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<document id="429F8EDD207EF107CFBE0F0D67DD95E8" ID-CLB-Dataset="55858" ID-DOI="10.5852/ejt.2022.816.1747" ID-GBIF-Dataset="5cfc08bb-f51f-47bb-8a02-754e4eda0162" ID-ISSN="2118-9773" ID-Zenodo-Dep="6484261" ID-ZooBank="C150007D-80F9-4C34-9F85-BDB1211B244D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1650913289141" checkinUser="felipe" docAuthor="Psarras, Christos, Merle, Didier &amp; Koskeridou, Efterpi" docDate="2022" docId="EE28878CDD065A2EFD8DFA2ECD57CFAA" docLanguage="en" docName="ejt-816_1-70.pdf" docOrigin="European Journal of Taxonomy 816" docSource="http://dx.doi.org/10.5852/ejt.2022.816.1747" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Conus (Lautoconus) Monterosato 1923" docType="treatment" docVersion="9" lastPageNumber="5" masterDocId="1211FFF4DD025A2BFF99FFAFCC26CE4A" masterDocTitle="Late Miocene Conidae (Mollusca: Gastropoda) of Crete (Greece). Part 2" masterLastPageNumber="70" masterPageNumber="1" pageNumber="5" updateTime="1699329182685" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="90DB20A5FE80D79C44CE8B930FA878A6">Late Miocene Conidae (Mollusca: Gastropoda) of Crete (Greece). Part 2</mods:title>
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<mods:namePart id="A4B915BF9C878A03C2057982E59FADED">Psarras, Christos</mods:namePart>
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<mods:affiliation id="093C26027A17CF8666A7F839C72F7C39">National and Kapodistrian University of Athens, Faculty of Geology and Geoenvironment, Department of Historical Geology-Paleontology, Athens, Greece.</mods:affiliation>
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<mods:namePart id="6BC59B0980D01D7C8CC49C7D389D1D44">Merle, Didier</mods:namePart>
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<mods:affiliation id="25F68CD19FB988F3B7EC0FCDF2E27376">Sorbonne Université - CR 2 P - MNHN, CNRS, UPMC-Paris 6 and Muséum national dhistoire naturelle, Département Origine et Évolution, 8 rue Buffon 75005 Paris, France.</mods:affiliation>
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<mods:affiliation id="33DFB32381097CC9F0EF5CDDE33B06BB">National and Kapodistrian University of Athens, Faculty of Geology and Geoenvironment, Department of Historical Geology-Paleontology, Athens, Greece.</mods:affiliation>
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<subSubSection id="2E9B6511DD065A2FFD8DFA2EC84DCBD1" box="[532,1131,1408,1435]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="663E369ADD065A2FFD8DFA2EC84DCBD1" blockId="4.[532,1131,1408,1435]" box="[532,1131,1408,1435]" pageId="4" pageNumber="5">
<heading id="3D7681F6DD065A2FFD8DFA2EC84DCBD1" box="[532,1131,1408,1435]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
Subgenus
<taxonomicName id="A1814D19DD065A2FFD14FA2EC84DCBD1" authority="Monterosato, 1923" authorityName="Monterosato" authorityYear="1923" box="[653,1131,1408,1435]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="subGenus" subGenus="Lautoconus">
<emphasis id="54F5EA88DD065A2FFD14FA2ECFA0CBD0" bold="true" box="[653,902,1408,1435]" italics="true" pageId="4" pageNumber="5">Conus (Lautoconus)</emphasis>
Monterosato, 1923
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2E9B6511DD065A2FFF7AFA6CC9A8C846" pageId="4" pageNumber="5" type="type_taxon">
<paragraph id="663E369ADD065A2FFF7AFA6CCD5BCB97" blockId="4.[227,381,1475,1501]" box="[227,381,1475,1501]" pageId="4" pageNumber="5">
<heading id="3D7681F6DD065A2FFF7AFA6CCD5BCB97" bold="true" box="[227,381,1475,1501]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="54F5EA88DD065A2FFF7AFA6CCD5BCB97" bold="true" box="[227,381,1475,1501]" pageId="4" pageNumber="5">
<typeStatus id="B93A8838DD065A2FFF7AFA6CCD06CB97" box="[227,288,1475,1501]" pageId="4" pageNumber="5">Type</typeStatus>
species
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<paragraph id="663E369ADD065A2FFF7AFA5CC9A8C846" blockId="4.[227,1422,1522,1549]" box="[227,1422,1522,1549]" pageId="4" pageNumber="5">
<taxonomicName id="A1814D19DD065A2FFF7AFA5CCE12C847" ID-CoL="XXTL" authority="Hwass" authorityName="Hwass" box="[227,564,1522,1549]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="mediterraneus">
<emphasis id="54F5EA88DD065A2FFF7AFA5CCDF8C846" box="[227,478,1522,1549]" italics="true" pageId="4" pageNumber="5">Conus mediterraneus</emphasis>
Hwass
</taxonomicName>
in
<treatmentCitation id="E720108BDD065A2FFDC3FA5DCF3FC846" box="[602,793,1522,1548]" pageId="4" pageNumber="5">Bruguière, 1792</treatmentCitation>
(Recent, Mediterranean Sea) by original designation.
</paragraph>
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<subSubSection id="2E9B6511DD065A2FFF7AF99BCFE2C947" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="663E369ADD065A2FFF7AF99BCD7AC804" blockId="4.[227,348,1588,1614]" box="[227,348,1588,1614]" pageId="4" pageNumber="5">
<heading id="3D7681F6DD065A2FFF7AF99BCD7AC804" bold="true" box="[227,348,1588,1614]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="54F5EA88DD065A2FFF7AF99BCD7AC804" bold="true" box="[227,348,1588,1614]" pageId="4" pageNumber="5">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="663E369ADD065A2FFF7AF9CBCFE2C947" blockId="4.[225,1437,1636,1806]" pageId="4" pageNumber="5">Moderately small to moderately large, turnip-shaped species. Medium-wide to wide shells and convex sides. Spire low to moderately high. Paucispiral protoconch. Early whorls ornamented with cords, which might persist on later spire whorls or appear as spiral threads. Subsutural flexure from very shallow to deep, usually moderately curved, moderately asymmetrical. colour pattern variable, displaying blotches, spiral rows of dots, dashes, and tents in multiple combinations.</paragraph>
</subSubSection>
<subSubSection id="2E9B6511DD065A2EFF7AF89ACD57CFAA" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="discussion">
<paragraph id="663E369ADD065A2FFF7AF89ACD73C905" blockId="4.[227,341,1845,1871]" box="[227,341,1845,1871]" pageId="4" pageNumber="5">
<heading id="3D7681F6DD065A2FFF7AF89ACD73C905" bold="true" box="[227,341,1845,1871]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="54F5EA88DD065A2FFF7AF89ACD73C905" bold="true" box="[227,341,1845,1871]" pageId="4" pageNumber="5">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="663E369ADD065A2EFF7AF8CBCD57CFAA" blockId="4.[225,1437,1892,2063]" lastBlockId="5.[226,1436,303,480]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<bibRefCitation id="02104B6BDD065A2FFF7AF8CBCDC3C935" author="Puillandre N. &amp; Bouchet P. &amp; Duda T. F. &amp; Kauferstein S. &amp; Kohn A. J. &amp; Olivera B. M. &amp; Watkins M. &amp; Meyer C." box="[227,485,1892,1919]" pageId="4" pageNumber="5" pagination="290 - 303" refId="ref31454" refString="Puillandre N., Bouchet P., Duda T. F., Kauferstein S., Kohn A. J., Olivera B. M., Watkins M. &amp; Meyer C. 2014. Molecular phylogeny and evolution of the cone snails (Gastropoda, Conoidea). Molecular Phylogenetics and Evolution 78: 290 - 303. https: // doi. org / 10.1016 / j. ympev. 2014.05.023" type="journal article" year="2014">
Puillandre
<emphasis id="54F5EA88DD065A2FFEF8F8CACDBCC934" box="[353,410,1892,1918]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
(2014
</bibRefCitation>
,
<bibRefCitation id="02104B6BDD065A2FFE76F8CBCE0DC935" author="Puillandre N. &amp; Duda T. F. &amp; Meyer C. &amp; Olivera B. M. &amp; Bouchet P." box="[495,555,1892,1919]" pageId="4" pageNumber="5" pagination="1 - 23" refId="ref31531" refString="Puillandre N., Duda T. F., Meyer C., Olivera B. M. &amp; Bouchet P. 2015. One, four or 100 genera? A new classification of the cone snails. The Journal of Molluscan Studies 81 (1): 1 - 23. https: // doi. org / 10.1093 / mollus / eyu 055" type="journal article" year="2015">2015</bibRefCitation>
) treated
<taxonomicName id="A1814D19DD065A2FFD17F8CACF31C934" authorityName="Monterosato" authorityYear="1923" box="[654,791,1893,1918]" class="Gastropoda" family="Conidae" genus="Lautoconus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
<emphasis id="54F5EA88DD065A2FFD17F8CACF31C934" box="[654,791,1893,1918]" italics="true" pageId="4" pageNumber="5">Lautoconus</emphasis>
</taxonomicName>
at the subgenus level. In this work we show two species that are large-sized (
<taxonomicName id="A1814D19DD065A2FFE44F826CF4FC9E8" authority="Psarras &amp; Merle &amp; Koskeridou, 2022" authorityName="Psarras &amp; Merle &amp; Koskeridou" authorityYear="2022" box="[477,873,1928,1955]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="damianakisi" status="sp. nov." subGenus="Lautoconus">
<emphasis id="54F5EA88DD065A2FFE44F826CE0EC9E9" box="[477,552,1929,1955]" italics="true" pageId="4" pageNumber="5">Conus</emphasis>
(
<emphasis id="54F5EA88DD065A2FFDA5F826CEE3C9E8" box="[572,709,1929,1954]" italics="true" pageId="4" pageNumber="5">Lautoconus</emphasis>
)
<emphasis id="54F5EA88DD065A2FFD43F827CF4FC9E8" box="[730,873,1928,1954]" italics="true" pageId="4" pageNumber="5">damianakisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC657F3DD065A2FFCEDF826CFF4C9E9" box="[884,978,1929,1955]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="A1814D19DD065A2FFB88F826C9BAC9E8" authority="Psarras &amp; Merle &amp; Koskeridou, 2022" authorityName="Psarras &amp; Merle &amp; Koskeridou" authorityYear="2022" box="[1041,1436,1928,1955]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="lauriatragei" status="sp. nov." subGenus="Lautoconus">
<emphasis id="54F5EA88DD065A2FFB88F826C87AC9E9" box="[1041,1116,1929,1955]" italics="true" pageId="4" pageNumber="5">Conus</emphasis>
(
<emphasis id="54F5EA88DD065A2FFBE8F826C8DCC9E8" box="[1137,1274,1929,1954]" italics="true" pageId="4" pageNumber="5">Lautoconus</emphasis>
)
<emphasis id="54F5EA88DD065A2FFA97F827C9BAC9E8" box="[1294,1436,1928,1954]" italics="true" pageId="4" pageNumber="5">lauriatragei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FC657F3DD065A2FFF7AF802CD62C98D" box="[227,324,1965,1991]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
). The species of the subgenus display an intraspecific morphological variability, as it is here shown for more than one species. The colour patterns are also variable, displaying patterns such as blotches, spiral rows of dots, dashes and tents in multiple combinations (see
<bibRefCitation id="02104B6BDD065A2FFBE8F85BC944C644" author="Monnier E. &amp; Limpalaer L. &amp; Robin A. &amp; Roux C." box="[1137,1378,2036,2063]" pageId="4" pageNumber="5" refId="ref31045" refString="Monnier E., Limpalaer L., Robin A. &amp; Roux C. 2018. A Taxonomic Iconography of the Living Conidae, Volumes 1 - 2. Conchbooks, Harxheim." type="book" year="2018">
Monnier
<emphasis id="54F5EA88DD065A2FFB46F85AC935C644" box="[1247,1299,2036,2062]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. 2018
</bibRefCitation>
). We follow
<bibRefCitation id="02104B6BDD075A2EFEAEFE80CEA9CF00" author="Harzhauser M. &amp; Landau B." box="[311,655,303,330]" pageId="5" pageNumber="6" pagination="1 - 178" refId="ref29682" refString="Harzhauser M. &amp; Landau B. 2016. A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea. Zootaxa 4210 (1): 1 - 178. https: // doi. org / 10.11646 / zootaxa. 4210.1.1" type="journal article" year="2016">Harzhauser &amp; Landau (2016)</bibRefCitation>
in excluding from this subgenus the species with tuberculate spire whorls. The oldest occurrence in
<collectingCountry id="1E96760ADD075A2EFDF6FEFACEE4CF25" box="[623,706,341,367]" name="Greece" pageId="5" pageNumber="6">Greece</collectingCountry>
is reported from the early Miocene (
<bibRefCitation id="02104B6BDD075A2EFBE0FEFBC948CF25" author="Thivaiou D. &amp; Harzhauser M. &amp; Koskeridou E." box="[1145,1390,340,367]" pageId="5" pageNumber="6" pagination="323 - 366" refId="ref32172" refString="Thivaiou D., Harzhauser M. &amp; Koskeridou E. 2019. Early Miocene Gastropods from the Felli Section (Proto-Mediterranean Sea, NW Greece). Geodiversitas 41 (8): 323 - 366. https: // doi. org / 10.5252 / geodiversitas 2019 v 41 a 8" type="journal article" year="2019">
Thivaiou
<emphasis id="54F5EA88DD075A2EFB76FEFAC904CF24" box="[1263,1314,340,366]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
. 2019
</bibRefCitation>
) by an incomplete specimen (AMPG(IV) 2473). However, the attribution to
<taxonomicName id="A1814D19DD075A2EFBB3FED4C895CFDE" authorityName="Monterosato" authorityYear="1923" box="[1066,1203,379,404]" class="Gastropoda" family="Conidae" genus="Lautoconus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">
<emphasis id="54F5EA88DD075A2EFBB3FED4C895CFDE" box="[1066,1203,379,404]" italics="true" pageId="5" pageNumber="6">Lautoconus</emphasis>
</taxonomicName>
seems doubtful and this specimen could possibly belong to
<taxonomicName id="A1814D19DD075A2EFD21FE0FC88FCFF0" authority="Tucker &amp; Tenorio, 2009" authorityName="Tucker &amp; Tenorio" authorityYear="2009" box="[696,1193,416,442]" class="Gastropoda" family="Conidae" genus="Conus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="6" phylum="Mollusca" rank="subGenus" subGenus="Eoconus">
<emphasis id="54F5EA88DD075A2EFD21FE0FCF25CFF0" box="[696,771,416,442]" italics="true" pageId="5" pageNumber="6">Conus</emphasis>
(
<emphasis id="54F5EA88DD075A2EFC8DFE0ECF5FCFF0" box="[788,889,417,442]" italics="true" pageId="5" pageNumber="6">Eoconus</emphasis>
)
<bibRefCitation id="02104B6BDD075A2EFC12FE0FC88FCFF0" author="Tucker J. K. &amp; Tenorio M. J." box="[907,1193,416,442]" pageId="5" pageNumber="6" refId="ref32426" refString="Tucker J. K. &amp; Tenorio M. J. 2009. Systematic Classification of Recent and Fossil Conoidean Gastropods: With Keys to the Genera of Cone Shells. ConchBooks, Harxheim." type="book" year="2009">Tucker &amp; Tenorio, 2009</bibRefCitation>
</taxonomicName>
(see more in
<bibRefCitation id="02104B6BDD075A2EFAD5FE0FCD43CFAA" author="Tracey S. &amp; Craig B. &amp; Belliard L. &amp; Gain O." pageId="5" pageNumber="6" pagination="1 - 38" refId="ref32361" refString="Tracey S., Craig B., Belliard L. &amp; Gain O. 2017. One, four or forty species? Early Conidae (Mollusca, Gastropoda) that led to a radiation and biodiversity peak in the late Lutetian Eocene of the Cotentin, NW France. Carnets de Voyages paleontologiques dans le Bassin Anglo-Parisien 3: 1 - 38." type="journal article" year="2017">
Tracey
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</paragraph>
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</treatment>
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