217 lines
21 KiB
XML
217 lines
21 KiB
XML
<document id="3556721F93360A3F5D1EA9128AB623DB" ENCODING="UTF-8" ID-GBIF-Dataset="b8f58d00-19e5-4cfa-a1f0-4c686286fc5e" ModsDocID="z01603p001" checkinTime="1247319475874" checkinUser="thomas" docAuthor="Juan J. Schmitter-Soto" docDate="2007" docId="57B1B68F1607DD82237A58874865713C" docLanguage="en" docName="2007_Schmitter-Soto_gg1.xml" docOrigin="Zootaxa 1603" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:AFFCB590-1FC7-4CD0-950C-D1D1A6E59F6C" docTitle="Cryptoheros cutteri Schmitter-Soto, 2007, n. comb." docType="treatment" docVersion="12" lastPageNumber="40" masterDocId="9F5EADD85808D5BD56FDE2FB757992C0" masterDocTitle="A systematic revision of the genus Archocentrus (Perciformes: Cichlidae), with the description of two new genera and six new species." masterLastPageNumber="78" masterPageNumber="1" pageNumber="39" updateTime="1696279408464" updateUser="admin">
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<mods:title id="BD285EDBCF25AFF9E578AF4BC2C04A7D">A systematic revision of the genus Archocentrus (Perciformes: Cichlidae), with the description of two new genera and six new species.</mods:title>
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<mods:namePart id="E1B076265AC4D6CD66515CE9057E431F">Juan J. Schmitter-Soto</mods:namePart>
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<mods:title id="7F20379ABBF5381703C49AAB54AA2B61">Zootaxa</mods:title>
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<mods:part id="97EB9B86A8B4C91A0C8B38030A5413ED">
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<mods:date id="EEF31BCAB8C3061C8383009956BE564C">2007</mods:date>
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<mods:classification id="05D843979E44D724E2C6BE1C015D043E">journal article</mods:classification>
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<treatment id="57B1B68F1607DD82237A58874865713C" ID-DOI="http://doi.org/10.5281/zenodo.6248768" ID-GBIF-Taxon="100117459" ID-Zenodo-Dep="6248768" LSID="urn:lsid:plazi:treatment:57B1B68F1607DD82237A58874865713C" httpUri="http://treatment.plazi.org/id/57B1B68F1607DD82237A58874865713C" lastPageNumber="40" pageNumber="39">
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<subSubSection id="66EA4C9A825762684DDDEA80A806CBB4" pageNumber="3" type="nomenclature">
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<paragraph id="FA89CB97EF7803009588A9A860023100" pageNumber="39">
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<taxonomicName id="5AF604FC158F95B2C0F8100F9EA543A1" ID-CoL="ZXMF" ID-ENA="762638" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="cutteri" status="n. comb.">
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Cryptoheros cutteri (
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<bibRefCitation id="F8AAAC79F36EFDED17DC592C65F14DD7" author="Fowler, H. W." pageNumber="3" pagination="379 - 385" refId="ref35237" refString="Fowler, H. W. (1932) Notes on the freshwater fishes from Central America. Proceedings of the Academy of Natural Sciences of Philadelphia, 64, 379 - 385." type="journal article" year="1932">Fowler, 1932</bibRefCitation>
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)
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</taxonomicName>
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,
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<taxonomicNameLabel id="532E1091FF54A13005DBFE773E7AEC64" pageNumber="3" rank="species">n. comb.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="92CEAB4BE60652E4E95625068A1EED35" pageNumber="3" type="description">
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<paragraph id="D9C9590F3E5B115DEC70354ADB84FE77" pageNumber="39">Figures 5, 11</paragraph>
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</subSubSection>
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<subSubSection id="C2D4A6371B61D7DD79C0DA8FDBB443FD" pageNumber="3" type="reference_group">
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<paragraph id="418525F44C52C7F6C93FB58867F82691" pageNumber="39">
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<taxonomicName id="292C0A88DA5D9E3BAF5ED84F6183DF0D" LSID-ZBK="urn:lsid:zoobank.org:act:AA1787B8-6C57-46B3-B745-27ED2F0BC0D8" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" lsidName-ZBK="Cichlasoma cutteri Fowler 1932:380" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="cutteri">
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Cichlasoma cutteri
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<bibRefCitation id="EBBEEC1643B821C35FC405CE92528ADF" author="Fowler, H. W." pageNumber="3" pagination="379 - 385" refId="ref35237" refString="Fowler, H. W. (1932) Notes on the freshwater fishes from Central America. Proceedings of the Academy of Natural Sciences of Philadelphia, 64, 379 - 385." type="journal article" year="1932">Fowler 1932: 380</bibRefCitation>
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</taxonomicName>
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(original description).
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</paragraph>
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<paragraph id="51557449CB67F7088C437FBBEC824010" pageNumber="39">
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<taxonomicName id="2D0AD47F4EB89058D0410700E3AD8B3B" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="cutteri">Archocentrus cutteri</taxonomicName>
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,
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<bibRefCitation id="AECF30920DE7DF6A362693AA3C366D4A" author="Allgayer, R." pageNumber="3" pagination="6 - 24" refId="ref34494" refString="Allgayer, R. (1994) Description d’une espece nouvelle du genre Archocentrus. Revue Francaise des Cichlidophiles, 135, 6 - 24." type="journal article" year="1994">Allgayer 1994: 15</bibRefCitation>
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(
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<taxonomicNameLabel id="E9C00119B8A1F5B8CA8552B185435F10" pageNumber="3" rank="species">new combination</taxonomicNameLabel>
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).
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</paragraph>
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<paragraph id="6AFCD62CF21A4343B6EF300A9E800A58" pageNumber="39">
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<taxonomicName id="B7AB824F4148ECBC40286593BA81D225" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="spilurus">Cryptoheros spilurus</taxonomicName>
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(part. et non
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<normalizedToken id="AE30DAC89003B5B8EAE7721B19A4F769" originalValue="Günther" pageNumber="3">Guenther</normalizedToken>
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),
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<bibRefCitation id="8D9BE3B3B4AF7695CE2E03FBAA9F818B" author="Allgayer, R." pageNumber="3" pagination="13 - 20" refId="ref34520" refString="Allgayer, R. (2001) Description d'un genre nouveau, Cryptoheros, d'Amerique Centrale et d'une espece nouvelle du Panama (Pisces: Cichlidae). L’An Cichlide, 1, 13 - 20." type="journal article" year="2001">Allgayer 2001: 14</bibRefCitation>
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(misidentification).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="1422548A954C47D71F6E36FE76508916" pageNumber="3" type="nomenclature">
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||
<paragraph id="56C27F0E86B3141DC6330AC5DC414FE3" pageNumber="39">
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<materialsCitation id="697611A1BB88065CD56E68CD3E67B017" ID-GBIF-Occurrence="923904206" collectingDate="1930-08-24" collectionCode="ANSP" collectorName="C. B. Worth" country="Honduras" location="Rio Lancetilla" pageNumber="3" specimenCode="ANSP 53930" stateProvince="Atlantida Dept." typeStatus="holotype">
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<typeStatus id="C49AC7647A570CC338451DC4BA594D89" pageNumber="3" type="holotype">Holotype</typeStatus>
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.
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<specimenCode id="DAF35ABEC36A1E12EE4FC6F58FEA0E9A" pageNumber="3">
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<collectionCode id="F323542AFD94C714C37D44F3BBCDC245" pageNumber="3">ANSP</collectionCode>
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53930
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</specimenCode>
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, 112 mm SL (Fig. 11),
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<collectorName id="E3778D306BB1E0C449D0FDE407B07F3D" pageNumber="3">C. B. Worth</collectorName>
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,
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<date id="2D17EAC67B8436E1249E9627E2BDD10E" day="24" month="08" pageNumber="3" value="1930-08-24" year="1930">
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<collectingDate id="6746429347B08E9092910B69FA5D4968" day="24" month="08" pageNumber="3" value="1930-08-24" year="1930">Aug. 24, 1930</collectingDate>
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</date>
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.
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<location id="5EB87952269E849633C05B811EBC278E" LSID="urn:lsid:plazi:treatment:57B1B68F1607DD82237A58874865713C:5EB87952269E849633C05B811EBC278E" country="Honduras" name="Rio Lancetilla" pageNumber="3" stateProvince="Atlantida Dept.">
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<normalizedToken id="91231DF936B56C52ACC7F11D173A6095" originalValue="Río" pageNumber="3">Rio</normalizedToken>
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Lancetilla
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</location>
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,
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<collectingRegion id="C3A4FCAE51319F1BC61049723E15F17A" pageNumber="3">
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<normalizedToken id="AD043E9AC511017D27EAF5167E2F21E6" originalValue="Atlántida" pageNumber="3">Atlantida</normalizedToken>
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Dept.
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</collectingRegion>
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,
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<collectingCountry id="88800E8D0F8524252BAE6B04128894F3" pageNumber="3">Honduras</collectingCountry>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="B365E63846083473A7D8E2C61FEAD2FC" pageNumber="39">
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<materialsCitation id="EB531A1AFEA2E2E9437E94F5EF828881" ID-GBIF-Occurrence="923904204" collectionCode="ANSP" country="Honduras" location="Rio Lancetilla" pageNumber="3" specimenCode="ANSP 53931-53933" stateProvince="Atlantida Dept." typeStatus="paratype">
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<typeStatus id="44944F5820A4CF869343F42043C5EE94" pageNumber="3" type="paratype">Paratypes</typeStatus>
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.
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<specimenCode id="4A59B45AAC1B904A0BD23B7896769CB1" pageNumber="3">
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<collectionCode id="2F7159420EE37BEB4E3E1A15524CE863" pageNumber="3">ANSP</collectionCode>
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53931-53933
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</specimenCode>
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(3, 38-108 mm SL),
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<typeStatus id="418F2B97C65972209C8F6AEC5E935820" pageNumber="3" type="paratopotype">paratopotypes</typeStatus>
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.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="592823264BB2AFFE01878D9303D68E03" pageNumber="3" type="multiple">
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||
<caption id="A1C7363D059AE57B33C2DD329618DF74" pageNumber="3">
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||
<paragraph id="7BAA1B5AC648BF569E93E8EC04A9D6AB" pageNumber="39">
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FIGURE 11.
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<taxonomicName id="4DF93AC1FE75A9A85F02C8F790EC6D7F" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="cutteri">Cryptoheros cutteri</taxonomicName>
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, holotype, ANSP 53930. Photo, K. Luckenbill.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="F15CE8407D7EFDDA07E5D5621CB74F03" pageNumber="3" type="diagnosis">
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<paragraph id="8B7FF9FF86CE7195E9CD7A6176355B19" pageNumber="40">
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<pageBreakToken id="30EEBD848BEDDC60E079E997286ECB84" pageNumber="3">Diagnosis</pageBreakToken>
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. Autapomorphy (Schmitter-Soto, in press): a wine-colored abdomen in life (in adults). Bars on sides of body with alternating intensity, the second one much lighter than first and third. Further distinguished from other species in the subgenus by arms of first epibranchial bone, which are divergent (vs. parallel); supraoccipital crest undulating (vs. straight); ventral angle of articular obtuse (vs. right-angled); absence (vs. presence) of an anteriorly directed pronounced convexity on ventral process of articular; procurrent rays of caudal fin 2 (vs. 3-4).
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</paragraph>
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</subSubSection>
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<subSubSection id="A74680A8BF616CCFECC9624E67879305" pageNumber="3" type="description">
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<paragraph id="5418785A6B87C4C45DD33E09CB76F201" pageNumber="40">
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Description. D. XVII-XIX,9-11; A. VIII-X,7-9 (one specimen of 60 with 11 spines). Gill rakers on lower limb of first arch modally 7. Scale rows on cheek 4-6; scales from lateral line to origin of dorsal fin 4- 6 (contra
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<bibRefCitation id="F1273E55060E7C6B633A116E8D1CCDED" author="Fowler, H. W." pageNumber="3" pagination="379 - 385" refId="ref35237" refString="Fowler, H. W. (1932) Notes on the freshwater fishes from Central America. Proceedings of the Academy of Natural Sciences of Philadelphia, 64, 379 - 385." type="journal article" year="1932">Fowler 1932</bibRefCitation>
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, who counted 7); scales from lateral line to base of first dorsal-fin ray 2.5-3.5; circumpeduncular scales modally 18 (further meristic data appear in Table 3).
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</paragraph>
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<paragraph id="ED852234F262E9C5BE3F7E665E29CDC3" pageNumber="40">Largest examined specimen, 112 mm SL. Body depth (44-54%, usually greater than 49% of SL). Head length 32-43% of SL (further morphometric data appear in Table 4). Head profile convex or straight, concave above orbits. Teeth embedded; pointed or bluntish, slightly labiolingually compressed and retrorse, bicuspid. Upper symphysial teeth subequal to adjacent teeth, not abruptly larger; lower symphysial teeth may be slightly shorter than adjacent teeth. Upper lip medially narrow; lower lip squarish or rounded at corner, its lower angle acute.</paragraph>
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<paragraph id="C1857DEF9B5E3664FAC4D952C2209752" pageNumber="40">Pectoral and pelvic fins always reaching caudad beyond 2nd anal-fin spine. Filamentous rays of dorsal fin at least to mid-caudal fin. Subsidiary pored scales on caudal fin always present, occasionally forming twoscale rows; scales between dorsal fin rays distally in two rows, up to 11 scales long.</paragraph>
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<paragraph id="A8EBBB9E9402B9BF76EA8097767FCED5" pageNumber="40">In specimens from ca. 35 mm SL and larger, gut simple with two intermediate loops; medial and anal loops twisted and directed dorsad. Peritoneum pigmented only dorsally (to half sides), mainly anteriorly; with isolated blotches posteriorly. Genital papilla rounded, as long as broad, or oval, longer than broad or broader than long; in some, the tip crenulated or notched; sunken; no pigment, except on margins, basal half, and posterior(caudal) side.</paragraph>
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<paragraph id="FB1B3BC5C899B21204F321BB7D8F7090" pageNumber="40">
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No vertical bar on head; no interorbital bands, but postorbital-nape region somewhat dark; a stripe from snout to eye, sometimes diffuse; suborbital streak straight, sharp-ended; no opercular spots, but opercle dark, except on postorbital region. Eyes blue-greenish. No longitudinal stripe. Bars on sides of body, medially much more intense; 1st bar I-shaped, somewhat inclined on head; intensity alternating: 1st, 3rd, 5th, 7th bars, especially 3rd, much more intense than 2nd, 4th, 6th; 2nd, 4th bars may be dorsally evanescent; 3rd bar and sometimes also 5th and 7th may extend slightly onto base of dorsal fin. An ocellus may be present on spinous dorsal fin of mature females. Soft dorsal and anal fins usually immaculate; sometimes light dots discernible at bases. About 13 rows of light spots on sides, smaller than scales; breast region yellowish-greenish or olive (or “wood brown” according to
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<bibRefCitation id="F402565C1E6583BB403A768F1E167CAF" author="Fowler, H. W." pageNumber="3" pagination="379 - 385" refId="ref35237" refString="Fowler, H. W. (1932) Notes on the freshwater fishes from Central America. Proceedings of the Academy of Natural Sciences of Philadelphia, 64, 379 - 385." type="journal article" year="1932">Fowler [1932]</bibRefCitation>
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). Axil of pectoral fin somewhat dark (part of 1st bar); base of pectoral fin yellowish or white. Caudal blotch two-thirds or more on peduncle, across lateral line, usually forming a saddle.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="176BFE4279D40F5CF601DB2FEAB6D876" pageNumber="3" type="distribution">
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<paragraph id="5948DD0AB4552B18B76923DA1CE2E94C" pageNumber="40">
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Distribution. Atlantic Honduras (
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<normalizedToken id="8C002FB80F3B220CB240E1F3AA6A845D" originalValue="Ríos" pageNumber="3">Rios</normalizedToken>
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||
<normalizedToken id="334BB4F556957D2962192E2D435F9ECB" originalValue="Celán" pageNumber="3">Celan</normalizedToken>
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||
, Lancetilla,
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<normalizedToken id="3D7DDEAC06D31685D7F6803D0F54816B" originalValue="Aguán" pageNumber="3">Aguan</normalizedToken>
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, Jutiapa, and others) north to Guatemala (
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<normalizedToken id="F47BDE1DBE2767611E6E41AF4B9B003E" originalValue="Ríos" pageNumber="3">Rios</normalizedToken>
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Achuelo, Matasano, and others) (Fig. 5).
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</paragraph>
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</subSubSection>
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<subSubSection id="42520CB908DAAC46340804132EF1B671" pageNumber="3" type="discussion">
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<paragraph id="9E958ED72D948C3B90CCD77757A693FF" pageNumber="40">
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Remarks. Synonymized with
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<taxonomicName id="395C4D41DE9C30086C027E41492F49E0" family="Cichlidae" genus="Cichlasoma" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="spilurum">Cichlasoma spilurum</taxonomicName>
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by
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<bibRefCitation id="46E6007B964EB3FF9D3894A1494BEFFD" author="Miller, R. R." pageNumber="3" pagination="773 - 802" refId="ref36564" refString="Miller, R. R. (1966) Geographical distribution of Central American freshwater fishes. Copeia 1966, 773 - 802." type="journal article" year="1966">Miller (1966)</bibRefCitation>
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||
with no explicit justification. This move was followed by
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<bibRefCitation id="64D3B338F423BA17E238C5E52C811171" author="Allgayer, R." pageNumber="3" pagination="13 - 20" refId="ref34520" refString="Allgayer, R. (2001) Description d'un genre nouveau, Cryptoheros, d'Amerique Centrale et d'une espece nouvelle du Panama (Pisces: Cichlidae). L’An Cichlide, 1, 13 - 20." type="journal article" year="2001">Allgayer (2001)</bibRefCitation>
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||
, who added that the Honduran so-called “
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<taxonomicName id="3905A28CF505F68F37277B50BE3561C9" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="sp. yojoa">
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||
« Archocentrus » sp. «
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<normalizedToken id="1FA7594A9AD9333D031C3777D4130E89" originalValue="Yojoá" pageNumber="3">Yojoa</normalizedToken>
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||
»
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||
</taxonomicName>
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” also belonged in
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<taxonomicName id="AD2FD813892EA388C480BB5154C901F2" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="spilurus">Cr. spilurus</taxonomicName>
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, but acknowledged that “[plusieurs formes locales, notamment du Honduras[,] sont connues.”
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</paragraph>
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<paragraph id="D33F98573F348889AC46BA5EA465D557" pageNumber="40">
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Fowler’s (1932: 381) figure is not accurate: it depicts the holotype as showing all the bars of approximately the same intensity, which is not the case (Fig. 11). The figure contains other errors, e.g. the size of the scales on the breast, which are much smaller and numerous than shown.
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<bibRefCitation id="17F90DFBC6521526C5012FCB38CB7A04" author="Fowler, H. W." pageNumber="3" pagination="379 - 385" refId="ref35237" refString="Fowler, H. W. (1932) Notes on the freshwater fishes from Central America. Proceedings of the Academy of Natural Sciences of Philadelphia, 64, 379 - 385." type="journal article" year="1932">Fowler (1932)</bibRefCitation>
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thought his new species was “[r]elated to
|
||
<taxonomicName id="8FB84510573991A10979EC7C16DDD1A4" LSID-ZBK="urn:lsid:zoobank.org:act:D76352FE-1F2C-4AFD-9081-678A86D6C797" family="Cichlidae" genus="Heros" kingdom="Animalia" lsidName-ZBK="Heros octofasciatus Regan 1903:417" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciatus">Heros octofasciatus</taxonomicName>
|
||
” and felt that he needed to compare
|
||
<taxonomicName id="C2F91E405E9B053C6934F2B4B35E2404" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="cutteri">Cr. cutteri</taxonomicName>
|
||
with
|
||
<taxonomicName id="56D57E47D5B2D49EE9D84454AA0FCC1B" family="Cichlidae" genus="Rocio" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="octofasciata">R. octofasciata</taxonomicName>
|
||
, not with
|
||
<taxonomicName id="4B399E0E3CAF87CBF665E167014AD0D9" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="spilurus">Cr. spilurus</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7D8CF4FE526B2E83DF16C7773F4688DB" pageNumber="40">
|
||
I was unable to examine specimens of the “
|
||
<normalizedToken id="3FBED1D6EA30C34C042B65AD15370364" originalValue="Río" pageNumber="3">Rio</normalizedToken>
|
||
Papaloteca”
|
||
<taxonomicName id="64F1FA81408060F7AA454057CFF82FE6" family="Cichlidae" genus="Archocentrus" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="genus">Archocentrus</taxonomicName>
|
||
(
|
||
<bibRefCitation id="5B298416800F010225815A82F2E20293" author="Velasco, F. T." pageNumber="3" pagination="206" refId="ref37515" refString="Velasco, F. T. (2001) Cutter’s cichlid, not! A review of the literature concerning a colorful and possibly undescribed Archocentrus from northern Honduras. Buntbarsche Bulletin, 2001, 206." type="journal article" year="2001">Velasco 2001</bibRefCitation>
|
||
) from northern Honduras, but nearby specimens (
|
||
<materialsCitation id="51B46FC4D7E6D6A8E88F5EB1D965EB8D" ID-GBIF-Occurrence="923904213" collectionCode="UMMZ" country="Honduras" location="Rio Jutiapa" pageNumber="3" specimenCode="UMMZ 228665">
|
||
from
|
||
<location id="A9D1534A482C2ADD98A4FE3D18523E56" LSID="urn:lsid:plazi:treatment:57B1B68F1607DD82237A58874865713C:A9D1534A482C2ADD98A4FE3D18523E56" country="Honduras" name="Rio Jutiapa" pageNumber="3">
|
||
<normalizedToken id="226FD78FDE684FEDBCBE9B2199A01A34" originalValue="Río" pageNumber="3">Rio</normalizedToken>
|
||
Jutiapa
|
||
</location>
|
||
,
|
||
<specimenCode id="46B68E12072052DAC6552D7731C7D3D4" pageNumber="3">
|
||
<collectionCode id="AC4EC4A84B3CEA59D973858AB0EE7E59" pageNumber="3">UMMZ</collectionCode>
|
||
228665
|
||
</specimenCode>
|
||
</materialsCitation>
|
||
) are identifiable as
|
||
<taxonomicName id="3CE05184B0DDCF4DC4C2D844BDE1AD0E" family="Cichlidae" genus="Cryptoheros" kingdom="Animalia" order="Perciformes" pageNumber="3" phylum="Chordata" rank="species" species="cutteri">Cr. cutteri</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |