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<document id="374459350D2DBC750D026589AF2EE85C" ID-CLB-Dataset="26022" ID-DOI="10.11646/zootaxa.4656.2.2" ID-GBIF-Dataset="62912763-9710-4c81-88ea-5d8c9f9a3bdb" ID-ISSN="1175-5326" ID-Zenodo-Dep="3368143" ID-ZooBank="1CD78AA3-EC4F-4E7F-9005-5ED81CB1F72E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565768352213" checkinUser="plazi" docAuthor="Pérez-Gutiérrez, León Andrés" docDate="2019" docId="BC6E87EBFFBF071424A6FD80FA47ED94" docLanguage="en" docName="zootaxa.4656.2.2.pdf" docOrigin="Zootaxa 4656 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dactylobasis Pérez-Gutiérrez 2019, gen. nov." docType="treatment" docVersion="11" lastPageNumber="234" masterDocId="4057FF93FFBD07162431FFEDFFCEEB02" masterDocTitle="Dactylobasis gen. nov. from Colombia, a new genus of Zygoptera with Teinobasini affinities (Odonata: Coenagrionidae)" masterLastPageNumber="242" masterPageNumber="232" pageNumber="234" updateTime="1732786376359" updateUser="ExternalLinkService">
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<mods:title id="7DD001D600DB64E25BCB4A158A92CA66">Dactylobasis gen. nov. from Colombia, a new genus of Zygoptera with Teinobasini affinities (Odonata: Coenagrionidae)</mods:title>
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<mods:namePart id="BAE45218CE1B2F24FBCD7F6051721FCD">Pérez-Gutiérrez, León Andrés</mods:namePart>
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<mods:date id="0C3AFB23017CB81E8BA5BF1E68A77981">2019</mods:date>
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<treatment id="BC6E87EBFFBF071424A6FD80FA47ED94" ID-DOI="http://doi.org/10.5281/zenodo.5930769" ID-GBIF-Taxon="159134095" ID-Zenodo-Dep="5930769" LSID="urn:lsid:plazi:treatment:BC6E87EBFFBF071424A6FD80FA47ED94" httpUri="http://treatment.plazi.org/id/BC6E87EBFFBF071424A6FD80FA47ED94" lastPageNumber="234" pageId="2" pageNumber="234" scope_family="Coenagrionidae" scope_order="Odonata">
<subSubSection id="7CDD6576FFBF071424A6FD80FE68E985" box="[151,422,621,648]" pageId="2" pageNumber="234" type="nomenclature">
<paragraph id="347836FDFFBF071424A6FD80FE68E985" blockId="2.[151,422,621,684]" box="[151,422,621,648]" pageId="2" pageNumber="234">
<heading id="6F308191FFBF071424A6FD80FE68E985" bold="true" box="[151,422,621,648]" fontSize="11" level="1" pageId="2" pageNumber="234" reason="1">
<emphasis id="06B3EAEFFFBF071424A6FD80FE68E985" bold="true" box="[151,422,621,648]" pageId="2" pageNumber="234">
<taxonomicName id="F3C74D7EFFBF071424A6FD80FEFEE98A" ID-CoL="9WZM6" authority="Pérez-Gutiérrez, 2019" authorityName="Pérez-Gutiérrez" authorityYear="2019" box="[151,304,621,648]" class="Insecta" genus="Dactylobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="06B3EAEFFFBF071424A6FD80FEFEE98A" bold="true" box="[151,304,621,648]" italics="true" pageId="2" pageNumber="234">Dactylobasis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1D805794FFBF07142509FD80FE68E985" box="[312,422,621,647]" pageId="2" pageNumber="234" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="7CDD6576FFBF071424A6FD7CFEC5E9AE" box="[151,267,657,684]" pageId="2" pageNumber="234" type="description">
<paragraph id="347836FDFFBF071424A6FD7CFEC5E9AE" blockId="2.[151,422,621,684]" box="[151,267,657,684]" pageId="2" pageNumber="234">
(
<figureCitation id="ACFC2A78FFBF071424AEFD7CFECDE9AE" box="[159,259,657,684]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="3.[151,250,1589,1614]" captionStartId-1="4.[151,250,1928,1953]" captionStartId-2="5.[151,250,1781,1806]" captionStartId-3="6.[151,250,1582,1607]" captionStartId-4="7.[151,250,1635,1660]" captionTargetBox-0="[151,1436,227,1569]" captionTargetBox-1="[151,1437,561,1909]" captionTargetBox-2="[166,1425,196,1751]" captionTargetBox-3="[151,1436,453,1563]" captionTargetBox-4="[340,1247,381,1610]" captionTargetId-0="figure@3.[151,1436,227,1569]" captionTargetId-1="figure@4.[151,1437,561,1909]" captionTargetId-2="figure@5.[151,1437,181,1760]" captionTargetId-3="figure@6.[151,1436,453,1563]" captionTargetId-4="figure@7.[340,1247,381,1610]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 2. Dactylobasis demarmelsi sp. nov. (ab) male head and prothorax, lateral view; (b) female head, dorsal view; (c) thorax showing left projection of propleuron, lateral view; (d) thoracic color pattern; (e) male prothorax, dorsal view showing projections of propleuron (f) male thorax, dorsal view. Pp: Propleuron projections." captionText-1="FIGURE 3. Dactylobasis demarmelsi sp. nov. (a) male S10, lateral view; (b) cercus, dorsal view; (c) male S10 posterior view; (d) male cercus and paraproct, left lateral view; (e) male, cercus inner view; (f) male S10, ventral view; (g) female S810, lateral view. vs: ventral articulated spur." captionText-2="FIGURE 4. Dactylobasis demarmelsi sp. nov. (a) fore and hind left wings of male; (b) fore and hind left wings of female; (c) genital ligula, ectal view; (d) latero ental view; (e) lateral view." captionText-3="FIGURE 5. Dactylobasis demarmelsi sp. nov., male at Quibdó, Salero, Chocó, Colombia. Unión Panamericana (5° 2200.17 N 76° 3603.11W) (Chocó, Colombia), 28 January 2013. Photo by LP." captionText-4="FIGURE 6. Dactylobasis demarmelsi sp. nov., female at Salero, Unión Panamericana, Chocó, Colombia. (5° 2200.17N 76° 3603.11 W) (Chocó, Colombia), 28 January 2013. Photo by LP." httpUri-0="https://zenodo.org/record/3368147/files/figure.png" httpUri-1="https://zenodo.org/record/3368149/files/figure.png" httpUri-2="https://zenodo.org/record/3368151/files/figure.png" httpUri-3="https://zenodo.org/record/3368153/files/figure.png" httpUri-4="https://zenodo.org/record/3368155/files/figure.png" pageId="2" pageNumber="234">Figs 26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7CDD6576FFBF071424A6FD3BFCA2E9EF" box="[151,876,724,750]" pageId="2" pageNumber="234" type="type_taxon">
<paragraph id="347836FDFFBF071424A6FD3BFCA2E9EF" blockId="2.[151,876,724,750]" box="[151,876,724,750]" pageId="2" pageNumber="234">
<typeStatus id="EB7C885FFFBF071424A6FD3BFF07E9EC" box="[151,201,726,750]" pageId="2" pageNumber="234">Type</typeStatus>
species
<taxonomicName id="F3C74D7EFFBF0714252EFD38FDEEE9EF" authority="Pérez-Gutiérrez, 2019" authorityName="Pérez-Gutiérrez" authorityYear="2019" box="[287,544,725,749]" class="Insecta" genus="Dactylobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="species" species="demarmelsi" status="sp. nov.">
<emphasis id="06B3EAEFFFBF0714252EFD38FDEEE9EF" box="[287,544,725,749]" italics="true" pageId="2" pageNumber="234">Dactylobasis demarmelsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1D805794FFBF07142617FD3BFDBBE9EF" box="[550,629,726,749]" pageId="2" pageNumber="234" rank="species">sp. nov.</taxonomicNameLabel>
by original designation.
</paragraph>
</subSubSection>
<subSubSection id="7CDD6576FFBF071424A6FCF1FF28E858" pageId="2" pageNumber="234" type="materials_examined">
<paragraph id="347836FDFFBF071424A6FCF1FF28E858" blockId="2.[151,1437,795,1686]" pageId="2" pageNumber="234">
<emphasis id="06B3EAEFFFBF071424A6FCF1FED0E834" bold="true" box="[151,286,796,822]" pageId="2" pageNumber="234">Etymology.</emphasis>
Daktylos (Greek): finger; the genus name refers to the articulated, digitiform ventral spur at base of cercus.
</paragraph>
</subSubSection>
<subSubSection id="7CDD6576FFBF071424F6FC8EFC12EF7B" pageId="2" pageNumber="234" type="description">
<paragraph id="347836FDFFBF071424F6FC8EFA97E8C7" blockId="2.[151,1437,795,1686]" pageId="2" pageNumber="234">
<emphasis id="06B3EAEFFFBF071424F6FC8EFE69E87C" bold="true" box="[199,423,867,894]" pageId="2" pageNumber="234">Description. Head.</emphasis>
Medium sized coenagrionid with angulate frons, deeply sunk between bases of antennae; occipital rim prolonged laterally into an anteriorly curved carina, postocular lobe not surpassing compound eye posteriorly; postocular spot and pale mark between lateral ocellus and base of corresponding antenna absent.
</paragraph>
<paragraph id="347836FDFFBF071424F6FC3DFC4DEF33" blockId="2.[151,1437,795,1686]" pageId="2" pageNumber="234">
<emphasis id="06B3EAEFFFBF071424F6FC3DFEE9E8E8" bold="true" box="[199,295,976,1002]" pageId="2" pageNumber="234">Thorax.</emphasis>
Propleuron with distally directed, spinulose conical projection. Mesanepisternum entirely metallic green, each with a long horn, in both sexes. Legs short, metatibial spines slightly shorter than intervals separating them, supplementary teeth of tarsal claws reduced to low swelling.
</paragraph>
<paragraph id="347836FDFFBF071424F6FBD6FC12EF7B" blockId="2.[151,1437,795,1686]" pageId="2" pageNumber="234">
<emphasis id="06B3EAEFFFBF071424F6FBD6FE7CEF54" bold="true" box="[199,434,1083,1110]" pageId="2" pageNumber="234">Caudal appendages</emphasis>
. Male cercus short and cap-like in lateral view, and with ventral spur articulated at membranous base; paraproct long with short ventral branch at half of its length.
</paragraph>
</subSubSection>
<subSubSection id="7CDD6576FFBF071424F6FB6EFDCAED28" pageId="2" pageNumber="234" type="diagnosis">
<paragraph id="347836FDFFBF071424F6FB6EFDCAED28" blockId="2.[151,1437,795,1686]" pageId="2" pageNumber="234">
<emphasis id="06B3EAEFFFBF071424F6FB6EFE6DEF9C" bold="true" box="[199,419,1155,1182]" pageId="2" pageNumber="234">Generic diagnosis.</emphasis>
<taxonomicName id="F3C74D7EFFBF0714259AFB69FDF3EF9C" authority="Pérez-Gutiérrez, 2019" authorityName="Pérez-Gutiérrez" authorityYear="2019" box="[427,573,1156,1182]" class="Insecta" genus="Dactylobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="06B3EAEFFFBF0714259AFB69FDF3EF9C" box="[427,573,1156,1182]" italics="true" pageId="2" pageNumber="234">Dactylobasis</emphasis>
</taxonomicName>
is considered to be a member of Teinobasini based on the presence of an articulated ventro-basal spur on male cercus shared with
<emphasis id="06B3EAEFFFBF07142731FB45FED6EFE4" italics="true" pageId="2" pageNumber="234">
<taxonomicName id="F3C74D7EFFBF07142731FB45FC5BEFC0" authorityName="Kennedy" authorityYear="1920" box="[768,917,1192,1218]" class="Insecta" family="Coenagrionidae" genus="Aceratobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">Aceratobasis</taxonomicName>
,
<taxonomicName id="F3C74D7EFFBF07142790FB45FB86EFC0" authorityName="De Marmels" authorityYear="2005" box="[929,1096,1192,1218]" class="Insecta" family="Coenagrionidae" genus="Bromeliagrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">Bromeliagrion</taxonomicName>
,
<taxonomicName id="F3C74D7EFFBF07142062FB45FB23EFC0" authorityName="Needham" authorityYear="1903" box="[1107,1261,1192,1218]" class="Insecta" family="Coenagrionidae" genus="Chromagrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">Chromagrion</taxonomicName>
,
<taxonomicName id="F3C74D7EFFBF071420C9FB45FA56EFC0" authorityName="Kennedy" authorityYear="1920" box="[1272,1432,1192,1218]" class="Insecta" family="Coenagrionidae" genus="Diceratobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">Diceratobasis</taxonomicName>
,
<taxonomicName id="F3C74D7EFFBF071424A6FB21FED6EFE4" authorityName="Selys" authorityYear="1850" box="[151,280,1228,1254]" class="Insecta" family="Coenagrionidae" genus="Nehalennia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">Nehalennia</taxonomicName>
</emphasis>
and
<taxonomicName id="F3C74D7EFFBF07142561FB21FE09EFE4" authorityName="De Marmels" authorityYear="2007" box="[336,455,1228,1254]" class="Insecta" family="Coenagrionidae" genus="Tepuibasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">
<emphasis id="06B3EAEFFFBF07142561FB21FE09EFE4" box="[336,455,1228,1254]" italics="true" pageId="2" pageNumber="234">Tepuibasis</emphasis>
</taxonomicName>
. A movable spur is absent in the
<emphasis id="06B3EAEFFFBF0714270FFB21FB11EFE4" box="[830,1247,1228,1254]" italics="true" pageId="2" pageNumber="234">
Ceriagrion-Telebasis -
<taxonomicName id="F3C74D7EFFBF07142005FB21FB11EFE4" baseAuthorityName="von Ellenrieder" baseAuthorityYear="2013" box="[1076,1247,1228,1254]" class="Insecta" family="Coenagrionidae" genus="Metaleptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">Metaleptobasis</taxonomicName>
</emphasis>
series (
<bibRefCitation id="50564B0CFFBF07142104FB26FF1DEE08" author="Kennedy, C. H." pageId="2" pageNumber="234" pagination="19 - 29" refId="ref4653" refString="Kennedy, C. H. (1920) The phylogeny of the zygopterous dragonflies as based on the evidence of the penes. Ohio Journal of Science, 21 (1), 19 - 29. https: // doi. org / 10.5962 / bhl. part. 1454 0" type="journal article" year="1920">Kennedy 1920</bibRefCitation>
).
<taxonomicName id="F3C74D7EFFBF071424D8FB1DFEB9EE08" box="[233,375,1264,1290]" class="Insecta" family="Coenagrionidae" genus="Schistolobos" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">
<emphasis id="06B3EAEFFFBF071424D8FB1DFEB9EE08" box="[233,375,1264,1290]" italics="true" pageId="2" pageNumber="234">Schistolobos</emphasis>
</taxonomicName>
and some
<taxonomicName id="F3C74D7EFFBF071425C7FB1DFD46EE08" authorityName="Kennedy" authorityYear="1920" box="[502,648,1264,1290]" class="Insecta" family="Coenagrionidae" genus="Aceratobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">
<emphasis id="06B3EAEFFFBF071425C7FB1DFD46EE08" box="[502,648,1264,1290]" italics="true" pageId="2" pageNumber="234">Aceratobasis</emphasis>
</taxonomicName>
also have a ventro-basal branch (
<bibRefCitation id="50564B0CFFBF0714203FFB1DFA5EEE08" author="von Ellenrieder, N. &amp; Garrison, R. W." box="[1038,1424,1263,1290]" pageId="2" pageNumber="234" pagination="1 - 22" refId="ref5022" refString="von Ellenrieder, N. &amp; Garrison, R. W. (2008) A redefinition of Telagrion Selys and Aceratobasis Kennedy stat. rev. and the description of Schistolobos gen. nov. for Telagrion boliviense Daigle (Odonata: Coenagrionidae). Transactions of the American Entomological Society, 134 (1 + 2), 1 - 22. https: // doi. org / 10.3157 / 0002 - 8320 (2008) 134 [1: AROTSA] 2.0. CO; 2" type="journal article" year="2008">von Ellenrieder &amp; Garrison 2008</bibRefCitation>
), but it is not movable nor articulated.
<taxonomicName id="F3C74D7EFFBF07142605FAF9FD08EE2C" authority="Pérez-Gutiérrez, 2019" authorityName="Pérez-Gutiérrez" authorityYear="2019" box="[564,710,1300,1326]" class="Insecta" genus="Dactylobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="06B3EAEFFFBF07142605FAF9FD08EE2C" box="[564,710,1300,1326]" italics="true" pageId="2" pageNumber="234">Dactylobasis</emphasis>
</taxonomicName>
differs from other genera of neotropical Teinobasini by the completely metallic green mesanepisternum, and the unusual development of propleuron bearing a posteriorly directed conical projection with spinulose texture (however, there is a similar structure in the propleuron of several species of
<taxonomicName id="F3C74D7EFFBF07142487FA6DFEAFEE98" baseAuthorityName="von Ellenrieder" baseAuthorityYear="2013" box="[182,353,1408,1434]" class="Insecta" family="Coenagrionidae" genus="Metaleptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">
<emphasis id="06B3EAEFFFBF07142487FA6DFEAFEE98" box="[182,353,1408,1434]" italics="true" pageId="2" pageNumber="234">Metaleptobasis</emphasis>
</taxonomicName>
;
<bibRefCitation id="50564B0CFFBF07142541FA6DFDACEE98" author="von Ellenrieder, N." box="[368,610,1407,1434]" pageId="2" pageNumber="234" pagination="1 - 155" refId="ref4965" refString="von Ellenrieder, N. (2013) A revision of Metaleptobasis Calvert (Odonata: Coenagrionidae) with seven synonymies and the description of eighteen new species from South America. Zootaxa, 3738 (1), 1 - 155. https: // doi. org / 10.11646 / zootaxa. 3738.1. 1" type="journal article" year="2013">von Ellenrieder 2013</bibRefCitation>
). Chitinized spinules on genital ligula, like those present in
<taxonomicName id="F3C74D7EFFBF07142132FA6DFA5BEE98" authorityName="Kennedy" authorityYear="1920" box="[1283,1429,1408,1434]" class="Insecta" family="Coenagrionidae" genus="Aceratobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">
<emphasis id="06B3EAEFFFBF07142132FA6DFA5BEE98" box="[1283,1429,1408,1434]" italics="true" pageId="2" pageNumber="234">Aceratobasis</emphasis>
</taxonomicName>
,
<taxonomicName id="F3C74D7EFFBF071424A6FA49FEF5EEBC" authorityName="De Marmels" authorityYear="2005" box="[151,315,1444,1470]" class="Insecta" family="Coenagrionidae" genus="Bromeliagrion" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">
<emphasis id="06B3EAEFFFBF071424A6FA49FEF5EEBC" box="[151,315,1444,1470]" italics="true" pageId="2" pageNumber="234">Bromeliagrion</emphasis>
</taxonomicName>
and
<taxonomicName id="F3C74D7EFFBF0714255EFA49FE28EEBC" authorityName="De Marmels" authorityYear="2007" box="[367,486,1444,1470]" class="Insecta" family="Coenagrionidae" genus="Tepuibasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">
<emphasis id="06B3EAEFFFBF0714255EFA49FE28EEBC" box="[367,486,1444,1470]" italics="true" pageId="2" pageNumber="234">Tepuibasis</emphasis>
</taxonomicName>
are absent in
<emphasis id="06B3EAEFFFBF0714264FFA49FCD6EEBC" box="[638,792,1444,1470]" italics="true" pageId="2" pageNumber="234">
<taxonomicName id="F3C74D7EFFBF0714264FFA49FCDAEEBC" authority="Pérez-Gutiérrez, 2019" authorityName="Pérez-Gutiérrez" authorityYear="2019" box="[638,788,1444,1470]" class="Insecta" genus="Dactylobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus" status="gen. nov.">Dactylobasis</taxonomicName>
.
</emphasis>
In
<taxonomicName id="F3C74D7EFFBF0714270BFA49FC7FEEBC" authorityName="De Marmels" authorityYear="2007" box="[826,945,1444,1470]" class="Insecta" family="Coenagrionidae" genus="Tepuibasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">
<emphasis id="06B3EAEFFFBF0714270BFA49FC7FEEBC" box="[826,945,1444,1470]" italics="true" pageId="2" pageNumber="234">Tepuibasis</emphasis>
</taxonomicName>
and
<taxonomicName id="F3C74D7EFFBF071427D4FA49FBA8EEBC" authorityName="Selys" authorityYear="1850" box="[997,1126,1444,1470]" class="Insecta" family="Coenagrionidae" genus="Nehalennia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">
<emphasis id="06B3EAEFFFBF071427D4FA49FBA8EEBC" box="[997,1126,1444,1470]" italics="true" pageId="2" pageNumber="234">Nehalennia</emphasis>
</taxonomicName>
the inner fold is well developed, but absent in
<taxonomicName id="F3C74D7EFFBF07142542FA25FDCBEEE0" authority="Pérez-Gutiérrez, 2019" authorityName="Pérez-Gutiérrez" authorityYear="2019" box="[371,517,1480,1506]" class="Insecta" genus="Dactylobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="06B3EAEFFFBF07142542FA25FDCBEEE0" box="[371,517,1480,1506]" italics="true" pageId="2" pageNumber="234">Dactylobasis</emphasis>
</taxonomicName>
as well as in
<emphasis id="06B3EAEFFFBF071426ACFA25FC8CEEE0" box="[669,834,1480,1506]" italics="true" pageId="2" pageNumber="234">
<taxonomicName id="F3C74D7EFFBF071426ACFA25FCF0EEE0" authorityName="Kennedy" authorityYear="1920" box="[669,830,1480,1506]" class="Insecta" family="Coenagrionidae" genus="Diceratobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">Diceratobasis</taxonomicName>
.
</emphasis>
In addition,
<taxonomicName id="F3C74D7EFFBF071427E0FA25FBADEEE0" authority="Pérez-Gutiérrez, 2019" authorityName="Pérez-Gutiérrez" authorityYear="2019" box="[977,1123,1480,1506]" class="Insecta" genus="Dactylobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="06B3EAEFFFBF071427E0FA25FBADEEE0" box="[977,1123,1480,1506]" italics="true" pageId="2" pageNumber="234">Dactylobasis</emphasis>
</taxonomicName>
has very long and bifurcate paraproct that contrasts with the very short cap-like cercus. Absence of a vulvar spine on female S8 is shared with
<taxonomicName id="F3C74D7EFFBF071424A6F9FDFEE7ED28" authorityName="Kennedy" authorityYear="1920" box="[151,297,1552,1578]" class="Insecta" family="Coenagrionidae" genus="Aceratobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">
<emphasis id="06B3EAEFFFBF071424A6F9FDFEE7ED28" box="[151,297,1552,1578]" italics="true" pageId="2" pageNumber="234">Aceratobasis</emphasis>
</taxonomicName>
and
<taxonomicName id="F3C74D7EFFBF07142551F9FDFE33ED28" authorityName="Kennedy" authorityYear="1920" box="[352,509,1552,1578]" class="Insecta" family="Coenagrionidae" genus="Diceratobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus">
<emphasis id="06B3EAEFFFBF07142551F9FDFE33ED28" box="[352,509,1552,1578]" italics="true" pageId="2" pageNumber="234">Diceratobasis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="7CDD6576FFBF071424F6F9DEFA47ED94" pageId="2" pageNumber="234" type="discussion">
<paragraph id="347836FDFFBF071424F6F9DEFA47ED94" blockId="2.[151,1437,795,1686]" pageId="2" pageNumber="234">
Based on the vein descending from quadrangle in hind wing forming a straight line to wing margin and dorsum of head entirely black,
<taxonomicName id="F3C74D7EFFBF071425A5F9B5FDE8ED70" authority="Pérez-Gutiérrez, 2019" authorityName="Pérez-Gutiérrez" authorityYear="2019" box="[404,550,1624,1650]" class="Insecta" genus="Dactylobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="06B3EAEFFFBF071425A5F9B5FDE8ED70" box="[404,550,1624,1650]" italics="true" pageId="2" pageNumber="234">Dactylobasis</emphasis>
</taxonomicName>
keys out to couplets 59 and
<quantity id="F33F9B18FFBF0714276FF9B5FC58ED73" box="[862,918,1623,1650]" metricMagnitude="0" metricUnit="m" metricValue="1.9812" pageId="2" pageNumber="234" unit="in" value="78.0">78 in</quantity>
the keys to males and females of
<taxonomicName id="F3C74D7EFFBF07142139F9BAFF70ED97" class="Insecta" family="Coenagrionidae" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="family">Coenagrionidae</taxonomicName>
genera in
<bibRefCitation id="50564B0CFFBF07142502F996FDE8ED94" author="Garrison, R. W. &amp; von Ellenrieder, N. &amp; Louton, J. A." box="[307,550,1659,1686]" pageId="2" pageNumber="234" refId="ref4565" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2010) Damselfly genera of the New World. An illustrated and annotated key to the Zygoptera. Johns Hopkins University Press, Baltimore, 490 pp." type="book" year="2010">
Garrison
<emphasis id="06B3EAEFFFBF071425ADF991FE1BED94" box="[412,469,1660,1686]" italics="true" pageId="2" pageNumber="234">et al.</emphasis>
(2010)
</bibRefCitation>
respectively. These keys can be modified as follows to include
<taxonomicName id="F3C74D7EFFBF071420DEF991FA4FED94" authority="Pérez-Gutiérrez, 2019" authorityName="Pérez-Gutiérrez" authorityYear="2019" box="[1263,1409,1660,1686]" class="Insecta" genus="Dactylobasis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="234" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="06B3EAEFFFBF071420DEF991FA4FED94" box="[1263,1409,1660,1686]" italics="true" pageId="2" pageNumber="234">Dactylobasis</emphasis>
</taxonomicName>
:
</paragraph>
</subSubSection>
</treatment>
</document>