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<document id="036C6D3F5FBB950957EBF34FC6F54912" ID-DOI="10.5252/cryptogamie-bryologie2022v43a5" ID-ISSN="1776-0992" ID-Zenodo-Dep="12215006" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719003215836" checkinUser="felipe" docAuthor="Gradstein, S. Robbert &amp; Reeb, Catherine" docDate="2022" docId="AE6387E6FF84FFD3F7563EF6821EFDCB" docLanguage="en" docName="CryptogamieBryologie.43.5.65-106.pdf" docOrigin="Cryptogamie, Bryologie 20 (5)" docSource="http://dx.doi.org/10.5252/cryptogamie-bryologie2022v43a5" docStyle="DocumentStyle:3F4A000F5B047ABA0EAB2943AA9373AE.3:CryptogamieBryologie.2019-.journal_article" docStyleId="3F4A000F5B047ABA0EAB2943AA9373AE" docStyleName="CryptogamieBryologie.2019-.journal_article" docStyleVersion="3" docTitle="Plagiochila vandenberghenii Gradst. 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="104" masterDocId="525AFF9EFFA3FFFAF4213E21806AFF9B" masterDocTitle="The genus Plagiochila (Dumort.) Dumort. (Marchantiophyta) in Madagascar" masterLastPageNumber="106" masterPageNumber="65" pageNumber="102" updateTime="1719017834308" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5469F25576B9C0AB9BB413C28DC38E32">The genus Plagiochila (Dumort.) Dumort. (Marchantiophyta) in Madagascar</mods:title>
</mods:titleInfo>
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<mods:namePart id="6880470743598DEA3281D9E52DD45744">Gradstein, S. Robbert</mods:namePart>
<mods:affiliation id="26BE7B2D583E2923C60C99D8F7456900">Meise Botanic Garden, 1060 Meise (Belgium) and Institut de Systématique, Évolution, Biodiversité (UMR 7205) Sorbonne Université (MNHN, EPHE, CNRS), Muséum national dHistoire naturelle 57 rue Cuvier, case postale 39, 75005 Paris (France) robbert. gradstein @ mnhn. fr (corresponding author)</mods:affiliation>
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<mods:namePart id="FF3199527071B9A0B1FC02605B05E936">Reeb, Catherine</mods:namePart>
<mods:affiliation id="5C263D733F8886EEB48793CAC75531C6">Institut de Systématique, Évolution, Biodiversité (UMR 7205) Sorbonne Université (MNHN, EPHE, CNRS), Muséum national dHistoire naturelle 57 rue Cuvier, case postale 39, 75005 Paris (France) catherine. reeb @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="CD8BCCEC89567D02CD6FF5C548906716" type="email">reeb@mnhn.fr</mods:nameIdentifier>
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<mods:title id="9BDF259C130B6C9FD9044E75AFB6F89B">Cryptogamie, Bryologie</mods:title>
</mods:titleInfo>
<mods:part id="D4CAC060D0EA01AE2F44B8DDC83C36F4">
<mods:date id="9E0023666BED30859E0DDA8026F2D500">2022</mods:date>
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<mods:number id="2CD890E96460930E844B033D1855C355">2022-05-16</mods:number>
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<mods:url id="1E12ACD2B3D9E2CABF5EF805B1F08798">http://dx.doi.org/10.5252/cryptogamie-bryologie2022v43a5</mods:url>
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<treatment id="AE6387E6FF84FFD3F7563EF6821EFDCB" ID-DOI="http://doi.org/10.5281/zenodo.12215042" ID-Zenodo-Dep="12215042" LSID="urn:lsid:plazi:treatment:AE6387E6FF84FFD3F7563EF6821EFDCB" httpUri="http://treatment.plazi.org/id/AE6387E6FF84FFD3F7563EF6821EFDCB" lastPageId="41" lastPageNumber="104" pageId="39" pageNumber="102">
<subSubSection id="6ED0657BFF84FFDDF7563EF6850FFF69" box="[887,1381,215,242]" pageId="39" pageNumber="102" type="nomenclature">
<paragraph id="267536F0FF84FFDDF7563EF6850FFF69" blockId="39.[887,1381,215,242]" box="[887,1381,215,242]" pageId="39" pageNumber="102">
<heading id="7D3D819CFF84FFDDF7563EF6850FFF69" box="[887,1381,215,242]" centered="true" fontSize="11" level="2" pageId="39" pageNumber="102" reason="4">
<taxonomicName id="E1CA4D73FF84FFDDF7563EF6856CFF6A" authority="Gradst." authorityName="Gradst." authorityYear="2022" box="[887,1286,215,242]" class="Jungermanniopsida" family="Plagiochilaceae" genus="Plagiochila" kingdom="Plantae" order="Jungermanniales" pageId="39" pageNumber="102" phylum="Marchantiophyta" rank="species" species="vandenberghenii" status="sp. nov.">
<emphasis id="14BEEAE2FF84FFDDF7563EF684C1FF69" bold="true" box="[887,1195,215,242]" italics="true" pageId="39" pageNumber="102">Plagiochila vandenberghenii</emphasis>
Gradst.
</taxonomicName>
,
<taxonomicNameLabel id="0F8D5799FF84FFDDF1313EF9850FFF69" box="[1296,1381,216,242]" pageId="39" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6ED0657BFF84FFDDF01E3ED685C4FD9A" pageId="39" pageNumber="102" type="description">
<paragraph id="267536F0FF84FFDDF01E3ED684F7FE8A" blockId="39.[1087,1181,247,273]" box="[1087,1181,247,273]" pageId="39" pageNumber="102">
<heading id="7D3D819CFF84FFDDF01E3ED684F7FE8A" box="[1087,1181,247,273]" centered="true" fontSize="11" level="3" pageId="39" pageNumber="102" reason="3">
(
<figureCitation id="BEF12A75FF84FFDDF0663ED684FEFE8A" box="[1095,1172,247,273]" captionStart="FIG" captionStartId="40.[132,143,1962,1980]" captionText="FIG. 23. ― Plagiochila vandenberghenii Gradst., sp. nov.: A, shoots; B, shoot in dorsal view with terminal branching; C, intercalary branching; D, shoot in ventral view; E, stem leaves; F, branch leaves; G, branch in dorsal view; H, I, leaf margin cells; J, leaf base; K, leaf apex; L, shoot with perianth, two leaves at perianth base removed to show the perianth stalk; M, female bracts. From Reeb CR17M610 (PC). Scale bars: A, 5 mm; B-G, J, K, 1 mm; H, I, L, 0.1 mm; M, 0.025 mm." pageId="39" pageNumber="102">Fig. 23</figureCitation>
)
</heading>
</paragraph>
<paragraph id="267536F0FF84FFDDF70C3F1185C4FD9A" blockId="39.[811,1457,304,513]" pageId="39" pageNumber="102">
<taxonomicName id="E1CA4D73FF84FFDDF70C3F118414FED3" authority="Gradst." authorityName="Gradst." authorityYear="2022" box="[813,1150,304,328]" class="Jungermanniopsida" family="Plagiochilaceae" genus="Plagiochila" kingdom="Plantae" order="Jungermanniales" pageId="39" pageNumber="102" phylum="Marchantiophyta" rank="species" species="vandenberghenii" status="sp. nov.">
<emphasis id="14BEEAE2FF84FFDDF70C3F118445FED3" box="[813,1071,304,328]" italics="true" pageId="39" pageNumber="102">Plagiochila vandenberghenii</emphasis>
Gradst.
</taxonomicName>
,
<taxonomicNameLabel id="0F8D5799FF84FFDDF0A73F1084BAFED3" box="[1158,1232,305,328]" pageId="39" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
(sect.
<emphasis id="14BEEAE2FF84FFDDF1293F10852AFED3" box="[1288,1344,305,328]" italics="true" pageId="39" pageNumber="102">Vagae</emphasis>
) stands out by irregularly pinnate to dichotomous branching by terminal and intercalary branches, and somewhat distant, horizontally spreading, narrowly oblong-rectangular, non-ampliate leaves (
<emphasis id="14BEEAE2FF84FFDDF1393FA1854BFE0C" box="[1304,1313,384,407]" italics="true" pageId="39" pageNumber="102">c</emphasis>
. 2.5-3× longer than wide) with the apex bifid by two large, lobe-like teeth, and a slightly bordered leaf margin with more elongate cells in 1-2 rows; 2-5 additional, smaller teeth are produced at the leaf apex and on the distal half of the ventral and dorsal margin. Underleaves are absent.
</paragraph>
</subSubSection>
<subSubSection id="6ED0657BFF84FFDDF70C3C3D847AFC9F" pageId="39" pageNumber="102" type="materials_examined">
<paragraph id="267536F0FF84FFDDF70C3C3D8390FD06" blockId="39.[811,1457,539,669]" pageId="39" pageNumber="102">
<materialsCitation id="96A23CADFF84FFDDF70C3C3D8390FD06" collectingDate="2004-07" collectionCode="EGR, GOET" country="Madagascar" county="Ranomafana Nat. Park" elevation="940" location="Namorona river" municipality="Ambodiamontana" pageId="39" pageNumber="102" specimenCount="3" stateProvince="Fianarantsoa" typeStatus="holotype">
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<typeStatus id="F9718852FF84FFDDF70C3C3D83F3FDA9" box="[813,921,540,563]" pageId="39" pageNumber="102" type="holotype">HOLOTYPE</typeStatus>
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. —
<emphasis id="14BEEAE2FF84FFDDF7ED3C3A842CFDA8" bold="true" box="[972,1094,539,563]" pageId="39" pageNumber="102">
<collectingCountry id="5EDD7660FF84FFDDF7ED3C3A842CFDA8" box="[972,1094,539,563]" name="Madagascar" pageId="39" pageNumber="102">Madagascar</collectingCountry>
</emphasis>
.
<emphasis id="14BEEAE2FF84FFDDF0743C3A857CFDAE" bold="true" box="[1109,1302,539,565]" pageId="39" pageNumber="102">
Prov.
<collectingRegion id="E40EF812FF84FFDDF0B33C3C857CFDAE" box="[1170,1302,541,565]" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="102">Fianarantsoa</collectingRegion>
</emphasis>
<collectingCounty id="CF144E7CFF84FFDDF1133C3A83E5FDD5" pageId="39" pageNumber="102">Ranomafana Nat. Park</collectingCounty>
, opposite park gate, near
<collectingMunicipality id="C611AC8AFF84FFDDF0AC3C178554FDD5" box="[1165,1342,566,590]" pageId="39" pageNumber="102">Ambodiamontana</collectingMunicipality>
settlement,
<location id="2315602BFF84FFDDF70C3C7083A1FDF3" LSID="urn:lsid:plazi:treatment:AE6387E6FF84FFD3F7563EF6821EFDCB:2315602BFF84FFDDF70C3C7083A1FDF3" box="[813,971,592,616]" country="Madagascar" county="Ranomafana Nat. Park" municipality="Ambodiamontana" name="Namorona river" pageId="39" pageNumber="102" stateProvince="Fianarantsoa">Namorona river</location>
valley, on bark in partially degraded lower montane rainforest,
<quantity id="E1329B15FF84FFDDF7E03C4B8457FD19" box="[961,1085,618,642]" metricMagnitude="2" metricUnit="m" metricValue="9.4" metricValueMax="10.15" metricValueMin="8.65" pageId="39" pageNumber="102" unit="m" value="940.0" valueMax="1015.0" valueMin="865.0">
<elevation id="ADE7D1C3FF84FFDDF7E03C4B8457FD19" box="[961,1085,618,642]" metricMagnitude="2" metricUnit="m" metricValue="9.4" metricValueMax="10.15" metricValueMin="8.65" pageId="39" pageNumber="102" unit="m" value="940.0" valueMax="1015.0" valueMin="865.0">865-1015 m</elevation>
</quantity>
,
<date id="52741030FF84FFDDF0643C4A84C9FD18" box="[1093,1187,619,643]" pageId="39" pageNumber="102" value="2004-07">
<collectingDate id="4230E9D8FF84FFDDF0643C4A84C9FD18" box="[1093,1187,619,643]" pageId="39" pageNumber="102" value="2004-07">VII.2004</collectingDate>
</date>
,
<emphasis id="14BEEAE2FF84FFDDF08B3C4A850CFD18" box="[1194,1382,618,643]" italics="true" pageId="39" pageNumber="102">Pócs et al. 04118/AY</emphasis>
(
<typeStatus id="F9718852FF84FFDDF1553C4B85C1FD19" box="[1396,1451,618,642]" pageId="39" pageNumber="102" type="holotype">holo-</typeStatus>
,
<collectionCode id="40DBAE35FF84FFDDF70A3CA48308FD06" box="[811,866,645,669]" country="Hungary" httpUri="http://grbio.org/cool/39sm-y0v8" name="Eszterházy Károly College" pageId="39" pageNumber="102" type="Herbarium">EGR</collectionCode>
!;
<typeStatus id="F9718852FF84FFDDF74E3CA483FFFD06" box="[879,917,645,669]" pageId="39" pageNumber="102" type="isotype">iso-</typeStatus>
,
<collectionCode id="40DBAE35FF84FFDDF7BE3CA48386FD06" box="[927,1004,645,669]" country="Germany" httpUri="http://grbio.org/cool/pq74-a94z" name="Universität Göttingen" pageId="39" pageNumber="102" type="Herbarium">GOET</collectionCode>
!).
</materialsCitation>
</paragraph>
<paragraph id="267536F0FF84FFDDF70C3C96847AFC9F" blockId="39.[813,1456,695,772]" pageId="39" pageNumber="102">
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. —
<emphasis id="14BEEAE2FF84FFDDF0B43C968563FD54" bold="true" box="[1173,1289,695,719]" pageId="39" pageNumber="102">
<collectingCountry id="5EDD7660FF84FFDDF0B43C968563FD54" box="[1173,1289,695,719]" name="Madagascar" pageId="39" pageNumber="102">Madagascar</collectingCountry>
</emphasis>
.
<emphasis id="14BEEAE2FF84FFDDF1353C968327FD72" bold="true" pageId="39" pageNumber="102">
Prov.
<collectingRegion id="E40EF812FF84FFDDF16B3C998327FD72" country="Madagascar" name="Fianarantsoa" pageId="39" pageNumber="102">Fianarantsoa</collectingRegion>
</emphasis>
• Ranomafana Nat. Park, near park gate,
<quantity id="E1329B15FF84FFDDF0D73CF0852DFD72" box="[1270,1351,721,745]" metricMagnitude="3" metricUnit="m" metricValue="1.067" pageId="39" pageNumber="102" unit="m" value="1067.0">1067 m</quantity>
, on twigs,
<emphasis id="14BEEAE2FF84FFDDF70F3CCD83BFFC9F" box="[814,981,748,772]" italics="true" pageId="39" pageNumber="102">
Reeb
<collectionCode id="40DBAE35FF84FFDDF7413CCD8314FC9F" box="[864,894,748,772]" country="Costa Rica" lsid="urn:lsid:biocol.org:col:15652" name="Museo Nacional de Costa Rica" pageId="39" pageNumber="102" type="Herbarium">CR</collectionCode>
17
<collectionCode id="40DBAE35FF84FFDDF7B93CCD83C5FC9F" box="[920,943,748,772]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="39" pageNumber="102" type="Herbarium">M</collectionCode>
610
</emphasis>
(
<collectionCode id="40DBAE35FF84FFDDF7C03CCD846FFC9F" box="[993,1029,748,772]" country="France" lsid="urn:lsid:biocol.org:col:15359" name="Museum National d'Histoire Naturelle, Paris (MNHN) - Non-vascular Plants and Fungi" pageId="39" pageNumber="102" type="Herbarium">PC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="6ED0657BFF84FFDDF70C3D3F8530FCF0" pageId="39" pageNumber="102" type="etymology">
<paragraph id="267536F0FF84FFDDF70C3D3F8530FCF0" blockId="39.[813,1456,798,875]" pageId="39" pageNumber="102">
<smallCapsWord id="2093A02CFF84FFDDF70C3D3F83CCFCAF" baselines="815,816" box="[813,934,798,821]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="39" pageNumber="102">ETYMOLOGY</smallCapsWord>
. — Named after the late Professor Constant Vanden Berghen, renowned Belgian botanist and bryologist who has laid the basis for our knowledge of
<taxonomicName id="E1CA4D73FF84FFDDF0783D7284AAFCF0" box="[1113,1216,851,875]" class="Jungermanniopsida" family="Plagiochilaceae" genus="Plagiochila" kingdom="Plantae" order="Jungermanniales" pageId="39" pageNumber="102" phylum="Marchantiophyta" rank="genus">
<emphasis id="14BEEAE2FF84FFDDF0783D7284AAFCF0" box="[1113,1216,851,875]" italics="true" pageId="39" pageNumber="102">Plagiochila</emphasis>
</taxonomicName>
in
<collectingCountry id="5EDD7660FF84FFDDF0C03D72853CFCF0" box="[1249,1366,851,875]" name="Madagascar" pageId="39" pageNumber="102">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="6ED0657BFF84FFDDF70C3DA485C1FC2C" pageId="39" pageNumber="102" type="distribution">
<paragraph id="267536F0FF84FFDDF70C3DA485C1FC2C" blockId="39.[813,1455,900,951]" pageId="39" pageNumber="102">
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<smallCapsWord id="2093A02CFF84FFDDF7D23DAB842FFC00" baselines="919" box="[1011,1093,906,923]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="habitat" pageId="39" pageNumber="102">HABITAT</smallCapsWord>
. — Only known from Ranomafana National Park (two collections) on
<collectingCountry id="5EDD7660FF84FFDDF0A43DBE8493FC2C" box="[1157,1273,927,951]" name="Madagascar" pageId="39" pageNumber="102">Madagascar</collectingCountry>
, growing on bark.
</paragraph>
</subSubSection>
<subSubSection id="6ED0657BFF84FFD3F70C3DF4821EFDCB" lastPageId="41" lastPageNumber="104" pageId="39" pageNumber="102" type="description">
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</heading>
</paragraph>
<paragraph id="267536F0FF84FFDDF70C3DD58522F8F7" blockId="39.[811,1457,1012,1900]" pageId="39" pageNumber="102">
Plants
<quantity id="E1329B15FF84FFDDF7533DD483ADFB94" box="[882,967,1013,1039]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="39" pageNumber="102" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3-4 mm</quantity>
wide, to
<quantity id="E1329B15FF84FFDDF0043DD48433FB94" box="[1061,1113,1013,1039]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="39" pageNumber="102" unit="cm" value="4.0">4 cm</quantity>
long, irregularly branched below and on broken shoots by intercalary branches, dichotomous above by terminal branches, rhizome-like creeping shoot present. Mature stem leaves distant to contiguous, widely and horizontally spreading (not ventrad), narrowly and subsymmetrically oblong-rectangular, 2.5-3× longer than wide, on branches up to 4× longer than wide, not caducous, not ampliate and not shouldered, margins almost parallel, unbordered or weakly bordered, with 5-8 teeth in the upper half made up of rectangular cells; leaf apex bifid by two long, lobe-like teeth, to 10-15 cells long, the two teeth separated by a wide rounded sinus, sometimes one of the two teeth is broken, a smaller tooth may be present in the sinus between the two large lobe-like teeth and occasionally the apex is trifid, tip cell of the teeth not much longer than the rest of the cells, distal portion of ventral and dorsal leaf margin with 1-3 smaller teeth, lower half of the margins and bases entire; leaf bases shortly and narrowly decurrent, the decurrent ventral part maximally
<quantity id="E1329B15FF84FFDDF7FA3813842AF9D6" box="[987,1088,1586,1613]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="39" pageNumber="102" unit="mm" value="0.15">0.15 mm</quantity>
long. Leaf cells subisodiametrical to slightly elongate, 1-1.5× longer than wide, 18-30 µm wide in midleaf, trigones small but not swollen, triangular with concave walls and frequently radiate along the longer walls, cells along the margin sometimes more elongate and with a thicker inner tangential wall, forming a weak border, leaf base without vitta-like area, cuticle smooth; oil bodies not observed. Underleaves absent. Androecia not seen. Perianth campanulate, slightly elevated above the bracts on a short, thickened stalk. Vegetative reproduction not seen.
</paragraph>
<paragraph id="267536F0FF84FFDDF70C39B38312F831" blockId="39.[813,888,1938,1964]" box="[813,888,1938,1964]" pageId="39" pageNumber="102">
<heading id="7D3D819CFF84FFDDF70C39B38312F831" box="[813,888,1938,1964]" fontSize="9" level="3" pageId="39" pageNumber="102" reason="5">
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</heading>
</paragraph>
<paragraph id="267536F0FF84FFD3F70C3990821BFE4B" blockId="39.[813,1456,1969,2027]" lastBlockId="41.[130,776,215,593]" lastPageId="41" lastPageNumber="104" pageId="39" pageNumber="102">
<taxonomicName id="E1CA4D73FF84FFDDF70C399084C6F850" authority="Gradst." authorityName="Gradst." authorityYear="2022" box="[813,1196,1969,1995]" class="Jungermanniopsida" family="Plagiochilaceae" genus="Plagiochila" kingdom="Plantae" order="Jungermanniales" pageId="39" pageNumber="102" phylum="Marchantiophyta" rank="species" species="vandenberghenii" status="sp. nov.">
<emphasis id="14BEEAE2FF84FFDDF70C39908438F850" box="[813,1106,1969,1995]" italics="true" pageId="39" pageNumber="102">Plagiochila vandenberghenii</emphasis>
Gradst.
</taxonomicName>
,
<taxonomicNameLabel id="0F8D5799FF84FFDDF09939938561F857" box="[1208,1291,1970,1996]" pageId="39" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
(sect.
<emphasis id="14BEEAE2FF84FFDDF172399385E4F850" box="[1363,1422,1970,1995]" italics="true" pageId="39" pageNumber="102">Vagae</emphasis>
) is readily distinguished from all other African members of the genus
<taxonomicName id="E1CA4D73FF8AFFD3F4EC3EF68129FF6A" box="[205,323,215,241]" class="Jungermanniopsida" family="Plagiochilaceae" genus="Plagiochila" kingdom="Plantae" order="Jungermanniales" pageId="41" pageNumber="104" phylum="Marchantiophyta" rank="genus">
<emphasis id="14BEEAE2FF8AFFD3F4EC3EF68129FF6A" box="[205,323,215,241]" italics="true" pageId="41" pageNumber="104">Plagiochila</emphasis>
</taxonomicName>
by the horizontally spreading, narrowly oblong-rectangular, non-ampliate leaves (
<emphasis id="14BEEAE2FF8AFFD3F6113ED98253FE8A" box="[560,569,248,273]" italics="true" pageId="41" pageNumber="104">c</emphasis>
. 2.5-3× longer than wide) with a strongly bifid apex, with two long teeth. In its horizontally spreading, bifid leaves, the new species cannot be confused with any other African species of
<taxonomicName id="E1CA4D73FF8AFFD3F6163F7682CCFEEA" box="[567,678,343,369]" class="Jungermanniopsida" family="Plagiochilaceae" genus="Plagiochila" kingdom="Plantae" order="Jungermanniales" pageId="41" pageNumber="104" phylum="Marchantiophyta" rank="genus">
<emphasis id="14BEEAE2FF8AFFD3F6163F7682CCFEEA" box="[567,678,343,369]" italics="true" pageId="41" pageNumber="104">Plagiochila</emphasis>
</taxonomicName>
and only resembles the Asian
<taxonomicName id="E1CA4D73FF8AFFD3F5743F568273FE0A" authority="Steph." authorityName="Steph." box="[341,537,375,401]" class="Jungermanniopsida" family="Plagiochilaceae" genus="Plagiochila" kingdom="Plantae" order="Jungermanniales" pageId="41" pageNumber="104" phylum="Marchantiophyta" rank="species" species="bicornuta">
<emphasis id="14BEEAE2FF8AFFD3F5743F5681A4FE0A" box="[341,462,375,401]" italics="true" pageId="41" pageNumber="104">P. bicornuta</emphasis>
Steph.
</taxonomicName>
(synonym:
<taxonomicName id="E1CA4D73FF8AFFD3F6B33F5680B0FE2B" authority="Schiffn." authorityName="Schiffn." class="Jungermanniopsida" family="Plagiochilaceae" genus="Plagiochila" kingdom="Plantae" order="Jungermanniales" pageId="41" pageNumber="104" phylum="Marchantiophyta" rank="species" species="laxissima">
<emphasis id="14BEEAE2FF8AFFD3F6B33F56836FFE0A" box="[658,773,375,401]" italics="true" pageId="41" pageNumber="104">P. laxissima</emphasis>
Schiffn.
</taxonomicName>
;
<emphasis id="14BEEAE2FF8AFFD3F4C63FB78166FE2B" box="[231,268,406,432]" italics="true" pageId="41" pageNumber="104">fide</emphasis>
<bibRefCitation id="425B4B01FF8AFFD3F5353FB68105FE2A" author="SO M. L." box="[276,367,407,433]" pageId="41" pageNumber="104" pagination="179 - 186" refId="ref27496" refString="SO M. L. 2001. - On Plagiochila sect. Cobanae Carl in Asia and Melanesia. Cryptogamie, Bryologie 22 (3): 179 - 186. https: // doi. org / 10.1016 / S 1290 - 0796 (01) 90003 - 3" type="journal article" year="2001">So 2001</bibRefCitation>
), but the apical teeth in latter species are smaller and terminal branching is lacking.
</paragraph>
<paragraph id="267536F0FF8BFFD2F7283EFB8134F9AA" pageId="40" pageNumber="103">
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D
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</paragraph>
<caption id="72B56678FF8BFFD2F4A5398B85C0F872" pageId="40" pageNumber="103" startId="40.[132,143,1962,1980]" targetType="figure">
<paragraph id="267536F0FF8BFFD2F4A5398B85C0F872" blockId="40.[132,1455,1962,2025]" pageId="40" pageNumber="103">
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. 23. ―
<taxonomicName id="E1CA4D73FF8BFFD2F4FF398B8264F827" authority="Gradst." authorityName="Gradst." authorityYear="2022" box="[222,526,1962,1980]" class="Jungermanniopsida" family="Plagiochilaceae" genus="Plagiochila" kingdom="Plantae" order="Jungermanniales" pageId="40" pageNumber="103" phylum="Marchantiophyta" rank="species" species="vandenberghenii" status="sp. nov.">
<emphasis id="14BEEAE2FF8BFFD2F4FF398B81ACF827" box="[222,454,1962,1980]" italics="true" pageId="40" pageNumber="103">Plagiochila vandenberghenii</emphasis>
Gradst.
</taxonomicName>
,
<taxonomicNameLabel id="0F8D5799FF8BFFD2F632398B8233F827" box="[531,601,1962,1980]" pageId="40" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
:
<emphasis id="14BEEAE2FF8BFFD2F67F398B8201F827" bold="true" box="[606,619,1962,1980]" pageId="40" pageNumber="103">A</emphasis>
, shoots;
<emphasis id="14BEEAE2FF8BFFD2F699398B82AFF827" bold="true" box="[696,709,1962,1980]" pageId="40" pageNumber="103">B</emphasis>
, shoot in dorsal view with terminal branching;
<emphasis id="14BEEAE2FF8BFFD2F066398B843FF827" bold="true" box="[1095,1109,1962,1980]" pageId="40" pageNumber="103">C</emphasis>
, intercalary branching;
<emphasis id="14BEEAE2FF8BFFD2F139398B854CF827" bold="true" box="[1304,1318,1962,1980]" pageId="40" pageNumber="103">D</emphasis>
, shoot in ventral view;
<emphasis id="14BEEAE2FF8BFFD2F49439E080ABF848" bold="true" box="[181,193,1985,2003]" pageId="40" pageNumber="103">E</emphasis>
, stem leaves;
<emphasis id="14BEEAE2FF8BFFD2F51839E0812EF848" bold="true" box="[313,324,1985,2003]" pageId="40" pageNumber="103">F</emphasis>
, branch leaves;
<emphasis id="14BEEAE2FF8BFFD2F5EF39E181B6F849" bold="true" box="[462,476,1984,2002]" pageId="40" pageNumber="103">G</emphasis>
, branch in dorsal view;
<emphasis id="14BEEAE2FF8BFFD2F68239E082DBF848" bold="true" box="[675,689,1985,2003]" pageId="40" pageNumber="103">H</emphasis>
,
<emphasis id="14BEEAE2FF8BFFD2F69A39E082ABF848" bold="true" box="[699,705,1985,2003]" pageId="40" pageNumber="103">I</emphasis>
, leaf margin cells;
<emphasis id="14BEEAE2FF8BFFD2F77C39E0830DF848" bold="true" box="[861,871,1985,2003]" pageId="40" pageNumber="103">J</emphasis>
, leaf base;
<emphasis id="14BEEAE2FF8BFFD2F7E939E083BFF848" bold="true" box="[968,981,1985,2003]" pageId="40" pageNumber="103">K</emphasis>
, leaf apex;
<emphasis id="14BEEAE2FF8BFFD2F01639E08428F848" bold="true" box="[1079,1090,1985,2003]" pageId="40" pageNumber="103">L</emphasis>
, shoot with perianth, two leaves at perianth base removed to show the perianth stalk;
<emphasis id="14BEEAE2FF8BFFD2F5C339F68199F872" bold="true" box="[482,499,2007,2025]" pageId="40" pageNumber="103">M</emphasis>
, female bracts. From
<emphasis id="14BEEAE2FF8BFFD2F68839F68352F872" box="[681,824,2007,2025]" italics="true" pageId="40" pageNumber="103">Reeb CR17M610</emphasis>
(PC). Scale bars: A, 5 mm; B-G, J, K, 1 mm; H, I, L, 0.1 mm; M, 0.025 mm.
</paragraph>
</caption>
<paragraph id="267536F0FF8AFFD3F4BA3FF7821EFDCB" blockId="41.[130,776,215,593]" pageId="41" pageNumber="104">
The perianths in the new species are campanulate.According to
<bibRefCitation id="425B4B01FF8AFFD3F4BE3FD78132FD8B" author="HEINRICHS J." box="[159,344,502,528]" pageId="41" pageNumber="104" pagination="1 - 184" refId="ref26203" refString="HEINRICHS J. 2002. - A taxonomic revision of Plagiochila sect. Hylacoetes, sect. Adiantoideae and sect. Fuscoluteae in the Neotropics, with a preliminary subdivision of neotropical Plagiochilaceae into nine lineages. Bryophytorum Bibliotheca 58: 1 - 184." type="journal article" year="2002">Heinrichs (2002)</bibRefCitation>
, campanulate perianths are characteristic of unfertilized gynoecia, but the perianths in
<taxonomicName id="E1CA4D73FF8AFFD3F6433C36816CFDCB" authority="Gradst." authorityName="Gradst." authorityYear="2022" class="Jungermanniopsida" family="Plagiochilaceae" genus="Plagiochila" kingdom="Plantae" order="Jungermanniales" pageId="41" pageNumber="104" phylum="Marchantiophyta" rank="species" species="vandenberghenii" status="sp. nov.">
<emphasis id="14BEEAE2FF8AFFD3F6433C3680C1FDCB" italics="true" pageId="41" pageNumber="104">P. vandenberghenii</emphasis>
Gradst.
</taxonomicName>
,
<taxonomicNameLabel id="0F8D5799FF8AFFD3F5313C168109FDCA" box="[272,355,567,593]" pageId="41" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
had mature sporophytes.
</paragraph>
</subSubSection>
</treatment>
</document>