285 lines
48 KiB
XML
285 lines
48 KiB
XML
<document id="9085D37AB409FC98181B67B03BC8EE1F" ID-CLB-Dataset="2709" ID-DOI="10.11646/zootaxa.5087.4.5" ID-GBIF-Dataset="8b4568fc-0433-401b-9786-63ddc003aa82" ID-ISSN="1175-5326" ID-Zenodo-Dep="5832883" ID-ZooBank="68B99D6A-0E64-466F-8D19-FCBDDBD8DBA5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641804066547" checkinUser="plazi" docAuthor="Szawaryn, Karol & Churata-Salcedo, Julissa" docDate="2022" docId="C5598790FFC63A32FF3DF9C3EEEFF88D" docLanguage="en" docName="zootaxa.5087.4.5.pdf" docOrigin="Zootaxa 5087 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Merma quadriguttata Szawaryn & Churata-Salcedo 2022, comb. nov." docType="treatment" docVersion="8" lastPageNumber="578" masterDocId="3960FFE8FFC33A35FFAAFFF1EC44FFFC" masterDocTitle="Revision of Malagasy species of the genus Merma Weise (Coccinellidae: Epilachnini)" masterLastPageNumber="582" masterPageNumber="571" pageNumber="576" updateTime="1699297914291" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="EABEBD6B8E3AF70A643C2446308AEFC3">
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<mods:title id="C2A73679979B54651B4314E4A6F75318">Revision of Malagasy species of the genus Merma Weise (Coccinellidae: Epilachnini)</mods:title>
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<mods:name id="F8B3DDEF42B786B516182071848B61A6" type="personal">
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<mods:roleTerm id="0FC33245ADCB1163C09B6D3781B6DE86">Author</mods:roleTerm>
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<mods:namePart id="F7C2DD5C4FE6BAC4C2B98509B5A82FCB">Szawaryn, Karol</mods:namePart>
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<mods:affiliation id="C338DDA66A82C303C3A32887E0816327">Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679, Warsaw, Poland.</mods:affiliation>
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<mods:name id="0D2E56771CC4EF7E361C248B4DD87793" type="personal">
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<mods:roleTerm id="CE525414A94BB486B1693A9BA9D80840">Author</mods:roleTerm>
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<mods:namePart id="25295CEB77E09FA68E68316BF5BE271A">Churata-Salcedo, Julissa</mods:namePart>
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<mods:affiliation id="9498FD99AAFF1ACB29A705858ED2F016">Laboratório de Sistemática e Bioecologaia de Coleoptera, Department of Zoology, Universidade Federal do Paraná, Caixa Postal 19030, 81581 - 980, Curitiba, Paraná, Brazil.</mods:affiliation>
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<mods:title id="2D559237B4FA1E2EEE12617753670890">Zootaxa</mods:title>
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<mods:part id="34C801E68D350D7399C437D75163DF71">
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<mods:date id="E31C64F8D3648433E135789D86CE4D8E">2022</mods:date>
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<mods:number id="B77D3F6628CA7B64CFAE474E3BC0AC13">2022-01-10</mods:number>
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<subSubSection id="05EA650DFFC63A30FF3DF9C3EE80F9B7" box="[151,708,1585,1612]" pageId="5" pageNumber="576" type="nomenclature">
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<paragraph id="4D4F3686FFC63A30FF3DF9C3EE80F9B7" blockId="5.[151,708,1585,1647]" box="[151,708,1585,1612]" pageId="5" pageNumber="576">
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<heading id="160781EAFFC63A30FF3DF9C3EE80F9B7" bold="true" box="[151,708,1585,1612]" fontSize="11" level="1" pageId="5" pageNumber="576" reason="1">
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<emphasis id="7F84EA94FFC63A30FF3DF9C3EE80F9B7" bold="true" box="[151,708,1585,1612]" pageId="5" pageNumber="576">
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<taxonomicName id="8AF04D05FFC63A30FF3DF9C3EE79F9B7" authority="(Sicard, 1909)" authorityName="Szawaryn & Churata-Salcedo" authorityYear="2022" baseAuthorityName="Sicard" baseAuthorityYear="1909" box="[151,573,1585,1612]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="quadriguttata" status="comb. nov.">
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<emphasis id="7F84EA94FFC63A30FF3DF9C3EDCBF9B0" bold="true" box="[151,399,1585,1612]" italics="true" pageId="5" pageNumber="576">Merma quadriguttata</emphasis>
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(
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<bibRefCitation id="29614B77FFC63A30FE34F9C0EE70F9B7" author="Sicard, A." box="[414,564,1585,1611]" pageId="5" pageNumber="576" pagination="63 - 165" refId="ref5633" refString="Sicard, A. (1909) Revision des Coccinellides de la faune Malgache. II. Coccinellidae Aphidiphagae. Annales de la Societe Entomologique de France, 78, 63 - 165." type="journal article" year="1909">Sicard, 1909</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="64B757EFFFC63A30FDEEF9C0EE80F9B7" box="[580,708,1585,1611]" pageId="5" pageNumber="576" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="05EA650DFFC63A30FF3DF9A4ECBCF993" box="[151,248,1621,1647]" pageId="5" pageNumber="576" type="description">
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<paragraph id="4D4F3686FFC63A30FF3DF9A4ECBCF993" blockId="5.[151,708,1585,1647]" box="[151,248,1621,1647]" pageId="5" pageNumber="576">
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<figureCitation id="D5CB2A03FFC63A30FF3DF9A4ECBCF993" box="[151,248,1621,1647]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="6.[151,250,1594,1618]" captionStartId-1="8.[151,250,1943,1967]" captionStartId-2="9.[151,250,1505,1529]" captionTargetBox-0="[191,1392,321,1555]" captionTargetBox-1="[178,1409,203,1919]" captionTargetBox-2="[151,1435,181,1480]" captionTargetId-0="figure-88@6.[180,1408,308,1570]" captionTargetId-1="figure-29@8.[178,1409,181,1919]" captionTargetId-2="figure-17@9.[151,1435,181,1481]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURE 4. Merma quadriguttata (Sicard, 1909), holotype (MNHN), female: A—habitus, dorsal, B—habitus, frontal, C—habitus, lateral, D—holotype label data, E—abdomen, F—female genitalia." captionText-1="FIGURE 5. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, frontal, B—habitus, lateral, C—habitus, dorsal, D—abdomen, male, E—penis, lateral, F—tegmen, inner, G—tegmen, lateral, H—sternite VIII, male, I—tergite VIII, male, J—male genital segments IX and X." captionText-2="FIGURE 6. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, ventral, B—mouthparts, C—head, prosternum and mesoventrite, D—legs, meso and metaventrite, E—tarsal claws." figureDoi-0="http://doi.org/10.5281/zenodo.5832891" figureDoi-1="http://doi.org/10.5281/zenodo.5832893" figureDoi-2="http://doi.org/10.5281/zenodo.5832895" httpUri-0="https://zenodo.org/record/5832891/files/figure.png" httpUri-1="https://zenodo.org/record/5832893/files/figure.png" httpUri-2="https://zenodo.org/record/5832895/files/figure.png" pageId="5" pageNumber="576">Figs 4–6</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="05EA650DFFC63A30FF3DF96BEF06F94E" box="[151,834,1690,1714]" pageId="5" pageNumber="576" type="reference_group">
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<paragraph id="4D4F3686FFC63A30FF3DF96BEF06F94E" blockId="5.[151,834,1690,1714]" box="[151,834,1690,1714]" pageId="5" pageNumber="576">
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<treatmentCitationGroup id="6DE011A8FFC63A30FF3DF96BEF06F94E" box="[151,834,1690,1714]" pageId="5" pageNumber="576">
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<treatmentCitation id="CC511097FFC63A30FF3DF96BEE72F94E" author="Sicard, A." box="[151,566,1690,1714]" page="114" pageId="5" pageNumber="576" year="1909">
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<taxonomicName id="8AF04D05FFC63A30FF3DF96BEE72F94E" authority="Sicard, 1909: 114" authorityName=", Sicard" authorityPageNumber="114" authorityYear="1909" box="[151,566,1690,1714]" class="Insecta" family="Coccinellidae" genus="Peralda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="quadriguttata">
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<emphasis id="7F84EA94FFC63A30FF3DF96BED3EF94E" box="[151,378,1690,1714]" italics="true" pageId="5" pageNumber="576">Peralda quadriguttata</emphasis>
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<bibRefCitation id="29614B77FFC63A30FE2BF96BEE72F94E" author="Sicard, A." box="[385,566,1690,1714]" pageId="5" pageNumber="576" pagination="63 - 165" refId="ref5633" refString="Sicard, A. (1909) Revision des Coccinellides de la faune Malgache. II. Coccinellidae Aphidiphagae. Annales de la Societe Entomologique de France, 78, 63 - 165." type="journal article" year="1909">Sicard, 1909: 114</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<typeStatus id="924B8824FFC63A30FD95F96BEE35F94E" box="[575,625,1690,1714]" pageId="5" pageNumber="576">Type</typeStatus>
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locality: Ambositra.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="05EA650DFFC63A30FF3DF911EE79F80A" pageId="5" pageNumber="576" type="diagnosis">
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<paragraph id="4D4F3686FFC63A30FF3DF911EE79F80A" blockId="5.[151,1437,1760,2038]" pageId="5" pageNumber="576">
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<emphasis id="7F84EA94FFC63A30FF3DF911ED56F906" bold="true" box="[151,274,1760,1786]" pageId="5" pageNumber="576">Diagnosis.</emphasis>
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Colouration of
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<taxonomicName id="8AF04D05FFC63A30FE6DF910EEFEF906" authorityName="Szawaryn & Churata-Salcedo" authorityYear="2022" baseAuthorityName="Sicard" baseAuthorityYear="1909" box="[455,698,1760,1786]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="quadriguttata">
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<emphasis id="7F84EA94FFC63A30FE6DF910EEFEF906" box="[455,698,1760,1786]" italics="true" pageId="5" pageNumber="576">Merma quadriguttata</emphasis>
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</taxonomicName>
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is identical to
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<taxonomicName id="8AF04D05FFC63A30FCC9F910EF98F906" authorityName="Szawaryn & Churata-Salcedo" authorityYear="2022" baseAuthorityName="Sicard" baseAuthorityYear="1907" box="[867,988,1760,1786]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="hovana">
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<emphasis id="7F84EA94FFC63A30FCC9F910EF98F906" box="[867,988,1760,1786]" italics="true" pageId="5" pageNumber="576">M. hovana</emphasis>
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</taxonomicName>
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, however, its body is smaller and more elongate, with lateral elytral margin not visible from above, moreover, it has terminal labial palpomere elongate, parallel sided, hypomeron and prosternum covered with distinct large punctae, elytral epipleuron narrow, less than two times as broad as width of metanepisternum, median part of metaventrite covered with large punctae, abdominal postcoxal lines distinctly recurved laterally, and male genitalia lightly sclerotized with penis guide simple with grooves in apical part, and penis long, strongly sinuate with long apical flagellum. From the African species that also have four maculae on elytra it can be distinguished by distinctly smaller spots, while in African species they occupy at least half of elytral surface.
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</paragraph>
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</subSubSection>
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<subSubSection id="05EA650DFFC53A32FF6DFF67E844FA19" lastPageId="7" lastPageNumber="578" pageId="6" pageNumber="577" type="description">
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<paragraph id="4D4F3686FFC53A33FF6DFF67EF8BFEE0" blockId="6.[151,1436,150,285]" pageId="6" pageNumber="577">
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<emphasis id="7F84EA94FFC53A33FF6DFF67ED1DFF4C" bold="true" box="[199,345,150,176]" pageId="6" pageNumber="577">Description.</emphasis>
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Length
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<quantity id="8A089B63FFC53A33FE1FFF67EE48FF4D" box="[437,524,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="6" pageNumber="577" unit="mm" value="3.3">3.3 mm</quantity>
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, width
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<quantity id="8A089B63FFC53A33FDF7FF67EEF1FF4D" box="[605,693,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="6" pageNumber="577" unit="mm" value="2.6">2.6 mm</quantity>
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. Body (
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<figureCitation id="D5CB2A03FFC53A33FCA0FF67EFCCFF4C" box="[778,904,150,176]" captionStart="FIGURE 4" captionStartId="6.[151,250,1594,1618]" captionTargetBox="[191,1392,321,1555]" captionTargetId="figure-88@6.[180,1408,308,1570]" captionTargetPageId="6" captionText="FIGURE 4. Merma quadriguttata (Sicard, 1909), holotype (MNHN), female: A—habitus, dorsal, B—habitus, frontal, C—habitus, lateral, D—holotype label data, E—abdomen, F—female genitalia." figureDoi="http://doi.org/10.5281/zenodo.5832891" httpUri="https://zenodo.org/record/5832891/files/figure.png" pageId="6" pageNumber="577">Figs 4A–C</figureCitation>
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,
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<figureCitation id="D5CB2A03FFC53A33FC39FF67EF93FF4D" box="[915,983,150,177]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[178,1409,203,1919]" captionTargetId="figure-29@8.[178,1409,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, frontal, B—habitus, lateral, C—habitus, dorsal, D—abdomen, male, E—penis, lateral, F—tegmen, inner, G—tegmen, lateral, H—sternite VIII, male, I—tergite VIII, male, J—male genital segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.5832893" httpUri="https://zenodo.org/record/5832893/files/figure.png" pageId="6" pageNumber="577">5A–C</figureCitation>
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) elongate oval, strongly convex, dorsum pubescent. Head, mouthparts, legs and abdomen reddish-orange. Pronotum chestnut brown with narrow reddish bordering of anterior pronotal margins. Elytra chestnut brown, with two orange or reddish maculae on each elytron, the first placed below humeral callus, the second in apical part of elytron.
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</paragraph>
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<caption id="198F660EFFC53A33FF3DF9CBEF3AF98A" ID-DOI="http://doi.org/10.5281/zenodo.5832891" ID-Zenodo-Dep="5832891" httpUri="https://zenodo.org/record/5832891/files/figure.png" pageId="6" pageNumber="577" startId="6.[151,250,1594,1618]" targetBox="[191,1392,321,1555]" targetPageId="6">
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<paragraph id="4D4F3686FFC53A33FF3DF9CBEF3AF98A" blockId="6.[151,1436,1594,1654]" pageId="6" pageNumber="577">
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<emphasis id="7F84EA94FFC53A33FF3DF9CBED57F9AE" bold="true" box="[151,275,1594,1618]" pageId="6" pageNumber="577">FIGURE 4.</emphasis>
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<taxonomicName id="8AF04D05FFC53A33FEB0F9CAEECAF9AE" authority="(Sicard, 1909)" authorityName="Szawaryn & Churata-Salcedo" authorityYear="2022" baseAuthorityName="Sicard" baseAuthorityYear="1909" box="[282,654,1594,1618]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="quadriguttata">
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<emphasis id="7F84EA94FFC53A33FEB0F9CAEDB0F9AE" box="[282,500,1594,1618]" italics="true" pageId="6" pageNumber="577">Merma quadriguttata</emphasis>
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(
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<bibRefCitation id="29614B77FFC53A33FDA8F9CBEEC3F9AE" author="Sicard, A." box="[514,647,1594,1618]" pageId="6" pageNumber="577" pagination="63 - 165" refId="ref5633" refString="Sicard, A. (1909) Revision des Coccinellides de la faune Malgache. II. Coccinellidae Aphidiphagae. Annales de la Societe Entomologique de France, 78, 63 - 165." type="journal article" year="1909">Sicard, 1909</bibRefCitation>
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)
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</taxonomicName>
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, holotype (MNHN), female: A—habitus, dorsal, B—habitus, frontal, C—habitus, lateral, D—holotype label data, E—abdomen, F—female genitalia.
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</paragraph>
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</caption>
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<paragraph id="4D4F3686FFC53A32FF6DF955E895FEBD" blockId="6.[151,1437,1700,2015]" lastBlockId="7.[151,1437,151,1905]" lastPageId="7" lastPageNumber="578" pageId="6" pageNumber="577">
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Head transverse, partially concealed in prothorax. Interocular distance about 0.6 times of the head width. Inner eye orbits slightly emarginate antero-medially, closest in the middle. Antenna (
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<figureCitation id="D5CB2A03FFC53A33FB82F939E8C6F91F" box="[1064,1154,1736,1763]" captionStart="FIGURE 6" captionStartId="9.[151,250,1505,1529]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-17@9.[151,1435,181,1481]" captionTargetPageId="9" captionText="FIGURE 6. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, ventral, B—mouthparts, C—head, prosternum and mesoventrite, D—legs, meso and metaventrite, E—tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5832895" httpUri="https://zenodo.org/record/5832895/files/figure.png" pageId="6" pageNumber="577">Fig. 6C</figureCitation>
|
||
) composed of 11 antennomeres, about as long as 0.6 head width; scape large, elongate, slightly swollen; pedicel distinctly narrower than scape, elongate; antennomere 3 elongate, about as long as pedicel; antennomeres 4–5 elongate, about twice as long as its width; antennomeres 6–7 subquadrate; antennomere 8 slightly transverse; antennomeres 9–11 forming a distinct asymmetrical club. Ventral antennal grooves (
|
||
<figureCitation id="D5CB2A03FFC53A33FD7DF8A9EF6BF88F" box="[727,815,1880,1907]" captionStart="FIGURE 6" captionStartId="9.[151,250,1505,1529]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-17@9.[151,1435,181,1481]" captionTargetPageId="9" captionText="FIGURE 6. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, ventral, B—mouthparts, C—head, prosternum and mesoventrite, D—legs, meso and metaventrite, E—tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5832895" httpUri="https://zenodo.org/record/5832895/files/figure.png" pageId="6" pageNumber="577">Fig. 6C</figureCitation>
|
||
) distinct in anterior part, moderately long and straight, along inner margin of eye. Clypeus short, parallel-sided, its anterior margin emarginate, smooth without groove. Labrum transverse, short, anterior margin emarginate, membranous. Mandible multidentate apically. Maxilla (
|
||
<figureCitation id="D5CB2A03FFC53A33FAC5F851ECFDF823" captionStart="FIGURE 6" captionStartId="9.[151,250,1505,1529]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-17@9.[151,1435,181,1481]" captionTargetPageId="9" captionText="FIGURE 6. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, ventral, B—mouthparts, C—head, prosternum and mesoventrite, D—legs, meso and metaventrite, E—tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5832895" httpUri="https://zenodo.org/record/5832895/files/figure.png" pageId="6" pageNumber="577">Fig. 6B</figureCitation>
|
||
) with cardo semicircular; with suture between basistipes and mediostipes well visible; lacinia simple, its mesal surface simply setose; galea oval, mostly sclerotized, its ventral surface sparsely pubescent, anterior surface covered with long setae and sensillae; terminal palpomere elongate, with sides subparallel. Submentum short, transverse, with suture well visible; mentum (
|
||
<figureCitation id="D5CB2A03FFC43A32FDB6FF2EEE37FF06" box="[540,627,223,250]" captionStart="FIGURE 6" captionStartId="9.[151,250,1505,1529]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-17@9.[151,1435,181,1481]" captionTargetPageId="9" captionText="FIGURE 6. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, ventral, B—mouthparts, C—head, prosternum and mesoventrite, D—legs, meso and metaventrite, E—tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5832895" httpUri="https://zenodo.org/record/5832895/files/figure.png" pageId="7" pageNumber="578">Fig. 6B</figureCitation>
|
||
) transverse, widest at base; prementum transverse with anterior margin emarginae medially, ligula covered with moderately long setae; labial palps (
|
||
<figureCitation id="D5CB2A03FFC43A32FC57FEF2E810FEE2" box="[1021,1108,259,286]" captionStart="FIGURE 6" captionStartId="9.[151,250,1505,1529]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-17@9.[151,1435,181,1481]" captionTargetPageId="9" captionText="FIGURE 6. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, ventral, B—mouthparts, C—head, prosternum and mesoventrite, D—legs, meso and metaventrite, E—tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5832895" httpUri="https://zenodo.org/record/5832895/files/figure.png" pageId="7" pageNumber="578">Fig. 6B</figureCitation>
|
||
) separated by distance about half of width of palpiger; apical palpomere distinctly shorter and narrower than penultimate one.
|
||
</paragraph>
|
||
<paragraph id="4D4F3686FFC43A32FF6DFEBAEFE0FE2D" blockId="7.[151,1437,151,1905]" pageId="7" pageNumber="578">
|
||
Prothorax. Hypomeron covered with large punctae. Prosternal process (
|
||
<figureCitation id="D5CB2A03FFC43A32FC41FEBAE806FE9A" box="[1003,1090,331,358]" captionStart="FIGURE 6" captionStartId="9.[151,250,1505,1529]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-17@9.[151,1435,181,1481]" captionTargetPageId="9" captionText="FIGURE 6. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, ventral, B—mouthparts, C—head, prosternum and mesoventrite, D—legs, meso and metaventrite, E—tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5832895" httpUri="https://zenodo.org/record/5832895/files/figure.png" pageId="7" pageNumber="578">Fig. 6C</figureCitation>
|
||
) smooth, without carinae, bordered laterally. Prosternum in front of coxa about as long as coxal longitudinal diameter; anterior margin, uniformly arcuate, bordered; posterior margin with bordering line starting from base of prosternal process, laterally reaching notosternal suture. Surface of prosternum covered with large punctae.
|
||
</paragraph>
|
||
<paragraph id="4D4F3686FFC43A32FF6DFE2AECBDFCE9" blockId="7.[151,1437,151,1905]" pageId="7" pageNumber="578">
|
||
Pterothorax. Mesoventrite (
|
||
<figureCitation id="D5CB2A03FFC43A32FE51FE2AEE17FE0A" box="[507,595,475,502]" captionStart="FIGURE 6" captionStartId="9.[151,250,1505,1529]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-17@9.[151,1435,181,1481]" captionTargetPageId="9" captionText="FIGURE 6. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, ventral, B—mouthparts, C—head, prosternum and mesoventrite, D—legs, meso and metaventrite, E—tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5832895" httpUri="https://zenodo.org/record/5832895/files/figure.png" pageId="7" pageNumber="578">Fig. 6D</figureCitation>
|
||
) with anterior edge weakly emarginate, anterior margin entirely raised and bordered, with distinct foveae for receiving midlegs (
|
||
<figureCitation id="D5CB2A03FFC43A32FD50FE0EEF13FDE6" box="[762,855,511,538]" captionStart="FIGURE 6" captionStartId="9.[151,250,1505,1529]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-17@9.[151,1435,181,1481]" captionTargetPageId="9" captionText="FIGURE 6. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, ventral, B—mouthparts, C—head, prosternum and mesoventrite, D—legs, meso and metaventrite, E—tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5832895" httpUri="https://zenodo.org/record/5832895/files/figure.png" pageId="7" pageNumber="578">Fig. 6A</figureCitation>
|
||
); mesoventral process about as wide as mesocoxal diameter, covered with small tubercles on entire surface (
|
||
<figureCitation id="D5CB2A03FFC43A32FC93FDD2EFD3FDC2" box="[825,919,547,574]" captionStart="FIGURE 6" captionStartId="9.[151,250,1505,1529]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-17@9.[151,1435,181,1481]" captionTargetPageId="9" captionText="FIGURE 6. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, ventral, B—mouthparts, C—head, prosternum and mesoventrite, D—legs, meso and metaventrite, E—tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5832895" httpUri="https://zenodo.org/record/5832895/files/figure.png" pageId="7" pageNumber="578">Fig. 6D</figureCitation>
|
||
); meso-metaventral suture straight. Scutellar shield triangular, about as long as broad. Elytra with
|
||
<specimenCount id="5BF6FD0FFFC43A32FD75FDB6EF0EFD9D" box="[735,842,583,609]" pageId="7" pageNumber="578" type="generic" typeStatus="types">two types</specimenCount>
|
||
of punctate; lateral margins not explanate, not visible from above. Epipleuron incomplete apically only (
|
||
<figureCitation id="D5CB2A03FFC43A32FD78FD9AEF69FD7A" box="[722,813,619,646]" captionStart="FIGURE 6" captionStartId="9.[151,250,1505,1529]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-17@9.[151,1435,181,1481]" captionTargetPageId="9" captionText="FIGURE 6. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, ventral, B—mouthparts, C—head, prosternum and mesoventrite, D—legs, meso and metaventrite, E—tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5832895" httpUri="https://zenodo.org/record/5832895/files/figure.png" pageId="7" pageNumber="578">Fig. 6A</figureCitation>
|
||
), narrow, less than two times of width of metanepisternum, with distinct depressions to accommodate mid and hind legs in repose, inner margin with narrow bordering line complete. Mesoventrite covered with sparse and larger punctae in median part, with smaller punctae laterally. Metaventral postcoxal lines (
|
||
<figureCitation id="D5CB2A03FFC43A32FE75FD26EE7CFD0E" box="[479,568,727,754]" captionStart="FIGURE 6" captionStartId="9.[151,250,1505,1529]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-17@9.[151,1435,181,1481]" captionTargetPageId="9" captionText="FIGURE 6. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, ventral, B—mouthparts, C—head, prosternum and mesoventrite, D—legs, meso and metaventrite, E—tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5832895" httpUri="https://zenodo.org/record/5832895/files/figure.png" pageId="7" pageNumber="578">Fig. 6D</figureCitation>
|
||
) joined on metaventral process forming straight line, recurved and complete laterally.
|
||
</paragraph>
|
||
<paragraph id="4D4F3686FFC43A32FF6DFCEEEE3CFC35" blockId="7.[151,1437,151,1905]" pageId="7" pageNumber="578">
|
||
Legs (
|
||
<figureCitation id="D5CB2A03FFC43A32FEA7FCEEED22FCC6" box="[269,358,799,826]" captionStart="FIGURE 6" captionStartId="9.[151,250,1505,1529]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-17@9.[151,1435,181,1481]" captionTargetPageId="9" captionText="FIGURE 6. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, ventral, B—mouthparts, C—head, prosternum and mesoventrite, D—legs, meso and metaventrite, E—tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5832895" httpUri="https://zenodo.org/record/5832895/files/figure.png" pageId="7" pageNumber="578">Fig. 6A</figureCitation>
|
||
) short and stout with apices of mid and hind femora not protruding from outer margin of elytral epipleuron. Trochanters subquadrate, angulately produced, with bordering line on their outer surfaces, and cavities on their inner surfaces for receiving tip of tibiae in repose. Protibiae slightly widened, all tibiae on outer edge with oblique carina near apex (
|
||
<figureCitation id="D5CB2A03FFC43A32FE10FC7AEE50FC5A" box="[442,532,907,934]" captionStart="FIGURE 6" captionStartId="9.[151,250,1505,1529]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-17@9.[151,1435,181,1481]" captionTargetPageId="9" captionText="FIGURE 6. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, ventral, B—mouthparts, C—head, prosternum and mesoventrite, D—legs, meso and metaventrite, E—tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5832895" httpUri="https://zenodo.org/record/5832895/files/figure.png" pageId="7" pageNumber="578">Fig. 6D</figureCitation>
|
||
), without distinct apical spurs. Tarsi pseudotrimerous with tarsal claws (
|
||
<figureCitation id="D5CB2A03FFC43A32FA95FC7AE9D1FC5A" box="[1343,1429,907,934]" captionStart="FIGURE 6" captionStartId="9.[151,250,1505,1529]" captionTargetBox="[151,1435,181,1480]" captionTargetId="figure-17@9.[151,1435,181,1481]" captionTargetPageId="9" captionText="FIGURE 6. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, ventral, B—mouthparts, C—head, prosternum and mesoventrite, D—legs, meso and metaventrite, E—tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5832895" httpUri="https://zenodo.org/record/5832895/files/figure.png" pageId="7" pageNumber="578">Fig. 6E</figureCitation>
|
||
) double, swollen at base, empodium absent.
|
||
</paragraph>
|
||
<paragraph id="4D4F3686FFC43A32FF6DFC22EDEAFB5D" blockId="7.[151,1437,151,1905]" pageId="7" pageNumber="578">
|
||
Abdomen with five ventrites in both sexes with ventrite 5 arcuate posteriorly (
|
||
<figureCitation id="D5CB2A03FFC43A32FB94FC22E8DDFC11" box="[1086,1177,979,1005]" captionStart="FIGURE 4" captionStartId="6.[151,250,1594,1618]" captionTargetBox="[191,1392,321,1555]" captionTargetId="figure-88@6.[180,1408,308,1570]" captionTargetPageId="6" captionText="FIGURE 4. Merma quadriguttata (Sicard, 1909), holotype (MNHN), female: A—habitus, dorsal, B—habitus, frontal, C—habitus, lateral, D—holotype label data, E—abdomen, F—female genitalia." figureDoi="http://doi.org/10.5281/zenodo.5832891" httpUri="https://zenodo.org/record/5832891/files/figure.png" pageId="7" pageNumber="578">Figs 4E</figureCitation>
|
||
,
|
||
<figureCitation id="D5CB2A03FFC43A32FB0FFC23E88FFC11" box="[1189,1227,978,1005]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[178,1409,203,1919]" captionTargetId="figure-29@8.[178,1409,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, frontal, B—habitus, lateral, C—habitus, dorsal, D—abdomen, male, E—penis, lateral, F—tegmen, inner, G—tegmen, lateral, H—sternite VIII, male, I—tergite VIII, male, J—male genital segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.5832893" httpUri="https://zenodo.org/record/5832893/files/figure.png" pageId="7" pageNumber="578">5D</figureCitation>
|
||
). Abdominal postcoxal lines (
|
||
<figureCitation id="D5CB2A03FFC43A32FEB4FC06ED30FBED" box="[286,372,1015,1041]" captionStart="FIGURE 4" captionStartId="6.[151,250,1594,1618]" captionTargetBox="[191,1392,321,1555]" captionTargetId="figure-88@6.[180,1408,308,1570]" captionTargetPageId="6" captionText="FIGURE 4. Merma quadriguttata (Sicard, 1909), holotype (MNHN), female: A—habitus, dorsal, B—habitus, frontal, C—habitus, lateral, D—holotype label data, E—abdomen, F—female genitalia." figureDoi="http://doi.org/10.5281/zenodo.5832891" httpUri="https://zenodo.org/record/5832891/files/figure.png" pageId="7" pageNumber="578">Fig. 4E</figureCitation>
|
||
,
|
||
<figureCitation id="D5CB2A03FFC43A32FED5FC07EDE6FBED" box="[383,418,1014,1041]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[178,1409,203,1919]" captionTargetId="figure-29@8.[178,1409,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, frontal, B—habitus, lateral, C—habitus, dorsal, D—abdomen, male, E—penis, lateral, F—tegmen, inner, G—tegmen, lateral, H—sternite VIII, male, I—tergite VIII, male, J—male genital segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.5832893" httpUri="https://zenodo.org/record/5832893/files/figure.png" pageId="7" pageNumber="578">5D</figureCitation>
|
||
) rounded, recurved, not parallel to the posterior margin of ventrite 1. In male apical margin of sternite VIII truncate (
|
||
<figureCitation id="D5CB2A03FFC43A32FE04FBEAEE43FBC9" box="[430,519,1050,1077]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[178,1409,203,1919]" captionTargetId="figure-29@8.[178,1409,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, frontal, B—habitus, lateral, C—habitus, dorsal, D—abdomen, male, E—penis, lateral, F—tegmen, inner, G—tegmen, lateral, H—sternite VIII, male, I—tergite VIII, male, J—male genital segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.5832893" httpUri="https://zenodo.org/record/5832893/files/figure.png" pageId="7" pageNumber="578">Fig. 5H</figureCitation>
|
||
), tergite VIII rounded (
|
||
<figureCitation id="D5CB2A03FFC43A32FCA0FBEAEF1CFBC9" box="[778,856,1050,1077]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[178,1409,203,1919]" captionTargetId="figure-29@8.[178,1409,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, frontal, B—habitus, lateral, C—habitus, dorsal, D—abdomen, male, E—penis, lateral, F—tegmen, inner, G—tegmen, lateral, H—sternite VIII, male, I—tergite VIII, male, J—male genital segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.5832893" httpUri="https://zenodo.org/record/5832893/files/figure.png" pageId="7" pageNumber="578">Fig. 5I</figureCitation>
|
||
); apodeme of sternum IX (
|
||
<figureCitation id="D5CB2A03FFC43A32FB2EFBEAE896FBC9" box="[1156,1234,1050,1077]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[178,1409,203,1919]" captionTargetId="figure-29@8.[178,1409,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, frontal, B—habitus, lateral, C—habitus, dorsal, D—abdomen, male, E—penis, lateral, F—tegmen, inner, G—tegmen, lateral, H—sternite VIII, male, I—tergite VIII, male, J—male genital segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.5832893" httpUri="https://zenodo.org/record/5832893/files/figure.png" pageId="7" pageNumber="578">Fig. 5J</figureCitation>
|
||
) rod-like, narrow. Tergite X transverse, truncate apically (
|
||
<figureCitation id="D5CB2A03FFC43A32FDFAFBCEEEE4FBA5" box="[592,672,1086,1113]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[178,1409,203,1919]" captionTargetId="figure-29@8.[178,1409,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, frontal, B—habitus, lateral, C—habitus, dorsal, D—abdomen, male, E—penis, lateral, F—tegmen, inner, G—tegmen, lateral, H—sternite VIII, male, I—tergite VIII, male, J—male genital segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.5832893" httpUri="https://zenodo.org/record/5832893/files/figure.png" pageId="7" pageNumber="578">Fig. 5J</figureCitation>
|
||
). In female apical margin of sternite VIII rounded, simply arcuate at base, longitudinally at middle not divided; tergite VIII rounded (
|
||
<figureCitation id="D5CB2A03FFC43A32FCC2FB92EFF9FB81" box="[872,957,1123,1149]" captionStart="FIGURE 4" captionStartId="6.[151,250,1594,1618]" captionTargetBox="[191,1392,321,1555]" captionTargetId="figure-88@6.[180,1408,308,1570]" captionTargetPageId="6" captionText="FIGURE 4. Merma quadriguttata (Sicard, 1909), holotype (MNHN), female: A—habitus, dorsal, B—habitus, frontal, C—habitus, lateral, D—holotype label data, E—abdomen, F—female genitalia." figureDoi="http://doi.org/10.5281/zenodo.5832891" httpUri="https://zenodo.org/record/5832891/files/figure.png" pageId="7" pageNumber="578">Fig. 4F</figureCitation>
|
||
). Proctiger (TX) large, transverse, truncate at apex, entire in middle.
|
||
</paragraph>
|
||
<paragraph id="4D4F3686FFC43A32FF6DFB5AEE04FA85" blockId="7.[151,1437,151,1905]" pageId="7" pageNumber="578">
|
||
Male genitalia (
|
||
<figureCitation id="D5CB2A03FFC43A32FEDDFB5AEDB4FB39" box="[375,496,1194,1221]" captionStart="FIGURE 5" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[178,1409,203,1919]" captionTargetId="figure-29@8.[178,1409,181,1919]" captionTargetPageId="8" captionText="FIGURE 5. Merma quadriguttata (Sicard, 1909), non type (CAS), male: A—habitus, frontal, B—habitus, lateral, C—habitus, dorsal, D—abdomen, male, E—penis, lateral, F—tegmen, inner, G—tegmen, lateral, H—sternite VIII, male, I—tergite VIII, male, J—male genital segments IX and X." figureDoi="http://doi.org/10.5281/zenodo.5832893" httpUri="https://zenodo.org/record/5832893/files/figure.png" pageId="7" pageNumber="578">Fig. 5E–G</figureCitation>
|
||
). Tegminal basal piece without distinct spines on inner margin near base of tegminal strut. Penis guide symmetrical in inner view, as long as parameres, with apical third subtriangular in shape, pointed, entire at apex; in lateral view outer edge covered with distinct small tubercles each bearing long seta, inner edge with distinct groove before apex. Parameres well developed, simple apically. Tegminal strut about as long as the rest of tegmen, narrow, simple at apex. Penis rod-like, strongly sinuate in apical half, ending with translucent flagellum; penis capsule with weakly developed.
|
||
</paragraph>
|
||
<paragraph id="4D4F3686FFC43A32FF6DFA72E844FA19" blockId="7.[151,1437,151,1905]" pageId="7" pageNumber="578">
|
||
Female genitalia (
|
||
<figureCitation id="D5CB2A03FFC43A32FE32FA72EDB6FA61" box="[408,498,1411,1437]" captionStart="FIGURE 4" captionStartId="6.[151,250,1594,1618]" captionTargetBox="[191,1392,321,1555]" captionTargetId="figure-88@6.[180,1408,308,1570]" captionTargetPageId="6" captionText="FIGURE 4. Merma quadriguttata (Sicard, 1909), holotype (MNHN), female: A—habitus, dorsal, B—habitus, frontal, C—habitus, lateral, D—holotype label data, E—abdomen, F—female genitalia." figureDoi="http://doi.org/10.5281/zenodo.5832891" httpUri="https://zenodo.org/record/5832891/files/figure.png" pageId="7" pageNumber="578">Fig. 4F</figureCitation>
|
||
). Sclerite anterior to coxites in membrane connecting paraprocts absent. Coxites small, distinctly transverse, with inner edge rounded, ventral surface smooth. Styli well visible. Bursa copulatrix without additional sclerite, simple, not divided, ending with common oviduct.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05EA650DFFC43A32FF6DFA1FEE14F9F5" box="[199,592,1518,1545]" pageId="7" pageNumber="578" type="distribution">
|
||
<paragraph id="4D4F3686FFC43A32FF6DFA1FEE14F9F5" blockId="7.[151,1437,151,1905]" box="[199,592,1518,1545]" pageId="7" pageNumber="578">
|
||
<emphasis id="7F84EA94FFC43A32FF6DFA1FED25F9F4" bold="true" box="[199,353,1518,1544]" pageId="7" pageNumber="578">Distribution.</emphasis>
|
||
<collectingCountry id="35E77616FFC43A32FECDFA1EEDAAF9F5" box="[359,494,1519,1545]" name="Madagascar" pageId="7" pageNumber="578">Madagascar</collectingCountry>
|
||
(
|
||
<figureCitation id="D5CB2A03FFC43A32FE54FA1EEE06F9F5" box="[510,578,1519,1545]" captionStart="FIGURE 7" captionStartId="10.[151,250,1307,1331]" captionTargetId="figure-29@10.[412,1176,181,1282]" captionTargetPageId="10" captionText="FIGURE 7. Distribution map of the Malagasy species of Merma Weise." figureDoi="http://doi.org/10.5281/zenodo.5834515" httpUri="https://zenodo.org/record/5834515/files/figure.png" pageId="7" pageNumber="578">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05EA650DFFC43A32FF6DF9E2EDCDF941" pageId="7" pageNumber="578" type="materials_examined">
|
||
<paragraph id="4D4F3686FFC43A32FF6DF9E2EDCDF941" blockId="7.[151,1437,151,1905]" pageId="7" pageNumber="578">
|
||
<emphasis id="7F84EA94FFC43A32FF6DF9E2EDA9F9D0" bold="true" box="[199,493,1554,1581]" pageId="7" pageNumber="578">Type material examined.</emphasis>
|
||
<materialsCitation id="FD983CDBFFC43A32FE58F9E2EE51F9AD" ID-GBIF-Occurrence="3434635306" collectionCode="MNHM" determinerName="L. A. Almeida" pageId="7" pageNumber="578" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="7F84EA94FFC43A32FE58F9E2EE2CF9D1" bold="true" box="[498,616,1555,1581]" pageId="7" pageNumber="578">
|
||
<typeStatus id="924B8824FFC43A32FE58F9E2EE27F9D1" box="[498,611,1555,1581]" pageId="7" pageNumber="578" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
“Madagascar, Ambositra/ TYPE/
|
||
<taxonomicName id="8AF04D05FFC43A32FC49F9E2E873F9D1" authorityName="Sicard" authorityYear="1909" box="[995,1079,1555,1581]" class="Insecta" family="Coccinellidae" genus="Peralda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="576" phylum="Arthropoda" rank="genus">Peralda</taxonomicName>
|
||
quadrigutata,
|
||
<determinerName id="B893DA42FFC43A32FB7FF9E2E929F9D1" box="[1237,1389,1555,1581]" pageId="7" pageNumber="578">L.A. Almeida</determinerName>
|
||
det. 2017”, female (
|
||
<collectionCode id="2BE1AE43FFC43A32FEEAF9C6EDDAF9AD" box="[320,414,1591,1617]" country="USA" httpUri="http://grbio.org/cool/q3kg-qwj4" name="John May Museum of Natural History" pageId="7" pageNumber="578">MNHM</collectionCode>
|
||
) (
|
||
<figureCitation id="D5CB2A03FFC43A32FE19F9C6EE48F9AD" box="[435,524,1591,1617]" captionStart="FIGURE 4" captionStartId="6.[151,250,1594,1618]" captionTargetBox="[191,1392,321,1555]" captionTargetId="figure-88@6.[180,1408,308,1570]" captionTargetPageId="6" captionText="FIGURE 4. Merma quadriguttata (Sicard, 1909), holotype (MNHN), female: A—habitus, dorsal, B—habitus, frontal, C—habitus, lateral, D—holotype label data, E—abdomen, F—female genitalia." figureDoi="http://doi.org/10.5281/zenodo.5832891" httpUri="https://zenodo.org/record/5832891/files/figure.png" pageId="7" pageNumber="578">Fig. 4D</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<emphasis id="7F84EA94FFC43A32FDB7F9C7EE91F9AC" bold="true" box="[541,725,1590,1616]" pageId="7" pageNumber="578">Other material.</emphasis>
|
||
<materialsCitation id="FD983CDBFFC43A32FD70F9C6EDC0F941" ID-GBIF-Occurrence="3434635304" accessionNumber="BLF03524" collectingDate="2001-01-04" collectionCode="CAS" collectorName="B. L. Fisher & C. E. Griswold " country="Madagascar" county="Parc National d'Ankarafantsika" elevation="130" latitude="-16.32083" location="Ampijoroa Station" longLatPrecision="1" longitude="46.81067" municipality="Ampijoroa Station Forestiere" pageId="7" pageNumber="578" specimenCount="1" specimenCount-male="1" stateProvince="Mahajanga">
|
||
<collectingCountry id="35E77616FFC43A32FD70F9C6EFE2F9AD" box="[730,934,1591,1617]" name="Madagascar" pageId="7" pageNumber="578">MADAGASCAR</collectingCountry>
|
||
:
|
||
<collectingRegion id="8F34F864FFC43A32FC04F9C6E869F9AD" box="[942,1069,1591,1617]" country="Madagascar" name="Mahajanga" pageId="7" pageNumber="578">Mahajanga</collectingRegion>
|
||
,
|
||
<collectingCounty id="A42E4E0AFFC43A32FB9DF9C6E9DCF9AD" box="[1079,1432,1591,1617]" pageId="7" pageNumber="578">Parc National d’Ankarafantsika</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="AD2BACFCFFC43A32FF3DF9AAEDA0F989" box="[151,484,1627,1653]" pageId="7" pageNumber="578">
|
||
<location id="482F605DFFC43A32FF3DF9AAED2CF989" LSID="urn:lsid:plazi:treatment:C5598790FFC63A32FF3DF9C3EEEFF88D:482F605DFFC43A32FF3DF9AAED2CF989" box="[151,360,1627,1653]" country="Madagascar" county="Parc National d'Ankarafantsika" latitude="-16.32083" longLatPrecision="1" longitude="46.81067" municipality="Ampijoroa Station Forestiere" name="Ampijoroa Station" pageId="7" pageNumber="578" stateProvince="Mahajanga">Ampijoroa Station</location>
|
||
Forestière
|
||
</collectingMunicipality>
|
||
,
|
||
<locationDeviation id="5CFD2FA4FFC43A32FE5AF9AAEF1CF989" box="[496,856,1627,1653]" location="Andranofasika" pageId="7" pageNumber="578">
|
||
<quantity id="8A089B63FFC43A32FE5AF9AAEE7CF989" box="[496,568,1627,1653]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="578" unit="km" value="40.0">40 km</quantity>
|
||
306° NW Andranofasika
|
||
</locationDeviation>
|
||
(
|
||
<geoCoordinate id="28C45041FFC43A32FCC2F9AAEF98F989" box="[872,988,1627,1653]" degrees="16.32083" direction="south" orientation="latitude" pageId="7" pageNumber="578" precision="1" value="-16.32083">-16.32083</geoCoordinate>
|
||
<geoCoordinate id="28C45041FFC43A32FC4FF9AAE817F98A" box="[997,1107,1627,1654]" degrees="46.81067" direction="east" orientation="longitude" pageId="7" pageNumber="578" precision="1" value="46.81067">46.81067</geoCoordinate>
|
||
), sample number BLF03524,
|
||
<elevation id="C6DDD1B5FFC43A32FF3DF98EED59F965" box="[151,285,1663,1689]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="578" unit="m" value="130.0">
|
||
<quantity id="8A089B63FFC43A32FF3DF98EEC9BF965" box="[151,223,1663,1689]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="578" unit="m" value="130.0">130 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
, 26.03-
|
||
<date id="394E1046FFC43A32FED8F98EEDB0F965" box="[370,500,1663,1689]" pageId="7" pageNumber="578" value="2001-01-04">
|
||
<collectingDate id="290AE9AEFFC43A32FED8F98EEDB0F965" box="[370,500,1663,1689]" pageId="7" pageNumber="578" value="2001-01-04">01.04.2001</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="E0055350FFC43A32FD93F98EEEF9F965" box="[569,701,1663,1689]" pageId="7" pageNumber="578">B.L. Fisher</collectorName>
|
||
;
|
||
<collectorName id="E0055350FFC43A32FD63F98EEF2BF965" box="[713,879,1663,1689]" pageId="7" pageNumber="578">C.E. Griswold</collectorName>
|
||
;
|
||
<collectorName id="E0055350FFC43A32FCD1F98EE8E8F965" box="[891,1196,1663,1689]" pageId="7" pageNumber="578">Malagasy Arthropod Team</collectorName>
|
||
, beating, tropical dry forest (
|
||
<specimenCount id="5BF6FD0FFFC43A32FF43F952ED73F941" box="[233,311,1699,1725]" pageId="7" pageNumber="578" type="male">1 male</specimenCount>
|
||
:
|
||
<collectionCode id="2BE1AE43FFC43A32FEEBF952ED38F941" box="[321,380,1699,1725]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="7" pageNumber="578" type="Museum">CAS</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="05EA650DFFC43A32FF6DF936EEEFF88D" pageId="7" pageNumber="578" type="discussion">
|
||
<paragraph id="4D4F3686FFC43A32FF6DF936EEEFF88D" blockId="7.[151,1437,151,1905]" pageId="7" pageNumber="578">
|
||
<emphasis id="7F84EA94FFC43A32FF6DF936ED7FF91D" bold="true" box="[199,315,1735,1761]" pageId="7" pageNumber="578">Remarks.</emphasis>
|
||
<taxonomicName id="8AF04D05FFC43A32FEE9F936EDBEF91D" authorityName="Szawaryn & Churata-Salcedo" authorityYear="2022" baseAuthorityName="Sicard" baseAuthorityYear="1909" box="[323,506,1735,1761]" class="Insecta" family="Coccinellidae" genus="Merma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="quadriguttata">
|
||
<emphasis id="7F84EA94FFC43A32FEE9F936EDBEF91D" box="[323,506,1735,1761]" italics="true" pageId="7" pageNumber="578">P. quadriguttata</emphasis>
|
||
</taxonomicName>
|
||
was originally described based on a single female specimen, the
|
||
<typeStatus id="924B8824FFC43A32FB4AF936E906F91D" box="[1248,1346,1735,1761]" pageId="7" pageNumber="578" type="holotype">holotype</typeStatus>
|
||
that we examined. We also found a second specimen (male) that is identical externally, however, it comes from a different location more than
|
||
<quantity id="8A089B63FFC43A32FED8F8FFED8CF8D5" box="[370,456,1806,1833]" metricMagnitude="5" metricUnit="m" metricValue="5.5" pageId="7" pageNumber="578" unit="km" value="550.0">550 km</quantity>
|
||
north from the type locality. As there are no external differences in shape, colouration, size or other features we consider this specimen conspecific. Nonetheless, examination of further specimens from the type locality would confirm our hypothesis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |