434 lines
33 KiB
XML
434 lines
33 KiB
XML
<document id="8945B97871A27CD9EAF989D41A31010A" ID-CLB-Dataset="302074" ID-DOI="10.3897/phytokeys.246.129476" ID-GBIF-Dataset="b381880a-e0bb-4b02-ad0f-2f90a0b6c995" ID-publisher-id="129476" URI-arpha="7631F16E-4CD8-5A76-9F28-45E5F398CE20" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1725298343909" checkinUser="pensoft" docAuthor="Meerts, Pierre J. & Paton, Alan J." docDate="2024" docId="9A165417EA3A541A9493B44BF30DA1A2" docLanguage="en" docName="PhytoKeys 246: 71-178" docOrigin="PhytoKeys 246" docSource="https://phytokeys.pensoft.net/article/129476/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.phytokeys" docStyleName="PensoftTaxPub.0000.journal_article.phytokeys" docTitle="Coleus homblei De Wild." docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="178" masterDocId="7631F16E4CD85A769F2845E5F398CE20" masterDocTitle="The genus Coleus (Lamiaceae) in Central Africa (Democratic Republic of the Congo, Rwanda, Burundi), with the description of 15 new species" masterLastPageNumber="178" masterPageNumber="71" pageNumber="71" updateTime="1725322260938" updateUser="ExternalLinkService">
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<mods:title id="CE6C6B12449B69DDD424689BBE56BA32">The genus Coleus (Lamiaceae) in Central Africa (Democratic Republic of the Congo, Rwanda, Burundi), with the description of 15 new species</mods:title>
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<mods:name id="9AC16F01ADE9C3BCD1C0F39FBCEDF6C9" type="personal">
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<mods:namePart id="40AE7E22975016D76888906850540C33">Meerts, Pierre J.</mods:namePart>
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<mods:nameIdentifier id="210B965669A0150A4DDAEE0F9E7C7952" type="ORCID">0000-0003-4215-027X</mods:nameIdentifier>
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<mods:affiliation id="F9913D99C2ED48435422D183E30CE40F">Meise Botanic Garden, Nieuwelaan 38, BE- 1860 Meise, Belgium & Université Libre de Bruxelles, Av. F. D. Roosevelt 50 CP 244, BE- 1050 Brussels, Belgium & Fédération Wallonie-Bruxelles, Service général de l’Enseignement supérieur et de la Recherche scientifique, Brussels, Belgium</mods:affiliation>
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<mods:namePart id="C3B041022A6929A7080E5B99222BA3AA">Paton, Alan J.</mods:namePart>
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<mods:nameIdentifier id="7616D7292E4ED259BFF8983892BFCD0D" type="ORCID">0000-0002-6052-6675</mods:nameIdentifier>
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<mods:affiliation id="91AC5E848513330A3E96175B4FF906E7">The Herbarium, Royal Botanic Gardens, Kew, Richmond TW 9 3 AB, UK</mods:affiliation>
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<mods:title id="537DC4E989DBAD6AF746A9AF9B56D928">PhytoKeys</mods:title>
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<mods:part id="5A025D7772C6D2A4F8056EEF22CC2581">
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<mods:date id="1AAF4B82B885DF83B2CCC7A538A9DD08">2024</mods:date>
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<mods:number id="D491E4B43A6FA1084FFE06784D38E48B">246</mods:number>
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<mods:identifier id="79F462D07C279605F382DA3DE3B8B8B2" type="DOI">10.3897/phytokeys.246.129476</mods:identifier>
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<treatment id="9A165417EA3A541A9493B44BF30DA1A2" ID-DOI="http://doi.org/10.5281/zenodo.13629815" ID-GBIF-Taxon="240061489" ID-Zenodo-Dep="13629815" ID-arpha="9A165417-EA3A-541A-9493-B44BF30DA1A2" LSID="urn:lsid:plazi:treatment:9A165417EA3A541A9493B44BF30DA1A2" httpUri="http://treatment.plazi.org/id/9A165417EA3A541A9493B44BF30DA1A2">
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<subSubSection id="454616FBF93337D7F4BB4DBAA4E6099D" type="nomenclature">
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<paragraph id="139B1A45A4B6432FA8FDA8C4FA6B5BB1">
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<treatmentCitation id="234F9CDD25EEFCD457015311BF086E63" author="De Wild., Contr. Fl." issue="1" journal="Ann. Soc. Sci. Bruxelles" page="49" score="156.5387788110742" volume="42" year="1921">
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||
<taxonomicName id="2C86F09E8C855FEA0C059537D9F07E9C" ID-CoL="X26H" authority="De Wild." authorityName="De Wild." class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="homblei">
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||
<emphasis id="259F330F7763FFFEC8B62172BAFC9DBF" italics="true">Coleus homblei</emphasis>
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||
De Wild.
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||
</taxonomicName>
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||
, Contr. Fl.
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<collectingRegion id="241ADF659A1D5253BD5E3CFAC73F145C" country="Republic of the Congo" name="Katanga">Katanga</collectingRegion>
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||
: 174. &
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<bibCitation id="848DFED508E81EBC062CC31B7BA6B939" issue="1" journal="Ann. Soc. Sci. Bruxelles" page="49" score="156.5387788110742" volume="42" year="1921">
|
||
Ann. Soc. Sci.
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<collectingRegion id="01468E5E478B9D481E25C4F03CC7F87A" country="Belgium" name="Region de Bruxelles-Capitale">Bruxelles</collectingRegion>
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42 (1): 49. 1921
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</bibCitation>
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||
</treatmentCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0A7F79C54A2307A252D9E3498DDBF3B1" type="reference_group">
|
||
<paragraph id="1192E768D867EA88B21BEFCCFE5E211B">
|
||
<treatmentCitationGroup id="E2B99B9BCC43CDA5B995BB249F12F501">
|
||
=
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<treatmentCitation id="44C6B4A33CEEDAF1A2F03FD1FD367C75" author="E. Afr." page="333" volumeTitle="Lamiac." year="2009">
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||
<taxonomicName id="73E1E8C73ACA91070F5851C9E73E099D" ID-CoL="4K4LX" authority="A. J. Paton" authorityName="A. J. Paton" class="Magnoliopsida" family="Lamiaceae" genus="Plectranthus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="sigmoideus">
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<emphasis id="91B68B9C8ACDB70413BB8171E49AD394" italics="true">Plectranthus sigmoideus</emphasis>
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A. J. Paton
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</taxonomicName>
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||
, Fl. Trop. E. Afr., Lamiac.: 333. 2009
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||
</treatmentCitation>
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||
., syn. nov.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="12BE9216F3C81D7E6FB75A2C7E3BCC97">
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||
<treatmentCitationGroup id="6E3388F44D0EAFD0A6C8AF5C9E112D73">
|
||
≡
|
||
<treatmentCitation id="B965B9D42885131AF3366FE8DE4161C7" author="A. J. Paton" httpUri="http://treatment.plazi.org/id/377EB9F4B94D51668924FBD7F2BBB624" journal="Phytokeys" page="97" score="9.729594737742866" volume="129" year="2019">
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<taxonomicName id="215FD4460790533D8AAAEBC90FFEB989" ID-CoL="8XY96" authority="(A. J. Paton) A. J. Paton" authorityName="A. J. Paton" baseAuthorityName="A. J. Paton" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="sigmoideus">
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||
<emphasis id="F3173F8195D6E9C2388E63A273D790D8" italics="true">Coleus sigmoideus</emphasis>
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||
(A. J. Paton) A. J. Paton
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</taxonomicName>
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||
, Phytokeys 129: 97. 2019
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||
</treatmentCitation>
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||
. Type:
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<collectingCountry id="363A0251413287F2CF58F57DB56B83D5" name="Zambia">Zambia</collectingCountry>
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, track opposite turning to Mbala (Abercorn) Club,
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<emphasis id="F24D804D6A82622E09DBD9CA7F29F931" italics="true">
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||
<collectionCode id="9CF22EB9CCEF58BFD30538608D431459" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" type="Herbarium">H</collectionCode>
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.
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||
<collectionCode id="CCF48074552CFB9DE6FEA052CEF310D7" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" type="Herbarium">M</collectionCode>
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||
. Richards 4353
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</emphasis>
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||
(
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<typeStatus id="95777716C5BC91184D7EE4A6BF57CBD2" type="holotype">holotype</typeStatus>
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<collectionCode id="7B4949960FC2DE046881CB0966439553" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
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||
[
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<specimenCode id="12AE3D0BFC2029A318708CB1931DE7AB">
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||
<collectionCode id="04997EFA0133C8697426F9024B9BEF01" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
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000070564
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</specimenCode>
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]), syn. nov.
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</treatmentCitationGroup>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="SECID0EAVAM" type="material">
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<paragraph id="4D325BF07A8A0734B1B03BF2CF37625E">
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||
<heading id="5F4932DD2E1C79CF9B64AD8DE9592DFD" reason="title">Type.</heading>
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</paragraph>
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<paragraph id="BBF436A57BA22D1959FB972EC3F71AC3">
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<collectingCountry id="39A61A09AF4867B41A9EBC38314DE9B9">DR. Congo</collectingCountry>
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, Elisabethville [Lubumbashi], brousse,
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<date id="08266D387AF3D673B37820C5D03A0420" value="1912-02">Feb. 1912</date>
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||
.
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<emphasis id="B0A469FDCFBEF733B6F452E7EB340883" italics="true">
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<collectionCode id="2679C4048BFFA09409C9969C7F3A1932" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" type="Herbarium">H</collectionCode>
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. Homblé 210
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</emphasis>
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(
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<typeStatus id="4FCACAF4F6A0D6969F64BE0425BE9397" type="lectotype">lectotype</typeStatus>
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<collectionCode id="A5345EEF3F07364F3615CA2364BA16D0" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
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||
[
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||
<specimenCode id="C31B931A92D3A2CA83AF3C679ADC8C7F">
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<collectionCode id="DB7EBF3E56321E1271E341D5EFBBD773" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
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0000006262181
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</specimenCode>
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], designated here).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="SECID0EOVAM" type="description">
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||
<paragraph id="C08F86EBDA7420F2FE7D82E8A524D0E4">
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<heading id="AABD8C17D3B90D3D84495D12A9E3C12F" reason="title">Description.</heading>
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</paragraph>
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<paragraph id="D8A1290BACEBD2894CA9D3E593F3C73D">
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Short-lived perennial herb 0.20–0.40 (– 0.65) m, rootstock a thin rhizome rooting at nodes, tubers occasionally observed (not collected in Central Africa). Stem erect, quadrangular, with short appressed retrorse hairs (also patent hairs outside Central Africa), yellowish to green, mostly unbranched, rarely with a few erect branches. Leaves opposite, in ca. 5 or 6 pairs, the uppermost pair often much reduced, petiolate, except uppermost pair, ascending, petiole 0.2–2.0 (– 2.5) cm with short retrorse hairs and a few long cilia, blade ovate, ovate-elliptic to subrhombic, 1.2–4.5 (– 5.0) × 0.8–2.5 (– 3.2) cm, apex acute, base cuneate and shortly attenuate into the petiole, margin narrowly recurved, with 4–8 sharp teeth on either side (
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<quantity id="1F96D1D78F8D51994262BBA18AE9E8B3" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
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deep on adaxial side), upper surface pubescent, with a mixed indumentum of antrorse hyaline hairs
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<quantity id="1EC4B99E19E074BBD5162C993B5F79D5" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" unit="mm" value="1.25" valueMax="2.0" valueMin="0.5">0.5–2 mm</quantity>
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long and very short papilliform hairs, lower surface very shortly appressed pubescent on veins (hairs often retrorse), also with red sessile glands, 3–5 pairs of secondary veins. Inflorescence lax, 5–13 (– 17) cm long, with 3–9 (– 15) verticils spaced
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<quantity id="0A10D37A580CF4292A2F267266C647EA" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.5" metricValueMin="0.5" unit="mm" value="15.0" valueMax="25.0" valueMin="5.0">5–25 mm</quantity>
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; bracts narrowly ovate, cucullate,
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<quantity id="B6B9C59CCB3E66CA7F81E1F5C3F0C9A5" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">3–7 mm</quantity>
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long, acuminate, forming an apical coma, caducous; cymes ascending to spreading, sessile, with 3–7 flowers on a
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<quantity id="59B4CE9AC385D9329C20166E3D29CD56" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" unit="mm" value="1.0" valueMax="2.0" valueMin="0.0">0–2 mm</quantity>
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long rachis; pedicels ca.
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<quantity id="BFB1FF791D4D760AA55CD0C39C600D50" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
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||
long at anthesis, spreading to slightly ascending. Flower: calyx campanulate, ca.
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||
<quantity id="B3CB5A91439B1C4FE0DAB5422EE14A17" metricMagnitude="-3" metricUnit="m" metricValue="3.0" unit="mm" value="3.0">3 mm</quantity>
|
||
long at anthesis, shortly pubescent, with red sessile glands, widely open; fruiting calyx
|
||
<quantity id="256F8911B7490AECB2BFF623AB89AEEC" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
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long, lateral teeth obtuse, ca.
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<quantity id="F093AE8E1F3A9092DBE00231D63D30C9" metricMagnitude="-3" metricUnit="m" metricValue="2.0" unit="mm" value="2.0">2 mm</quantity>
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long, lower teeth fused in a ca.
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<quantity id="2722E56CD129B8E69897133A15E66DFC" metricMagnitude="-3" metricUnit="m" metricValue="4.0" unit="mm" value="4.0">4 mm</quantity>
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long very narrow lip, with two acute tips; corolla pale blue or white, with red sessile glands, ca.
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<quantity id="493B001E145CB247614B6C30E48F77B6" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" unit="mm" value="11.0" valueMax="13.0" valueMin="9.0">9–13 mm</quantity>
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long, tube ca.
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<quantity id="795CCDD4D219A5A6E129ABA095CB3A11" metricMagnitude="-3" metricUnit="m" metricValue="5.0" unit="mm" value="5.0">5 mm</quantity>
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long, strongly sigmoid, lower lip ca.
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<quantity id="D6099661A34B2F25111418E73BE154E9" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
|
||
long,
|
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<quantity id="C315506EE8C3E35474269420E3D959D7" metricMagnitude="-3" metricUnit="m" metricValue="4.0" unit="mm" value="4.0">4 mm</quantity>
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deep, upper lip ca.
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<quantity id="0114BD21CAF1799BA9AA42F5AC62FD1F" metricMagnitude="-3" metricUnit="m" metricValue="2.0" unit="mm" value="2.0">2 mm</quantity>
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||
long; staminal filaments fused, anther ca.
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<quantity id="F1A27A6610BB9D44887F259DE8E20D4F" metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
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long. Nutlets brown, red-speckled, ovoid, ca.
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<quantity id="820FC3A4D0B11FBA09E48D3A70C75CE9" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
|
||
long.
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||
</paragraph>
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</subSubSection>
|
||
<subSubSection id="SECID0EAWAM" type="distribution">
|
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<paragraph id="F379FA69355F18BDDFBC9721B9AFFF46">
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||
<heading id="97A40D8BBA318AA20201D847B64B4A75" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="00C901C9A7313582998DFC032E3B6A2E">
|
||
<collectionCode id="E02E50FE890A9A55B3DCF7F1BD3F8473">SW</collectionCode>
|
||
<collectingCountry id="972BD4C18C28A91AE3A2FAC5D19826ED" name="Tanzania">Tanzania</collectingCountry>
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||
to
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<collectingCountry id="A6436A4B3BF3A68D41AD3D0A6A330903" name="Zambia">Zambia</collectingCountry>
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and SE
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<collectingCountry id="AC5B96E99ACF6968DC19A1AE4A67A16C">DR. Congo</collectingCountry>
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.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EMWAM" type="habitat">
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||
<paragraph id="120765E774F27FC259227E4DF05765D9">
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<heading id="FBA0DCC836E3BC4F7F7389EB05CDE092" reason="title">Habitat and ecology.</heading>
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</paragraph>
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<paragraph id="553AE3BA9B80C4BCB8DDE9AD80397FCB">
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Miombo woodlands, often with
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<taxonomicName id="0713C36A48F4471EED4E8DDA243A795E" class="Magnoliopsida" family="Fabaceae" genus="Brachystegia" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="microphylla">
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||
<emphasis id="A7F9AA7DD2E05F6D6A70D1E4296CBAEA" italics="true">Brachystegia microphylla</emphasis>
|
||
</taxonomicName>
|
||
, on shallow rocky soil, termite mounds; ca.
|
||
<quantity id="B2FDAC91862EDB442488102999848996" metricMagnitude="3" metricUnit="m" metricValue="1.385" metricValueMax="1.52" metricValueMin="1.25" unit="m" value="1385.0" valueMax="1520.0" valueMin="1250.0">1250–1520 m</quantity>
|
||
elev.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E4WAM" type="Additional specimens">
|
||
<paragraph id="1E25BA90B21906424E195C42CC62B8F1">
|
||
<heading id="7E399043AEB4EEBD6C06EF96E008C03A" reason="title">Additional specimens.</heading>
|
||
</paragraph>
|
||
<paragraph id="1FC684E3BC175BD819770D0AFFC44ED3">
|
||
<collectingCountry id="77A61E80246165782E0B7B72FFB0E12D">DR. Congo</collectingCountry>
|
||
,
|
||
<emphasis id="0D20BDCB82743E985EE6AD5E39B28253" bold="true" italics="true">
|
||
<collectingRegion id="3CC25F73FB5AB7448862C0E5A60A86A8">Haut-Katanga</collectingRegion>
|
||
</emphasis>
|
||
, Près de Lubumbashi, colline Kiswishi,
|
||
<date id="CFB5AFDE3DF559E8A0C345AEF6E30C25" value="1987-02-24">24 Feb 1987</date>
|
||
,
|
||
<emphasis id="E0D3A66873D4710CE8C653640C9ADE62" italics="true">
|
||
F. Billet &
|
||
<collectionCode id="1C4214404D4BE18592256741508C8A4D" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" type="Herbarium">B</collectionCode>
|
||
. Jadin 4227
|
||
</emphasis>
|
||
(
|
||
<collectionCode id="259005F5B4756D1E95B1CB940E7DB972" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
|
||
,
|
||
<collectionCode id="B880C1383C0195CB20ACA9FDB062FF10" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15542" lsid="urn:lsid:biocol.org:col:15542" name="Uppsala University, Museum of Evolution, Botany Section (Fytoteket)" type="Herbarium">UPS</collectionCode>
|
||
); Dilolo, 1935,
|
||
<emphasis id="4D02587A3C8AD94DC1EF4C1C979BFE93" italics="true">de Wouters d’Oplinter 3</emphasis>
|
||
(
|
||
<collectionCode id="E09F47C8036E52E6919B63AD8A2997FF" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
|
||
); Lubumbashi, route de la mine de l’Etoile, km 12,
|
||
<date id="D3A872AEA96E29DB7882EEA991C7C2B5" value="1957-04-19">19 Apr 1957</date>
|
||
<emphasis id="7A4CB35D6A19A51339EC93AB02F7E84A" italics="true">P. Duvigneaud 2847 Co</emphasis>
|
||
(
|
||
<collectionCode id="9B60C8B67441C6A51A9EDF48E3FAEA7B" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14449" lsid="urn:lsid:biocol.org:col:14449" name="Université Libre de Bruxelles" type="Herbarium">BRLU</collectionCode>
|
||
); Entre Welgelegen et Kasumbalesa,
|
||
<date id="47C701B0BA5F634071A153201A9E18A3" value="1960-01-29">29 Jan 1960</date>
|
||
,
|
||
<emphasis id="98481EEF22198F96703E4B80DB7ADB34" italics="true">
|
||
P. Duvigneaud 5302
|
||
<collectionCode id="6A1CC5E4A4DDD32165675B8EB5753469" country="Denmark" name="University of Copenhagen" type="Herbarium">C</collectionCode>
|
||
</emphasis>
|
||
(
|
||
<collectionCode id="B261433DB94B68B38676337E616C40E7" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14449" lsid="urn:lsid:biocol.org:col:14449" name="Université Libre de Bruxelles" type="Herbarium">BRLU</collectionCode>
|
||
); Kasombo,
|
||
<date id="6AC259E31DCC4BA2A5D69532440200E7" value="1960-02-03">3 Feb 1960</date>
|
||
,
|
||
<emphasis id="CB2D47264DCC351AC82A3E56FED04982" italics="true">P. Duvigneaud 5384 Col</emphasis>
|
||
(
|
||
<collectionCode id="5C2D6882345493A0736FAF0BFB361E90" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14449" lsid="urn:lsid:biocol.org:col:14449" name="Université Libre de Bruxelles" type="Herbarium">BRLU</collectionCode>
|
||
);
|
||
<quantity id="518C9A2885A6EDE80992562DC2E99431" metricMagnitude="4" metricUnit="m" metricValue="1.2" unit="km" value="12.0">12 km</quantity>
|
||
NW de Lubumbashi,
|
||
<date id="715BF57CC0C112A2DF52F34A3A461D4D" value="1958-03-12">12 Mar 1958</date>
|
||
,
|
||
<emphasis id="789DBC08C970DBC51A72C6F99AB05D3A" italics="true">A. Gathy 300</emphasis>
|
||
(
|
||
<collectionCode id="0DD569DF686AB69DD85F3F0B617AFCB6" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14449" lsid="urn:lsid:biocol.org:col:14449" name="Université Libre de Bruxelles" type="Herbarium">BRLU</collectionCode>
|
||
); Elisabethville [Lubumbashi],
|
||
<date id="6B1178618D4BD2BDB37F61992C2B5620" value="1912-03">Mar 1912</date>
|
||
,
|
||
<emphasis id="E74883423C4CDAE45638F61727E0006B" italics="true">
|
||
<collectionCode id="C6250AE39CAEBE3B1B6DF97E75B33F13" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" type="Herbarium">H</collectionCode>
|
||
. Homblé 220
|
||
</emphasis>
|
||
(
|
||
<collectionCode id="C4F9CDE99EEC1DE4F59B74F1149F6E7E" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
|
||
) (about this specimen, see note under
|
||
<taxonomicName id="EE4D2BA36841D24B20222E2DDCDED9BB" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="heterotrichus">
|
||
<emphasis id="89DBB6922A3A95417BEA2447D001489D" italics="true">C. heterotrichus</emphasis>
|
||
</taxonomicName>
|
||
); Kasapa,
|
||
<date id="92FD043A6B0660342C8390F87AFDEF05" value="1966-02-26">26 Feb 1966</date>
|
||
,
|
||
<emphasis id="27A96B232485772916F15E12F90EBD7B" italics="true">F. Malaisse 4012</emphasis>
|
||
(
|
||
<collectionCode id="E9642686D391E6CE5B2BD78D70817ED7" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
|
||
,
|
||
<collectionCode id="FE06BAA25636671B13E4A961D274B245" country="Democratic Republic of the Congo" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14451" lsid="urn:lsid:biocol.org:col:14451" name="Université Nationale du Zaïre" type="Herbarium">LSHI</collectionCode>
|
||
); Ferme prince Léopold,
|
||
<date id="3EF459A830B376DE99694C26BB3FF650" value="1927-02-21">21 Feb 1927</date>
|
||
,
|
||
<emphasis id="66B3B216CF8065DCD3FE8ECEFD6D6BE4" italics="true">P. Quarré 1032</emphasis>
|
||
(
|
||
<collectionCode id="1401E1FA0465DC26526F30BA62340BB0" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
|
||
); Keyberg,
|
||
<quantity id="DD17371B7F90B1D6E565638949CF7DB6" metricMagnitude="3" metricUnit="m" metricValue="9.0" unit="km" value="9.0">9 km</quantity>
|
||
<collectionCode id="7AC6FB909FF4697C99F9ADBF96D35AB3">SW</collectionCode>
|
||
of Lubumbashi,
|
||
<date id="A4DB7871861C79B9B8AEA9A97433A8DA" value="1948-02-27">27 Feb 1948</date>
|
||
,
|
||
<emphasis id="E5377BE56E8048D5EB9977DA15C384B8" italics="true">A. Schmitz 1392</emphasis>
|
||
(
|
||
<collectionCode id="CCF3777E04A913154EDAE008ED00C9B6" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
|
||
);
|
||
<quantity id="C8B92EEBEB157C85ACBC8B2238FE3ABB" metricMagnitude="4" metricUnit="m" metricValue="1.4" unit="km" value="14.0">14 km</quantity>
|
||
from Lubumbashi,
|
||
<date id="B70593505D09934708CBDE525631E2AD" value="1966-02-17">17 Feb 1966</date>
|
||
,
|
||
<emphasis id="36F6D31F67896E33FB76CB79D58B8C54" italics="true">J. - J. Symoens 12233</emphasis>
|
||
(
|
||
<collectionCode id="DA4351C39E9F1080FE4C5A1DE3B3FECE" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
|
||
,
|
||
<collectionCode id="63C589F5D48C278062FA4B527B0944F3" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" type="Herbarium">K</collectionCode>
|
||
,
|
||
<collectionCode id="E0054117E3527095250648DE1E6D8393" country="Democratic Republic of the Congo" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14451" lsid="urn:lsid:biocol.org:col:14451" name="Université Nationale du Zaïre" type="Herbarium">LSHI</collectionCode>
|
||
); Likasi, Panda,
|
||
<date id="DBBA1760B7BB06EF78446767C50A2F3F" value="1970-03-25">25 Mar 1970</date>
|
||
,
|
||
<emphasis id="DAA8D8E24D05ABE5F7DA9523AD4B6AE1" italics="true">S. Lisowski 23382</emphasis>
|
||
(
|
||
<collectionCode id="E61A935BEE637A1960318F2C2F83D6C4" country="Poland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15257" lsid="urn:lsid:biocol.org:col:15257" name="Adam Mickiewicz University" type="Herbarium">POZG</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EPYAM" type="notes">
|
||
<paragraph id="86B09514230EFB47475C40269FA77B36">
|
||
<heading id="E61C698B3E2ACB98607F5A203E34F298" reason="title">Notes.</heading>
|
||
</paragraph>
|
||
<paragraph id="4AF65D26C124DC87F6487A2A2741C9CD">
|
||
1.
|
||
<taxonomicName id="46D11472C37F975845CB14411CE07011" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="homblei">
|
||
<emphasis id="3C79749267D09F57B381C1039AC47D8A" italics="true">C. homblei</emphasis>
|
||
</taxonomicName>
|
||
is superficially similar to
|
||
<taxonomicName id="00A29E026C3A022F1FE4CA03EB4B37DE" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="bojeri">
|
||
<emphasis id="042A6E6B64985AC6B45DB30A79A57C5E" italics="true">C. bojeri</emphasis>
|
||
</taxonomicName>
|
||
and was synonymised with it by
|
||
<bibRefCitation id="7890F83F46D24FE91DAFE7351E037750" author="Paton" editor="Beentje HJ & Ghazanfar SA & Polhill RM" etAl="et al." firstAuthor="Paton" journalOrPublisher="Royal Botanic Gardens, Kew, Richmond" pagination="1-430" refId="B20" refString="Paton AJ, Bramley G, Ryding O, Polhill RM, Harvey YB, Iwarsson M, Willis F, Phillipson PB, Balkwill K, Lukhoba CW, Otieno D, Harley RM (2009) Lamiaceae (Labiatae). In: Beentje HJ, Ghazanfar SA, Polhill RM (Eds) Flora of Tropical East Africa. Royal Botanic Gardens, Kew, Richmond, 1–430." title="Lamiaceae (Labiatae)." volumeTitle="Flora of Tropical East Africa." year="2009">Paton et al. (2009</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="2D999AD57B35496B6601FE31D29307E2" author="Paton" editor="Timberlake J" firstAuthor="Paton" journalOrPublisher="Royal Botanic Gardens, Kew, Richmond" pagination="1-346" refId="B21" refString="Paton AJ, Bramley G, Ryding O, Polhill RM, Harvey YB, Iwarsson M, Willis F, Phillipson PB, Balkwill K, Lukhoba CW, Otieno D, Harley RM (2013) Lamiaceae. In: Timberlake J (Ed.) Flora Zambesiaca 8 (8). Royal Botanic Gardens, Kew, Richmond, 1–346." title="Lamiaceae." volumeTitle="Flora Zambesiaca 8 (8)." year="2013">2013</bibRefCitation>
|
||
). It differs from it in the perennial habit (this often difficult to observe), the more sharply serrate leaf margin, verticils with fewer flowers, longer pedicels and longer calyx at anthesis. The generally unbranched shoot is also typical with only ca. 5 pairs of leaves, the uppermost pair sessile and bracteiform. The species is widespread in miombo woodlands in the region of Lubumbashi.
|
||
</paragraph>
|
||
<paragraph id="DF75BE446E897FB2C4CC121F50A677D0">
|
||
2. Lectotypification of
|
||
<taxonomicName id="52B92DF05F31EEF68D71C107359156D2" authority="De Wild. De Wildeman (1921 b)" authorityName="De Wild. De Wildeman" authorityYear="1921" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="homblei">
|
||
<emphasis id="E46F8A0502BF03A672DC0957747447D8" italics="true">Coleus homblei</emphasis>
|
||
De Wild.
|
||
<bibRefCitation id="2ADD3E0B1D8885F61DC6E7CCCF94C2F2" DOI="10.2307/3666466" author="De Wildeman E" issue="2" journalOrPublisher="Annales de la Société scientifique de Bruxelles" pagination="14-88" refId="B10" refString="De Wildeman E (1921 b) Notes sur la Flore du Katanga. 7. Annales de la Société scientifique de Bruxelles 41 (2): 14–88. https://doi.org/10.2307/3666466" title="Notes sur la Flore du Katanga. 7." volume="41">De Wildeman (1921 b</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
cited
|
||
<specimenCount id="CA0511A8A555AE36BD4E82BE94D478A1" count="2" type="generic" typeStatus="syntype">two syntypes</specimenCount>
|
||
, i. e.
|
||
<emphasis id="14289E97E810D1CE6007090F0B2CB8D8" italics="true">
|
||
<collectionCode id="5E5A33ADECFF908BB21095F028B96838" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" type="Herbarium">H</collectionCode>
|
||
. Homblé 210
|
||
</emphasis>
|
||
(
|
||
<typeStatus id="7992913A756243D6EC618A2DBA768818" type="syntype">syntype</typeStatus>
|
||
<collectionCode id="D3C1F8E29B78DFA0BCD164651CF6B7D7" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
|
||
[
|
||
<specimenCode id="E48D3B55309ECBB9648949F55D6FC67D">
|
||
<collectionCode id="52036CF6EABE177E0C70A0F05AC83B78" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
|
||
0000006262181
|
||
</specimenCode>
|
||
], [
|
||
<specimenCode id="54FDEAC5AD300E553F7111E3B10CEF6B">
|
||
<collectionCode id="C83333398B69264EFD7C7B9EC698FA1D" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
|
||
0000006262518
|
||
</specimenCode>
|
||
]),
|
||
<collectingCountry id="768DE6C5DAAFCD54188B2B94970090FA">DR. Congo</collectingCountry>
|
||
, Elisabethville [Lubumbashi], brousse,
|
||
<date id="74F128DDADB399109DCBC01E42338BF5" value="1912-02">Feb 1912</date>
|
||
&
|
||
<emphasis id="006C2879B613D799EAE45E69CA925C2B" italics="true">
|
||
<collectionCode id="126A6BF3522045CC5E1EF731C33F2F51" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" type="Herbarium">H</collectionCode>
|
||
. Homblé 1259
|
||
</emphasis>
|
||
(
|
||
<typeStatus id="65EED3423CFE42B0B290FD2E71CC9B5E" type="syntype">syntype</typeStatus>
|
||
<collectionCode id="7527698697B91F323A76BEB83341BB7D" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
|
||
[
|
||
<specimenCode id="A3B1DC892DD6C0AA73AF97548625C312">
|
||
<collectionCode id="A795FB70F0BB1C23E5C7C48967AC72CC" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
|
||
0000005201518
|
||
</specimenCode>
|
||
], [
|
||
<specimenCode id="4B7A019EC945EA0C25C30CD3C2121480">
|
||
<collectionCode id="4BFCCA4C6A8943FBE281D98BCE8F301A" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
|
||
0000005201846
|
||
</specimenCode>
|
||
]), Plateau Biano, Tshisinka, partie boisée,
|
||
<date id="D45E51C4652278939BAE71334C983C6B" value="1913-02">Feb 1913</date>
|
||
. Both
|
||
<typeStatus id="1064AF30EBD99934B96F530A32D25885" type="syntype">syntypes</typeStatus>
|
||
are unusual in having shoots branched from the base. Both match the protologue.
|
||
<emphasis id="E6AE73EE9732DCDCAF9F3834F106C591" italics="true">
|
||
<collectionCode id="3AB4ACA4EA1C4EE279F307EDC6EC354A" country="Finland" lsid="urn:lsid:biocol.org:col:15618" name="University of Helsinki" type="Herbarium">H</collectionCode>
|
||
. Homblé
|
||
</emphasis>
|
||
210 is more representative of the sharply serrate leaf margin and sheet [
|
||
<specimenCode id="D4846F9F196B95F7498EDA6922DE2935">
|
||
<collectionCode id="98191479BE04DE01893E7041775F29DE" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
|
||
0000006262181
|
||
</specimenCode>
|
||
] is designated as the
|
||
<typeStatus id="5A3AA94A3127CE45DC0CCF3DC05917AD" type="lectotype">lectotype</typeStatus>
|
||
because the label has “
|
||
<taxonomicName id="22962CFD74FE19E7BF11E8EED5AC7A6D" authorityName="De Wild. De Wildeman" authorityYear="1921" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="homblei">
|
||
<emphasis id="11AB05C61B9B054B60310F93346D9D60" italics="true">Coleus homblei</emphasis>
|
||
</taxonomicName>
|
||
” in De Wildeman’s handwriting.
|
||
</paragraph>
|
||
<paragraph id="08B3126DA139181CE94CCA2A3C949E09">
|
||
3. The type materials of
|
||
<taxonomicName id="9690ED28240FBD455D9CBA181F0E945D" authority="A. J. Paton" authorityName="A. J. Paton" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="sigmoideus">
|
||
<emphasis id="21ADD556F0A8DE0B675E754B7F544034" italics="true">Coleus sigmoideus</emphasis>
|
||
A. J. Paton
|
||
</taxonomicName>
|
||
show the diagnostic traits of
|
||
<taxonomicName id="567D897C3CF4367A9DDEC87B71438359" authority="De Wild." authorityName="De Wild." class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="homblei">
|
||
<emphasis id="2283C0928BE08A5BB33A4ADEFA53D395" italics="true">Coleus homblei</emphasis>
|
||
De Wild.
|
||
</taxonomicName>
|
||
, differing only in the presence of tubers, while tubers have not been collected in materials from
|
||
<collectingCountry id="E0EABBF1B58A1AAFC48672B4163D261E">DR. Congo</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7A209DCE84D522C692CDD3D0CBBECC74">
|
||
4.
|
||
<collectingRegion id="EED1DD7EB488E140ED9E7883ED2796F8" country="South Sudan" name="Lakes">See</collectingRegion>
|
||
also note under
|
||
<taxonomicName id="BC4C34DBC5AB257EB0AB3E20915E61A6" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="heterotrichus">
|
||
C.
|
||
<emphasis id="C38EDFCFE08FDB06B8E287A89121B167" italics="true">heterotrichus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |