treatments-xml/data/60/62/82/6062828ADDB65E4BBC565D0DBD10AC0E.xml

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<mods:title id="4298CBF88BF77810EA6458BE8DE60D2D">A new genus and five new species of pseudoscorpions (Arachnida, Pseudoscorpiones, Withiidae) from Colombia</mods:title>
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<mods:namePart id="B94C6DA49A9C1759CD8C054F39AEA358">Romero-Ortiz, Catalina</mods:namePart>
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<mods:affiliation id="0270AC52F175D7ACB883285927E7427D">Laboratorio de Sistematica y Biologia Comparada de Insectos, Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota, Colombia</mods:affiliation>
<mods:nameIdentifier id="3140AF625F83A5AA771DB87CF03935EA" type="email">icromeroo@unal.edu.co</mods:nameIdentifier>
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<mods:namePart id="A3764F2D3D8BD3886DA568F0768F34D3">Sarmiento, Carlos E.</mods:namePart>
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<mods:affiliation id="AB1B9DCEAE5E5E14B38B09860D5B401F">Laboratorio de Sistematica y Biologia Comparada de Insectos, Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota, Colombia</mods:affiliation>
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<mods:namePart id="839F27666E3A44E741DE5C16A85170F6">Harvey, Mark S.</mods:namePart>
<mods:nameIdentifier id="260D7D201C257A8A4103A372FD5AB4FE" type="ORCID">https://orcid.org/0000-0003-1482-0109</mods:nameIdentifier>
<mods:affiliation id="068409132F07D410C91CEF9A31964B71">Collections and Research, Western Australian Museum, Welshpool, Western Australia 6106, Australia &amp; School of Biological Sciences, The University of Western Australia, Crawley, Western Australia 6009, Australia</mods:affiliation>
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<mods:date id="E5F28A10CA3CA61E84B6F638B866463E">2023</mods:date>
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<paragraph id="94A2E1261EE588F18CA45E9BA33DD2BA" pageId="0" pageNumber="301">
<taxonomicName id="5546CC9D1F1F478108257071D4262A27" LSID="https://zoobank.org/A1720398-A29E-4F9E-8E8E-6D9DD9EDFB71" authority="Romero-Ortiz &amp; Sarmiento &amp; Harvey, 2023" authorityName="Romero-Ortiz &amp; Sarmiento &amp; Harvey" authorityYear="2023" class="Arachnida" family="Withiidae" genus="Parawithius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parawithius bromelicola" order="Pseudoscorpiones" pageId="0" pageNumber="301" phylum="Arthropoda" rank="species" species="bromelicola" status="sp. nov.">Parawithius bromelicola</taxonomicName>
<taxonomicNameLabel id="98F0AA20F05753E46B8583C9A42A3264" pageId="0" pageNumber="301">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="C38B8F49F859583E970A8653A94F371B" pageId="0" pageNumber="301" type="description">
<paragraph id="6B3C29E9D142565B8CC11645063AB21A" pageId="0" pageNumber="301">
<figureCitation id="5E9BD0D18F2C688BAD30FFF3EF200EE6" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of the new species (males) A Paciwithius chimbilacus sp. nov. B Cystowithius florezi sp. nov. C Parawithius bromelicola sp. nov. D Oligowithius achagua sp. nov. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1184.106698.figure1" httpUri="https://binary.pensoft.net/fig/940870" pageId="0" pageNumber="301">Figs 1C</figureCitation>
<figureCitation id="43FE6D63B4E1E7B1A38FA97B70D5A5AA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of the new species and the new record of Withiidae from Colombia." figureDoi="10.3897/zookeys.1184.106698.figure2" httpUri="https://binary.pensoft.net/fig/940871" pageId="0" pageNumber="301">, 2</figureCitation>
<figureCitation id="0AC3CA11F0E203EF02C2B132F6E1E663" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Parawithius bromelicola sp. Nov. (male holotype, ICN-Aps- 082) A carapace B chelicera C rallum D right chela E left leg I F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, D-F, H); 0.05 mm (B); 0.0125 mm (C, I)." figureDoi="10.3897/zookeys.1184.106698.figure7" httpUri="https://binary.pensoft.net/fig/940876" pageId="0" pageNumber="301">, 7</figureCitation>
<figureCitation id="8EAD658524718484C4BF858DE8C17E90" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glandular setae patches of the new species A Paciwithius chimbilacus sp. nov. (holotype, ICN-Aps- 076) B Parawithius bromelicola sp. nov. (holotype, ICN-Aps- 082) C Paciwithius valduparensis sp. nov. (holotype, ICN-Aps- 597) D Cystowithius florezi sp. nov. (holotype, ICN-Aps- 07) E, F Oligowithius achagua sp. nov. E male paratype (ICN-Aps- 388) F male holotype (ICN-Aps- 388). Scale bars: 0.1 mm (A-C, E, F); 0.05 mm (D)." figureDoi="10.3897/zookeys.1184.106698.figure9" httpUri="https://binary.pensoft.net/fig/940878" pageId="0" pageNumber="301">, 9B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8F3B81D78DD240AD0C0578A07068D7A1" pageId="0" pageNumber="301" type="materials_examined">
<paragraph id="14707583BC1A6605E1E43A6B7C5782FE" pageId="0" pageNumber="301">Material examined.</paragraph>
<paragraph id="299A293A7FD0AB88F05D114CFDC88090" pageId="0" pageNumber="301">
<materialsCitation id="27C3F0565AD3FFBD5C4D20DF6DA57F45" collectingDate="2004-03-07" collectorName="AL Leon" country="Colombia" county="Cogua" elevation="2900" latitude="5.19275" location="Llano Grande" longLatPrecision="1" longitude="-73.89828" municipality="Embalse del Neusa Tausa" specimenCount="♂" stateProvince="Cundinamarca" typeStatus="Holotype">
<emphasis id="68DD858678D72DD87A628945585F1B05" bold="true" pageId="0" pageNumber="301">
<emphasis id="C4473881BE4EF875AB710BDF5958902B" italics="true" pageId="0" pageNumber="301">
<typeStatus id="8D062AFB20D8BD93FE9F6220C33FE723">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="379735B4E3C2927D6D31E05F52050D40" name="Colombia">Colombia</collectingCountry>
• 
<specimenCount id="D94692BA2ED162DC84E15FCD35ABFD51">♂</specimenCount>
; 
<collectingRegion id="DB771474ADA4BBF73CA6F631BC494E19" country="Colombia" name="Cundinamarca">Cundinamarca</collectingRegion>
, 
<collectingCounty id="0E67A8A6503C85715A786A787D571AE6">Cogua</collectingCounty>
, 
<collectingMunicipality id="86A65762D6C8F22DC4C4CB67497DBFB2">Embalse del Neusa Tausa</collectingMunicipality>
, 
<location id="688A20A14DB093BA9E9821AB3B2C0010" LSID="urn:lsid:plazi:treatment:6062828ADDB65E4BBC565D0DBD10AC0E:688A20A14DB093BA9E9821AB3B2C0010" country="Colombia" county="Cogua" latitude="5.19275" longLatPrecision="1" longitude="-73.89828" municipality="Embalse del Neusa Tausa" name="Llano Grande" stateProvince="Cundinamarca">Llano Grande</location>
; [
<geoCoordinate id="F77BFCF43F7CC5E324DC0C5E06C8C204" degrees="5" direction="north" minutes="11" orientation="latitude" precision="1" seconds="33.9" value="5.19275">5°11'33.9&quot;N</geoCoordinate>
, 
<geoCoordinate id="43616A15951EA5B295248A82B704EC9F" degrees="73" direction="west" minutes="53" orientation="longitude" precision="1" seconds="53.8" value="-73.89828">73°53'53.8&quot;W</geoCoordinate>
]; 
<quantity id="D17DC607BAD89B54139E9FF34D5D5DDF" metricMagnitude="3" metricUnit="m" metricValue="2.9" unit="m" value="2900.0">
<elevation id="9DB4E7959BDA71F46263196C5D4110B7" metricMagnitude="3" metricUnit="m" metricValue="2.9" unit="m" value="2900.0">2900 m</elevation>
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; 
<collectingDate id="D4CF0D6689C826FE0A41CE11BC4E9A11" value="2004-03-07">7 Mar. 2004</collectingDate>
; 
<collectorName id="1EC6BCDE34284E64A3EE111F57F13F8D">AL Leon</collectorName>
leg.; under tree bark; ICN-APs-082.
</materialsCitation>
</paragraph>
<paragraph id="F069624FF81360E31D003893F2AFE0DB" pageId="0" pageNumber="301">
<materialsCitation id="6E35955C73D61E45BA667E5251ADD46F" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="AL Leon" country="Colombia" county="Cogua" elevation="2900" latitude="5.19275" location="Cundinamarca" longLatPrecision="1" longitude="-73.89828" municipality="Embalse del Neusa Tausa" specimenCount="1" specimenCount-male="1" stateProvince="Cundinamarca" typeStatus="Paratypes">
<emphasis id="F3549F7991E7D0A9F7D5401A8080C467" bold="true" pageId="0" pageNumber="301">
<emphasis id="79B9E4E573636037ECD0BC7BF65B2040" italics="true" pageId="0" pageNumber="301">
<typeStatus id="13A76346731FC4C027577FDECA8EA624">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="810FC5E6712FA8A9366E17C9BD8827E6" name="Colombia">Colombia</collectingCountry>
• 
<specimenCount id="A3C90521BBCC54216DB7EBC539AB1EFC" type="male">1 ♂</specimenCount>
; 
<collectingRegion id="4AE13E7B07F83C51AD72DB2F32EF6EDE" country="Colombia" name="Cundinamarca">Cundinamarca</collectingRegion>
, same data as for the holotype
</materialsCitation>
. • 
<materialsCitation id="CFC45F726B4911A355723B3B2AADD2D3" collectingDate="2019-09-08" collectorName="C. Romero-Ortiz, F. Garcia, J. J. Lagos, A. Carvajal" country="Colombia" county="Sesquile" elevation="2740" latitude="5.0380554" location="Cerro de Las Tres Viejas" longLatPrecision="1" longitude="-73.78694" municipality="Camino" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="AEEAFFEE7A6D6A21CE161FC81C996BEC" type="female">1 ♀</specimenCount>
, 
<specimenCount id="D3BFE2E45A6C34DE9E69DDA8092742FE" type="male">1 ♂</specimenCount>
; 
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, 
<collectingMunicipality id="FDD2DBE67DC0BE76ED3D68493673DC6A">Camino</collectingMunicipality>
al 
<location id="8AE368B572BC4D43AD6452FF6962D6D6" LSID="urn:lsid:plazi:treatment:6062828ADDB65E4BBC565D0DBD10AC0E:8AE368B572BC4D43AD6452FF6962D6D6" country="Colombia" county="Sesquile" latitude="5.0380554" longLatPrecision="1" longitude="-73.78694" municipality="Camino" name="Cerro de Las Tres Viejas">Cerro de Las Tres Viejas</location>
; 
<geoCoordinate id="2B3520F8E984153398A3597B78CA35BE" degrees="5" direction="north" minutes="02" orientation="latitude" precision="1" seconds="17.0" value="5.0380554">5°02'17.0&quot;N</geoCoordinate>
, 
<geoCoordinate id="59B62603D520DDE425C1428D96BEABFA" degrees="73" direction="west" minutes="47" orientation="longitude" precision="1" seconds="13.0" value="-73.78694">73°47'13.0&quot;W</geoCoordinate>
; 
<quantity id="35D9F235F7F68D8BE62DA84305BD5580" metricMagnitude="3" metricUnit="m" metricValue="2.7399999999999998" unit="m" value="2740.0">
<elevation id="BF8371275C04D00722F4F3A19D967082" metricMagnitude="3" metricUnit="m" metricValue="2.7399999999999998" unit="m" value="2740.0">2740 m</elevation>
</quantity>
; 
<collectingDate id="853299FE7B3F28F347503B10E6434E5F" value="2019-09-08">8 Sep. 2019</collectingDate>
; 
<collectorName id="A2E5475C58BD3E22ABC359FE4CF1753E">C. Romero-Ortiz</collectorName>
, 
<collectorName id="3A87A7F96B95AC00955DCBBBEA1C0C64">F. Garcia</collectorName>
, 
<collectorName id="3CC7942F1C57AE2480D7BCB05AC4CCBD">J.J. Lagos</collectorName>
, 
<collectorName id="D390238F520215AD9E20A174AD63CBFC">A. Carvajal</collectorName>
, D. Mayorga-Ch leg.; on bromeliad; ICN-APs-836
</materialsCitation>
.
</paragraph>
<paragraph id="D53BE66D9DBAC37893C74F26F9624978" pageId="0" pageNumber="301">
<emphasis id="1034C6B73C9EF11521C4EEF8BEC07F74" bold="true" pageId="0" pageNumber="301">
<emphasis id="EBD6C963340541341B2B5D8EC18176D4" italics="true" pageId="0" pageNumber="301">Other material</emphasis>
.
</emphasis>
<materialsCitation id="5DA892F7B029035119E0CFE51A7D481E" collectingDate="2020-01-01" collectorName="C. Romero-Ortiz, J. J. Lagos" country="Colombia" elevation="2620" latitude="6.706583" location="Bucaramanga" longLatPrecision="1" longitude="-72.74794" municipality="Malaga" specimenCount="1" specimenCount-female="1" stateProvince="Santander">
<collectingCountry id="4468A473FF9739F655890C89E14241EE" name="Colombia">Colombia</collectingCountry>
• 
<specimenCount id="522BD359CB47897809AF9575C81EE747" type="female">1 ♀</specimenCount>
; 
<collectingRegion id="F62E60518DBE4B50797B720B28491AD5" country="Colombia" name="Santander">Santander</collectingRegion>
, 
<normalizedToken id="CE0EF0324D4983BF7C1B94C20C7DCB88" originalValue="Málaga">
<collectingMunicipality id="9F1EE1B3FE8BB1863AF5BBFCA8707648">Malaga</collectingMunicipality>
</normalizedToken>
, Vda. Buenavista, km 7 
<normalizedToken id="873ED53A68DE81678BB215E15144F198" originalValue="vía">via</normalizedToken>
<location id="500C6013F75521176CFD1D2EB20D6DC0" LSID="urn:lsid:plazi:treatment:6062828ADDB65E4BBC565D0DBD10AC0E:500C6013F75521176CFD1D2EB20D6DC0" country="Colombia" latitude="6.706583" longLatPrecision="1" longitude="-72.74794" municipality="Malaga" name="Bucaramanga" stateProvince="Santander">Bucaramanga</location>
; 
<geoCoordinate id="B61A2D80C209ABA28E6E613FC8F70E23" degrees="6" direction="north" minutes="42" orientation="latitude" precision="1" seconds="23.7" value="6.706583">6°42'23.7&quot;N</geoCoordinate>
, 
<geoCoordinate id="6AD65D1D96C63435F635D133C1F02849" degrees="72" direction="west" minutes="44" orientation="longitude" precision="1" seconds="52.6" value="-72.74794">72°44'52.6&quot;W</geoCoordinate>
; 
<quantity id="467A0B586A8DACF132CB2F4C9DB2943F" metricMagnitude="3" metricUnit="m" metricValue="2.62" unit="m" value="2620.0">
<elevation id="3AC3D9402B7D8CD7EA36F01BA616F40B" metricMagnitude="3" metricUnit="m" metricValue="2.62" unit="m" value="2620.0">2620 m</elevation>
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; 
<collectingDate id="1A816C152B3908B708850700CAAD1CD3" value="2020-01-01">1 Jan. 2020</collectingDate>
; 
<collectorName id="CB6F530394B8B8BFEF4BB19E398C57B2">C. Romero-Ortiz</collectorName>
, 
<collectorName id="8CFF54576AFD582BA08A7D6CF7D523C8">J.J. Lagos</collectorName>
leg.; on bromeliad under 
<normalizedToken id="A82AC1FBE2781EC76A499DFD91D99C28" originalValue="“Loqueto”">&quot;Loqueto&quot;</normalizedToken>
tree 
<taxonomicName id="CD688006A14934B7F8A0E8F729D84A2D" class="Magnoliopsida" family="Escalloniaceae" genus="Escallonia" higherTaxonomySource="GBIF,IPNI" kingdom="Plantae" lsidName="Escallonia pendula" order="Escalloniales" pageId="0" pageNumber="301" phylum="Magnoliophyta" rank="species" species="pendula">
<emphasis id="01D6A6102C5F0E9362FB0F61E336D006" italics="true" pageId="0" pageNumber="301">Escallonia pendula</emphasis>
</taxonomicName>
(Ruiz &amp; Pav.) Pers.; ICN-APs-847
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="855BBF14869838D805D8197423A3ABB9" pageId="0" pageNumber="301" type="diagnosis">
<paragraph id="688774CDAF36296FB8A5FC35C66A4539" pageId="0" pageNumber="301">Diagnosis.</paragraph>
<paragraph id="D0594958B37573BB43F17AA7E808D258" pageId="0" pageNumber="301">
<taxonomicName id="FEE9AFAC066FA0D3D74D1C0175CDE088" authorityName="Romero-Ortiz &amp; Sarmiento &amp; Harvey" authorityYear="2023" class="Arachnida" family="Withiidae" genus="Parawithius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parawithius bromelicola" order="Pseudoscorpiones" pageId="0" pageNumber="301" phylum="Arthropoda" rank="species" species="bromelicola">
<emphasis id="334009975E93DE0E5E5404F6F7D6F480" italics="true" pageId="0" pageNumber="301">Parawithius bromelicola</emphasis>
</taxonomicName>
sp. nov. can be separated from 
<taxonomicName id="413E8E5BEECEC67D0DBF51B4B31DC752" lsidName="P. nobilis" pageId="0" pageNumber="301" rank="species" species="nobilis">
<emphasis id="1E9FFBF2FCCCE8E716759A9A0B61123E" italics="true" pageId="0" pageNumber="301">P. nobilis</emphasis>
</taxonomicName>
(With, 1908) by the stouter pedipalpal segments (i.e. patella 3.36-3.44 
<normalizedToken id="DFB3B62C9359C449C7F173FC1A7BB7E0" originalValue="×">x</normalizedToken>
longer than broad compared with 3.24-3.30 
<normalizedToken id="F01CBF878ECE197AA10E1081AC81AFC5" originalValue="×">x</normalizedToken>
longer than broad, and the chela without pedicel 3.39-3.47 
<normalizedToken id="859E909445E7F794B6B020FF6A7EEB9A" originalValue="×">x</normalizedToken>
longer than broad, compared to 3.58-3.85 
<normalizedToken id="80DE3C20D8AC89F9B10C629B0FE5C1CD" originalValue="×">x</normalizedToken>
longer than broad); the extension of the strongly clavate setae on the dorsal surface of fixed chelal finger (i.e., distal to 
<emphasis id="E105481E8601E8D16670F78EC3E664C9" italics="true" pageId="0" pageNumber="301">it</emphasis>
and 
<emphasis id="6F5BC6ED7DF491C8BFCDD3DB5AD0F30B" italics="true" pageId="0" pageNumber="301">est</emphasis>
compared to proximal to 
<emphasis id="ADF03B50C46A8F786711C9AA1548B869" italics="true" pageId="0" pageNumber="301">it</emphasis>
and 
<emphasis id="005EC5418D2A8BEE78522AD51CC5A740" italics="true" pageId="0" pageNumber="301">est</emphasis>
); from 
<taxonomicName id="1223DC7620E6A629720D42054A8BF2E6" lsidName="P. iunctus" pageId="0" pageNumber="301" rank="species" species="iunctus">
<emphasis id="B16998551F934EC5BAC551E2DB247FE9" italics="true" pageId="0" pageNumber="301">P. iunctus</emphasis>
</taxonomicName>
Beier, 1932 and 
<taxonomicName id="A6FEB04847E04323ADD33E9727E8717D" lsidName="P. pseudorufus" pageId="0" pageNumber="301" rank="species" species="pseudorufus">
<emphasis id="054503DD46EA176F3F28A462F8D5674E" italics="true" pageId="0" pageNumber="301">P. pseudorufus</emphasis>
</taxonomicName>
Beier, 1932 by the presence of pale spots on the carapace metazone, in addition to the size of chelal fingers compared to the palpal hand (i.e. fingers shorter to the hand in 
<taxonomicName id="277FCDEC5BCC31CF20EF4068FB0F30FB" lsidName="P. bromelicola" pageId="0" pageNumber="301" rank="species" species="bromelicola">
<emphasis id="8DFAFDBB92A3F97910028EEC15021774" italics="true" pageId="0" pageNumber="301">P. bromelicola</emphasis>
</taxonomicName>
sp. nov., 
<taxonomicName id="0A003C1F54AC3F67C673395C7A43644B" lsidName="P. nobilis" pageId="0" pageNumber="301" rank="species" species="nobilis">
<emphasis id="8F352873F4C080D054F36BCD6B52CB95" italics="true" pageId="0" pageNumber="301">P. nobilis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="3B4752CB7113C4D1987091FE8675E53B" lsidName="P. iunctus" pageId="0" pageNumber="301" rank="species" species="iunctus">
<emphasis id="0E4A8EBEFDE096F4C4581A197F60808A" italics="true" pageId="0" pageNumber="301">P. iunctus</emphasis>
</taxonomicName>
and longer in 
<taxonomicName id="912816028E93BB264ABFFA1658D8C7AA" lsidName="P. pseudorufus" pageId="0" pageNumber="301" rank="species" species="pseudorufus">
<emphasis id="5A9DC145A16DF8400FC3FD04B20B34BE" italics="true" pageId="0" pageNumber="301">P. pseudorufus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="6F01595CFACE1632B98AB040E7C76512" pageId="0" pageNumber="301" type="description">
<paragraph id="362695F4A330DD408EBC8E4AE171C9B2" pageId="0" pageNumber="301">Description.</paragraph>
<paragraph id="5B76310C17BF4B4077A8115DAEFE0A73" pageId="0" pageNumber="301">
<emphasis id="51F9DC2444200A146D7F40A2F634A908" bold="true" pageId="0" pageNumber="301">
Adults. 
<emphasis id="AC1860BD939AE6B6D33EF155BFEF46C8" italics="true" pageId="0" pageNumber="301">Color</emphasis>
</emphasis>
: yellowish brown, carapace darker than body, carapace metazone with paired pale spots; pedipalps reddish brown, uniform in color, very granulated; tergites yellow-brown; big leg segments darker at posterior margin (Fig. 
<figureCitation id="8E16D3EFD7D55640600BD2B0D6A820CF" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of the new species (males) A Paciwithius chimbilacus sp. nov. B Cystowithius florezi sp. nov. C Parawithius bromelicola sp. nov. D Oligowithius achagua sp. nov. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1184.106698.figure1" httpUri="https://binary.pensoft.net/fig/940870" pageId="0" pageNumber="301">1C</figureCitation>
).
</paragraph>
<paragraph id="14B4A3EC0918980A2AFB643C1E387B2A" pageId="0" pageNumber="301">
<emphasis id="13E812AEABC01B98581F235179D2046B" bold="true" italics="true" pageId="0" pageNumber="301">Dimensions</emphasis>
(mm): male: holotype (followed by male paratypes): body length 2.46 (2.58, 2.46). Pedipalps: trochanter 0.38/0.21 (0.37/0.23, 0.38/0.22), femur 0.83/0.19 (0.84/0.19, 0.77/0.18), patella 0.74/0.22 (0.75/0.22, 0.71/0.21), chela (with pedicel) 1.18/0.30 (1.24/0.31, 1.12/0.30), chela (without pedicel) 1.14 (1.18, 1.06), hand (without pedicel) length 0.59 (0.57, 0.53), movable finger length 0.56 (0.60, 0.53). Chelicera 0.21, movable finger length 0.17. Carapace 0.82/0.58 (0.85/0.72, 0.82/0.74); eyespot diameter 0.07. Leg I: femur 0.16/0.16, patella 0.33/0.14, tibia 0.35/0.10, tarsus 0.32/0.06. Leg IV: femur + patella 0.66/0.19, tibia 0.51/0.12, tarsus 0.37/0.07, TS 0.73.
</paragraph>
<paragraph id="13C2AC52C3BF7D1CDA5A0270F5DD240F" pageId="0" pageNumber="301">
<emphasis id="E7C00C5B345D19C971D0140F73B6972C" bold="true" pageId="0" pageNumber="301">Female</emphasis>
: paratype (followed by female other material): body length 2.68 (2.70). Pedipalps: trochanter 0.41/0.23 (0.36/0.21), femur 0.78/0.21 (0.73/0.18), patella 0.74/0.23 (0.67/0.19), chela (with pedicel) 1.24/0.35 (1.14/0.30), chela (without pedicel) 1.16 (1.08), hand (without pedicel) length 0.59 (0.57), movable finger length 0.60 (0.49). Carapace 0.88/0.68 (0.74/0.64) (width at medial area).
</paragraph>
<paragraph id="326A87F736C0F28B9F5D272F53CE9D28" pageId="0" pageNumber="301">
<emphasis id="ED54C789FCEB640E3EFEA766BE6F13C7" bold="true" italics="true" pageId="0" pageNumber="301">Carapace</emphasis>
(Fig. 
<figureCitation id="4E46E8A873464EB1516052AE86B0255F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Parawithius bromelicola sp. Nov. (male holotype, ICN-Aps- 082) A carapace B chelicera C rallum D right chela E left leg I F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, D-F, H); 0.05 mm (B); 0.0125 mm (C, I)." figureDoi="10.3897/zookeys.1184.106698.figure7" httpUri="https://binary.pensoft.net/fig/940876" pageId="0" pageNumber="301">7A</figureCitation>
): 1.42 
<normalizedToken id="9D6677E61EC9C493F46324EF6B263B89" originalValue="×">x</normalizedToken>
(1.18-1.11) (♂), 1.29 
<normalizedToken id="7E75DE297DD58F7C36D36E0F21A474EC" originalValue="×">x</normalizedToken>
(1.16) (♀) 
<normalizedToken id="56DE232C77BCE66273D6CA689D14F670" originalValue="×">x</normalizedToken>
longer than broad; lateral margins convex, not posteriorly widened; with two non-corneate eyes; with 59 (♂) setae, including four (♂) near anterior margin, seven (♂) near posterior margin, 16 in the medial zone and 36 in the anterior region; with two distinct furrows; posterior furrow slightly closer to posterior carapace margin than to median furrow (Fig. 
<figureCitation id="45F621A2AC859AAFE6730829833C961C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Parawithius bromelicola sp. Nov. (male holotype, ICN-Aps- 082) A carapace B chelicera C rallum D right chela E left leg I F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, D-F, H); 0.05 mm (B); 0.0125 mm (C, I)." figureDoi="10.3897/zookeys.1184.106698.figure7" httpUri="https://binary.pensoft.net/fig/940876" pageId="0" pageNumber="301">7A</figureCitation>
).
</paragraph>
<caption id="06FD841E91BCCB959A71D476A4FE3AE7" doi="10.3897/zookeys.1184.106698.figure7" httpUri="https://binary.pensoft.net/fig/940876" pageId="0" pageNumber="301" start="Figure 7" startId="F7">
<paragraph id="1B4797E68D4FD27EF0FF5AC6A9E862BA" pageId="0" pageNumber="301">
<emphasis id="68F855F5F34B71491BCC3DDF38C39820" bold="true" pageId="0" pageNumber="301">Figure 7.</emphasis>
<taxonomicName id="A950CBDFF040904CAED75E7827B8C931" authorityName="Romero-Ortiz &amp; Sarmiento &amp; Harvey" authorityYear="2023" class="Arachnida" family="Withiidae" genus="Parawithius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parawithius bromelicola" order="Pseudoscorpiones" pageId="0" pageNumber="301" phylum="Arthropoda" rank="species" species="bromelicola">
<emphasis id="B16A273B6FCAC1564666BC19EDF1AC00" italics="true" pageId="0" pageNumber="301">Parawithius bromelicola</emphasis>
</taxonomicName>
sp. Nov. (male holotype, ICN-Aps-082) 
<emphasis id="D7306DAD9D8D4E6673529BE53C9C150B" bold="true" pageId="0" pageNumber="301">A</emphasis>
carapace 
<emphasis id="0BE502099633B5E35AF604758E91520F" bold="true" pageId="0" pageNumber="301">B</emphasis>
chelicera 
<emphasis id="DE31141D57CCF64A9F06E68309BF0AF0" bold="true" pageId="0" pageNumber="301">C</emphasis>
rallum 
<emphasis id="7B224C73B0703EF50D41D0E13C9FB578" bold="true" pageId="0" pageNumber="301">D</emphasis>
right chela 
<emphasis id="875DC7BD8D044A64AAA1219327CF7DA1" bold="true" pageId="0" pageNumber="301">E</emphasis>
left leg I 
<emphasis id="C89C80B897F72CDAFB1291A620FA9D13" bold="true" pageId="0" pageNumber="301">F</emphasis>
left leg IV 
<emphasis id="C5E53E7649BE465E47F6C745CD286878" bold="true" pageId="0" pageNumber="301">G</emphasis>
left pedipalp 
<emphasis id="461C32FCC07C25C5651C9F9D2D08EEC8" bold="true" pageId="0" pageNumber="301">H</emphasis>
right chelal fingers (arrows show sense-spots) 
<emphasis id="26D73B83308F9181EA53BB67B617A1FB" bold="true" pageId="0" pageNumber="301">I</emphasis>
rallum. Abbreviations: Pedipalp movable finger: 
<emphasis id="9F88E0BB83EAFA3AD5B39A413BA043C2" bold="true" italics="true" pageId="0" pageNumber="301">b</emphasis>
= basal; 
<emphasis id="DF0A5AA671E0A655647AC174767293E9" bold="true" italics="true" pageId="0" pageNumber="301">sb</emphasis>
= sub-basal; 
<emphasis id="0D9D9D14AC2975CB78AE932114A7FB55" bold="true" italics="true" pageId="0" pageNumber="301">st</emphasis>
= sub-terminal; 
<emphasis id="5104E403993C5F97FB1949A74BCCFEB4" bold="true" italics="true" pageId="0" pageNumber="301">t</emphasis>
= terminal; fixed finger: 
<emphasis id="A81407B475C2EE5F28A918218BFDF3A1" bold="true" italics="true" pageId="0" pageNumber="301">eb</emphasis>
= exterior basal; 
<emphasis id="A6796ED28E1B4A8D76A94180605871F3" bold="true" italics="true" pageId="0" pageNumber="301">esb</emphasis>
= exterior sub-basal; 
<emphasis id="D0C9233528EA3B19399F67768CADF2FE" bold="true" italics="true" pageId="0" pageNumber="301">est</emphasis>
= exterior sub-terminal; 
<emphasis id="7A0DD6366BA5320DFFCFAE35417D03C9" bold="true" italics="true" pageId="0" pageNumber="301">et</emphasis>
= exterior terminal; 
<emphasis id="43A1E193C228B9B8D06092077F30AF36" bold="true" italics="true" pageId="0" pageNumber="301">ib</emphasis>
= interior basal; 
<emphasis id="A07D91F1924ED794C0268BB7019A0520" bold="true" italics="true" pageId="0" pageNumber="301">isb</emphasis>
= interior sub-basal; 
<emphasis id="FCE847E6448C39E48D12E0CA4CBEE36C" bold="true" italics="true" pageId="0" pageNumber="301">ist</emphasis>
= interior sub-terminal; 
<emphasis id="6B803CA0EA90450531423C95B3D6660F" bold="true" italics="true" pageId="0" pageNumber="301">it</emphasis>
= interior terminal. Cheliceral setae: 
<emphasis id="F8B7F71D0CE255729EB6EF1C73A2A307" bold="true" italics="true" pageId="0" pageNumber="301">b</emphasis>
= basal seta; 
<emphasis id="05A8BEF36892CB7B14AFEED138F49747" bold="true" italics="true" pageId="0" pageNumber="301">es</emphasis>
= exterior seta; 
<emphasis id="4B5499B6A5DEB5B4BFA8FA1CAC8A699B" bold="true" italics="true" pageId="0" pageNumber="301">is</emphasis>
= interior seta; 
<emphasis id="67459FE650E42F1194F975BEDD22687A" bold="true" italics="true" pageId="0" pageNumber="301">ls</emphasis>
= laminal seta. Scale bars: 0.5 mm (
<emphasis id="1293B67A9D55C5BF9B21E504A0DB67D4" bold="true" pageId="0" pageNumber="301">G</emphasis>
); 0.1 mm (
<emphasis id="FF3E846185D00FF25640370433BC0505" bold="true" pageId="0" pageNumber="301">A, D-F, H</emphasis>
); 0.05 mm (
<emphasis id="644AF9FDBE2C7FAA0FA918D446D063C9" bold="true" pageId="0" pageNumber="301">B</emphasis>
); 0.0125 mm (
<emphasis id="F44B3EAF0FC8CCB15730217AF7788CED" bold="true" pageId="0" pageNumber="301">C, I</emphasis>
).
</paragraph>
</caption>
<caption id="F895821BA43BBB60AA82810C89AD478B" doi="10.3897/zookeys.1184.106698.figure8" httpUri="https://binary.pensoft.net/fig/940877" pageId="0" pageNumber="301" start="Figure 8" startId="F8">
<paragraph id="C03A6838DA6951B81BB9728432028D60" pageId="0" pageNumber="301">
<emphasis id="5328957DE7D61EBD0846D790367C262A" bold="true" pageId="0" pageNumber="301">Figure 8.</emphasis>
<taxonomicName id="3D627A7AF4C1931C22EA331CD1C5CFDA" authorityName="Romero-Ortiz &amp; Sarmiento &amp; Harvey" authorityYear="2023" class="Arachnida" family="Withiidae" genus="Oligowithius" kingdom="Animalia" lsidName="Oligowithius achagua" order="Pseudoscorpiones" pageId="0" pageNumber="301" rank="species" species="achagua">
<emphasis id="0963949EFBF2EE48B4306D98DDA88A1D" italics="true" pageId="0" pageNumber="301">Oligowithius achagua</emphasis>
</taxonomicName>
sp. nov. (male holotype, ICN-Aps-388) 
<emphasis id="59DC0E399E497D0B56B8F697FA707F64" bold="true" pageId="0" pageNumber="301">A</emphasis>
carapace 
<emphasis id="66D96B450CB42A082993A8746B972EAF" bold="true" pageId="0" pageNumber="301">B</emphasis>
chelicera 
<emphasis id="EF53F7037573200177E8EC7D3280550F" bold="true" pageId="0" pageNumber="301">C</emphasis>
left pedipalp 
<emphasis id="0516B6F2CD23C3CC107E0816620FCFA9" bold="true" pageId="0" pageNumber="301">D</emphasis>
abdomen, sternites 
<emphasis id="BCA600B224D7CA2C65E04B85118C74C4" bold="true" pageId="0" pageNumber="301">E</emphasis>
rallum 
<emphasis id="6285D1D0A482838949751DB85EE8F40F" bold="true" pageId="0" pageNumber="301">F</emphasis>
left leg I 
<emphasis id="DBA26C3585D2F89FB8498B07599D6ABC" bold="true" pageId="0" pageNumber="301">G</emphasis>
left leg IV 
<emphasis id="3D89244291D1CEB1AE3ED98ECDAC409F" bold="true" pageId="0" pageNumber="301">H</emphasis>
right chela (arrow shows sense-spots). Abbreviations: Pedipalp movable finger: 
<emphasis id="DD3A32C3C019710B0AAD9EFD869A9EB7" bold="true" italics="true" pageId="0" pageNumber="301">b</emphasis>
= basal; 
<emphasis id="0A0A6451B471C3CC2BD83247FD4EFF2A" bold="true" italics="true" pageId="0" pageNumber="301">sb</emphasis>
= sub-basal; 
<emphasis id="00BA5823D1F3C03C2558404649300DDA" bold="true" italics="true" pageId="0" pageNumber="301">st</emphasis>
= sub-terminal; 
<emphasis id="F3435C6BFED2D02C6C2B8ACEA1F792F4" bold="true" italics="true" pageId="0" pageNumber="301">t</emphasis>
= terminal; fixed finger: 
<emphasis id="0418D2047DA06F96CB89A96731E87B93" bold="true" italics="true" pageId="0" pageNumber="301">eb</emphasis>
= exterior basal; 
<emphasis id="843EE132F8243588DAD0AEE2D5F0A8B7" bold="true" italics="true" pageId="0" pageNumber="301">esb</emphasis>
= exterior sub-basal; 
<emphasis id="1333348B18A323380F9FA247507AB3BD" bold="true" italics="true" pageId="0" pageNumber="301">est</emphasis>
= exterior sub-terminal; 
<emphasis id="E1CF66E25ABB2E58BD0948102812ED39" bold="true" italics="true" pageId="0" pageNumber="301">et</emphasis>
= exterior terminal; 
<emphasis id="98F9114EFCBB3B9490F1F8C1AAC51B54" bold="true" italics="true" pageId="0" pageNumber="301">ib</emphasis>
= interior basal; 
<emphasis id="A54AA6443A7FFE4D608E81CACF4ABC94" bold="true" italics="true" pageId="0" pageNumber="301">isb</emphasis>
= interior sub-basal; 
<emphasis id="E88D86C63AD4727C550D4DC4ECE9D41C" bold="true" italics="true" pageId="0" pageNumber="301">ist</emphasis>
= interior sub-terminal; 
<emphasis id="1228FEA4585CE5DAC9555100832C8461" bold="true" italics="true" pageId="0" pageNumber="301">it</emphasis>
= interior terminal. Cheliceral setae: 
<emphasis id="1F6514D1AEF9C52969FB77688A53FD4D" bold="true" italics="true" pageId="0" pageNumber="301">b</emphasis>
= basal seta; 
<emphasis id="C4A55CCF16B8A2C1684C90BA058682E0" bold="true" italics="true" pageId="0" pageNumber="301">es</emphasis>
= exterior seta; 
<emphasis id="C06F3629F5472B46D183F80CCFD3913B" bold="true" italics="true" pageId="0" pageNumber="301">is</emphasis>
= interior seta; 
<emphasis id="596927098EE5FD995B2173B2DCED6325" bold="true" italics="true" pageId="0" pageNumber="301">ls</emphasis>
= laminal seta. Scale bars: 8 
<normalizedToken id="92C918C7F7E4B178F066CC4CA20F200B" originalValue="µm">µm</normalizedToken>
(
<emphasis id="88323EDBBFDF9AFA356A0A6C66AD6003" bold="true" pageId="0" pageNumber="301">A, C, D, F-H</emphasis>
); 2 
<normalizedToken id="2473A79BCA164D0CA120205877F7A1A1" originalValue="µm">µm</normalizedToken>
(
<emphasis id="31E605D8D73BBFCC8D28B669D143AB35" bold="true" pageId="0" pageNumber="301">B, E</emphasis>
). Granulation is shown in detail on each segment in a small section but is meant to cover all the structure.
</paragraph>
</caption>
<paragraph id="8E21AE0A478DE45BB502CAEFC0573640" pageId="0" pageNumber="301">
<emphasis id="09BB3DC6221148051F468F3CBF221400" bold="true" italics="true" pageId="0" pageNumber="301">Chelicera</emphasis>
(Fig. 
<figureCitation id="52756D4E7B26D673A75ABBF408197666" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Parawithius bromelicola sp. Nov. (male holotype, ICN-Aps- 082) A carapace B chelicera C rallum D right chela E left leg I F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, D-F, H); 0.05 mm (B); 0.0125 mm (C, I)." figureDoi="10.3897/zookeys.1184.106698.figure7" httpUri="https://binary.pensoft.net/fig/940876" pageId="0" pageNumber="301">7B</figureCitation>
): with five setae on hand, 
<emphasis id="5B29A2F60C2C91EB505EA15D33A44965" italics="true" pageId="0" pageNumber="301">sb</emphasis>
and 
<emphasis id="AB9DFF417F2C69CB779C110CE33C4716" italics="true" pageId="0" pageNumber="301">b</emphasis>
denticulate, all others acuminate; movable finger with one subdistal seta; galea of male with three or four very small terminal rami; rallum of four blades, the most distal blade with several serrations on leading edge, other blades smooth (Fig. 
<figureCitation id="25BDAF16C3269B6999EFC4CC2526C66D" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Parawithius bromelicola sp. Nov. (male holotype, ICN-Aps- 082) A carapace B chelicera C rallum D right chela E left leg I F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, D-F, H); 0.05 mm (B); 0.0125 mm (C, I)." figureDoi="10.3897/zookeys.1184.106698.figure7" httpUri="https://binary.pensoft.net/fig/940876" pageId="0" pageNumber="301">7C</figureCitation>
); two dorsal lyrifissures.
</paragraph>
<paragraph id="228479A283B62FCD2183E1D03A5FA7FF" pageId="0" pageNumber="301">
<emphasis id="86291258657D42A1A9A7AC67D538591A" bold="true" italics="true" pageId="0" pageNumber="301">Pedipalp</emphasis>
(Fig. 
<figureCitation id="27BF95B421A76B398A640F0B372F9126" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Parawithius bromelicola sp. Nov. (male holotype, ICN-Aps- 082) A carapace B chelicera C rallum D right chela E left leg I F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, D-F, H); 0.05 mm (B); 0.0125 mm (C, I)." figureDoi="10.3897/zookeys.1184.106698.figure7" httpUri="https://binary.pensoft.net/fig/940876" pageId="0" pageNumber="301">7D, G, H</figureCitation>
): trochanter, femur, patella and chelal hand granulate, chelal fingers smooth; dorsal setae clavate and denticulate; trochanter 1.81 
<normalizedToken id="57699A94524B8BF069D1E4F0BA7007B5" originalValue="×">x</normalizedToken>
(1.59-1.74) (♂), 1.76 
<normalizedToken id="57BB36B690E0805C5C35A48FE5129685" originalValue="×">x</normalizedToken>
(1.73) (♀), femur 4.33 
<normalizedToken id="ED2C1E74BAD37E0251F6EBA46A1408F8" originalValue="×">x</normalizedToken>
(4.17-4.38) (♂), 3.77 
<normalizedToken id="4469ED0332039762FC19F428AD0D4C11" originalValue="×">x</normalizedToken>
(4.14) (♀), patella 3.44 
<normalizedToken id="02B54B7075A64FA8C86F4A4B8B1CFABC" originalValue="×">x</normalizedToken>
(3.36-3.42) (♂), 3.17 
<normalizedToken id="4AAAFE5BB76659A6CBBC9EBD3201313A" originalValue="×">x</normalizedToken>
(3.50) (♀), chela (with pedicel) 3.99 
<normalizedToken id="CAE690B11DD303A3359F3486E80B4A84" originalValue="×">x</normalizedToken>
(3.78-3.97) (♂), 3.52 
<normalizedToken id="69DFCCC561205F1BB17979740ECF3D50" originalValue="×">x</normalizedToken>
(3.85) (♀), chela (without pedicel) 3.85 
<normalizedToken id="A4242A8A5B5F4AAB7DDC61881B2099AE" originalValue="×">x</normalizedToken>
(3.58-3.78) (♂), 3.29 
<normalizedToken id="3CBFB6F059A543BEF2350EE674B69B57" originalValue="×">x</normalizedToken>
(3.65) (♀), hand 2.0 
<normalizedToken id="7194BD9177D64CC1A0C8294330716022" originalValue="×">x</normalizedToken>
(1.78-1.82) (♂), 1.68 
<normalizedToken id="7FDEAB3314FA43E6412908CB6035154D" originalValue="×">x</normalizedToken>
(1.92) (♀)longer than broad, movable finger 0.95 
<normalizedToken id="E72E34D37A60BDC215C881E843C7D973" originalValue="×">x</normalizedToken>
(1.00-1.06) (♂), 1.01 
<normalizedToken id="8391CC3123EDD6508D48B089FE26246D" originalValue="×">x</normalizedToken>
(0.86) (♀)longer than hand. Fixed chelal finger with eight trichobothria, movable chelal finger with four trichobothria (Fig. 
<figureCitation id="44EC590F232B7A8951C3ACBF286A0E77" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Parawithius bromelicola sp. Nov. (male holotype, ICN-Aps- 082) A carapace B chelicera C rallum D right chela E left leg I F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, D-F, H); 0.05 mm (B); 0.0125 mm (C, I)." figureDoi="10.3897/zookeys.1184.106698.figure7" httpUri="https://binary.pensoft.net/fig/940876" pageId="0" pageNumber="301">7D, H</figureCitation>
): 
<emphasis id="2DB9641E32ECF5D543181196DC88A15F" italics="true" pageId="0" pageNumber="301">eb</emphasis>
and 
<emphasis id="9718892278657028C8A695A6B551ADF1" italics="true" pageId="0" pageNumber="301">esb</emphasis>
situated basally, 
<emphasis id="E03001CBA2FAE5DB3E6A5ABA8C89BD39" italics="true" pageId="0" pageNumber="301">est</emphasis>
midway between 
<emphasis id="9E93F17BD034A3027DBA3E9A47B83DE8" italics="true" pageId="0" pageNumber="301">eb</emphasis>
and the fingertip, 
<emphasis id="CE02E08CE50BBC671171012349AD4AD9" italics="true" pageId="0" pageNumber="301">et</emphasis>
situated distally; 
<emphasis id="555A5E7D89AEBA89FC7971B9365A45A2" italics="true" pageId="0" pageNumber="301">ib</emphasis>
and 
<emphasis id="648EFE3CB37DBD78B29E90F5BDC7EE9A" italics="true" pageId="0" pageNumber="301">ist</emphasis>
situated basally, 
<emphasis id="09D54D24AD23AF8B220694A92B3EA6F5" italics="true" pageId="0" pageNumber="301">it</emphasis>
directly behind 
<emphasis id="3E623A40D902144B71BEA59745F8B4F9" italics="true" pageId="0" pageNumber="301">est</emphasis>
and close distal to 
<emphasis id="9554A0F1FE68A0076822FF69A8E5B54F" italics="true" pageId="0" pageNumber="301">isb</emphasis>
; 
<emphasis id="A9BE4F1DCE3E0BF04229D4D7175BE003" italics="true" pageId="0" pageNumber="301">b</emphasis>
and 
<emphasis id="62E3EC66701D89BB22CCFB007CC06ABB" italics="true" pageId="0" pageNumber="301">sb</emphasis>
situated near one another; 
<emphasis id="280B9D245500D1112814E86335811D2A" italics="true" pageId="0" pageNumber="301">st</emphasis>
closer to 
<emphasis id="2796191C41040064A18604FFF7EA1FD4" italics="true" pageId="0" pageNumber="301">sb</emphasis>
than to 
<emphasis id="E72A332C79EB71E7DA00A28F33A9289A" italics="true" pageId="0" pageNumber="301">t</emphasis>
; 
<emphasis id="6D247C6B4C4D34B4653534D4071F6BB2" italics="true" pageId="0" pageNumber="301">t</emphasis>
parallel to 
<emphasis id="75A9C26AA5B01170E7588B79F9BD22F9" italics="true" pageId="0" pageNumber="301">est</emphasis>
. Venom apparatus not visible, venom ducts not visible in ♂ and ♀. Retrolateral margin of fixed finger with three sense-spots (Fig. 
<figureCitation id="4FF8A5A655F4E0F6DBB2DFF5830610C3" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Parawithius bromelicola sp. Nov. (male holotype, ICN-Aps- 082) A carapace B chelicera C rallum D right chela E left leg I F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, D-F, H); 0.05 mm (B); 0.0125 mm (C, I)." figureDoi="10.3897/zookeys.1184.106698.figure7" httpUri="https://binary.pensoft.net/fig/940876" pageId="0" pageNumber="301">7H</figureCitation>
) situated distal to 
<emphasis id="32E56A50A6C782CBF528837906181E9E" italics="true" pageId="0" pageNumber="301">esb</emphasis>
and linear in disposition. Chelal teeth very small and almost not developed or differentiated for a clear count; accessory teeth absent.
</paragraph>
<paragraph id="119972EC5C16F07051F2D9D951AC1BA0" pageId="0" pageNumber="301">
<emphasis id="3C00898ADCA9C1D2E20017E8E50AA93E" bold="true" italics="true" pageId="0" pageNumber="301">Coxal region</emphasis>
: coxal chaetotaxy: ♂, 10: 10: 5: 7; maxilla with 23 setae including three apical setae (one tactile setae) and one very small internal, sub-oral seta; median maxillary lyrifissure medial in position, posterior lyrifissure present.
</paragraph>
<paragraph id="D6CC010DED98FDE1E1F6D73CD724570C" pageId="0" pageNumber="301">
<emphasis id="763D0EE60C44CA07C0BB68994C7A867A" bold="true" italics="true" pageId="0" pageNumber="301">Legs</emphasis>
(Fig. 
<figureCitation id="6D317F316BB5DB9EC360E3F2DB32E72A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Parawithius bromelicola sp. Nov. (male holotype, ICN-Aps- 082) A carapace B chelicera C rallum D right chela E left leg I F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, D-F, H); 0.05 mm (B); 0.0125 mm (C, I)." figureDoi="10.3897/zookeys.1184.106698.figure7" httpUri="https://binary.pensoft.net/fig/940876" pageId="0" pageNumber="301">7E, F</figureCitation>
): junction between femora and patellae I and II parallel as well as junction in legs III and IV; tarsal tactile seta of leg IV situated distally, 0.73 
<normalizedToken id="49B6D01FB45C722676A09AB69342C178" originalValue="×">x</normalizedToken>
(♂) of tarsus length (Fig. 
<figureCitation id="DEAB61022F9E544CD05231E3B172424F" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Parawithius bromelicola sp. Nov. (male holotype, ICN-Aps- 082) A carapace B chelicera C rallum D right chela E left leg I F left leg IV G left pedipalp H right chelal fingers (arrows show sense-spots) I rallum. Abbreviations: Pedipalp movable finger: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; fixed finger: eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal. Cheliceral setae: b = basal seta; es = exterior seta; is = interior seta; ls = laminal seta. Scale bars: 0.5 mm (G); 0.1 mm (A, D-F, H); 0.05 mm (B); 0.0125 mm (C, I)." figureDoi="10.3897/zookeys.1184.106698.figure7" httpUri="https://binary.pensoft.net/fig/940876" pageId="0" pageNumber="301">7F</figureCitation>
); two subterminal tarsal setae distal form TS arcuate and acute; arolium shorter than claws. Ratios: leg I: femur 1 
<normalizedToken id="2A3FB403844F18DC3D51E390DA23CB59" originalValue="×">x</normalizedToken>
, patella 2.28 
<normalizedToken id="639E8F56E779E8FB49301123BFC0A557" originalValue="×">x</normalizedToken>
, tibia 3.67 
<normalizedToken id="541422FEB73624FACB7E1888AF349F46" originalValue="×">x</normalizedToken>
, tarsus 5.71 
<normalizedToken id="AFCA67E16E60800BF95EFD573407CE04" originalValue="×">x</normalizedToken>
deeper than broad; leg IV: femur + patella 3.46 
<normalizedToken id="D910270A263EA0D1F1DC5259F3B5A510" originalValue="×">x</normalizedToken>
, tibia 4.27 
<normalizedToken id="FB34F13110CF473B541752FBEBB20933" originalValue="×">x</normalizedToken>
, tarsus 5.11 
<normalizedToken id="72860890A0D22216D8421E01798AE74C" originalValue="×">x</normalizedToken>
deeper than broad.
</paragraph>
<paragraph id="6E693E5A08401900CCD68DD18DB01A58" pageId="0" pageNumber="301">
<emphasis id="1B347358FA1712EC74A65F363B62E50A" bold="true" italics="true" pageId="0" pageNumber="301">Abdomen</emphasis>
: all tergites divided with a medial suture, with a row of spots in the middle of each hemitergite; sternites entire. Tergal chaetotaxy: ♂, 8: 9: 9: 10: 13: 13: 16: 16: 18: 12: 10 (including two tactile setae): 2; all setae foliate. Sternal chaetotaxy: ♂, 15: (2) 11 (2): (2) 13 (2): 16: 14: 16 + 21 
<emphasis id="5F5B26641780FC1C28F9CEF3BFE065E6" italics="true" pageId="0" pageNumber="301">gls</emphasis>
: 13 + 86 
<emphasis id="C435E8FF273F6990129720C99FDAAC55" italics="true" pageId="0" pageNumber="301">gls</emphasis>
: 11 + 2 
<emphasis id="FBCCBB0B5C58DDC5842F192ABDEDB138" italics="true" pageId="0" pageNumber="301">gls</emphasis>
: 10 (including two tactile setae): 8 (including two tactile setae and some dentate): 2; sternites VII and VIII of ♂ with patches of glandular setae (Fig. 
<figureCitation id="7BEAB5B33A8402FC49DB1E54A5972FED" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Glandular setae patches of the new species A Paciwithius chimbilacus sp. nov. (holotype, ICN-Aps- 076) B Parawithius bromelicola sp. nov. (holotype, ICN-Aps- 082) C Paciwithius valduparensis sp. nov. (holotype, ICN-Aps- 597) D Cystowithius florezi sp. nov. (holotype, ICN-Aps- 07) E, F Oligowithius achagua sp. nov. E male paratype (ICN-Aps- 388) F male holotype (ICN-Aps- 388). Scale bars: 0.1 mm (A-C, E, F); 0.05 mm (D)." figureDoi="10.3897/zookeys.1184.106698.figure9" httpUri="https://binary.pensoft.net/fig/940878" pageId="0" pageNumber="301">9B</figureCitation>
); setae uniseriate and acuminate; ♂ without paired invaginations on anterior margins of sternites.
</paragraph>
<caption id="DADB2003DA7753F1D5DE54AFE13E3A80" doi="10.3897/zookeys.1184.106698.figure9" httpUri="https://binary.pensoft.net/fig/940878" pageId="0" pageNumber="301" start="Figure 9" startId="F9">
<paragraph id="F6E43A71FB411053ADBF998C1C98B30F" pageId="0" pageNumber="301">
<emphasis id="04D5C394805E4CF21EB46AD354AE7883" bold="true" pageId="0" pageNumber="301">Figure 9.</emphasis>
Glandular setae patches of the new species 
<emphasis id="213A5414C02D5CA9A47547840DBA5C2A" bold="true" pageId="0" pageNumber="301">A</emphasis>
<taxonomicName id="2A17B2C29C50C0A0919F9EF0742BEAC5" authorityName="Romero-Ortiz &amp; Sarmiento &amp; Harvey" authorityYear="2023" family="Withiidae" genus="Paciwithius" kingdom="Animalia" lsidName="Paciwithius chimbilacus" order="Pseudoscorpiones" pageId="0" pageNumber="301" rank="species" species="chimbilacus">
<emphasis id="05A4B2F59F33BDF1E2D1497B911F7A72" italics="true" pageId="0" pageNumber="301">Paciwithius chimbilacus</emphasis>
</taxonomicName>
sp. nov. (holotype, ICN-Aps-076) 
<emphasis id="206BF6B759E4FDFF40F7155CC76D7CCA" bold="true" pageId="0" pageNumber="301">B</emphasis>
<taxonomicName id="6A2DCDF1D5D9FE043057C783685B92E6" authorityName="Romero-Ortiz &amp; Sarmiento &amp; Harvey" authorityYear="2023" class="Arachnida" family="Withiidae" genus="Parawithius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parawithius bromelicola" order="Pseudoscorpiones" pageId="0" pageNumber="301" phylum="Arthropoda" rank="species" species="bromelicola">
<emphasis id="128AA880E09519B17060D293C04CC6D3" italics="true" pageId="0" pageNumber="301">Parawithius bromelicola</emphasis>
</taxonomicName>
sp. nov. (holotype, ICN-Aps-082) 
<emphasis id="8B1C1F2305A7E9102DBFF09304F9B509" bold="true" pageId="0" pageNumber="301">C</emphasis>
<taxonomicName id="7EE1C6BB5B068F0AF4B504FF2EB2F85E" authorityName="Romero-Ortiz &amp; Sarmiento &amp; Harvey" authorityYear="2023" family="Withiidae" genus="Paciwithius" kingdom="Animalia" lsidName="Paciwithius valduparensis" order="Pseudoscorpiones" pageId="0" pageNumber="301" rank="species" species="valduparensis">
<emphasis id="C65383F1AA4BA32027CF288FB0FB0B97" italics="true" pageId="0" pageNumber="301">Paciwithius valduparensis</emphasis>
</taxonomicName>
sp. nov. (holotype, ICN-Aps-597) 
<emphasis id="32DE60107264FB12537A258DEB5EFFEE" bold="true" pageId="0" pageNumber="301">D</emphasis>
<taxonomicName id="34B7D28594A08A637A2798AF219F87E9" authorityName="Romero-Ortiz &amp; Sarmiento &amp; Harvey" authorityYear="2023" class="Arachnida" family="Withiidae" genus="Cystowithius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cystowithius florezi" order="Pseudoscorpiones" pageId="0" pageNumber="301" phylum="Arthropoda" rank="species" species="florezi">
<emphasis id="FBCF70AF5410BDADBDB5077BC0425117" italics="true" pageId="0" pageNumber="301">Cystowithius florezi</emphasis>
</taxonomicName>
sp. nov. (holotype, ICN-Aps-07) 
<emphasis id="EB0323D20187A0AB95F2FBDB017640D6" bold="true" pageId="0" pageNumber="301">E, F</emphasis>
<taxonomicName id="BDDE582F0D5946576D8D672E9FB7F1DD" authorityName="Romero-Ortiz &amp; Sarmiento &amp; Harvey" authorityYear="2023" class="Arachnida" family="Withiidae" genus="Oligowithius" kingdom="Animalia" lsidName="Oligowithius achagua" order="Pseudoscorpiones" pageId="0" pageNumber="301" rank="species" species="achagua">
<emphasis id="7B88B0029AF3067F2787CFC3F8D25E1D" italics="true" pageId="0" pageNumber="301">Oligowithius achagua</emphasis>
</taxonomicName>
sp. nov. 
<emphasis id="F9F1F47D6AA0640A83FCB93EA59AF949" bold="true" pageId="0" pageNumber="301">E</emphasis>
male paratype (ICN-Aps-388) 
<emphasis id="05FB4D82EAF44B9A1BEAE3BED2E1C62E" bold="true" pageId="0" pageNumber="301">F</emphasis>
male holotype (ICN-Aps-388). Scale bars: 0.1 mm (
<emphasis id="9F17F7E089378CC3BA4EC59FA1518FC4" bold="true" pageId="0" pageNumber="301">A-C, E, F</emphasis>
); 0.05 mm (
<emphasis id="D0F1D184972F0B64703814C3430D07BC" bold="true" pageId="0" pageNumber="301">D</emphasis>
).
</paragraph>
</caption>
<paragraph id="544902CDF28B7F0DDB56B89FD305708D" pageId="0" pageNumber="301">
<emphasis id="8954BE2C7FDEC8DD5DF8FB2285BED0D0" bold="true" italics="true" pageId="0" pageNumber="301">Genitalia</emphasis>
: see 
<bibRefCitation id="4A9977327BA581580B6C58E5269E53A6" DOI="https://doi.org/10.1636/JoA-S-19-068" author="Romero-Ortiz, C" journalOrPublisher="The Journal of Arachnology" pageId="0" pageNumber="301" pagination="108 - 121" refId="B23" refString="Romero-Ortiz, C, Sarmiento, CE, 2021. A comparative study of the male genitalia of the Cacodemoniini (Pseudoscorpiones: Withiidae). The Journal of Arachnology 49 (1): 108 - 121, DOI: https://doi.org/10.1636/JoA-S-19-068" title="A comparative study of the male genitalia of the Cacodemoniini (Pseudoscorpiones: Withiidae)." url="https://doi.org/10.1636/JoA-S-19-068" volume="49" year="2021">Romero-Ortiz and Sarmiento 2021</bibRefCitation>
as &quot; 
<taxonomicName id="7585AD21AB15DB4BCD7D716EBE75E773" authorityName="Chamberlin" authorityYear="1931" class="Arachnida" family="Withiidae" genus="Parawithius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Parawithius" order="Pseudoscorpiones" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis id="BE4A1A214B7681E1C9D3DAE6C00CEBC3" italics="true" pageId="0" pageNumber="301">Parawithius</emphasis>
</taxonomicName>
msp1&quot;.
</paragraph>
</subSubSection>
<subSubSection id="E32495C01B7580AAB7368AD690D68DF1" pageId="0" pageNumber="301" type="etymology">
<paragraph id="3530A27BE3F40735B65930ADF8AC9C27" pageId="0" pageNumber="301">Etymology.</paragraph>
<paragraph id="08235711EA8BFFF13BE3B71B371836DE" pageId="0" pageNumber="301">This species is named after bromeliad plants, due to its close association with them. Most of the specimens were collected by sifting bromeliads on white sheets.</paragraph>
</subSubSection>
</treatment>
</document>