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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.115.1163" ID-GBIF-Dataset="16898250-8f0c-44c7-aa0b-23e070214ec3" ID-PMC="PMC3187662" ID-Pensoft-Pub="1313-2970-115-1" ID-PubMed="21976997" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-115-1" ModsDocOrigin="ZooKeys 115" ModsDocTitle="Description of two new species of Rissoella Gray, 1847 (Mollusca, Gastropoda, Heterobranchia) from Venezuela, with a key to the Caribbean species known for the genus" checkinTime="1451250215506" checkinUser="pensoft" docAuthor="Caballer, Manuel, Ortea, Jesus &amp; Narciso, Samuel" docDate="2011" docId="3CB0C1C584433E20E7D8376EB3347C41" docLanguage="en" docName="ZooKeys 115: 1-18" docOrigin="ZooKeys 115" docSource="http://dx.doi.org/10.3897/zookeys.115.1163" docTitle="Rissoella venezolanicola Caballer, Ortea &amp; Narciso, 2011, sp. n." docType="treatment" docVersion="3" lastPageNumber="9" masterDocId="EF0BFFD9214D3D64FFDFFFFE0C5A174C" masterDocTitle="Description of two new species of Rissoella Gray, 1847 (Mollusca, Gastropoda, Heterobranchia) from Venezuela, with a key to the Caribbean species known for the genus" masterLastPageNumber="18" masterPageNumber="1" pageNumber="7" updateTime="1668151816075" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Description of two new species of Rissoella Gray, 1847 (Mollusca, Gastropoda, Heterobranchia) from Venezuela, with a key to the Caribbean species known for the genus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Caballer, Manuel</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ortea, Jesus</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Narciso, Samuel</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>115</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>18</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.115.1163</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.115.1163</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-115-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152030126" LSID="urn:lsid:plazi:treatment:3CB0C1C584433E20E7D8376EB3347C41" httpUri="http://treatment.plazi.org/id/3CB0C1C584433E20E7D8376EB3347C41" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection pageId="6" pageNumber="7" type="nomenclature">
<paragraph pageId="6" pageNumber="7">
<taxonomicName LSID="urn:lsid:zoobank.org:act:69FA6D90-9365-4ADD-92DA-820BC27EC917" class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella venezolanicola" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="venezolanicola">Rissoella venezolanicola</taxonomicName>
<taxonomicNameLabel pageId="6" pageNumber="7">sp. n.</taxonomicNameLabel>
Figs 6
<normalizedToken originalValue="–811–1218–">-811-1218-</normalizedToken>
22Tabs 134
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="description">
<paragraph pageId="6" pageNumber="7">Description.</paragraph>
<paragraph pageId="6" pageNumber="7">Shell very small (Tab. 3), smooth and translucent (Fig. 18); protoconch about half a whorl after the nucleus (Fig. 19); teleoconch of two whorls and three quarters; aperture oval, with the columella slightly bowed and arched; umbilicus narrow and deep, slightly closed by the expansion of the columellar edge; preumbilical cord extended, with an angle delimiting and closing the umbilicus forming a trapezoid (Figs 11-12, 20); average height of the last whorl, 82.9% of shell length; average height of the aperture, 49.6% of shell length (Tab. 3); Shell length/width ratio = 1.52; spiral angle = 58°; umbilical angle = 23°-25°.</paragraph>
<caption pageId="6" pageNumber="7">
<paragraph pageId="6" pageNumber="7">
Figures 18-22.
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella venezolanicola" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="venezolanicola">Rissoella venezolanicola</taxonomicName>
sp. n.: shell (18), protoconch (19), detail of the umbilicus (20), odontophoral cartilages (21), radular teeth (22).
</paragraph>
</caption>
<caption pageId="6" pageNumber="7">
<paragraph pageId="6" pageNumber="7">
Figures 23-31. Color illustrations adapted from the original descriptions (
<bibRefCitation pageId="6" pageNumber="7">Ortea and Espinosa 2001</bibRefCitation>
;
<bibRefCitation author="Ortea, J" journalOrPublisher="Avicennia" pageId="11" pageNumber="12" pagination="77 - 94" title="Una combinacion de ciencia, arte y naturaleza: especies nuevas del genero Rissoella J. E. Gray, 1847 (Gastropoda, Heterobranchia) descritas en homenaje a las artistas de la plastica cubana." volume="17" year="2004">2004</bibRefCitation>
;
<bibRefCitation author="Espinosa, J" journalOrPublisher="Revista de la Academia Canaria de Ciencias" pageId="11" pageNumber="12" pagination="15 - 17" title="Nueva especie de Rissoella J. E. Gray, 1847 (Gastropoda: Heterobranchia) de la Peninsula de Guanahacabibes, Pinar del Rio, Cuba" volume="XX" year="2009">Espinosa and Ortea 2009</bibRefCitation>
).
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella gandocaensis" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="gandocaensis">Rissoella gandocaensis</taxonomicName>
(23),
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella ameliae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="ameliae">Rissoella ameliae</taxonomicName>
(24),
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella belkisae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="belkisae">Rissoella belkisae</taxonomicName>
(25),
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella dianae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="dianae">Rissoella dianae</taxonomicName>
(26),
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella zaidae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="zaidae">Rissoella zaidae</taxonomicName>
(27),
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella florae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="florae">Rissoella florae</taxonomicName>
(28),
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella elsae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="elsae">Rissoella elsae</taxonomicName>
(29),
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella taniae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="taniae">Rissoella taniae</taxonomicName>
(30),
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella aliciae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="aliciae">Rissoella aliciae</taxonomicName>
(31).
</paragraph>
</caption>
<paragraph pageId="6" pageNumber="7">Operculum oval, translucent with an amber tinge, membranous consistence; head, flanks, oral lobes and cephalic tentacles translucent white with scattered black to greenish brown dots (Fig. 6); eyes black; foot translucent white (Fig. 7); hypobranchial gland translucent white with scattered blotches black to greenish brown and a distinctive yellow design quaver-shaped (Fig. 8); visceral mass in the first whorls, black; protoconch with fuchsia highlights in live animals.</paragraph>
<paragraph pageId="6" pageNumber="7">
Odontophoral cartilages kidney-shaped, with large uncini placed longitudinally (Fig. 21); radular formula of an specimen 0.96 mm shell length, 16
<normalizedToken originalValue="×">x</normalizedToken>
1.1.R.1.1; rachidian tooth wide, with bilobed apex forming two smooth cusps (Fig. 22), slightly larger than marginal tooth; lateral tooth triangular, 28.3
<normalizedToken originalValue="µm">µm</normalizedToken>
long, with the apex hooked and smooth; imbricated with the opposite tooth above the rachidian; marginal tooth triangular and curved, 16.3
<normalizedToken originalValue="µm">µm</normalizedToken>
long, with blunt apex.
</paragraph>
<caption ID-Table-UUID="371D23D744C4B28ADE15D4C307F85EC6" httpUri="http://table.plazi.org/id/371D23D744C4B28ADE15D4C307F85EC6" pageId="6" pageNumber="7">
<paragraph pageId="6" pageNumber="7">
Table 3.
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella venezolanicola" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="venezolanicola">Rissoella venezolanicola</taxonomicName>
sp. n. Measurements of specimens. (%) means percentage compared to the total length of the shell.
</paragraph>
</caption>
<paragraph pageId="6" pageNumber="7">
<table pageId="6" pageNumber="7">
<tr pageId="6" pageNumber="7">
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">Protoconch (whorls)</th>
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">Last whorl (mm)</th>
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">Aperture(mm)</th>
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">Length /width</th>
</tr>
</table>
</paragraph>
<caption pageId="6" pageNumber="7">
<paragraph pageId="6" pageNumber="7">
Table 4. Synthesis of characters of the Caribbean species of
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">Rissoella</taxonomicName>
Gray, 1847 based on
<bibRefCitation author="Robertson, R" journalOrPublisher="The Nautilus" pageId="11" pageNumber="12" pagination="131 - 136" title="A second Western Atlantic Rissoella and a list of the species in the Rissoellidae." volume="74" year="1961">Robertson (1961)</bibRefCitation>
,
<bibRefCitation author="Wise, JB" journalOrPublisher="The Nautilus" pageId="11" pageNumber="12" pagination="13 - 21" title="Morphology and systematic position of Rissoella caribaea Rehder, 1943 (Gastropoda: Heterobranchia: Rissoellidae)." volume="111" year="1998">Wise (1998)</bibRefCitation>
,
<bibRefCitation pageId="6" pageNumber="7">Ortea and Espinosa (2001</bibRefCitation>
;
<bibRefCitation author="Ortea, J" journalOrPublisher="Avicennia" pageId="11" pageNumber="12" pagination="77 - 94" title="Una combinacion de ciencia, arte y naturaleza: especies nuevas del genero Rissoella J. E. Gray, 1847 (Gastropoda, Heterobranchia) descritas en homenaje a las artistas de la plastica cubana." volume="17" year="2004">2004</bibRefCitation>
),
<bibRefCitation author="Espinosa, J" journalOrPublisher="Revista de la Academia Canaria de Ciencias" pageId="11" pageNumber="12" pagination="15 - 17" title="Nueva especie de Rissoella J. E. Gray, 1847 (Gastropoda: Heterobranchia) de la Peninsula de Guanahacabibes, Pinar del Rio, Cuba" volume="XX" year="2009">Espinosa and Ortea (2009)</bibRefCitation>
. Aa = Average height of the aperture vs shell length, Al = Average height of the last whorl vs shell length, E = Eyes, H = Habitat, HG = Hypobranchial gland, LW = Shell length/width ratio, Pc = Preumbilical cord, Pf = Pigmentation of the body in the firsts whorls, Rf = Radular formula, Rt = lateral and marginal teeth, Sa = Spiral angle, U = Umbilicus, Ua = Umbilical angle.
</paragraph>
</caption>
<paragraph pageId="6" pageNumber="7">
<table pageId="6" pageNumber="7">
<tr pageId="6" pageNumber="7">
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">Shell</th>
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">Al</th>
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">Aa</th>
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">LW</th>
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">Sa</th>
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">Ua</th>
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">U</th>
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">Pc</th>
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">E</th>
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">HG</th>
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">Band around HG</th>
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">Pf</th>
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">Rf</th>
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">Rt</th>
<th colspan="1" pageId="6" pageNumber="7" rowspan="1">H</th>
</tr>
<tr pageId="6" pageNumber="7">
<td colspan="1" pageId="6" pageNumber="7" rowspan="1">
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella caribaea" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="caribaea">Rissoella caribaea</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="7">
<td colspan="1" pageId="6" pageNumber="7" rowspan="1">
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella galba" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="galba">Rissoella galba</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="7">
<td colspan="1" pageId="6" pageNumber="7" rowspan="1">
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella gandocaensis" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="gandocaensis">Rissoella gandocaensis</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="7">
<td colspan="1" pageId="6" pageNumber="7" rowspan="1">
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella ameliae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="ameliae">Rissoella ameliae</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="7">
<td colspan="1" pageId="6" pageNumber="7" rowspan="1">
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella belkisae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="belkisae">Rissoella belkisae</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="7">
<td colspan="1" pageId="6" pageNumber="7" rowspan="1">
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella dianae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="dianae">Rissoella dianae</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="7">
<td colspan="1" pageId="6" pageNumber="7" rowspan="1">
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella zaidae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="zaidae">Rissoella zaidae</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="7">
<td colspan="1" pageId="6" pageNumber="7" rowspan="1">
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella florae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="florae">Rissoella florae</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="7">
<td colspan="1" pageId="6" pageNumber="7" rowspan="1">
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella elsae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="elsae">Rissoella elsae</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="7">
<td colspan="1" pageId="6" pageNumber="7" rowspan="1">
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella taniae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="taniae">Rissoella taniae</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="7">
<td colspan="1" pageId="6" pageNumber="7" rowspan="1">
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella aliciae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="aliciae">Rissoella aliciae</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="7">
<td colspan="1" pageId="6" pageNumber="7" rowspan="1">
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella morrocoyensis" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="morrocoyensis">Rissoella morrocoyensis</taxonomicName>
</td>
</tr>
<tr pageId="6" pageNumber="7">
<td colspan="1" pageId="6" pageNumber="7" rowspan="1">
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella venezolanicola" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="venezolanicola">Rissoella venezolanicola</taxonomicName>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="types">
<paragraph pageId="6" pageNumber="7">Types.</paragraph>
<paragraph pageId="6" pageNumber="7">
Holotype, 1.02
<normalizedToken originalValue="µm">µm</normalizedToken>
<normalizedToken originalValue="×">x</normalizedToken>
0.66
<normalizedToken originalValue="µm">µm</normalizedToken>
, collected alive (August 2, 2010), preserved dry and deposited in FUDENA (CFRFSIA0002). Paratype 1, 0.96
<normalizedToken originalValue="×">x</normalizedToken>
0.64
<normalizedToken originalValue="µm">µm</normalizedToken>
, SOM-IVIC (IVICCMT008), used to get the radula, shell preserved dry. Paratype 2, 0.54
<normalizedToken originalValue="×">x</normalizedToken>
0.46
<normalizedToken originalValue="µm">µm</normalizedToken>
, preserved dry, SOM-IVIC (IVICCMT009).
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="type locality">
<paragraph pageId="6" pageNumber="7">Type locality.</paragraph>
<paragraph pageId="6" pageNumber="7">
Leeward patch reef, Isla de Aves, Venezuela (
<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.665055">15°39'54.2&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-63.621555">63°37'17.6&quot;W</geoCoordinate>
), on
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Dictyota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dictyota" order="Orthoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Dictyota</taxonomicName>
spp., 10 m depth.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="further localities">
<paragraph pageId="6" pageNumber="7">Further localities.</paragraph>
<paragraph pageId="6" pageNumber="7">
Paratypes: Northern end, Isla de Aves, Venezuela, (
<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.673528">15°40'24.7&quot;N</geoCoordinate>
,
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-63.61972">63°37'11&quot;W</geoCoordinate>
), on
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Dictyota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dictyota" order="Orthoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Dictyota</taxonomicName>
spp., 10 m depth.
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="7" type="etymology">
<paragraph pageId="6" pageNumber="7">Etymology.</paragraph>
<paragraph pageId="6" pageNumber="7">venezolanicola latinization of venezolana, inhabitant of Venezuela.</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="remarks">
<paragraph pageId="6" pageNumber="7">Remarks.</paragraph>
<paragraph lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
Due to the presence of the preumbilical cord,
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella venezolanicola" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="venezolanicola">Rissoella venezolanicola</taxonomicName>
sp. n., is comparable to
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella zaidae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="zaidae">Rissoella zaidae</taxonomicName>
,
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella florae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="florae">Rissoella florae</taxonomicName>
,
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella elsae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="elsae">Rissoella elsae</taxonomicName>
,
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella taniae" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="taniae">Rissoella taniae</taxonomicName>
(Ortea and Espinosa 2004) (listed above) and
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella morrocoyensis" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="morrocoyensis">Rissoella morrocoyensis</taxonomicName>
sp. n. But it is different because in none of them the preumbilical cord delimits and closes the umbilicus, neither the hypobranchial gland
<pageBreakToken pageId="7" pageNumber="8" start="start">has</pageBreakToken>
a yellow quaver-shaped design. Additionally,
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella venezolanicola" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="venezolanicola">Rissoella venezolanicola</taxonomicName>
sp. n. differs from all these species by:
</paragraph>
<paragraph lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
-
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella zaidae" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="zaidae">Rissoella zaidae</taxonomicName>
: (shell length/width ratio = 1.63, spiral angle = 50°, umbilical angle = 30°) the whorls of the spire are more angled and tilted, the umbilicus is in the middle of the last whorl, the head, oral lobes, cephalic tentacles and the visceral mass in the first whorls are lilac and the hypobranchial gland is kidney-
<pageBreakToken pageId="8" pageNumber="9" start="start">shaped</pageBreakToken>
, pale violet with lilac blotches and bounded by a yellow ribbon with transverse bands.
</paragraph>
<paragraph pageId="8" pageNumber="9">
-
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella florae" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="florae">Rissoella florae</taxonomicName>
: (shell length/width ratio = 1.35, umbilical angle = 18°) the head and oral lobes are black, the cephalic tentacles are different in colour than the oral lobes, the eyes are located in lighter areas of orange hue, the visceral mass in the first whorls is brown or orange and the hypobranchial gland is lemon-yellow without any design on it.
</paragraph>
<paragraph pageId="8" pageNumber="9">
-
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella elsae" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="elsae">Rissoella elsae</taxonomicName>
: (shell length/width ratio = 1.69, spiral angle = 45°) the shell is more conical, the preumbilical cord is higher than the columellar wall, the head, oral lobes and cephalic tentacles are lilac, the mantle is green and hides the eyes, and the hypobranchial gland has large patches of dark green and is enclosed by a white ribbon with transverse bands.
</paragraph>
<paragraph pageId="8" pageNumber="9">
-
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella taniae" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="taniae">Rissoella taniae</taxonomicName>
: (shell length/width ratio = 1.4, spiral angle = 65°, umbilical angle = 43°) the shell is spherical with slightly globose whorls, the umbilicus is very open, the head, oral lobes, cephalic tentacles and visceral mass in the first whorls are dark green, there are no eyes apparently and the hypobranchial gland is white with green spots.
</paragraph>
<paragraph pageId="8" pageNumber="9">
-
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella morrocoyensis" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="morrocoyensis">Rissoella morrocoyensis</taxonomicName>
sp. n.: (shell length/width ratio = 1.32) the umbilicus is deeper and more closed by the expansion of the columellar edge and the head, oral lobes and cephalic tentacles are black.
</paragraph>
<paragraph pageId="8" pageNumber="9">
Two species from Africa have a design on the hypobranchial gland similar to
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella venezolanicola" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="venezolanicola">Rissoella venezolanicola</taxonomicName>
sp. n.:
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella luteonigra" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="luteonigra">Rissoella luteonigra</taxonomicName>
<normalizedToken originalValue="Rolán">Rolan</normalizedToken>
&amp; Rubio, 2001 and
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella trigoi" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="trigoi">Rissoella trigoi</taxonomicName>
<normalizedToken originalValue="Rolán">Rolan</normalizedToken>
&amp;
<normalizedToken originalValue="Hernández">Hernandez</normalizedToken>
, 2004.
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella venezolanicola" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="venezolanicola">Rissoella venezolanicola</taxonomicName>
sp. n. differs from these species in:
</paragraph>
<paragraph pageId="8" pageNumber="9">
-
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella luteonigra" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="luteonigra">Rissoella luteonigra</taxonomicName>
;is bigger (1.8 mm), with a narrower and longer shell, lacking preumbilical cord. Animals are black homogeneous (
<bibRefCitation author="Rolan, E" journalOrPublisher="Novapex" pageId="11" pageNumber="12" pagination="133 - 136" title="New species of the genera Elachisina and Rissoella (Mollusca, Gastropoda) from the Cape Verde Archipelago." volume="2" year="2001">
<normalizedToken originalValue="Rolán">Rolan</normalizedToken>
and Rubio 2001
</bibRefCitation>
).
</paragraph>
<paragraph pageId="8" pageNumber="9">
-
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella trigoi" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="trigoi">Rissoella trigoi</taxonomicName>
; has a rough protoconch with cavities in the suture, the preumbilical cord parallel to the expansion of the columellar edge until it converges (divergent in
<taxonomicName class="Gastropoda" family="Rissoellidae" genus="Rissoella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rissoella venezolanicola" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="venezolanicola">Rissoella venezolanicola</taxonomicName>
sp. n.) and the animals are black with a white drawing in the hypobranchial gland (
<bibRefCitation author="Rolan, E" journalOrPublisher="Revista de la Academia Canaria de Ciencias" pageId="11" pageNumber="12" pagination="229 - 235" title="Dos nuevas especies de Rissoella (Mollusca: Rissoellidae) de las islas Canarias y Senegal" volume="XV" year="2004">
<normalizedToken originalValue="Rolán">Rolan</normalizedToken>
and
<normalizedToken originalValue="Hernández">Hernandez</normalizedToken>
2004
</bibRefCitation>
), very simple and very different.
</paragraph>
</subSubSection>
</treatment>
</document>
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