off-white) ventral surface from head to anal fin. Fins with brown rays; caudal fin with broad basal dark brown band and a distal dark brown band.</paragraph>
</subSubSection>
<subSubSection id="EDC63D1F47B48076056EB62CEF076380" type="distribution">
<paragraph id="466E973A0BF026B9591E151D32E8042C" pageNumber="45">Distribution. Known only from the Mpoulou River, a tributary of the Kouilou River within the Dimonika Biosphere Reserve, Mayombe, Republic of Congo (Fig. 3).</paragraph>
</subSubSection>
<subSubSection id="2EC973C794213906B1AE1AFB127ADE06" type="etymology">
<paragraph id="2CFC404E5DD0E04F3C65AA040EDEFDEA" pageNumber="45">Etymology. The name dimonikensis refers to the Dimonika Biosphere Reserve, Mayombe, Republic of Congo, from where this species is likely to be endemic.</paragraph>
</subSubSection>
</treatment>
</document>