treatments-xml/data/25/AC/64/25AC64DF00CF5404A087DD1248A654FD.xml

<document id="FA060D142907F9EEDACD257C3A8E96D2" ID-CLB-Dataset="307585" ID-DOI="10.3897/mycokeys.112.141766" ID-GBIF-Dataset="e1093352-070e-4a36-9a4e-7b101f33fdcf" ID-publisher-id="141766" URI-arpha="74A2726B-45CB-5F13-B2FF-E919480EB400" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" article-type="research-article" checkinTime="1737037778530" checkinUser="pensoft" docAuthor="Li, Qirui, Haqnawaz, Muhammad, Niazi, Abdul Rehman &amp; Khalid, Abdul Nasir" docDate="2025" docId="25AC64DF00CF5404A087DD1248A654FD" docLanguage="en" docName="MycoKeys 112: 165-182" docOrigin="MycoKeys 112" docSource="https://mycokeys.pensoft.net/article/141766/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.mycokeys" docStyleName="PensoftTaxPub.0000.journal_article.mycokeys" docTitle="Candolleomyces virgatus Haqnawaz, Niazi &amp; Khalid 2025, sp. nov." docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="182" masterDocId="74A2726B45CB5F13B2FFE919480EB400" masterDocTitle="Phylogenetic studies on the genus Candolleomyces (Psathyrellaceae, Basidiomycota) occurring in the bed of the Indus River, Punjab, Pakistan, reveal three new species" masterLastPageNumber="182" masterPageNumber="165" pageNumber="165" updateTime="1737046277866" updateUser="admin">
<mods:mods id="4154128EB28DBBF41A599C80629E9665" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E40BA164A4363F747A06D90DF66D9CB8">
<mods:title id="5AC712F2CA2B8905BD5227C96836242D">Phylogenetic studies on the genus Candolleomyces (Psathyrellaceae, Basidiomycota) occurring in the bed of the Indus River, Punjab, Pakistan, reveal three new species</mods:title>
</mods:titleInfo>
<mods:name id="41642254DC3F71352388164D617FE33D" type="personal">
<mods:role id="37E73FA3A2F46696C787D42637333EF6">
<mods:roleTerm id="2FA405B2FCF545EDC234D5F36B698B8D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="960ECE80F6FD87576C6563396A562EB3">Li, Qirui</mods:namePart>
<mods:nameIdentifier id="8CCAA1207A42C14BF909EC400497982C" type="ORCID">0000-0001-8735-2890</mods:nameIdentifier>
<mods:affiliation id="E30B831DD389039928A088B68B1D5A66">The High Efficacy Application of Natural Medicinal Resources Engineering Centre of Guizhou Province (The Key Laboratory of Optimal Utilization of Natural Medicine Resources), School of Pharmaceutical Sciences, Guizhou Medical University, Guiyang, Guizhou, 550004, China</mods:affiliation>
</mods:name>
<mods:name id="5065AEAE9B86F5AE275FAAF0DB7D69F7" type="personal">
<mods:role id="857F4E3C5725F4C8BF83E70C1D6AB05A">
<mods:roleTerm id="036225067CE4A7ECE9B5A83C6C8FAB44">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4E51FBF66A63A0B66D84D20A2C19C76B">Haqnawaz, Muhammad</mods:namePart>
<mods:nameIdentifier id="FD5AC0D73BB5A967B44D96594BA68298" type="ORCID">0000-0003-4667-242X</mods:nameIdentifier>
<mods:affiliation id="5C1B059F643F51BD29223C72576C2662">Fungal Biology and Systematics Research Laboratory, Institute of Botany, University of the Punjab, Quaid-e-Azam Campus 54590, Lahore, Pakistan</mods:affiliation>
</mods:name>
<mods:name id="D05FF46B5F15257A80823065C7C219EB" type="personal">
<mods:role id="F999DC59CB24B8B30AF97584A6675E6C">
<mods:roleTerm id="8C563279CDA35F36C5EA81CC4A33C788">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="402A415DA395F5B3362B9B799C07E59A">Niazi, Abdul Rehman</mods:namePart>
<mods:nameIdentifier id="C2CCC925E1589C7238340D1F1228657B" type="ORCID">0000-0002-1118-1148</mods:nameIdentifier>
<mods:affiliation id="874280F5FD7FA4BAD9FDE7F85165DE7D">Fungal Biology and Systematics Research Laboratory, Institute of Botany, University of the Punjab, Quaid-e-Azam Campus 54590, Lahore, Pakistan</mods:affiliation>
</mods:name>
<mods:name id="745BC2386A893B3902D4975C2DA1A492" type="personal">
<mods:role id="E9EFA52EB1258B1825C63B7795B739D6">
<mods:roleTerm id="50837D8AB30B2E7F7B699CB5D0C067FC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6B5BF53C7CB9C2B548188C965537DF55">Khalid, Abdul Nasir</mods:namePart>
<mods:nameIdentifier id="F116FA9ED27B5F3183F0B3BA6C86766C" type="ORCID">0000-0002-5635-8031</mods:nameIdentifier>
<mods:affiliation id="83555695F39A802AD230206EC7EFCA6F">Fungal Biology and Systematics Research Laboratory, Institute of Botany, University of the Punjab, Quaid-e-Azam Campus 54590, Lahore, Pakistan</mods:affiliation>
</mods:name>
<mods:typeOfResource id="FA1C0677E4A8AE9815B2640825EC0DE0">text</mods:typeOfResource>
<mods:relatedItem id="1BD964B6F02AD0AA18AEB0D229664849" type="host">
<mods:titleInfo id="4748C29EF3279712ADFD355F8E6A26F0">
<mods:title id="48A554AE7958ECC96A89E580FA37C3B7">MycoKeys</mods:title>
</mods:titleInfo>
<mods:part id="8FE23A5CA303A819959FFC35BED674B9">
<mods:date id="41C7F9474CBFFD0E3750C5510C84CC4A">2025</mods:date>
<mods:detail id="2A40FB26F79A5AA53C23A8B735B06E58" type="pubDate">
<mods:number id="6A377CF77E766D5A7DA90451DC514578">2025-01-16</mods:number>
</mods:detail>
<mods:detail id="D7C0E543BB92FADE1FD41D4019CFEEF9" type="volume">
<mods:number id="6B75A677DF7C2C271B1253D3F4C9EAD7">112</mods:number>
</mods:detail>
<mods:extent id="C246A9A16100A420FB86D148C2733979" unit="page">
<mods:start id="AF24BD181BC77299D5F119746359966A">165</mods:start>
<mods:end id="143BA9DE69A98C5A331C45BEA5B76CA3">182</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="BFB161C1C0FFF2623AD8321879BDF602">journal article</mods:classification>
<mods:identifier id="D4142F88A8691BBBB7DA8FE10E6CB4F9" type="CLB-Dataset">307585</mods:identifier>
<mods:identifier id="58F7F1AF9ED58679A61CA5D55FE79CB2" type="DOI">10.3897/mycokeys.112.141766</mods:identifier>
<mods:identifier id="EC5C2F8EECC18DB5DD787F803A58C910" type="GBIF-Dataset">e1093352-070e-4a36-9a4e-7b101f33fdcf</mods:identifier>
</mods:mods>
<treatment id="25AC64DF00CF5404A087DD1248A654FD" ID-DOI="http://doi.org/10.5281/zenodo.14673745" ID-GBIF-Taxon="252177807" ID-Zenodo-Dep="14673745" ID-arpha="25AC64DF-00CF-5404-A087-DD1248A654FD" ID-mycobank="855776" LSID="urn:lsid:plazi:treatment:25AC64DF00CF5404A087DD1248A654FD" httpUri="http://treatment.plazi.org/id/25AC64DF00CF5404A087DD1248A654FD" scope_family="Psathyrellaceae" scope_phylum="Basidiomycota">
<subSubSection id="6EFCF85C7507A8FC420F48A41A368E01" type="nomenclature">
<paragraph id="CFE6EEDE32DB101EBBC5E3848073B300">
<taxonomicName id="0293553EFBC9C5AD805555A79A002A4B" authority="Haqnawaz, Niazi &amp; Khalid" authorityName="Haqnawaz, Niazi &amp; Khalid" authorityYear="2025" class="Agaricomycetes" family="Psathyrellaceae" genus="Candolleomyces" higherTaxonomySource="CoL,GBIF" kingdom="Fungi" order="Agaricales" phylum="Basidiomycota" rank="species" species="virgatus" status="sp. nov.">
<emphasis id="A7600BC48C6F75299A90735ECE08BC1E" italics="true">Candolleomyces virgatus</emphasis>
Haqnawaz, Niazi &amp; Khalid
</taxonomicName>
<taxonomicNameLabel id="0710B96FDAC5C0AFB37D28B595D5447C" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="2C5A0F7F59126B328BCFE7FE9C0BADBC" type="description">
<paragraph id="E4F4F7E1AFB5ECEC478202D413BA53E9">
<figureCitation id="748A716B42FD4BF823BA1399957C5F23" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A – C basidiomata of Candolleomyces virgatus sp. nov. (Holotype LAH 38259). Scale bars: 10 mm." figureDoi="10.3897/mycokeys.112.141766.figure7" httpUri="https://binary.pensoft.net/fig/1221199">Figs 7</figureCitation>
, 
<figureCitation id="B40DD303F8EA8DB310095BCAA0C1223E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A – D microscopic structures of Candolleomyces virgatus (Holotype. LAH 38259) A basidia B basidiospores C cheilocystidia D pileipellis. Scale bars: 10 µm (A – C); (D) 20 µm (D)." figureDoi="10.3897/mycokeys.112.141766.figure8" httpUri="https://binary.pensoft.net/fig/1221200">8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUMAI" type="etymology">
<paragraph id="2617CF5F80417408114FF040412DAD3A">
<heading id="5D4BA1886976A4A1696D768C48291B11" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="C218F9FCB520046CC8CCBB9F5F4B5B9D">
Species name “ 
<taxonomicName id="7AF1D4F2BA1DF1C9D559CEEEEF7CB803" authority="(Latin)" authorityName="Haqnawaz, Niazi &amp; Khalid" authorityYear="2025" baseAuthorityName="Latin" class="Agaricomycetes" family="Psathyrellaceae" genus="Candolleomyces" higherTaxonomySource="CoL,GBIF" kingdom="Fungi" order="Agaricales" phylum="Basidiomycota" rank="species" species="virgatus">
<emphasis id="DE377DCB6B080F70581BEC48CB0B2D28" italics="true">virgatus</emphasis>
</taxonomicName>
” (Latin) refers to the virgate pileus surface.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EENAI" type="Holotype">
<paragraph id="7047FE320FDC4F62A82B049B7192B344">
<heading id="42576D4090AAA98441FA9A33C51C9D4E" reason="title">
<typeStatus id="E30299B4739C70858F23BAD51AE823BA" type="holotype">Holotype</typeStatus>
.
</heading>
</paragraph>
<paragraph id="22A2B251CE9C895CAA08AFBCE3FAEAC3">
<materialsCitation id="E5D09C0CFA8C995ABD7F1C10615B2E1A" ID-GBIF-Occurrence="5018575302" collectedFrom="on soil, rich in organic matter" collectingDate="2020-08-29" collectionCode="ITS, LSU" collectorName="Muhammad Haqnawaz" country="Pakistan" elevation="134" latitude="30.245556" location="Indus Riverbed" longLatPrecision="20" longitude="70.85111" municipality="Kot Addu District" specimenCode="LAH 38259, PQ 329549, PQ 369430" specimenCount="1" stateProvince="Punjab Province" typeStatus="holotype">
<collectingCountry id="A0E9A5A51744EF751DA1EB7E338AE9CF" name="Pakistan">Pakistan</collectingCountry>
• 
<collectingRegion id="CB47EE6F186D52FA437D5D3794EF79CE" country="Pakistan" name="Punjab">Punjab Province</collectingRegion>
: 
<collectingMunicipality id="CCEA690317A4F553AB7C00F94D617A78">Kot Addu District</collectingMunicipality>
, 
<location id="5FC6DA870FFD1992EEE16267979C8A7C" LSID="urn:lsid:plazi:treatment:25AC64DF00CF5404A087DD1248A654FD:5FC6DA870FFD1992EEE16267979C8A7C" country="Pakistan" latitude="30.245556" longLatPrecision="20" longitude="70.85111" municipality="Kot Addu District" name="Indus Riverbed" stateProvince="Punjab Province">Indus Riverbed</location>
, (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENNAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[70.851111,30.245556]}">
<geoCoordinate id="10BD9118E797AE0A9B5B6149DDE6731D" degrees="30" direction="north" minutes="14" orientation="latitude" precision="15" seconds="44" value="30.245556">30°14'44&quot;N</geoCoordinate>
, 
<geoCoordinate id="911FE424551BEC025FE7E5C6068F4D3F" degrees="70" direction="east" minutes="51" orientation="longitude" precision="15" seconds="04" value="70.85111">70°51'04&quot;E</geoCoordinate>
</named-content>
, 
<elevation id="B34C01C26507A53D5A4E020619C4169F" metricMagnitude="2" metricUnit="m" metricValue="1.34" unit="m" value="134.0">
<quantity id="B26AAB206FD7372FAF4634ACDD67766E" metricMagnitude="2" metricUnit="m" metricValue="1.34" unit="m" value="134.0">134 m</quantity>
a. s. l.
</elevation>
), 
<collectedFrom id="82E20D7ABE6A58B233AE0BA0E2F88747">on soil, rich in organic matter</collectedFrom>
, 
<date id="F49ACD553E91259810A0CE4ADFD18C99" value="2020-08-29">
<collectingDate id="90EDCDE9758AF34E4105F2696B631F97" value="2020-08-29">29 Aug 2020</collectingDate>
</date>
, 
<collectorName id="9DAEC69A5BB70E8A9CD99B4BA37A5A40">Muhammad Haqnawaz</collectorName>
, EF – 17 (
<emphasis id="188391A91C4C079199FF3C58730DC9B9" bold="true" italics="true">
<typeStatus id="B09566BDF3002AD559BCC14E000DB3FD" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCode id="5BF82070A59DCD0E0FD094CB3051A8DD" collectionCode="LAH" country="Pakistan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14637" lsid="urn:lsid:biocol.org:col:14637" name="University of the Punjab" type="Herbarium">LAH 38259</specimenCode>
). GenBank: 
<ext-link id="588A74D07812B47A468D9D7EC4320B39" ext-link-type="gen" xlink_href="PQ329543" xlink_type="simple">PQ 329543</ext-link>
[
<collectionCode id="82143E5E85241DC31EEAEC3633A777EF">ITS</collectionCode>
], 
<ext-link id="BE31CAB3D5754AA8046A2D368B74BA9A" ext-link-type="gen" xlink_href="PQ329549" xlink_type="simple">
<specimenCode id="E25F09F566D29EA407AA196DEED05648" collectionCode="ITS">PQ 329549</specimenCode>
</ext-link>
[
<collectionCode id="4307F9249D12DD6B04CAEC7FC948988D" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15590" lsid="urn:lsid:biocol.org:col:15590" name="Louisiana State University - Herbarium" type="Herbarium">LSU</collectionCode>
], 
<ext-link id="489536C544D1AF765C9E5C553CAE79CB" ext-link-type="gen" xlink_href="PQ369430" xlink_type="simple">
<specimenCode id="8C3849BC2D613052421A278BAF3BD7EA" collectionCode="LSU" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15590" lsid="urn:lsid:biocol.org:col:15590" name="Louisiana State University - Herbarium" type="Herbarium">PQ 369430</specimenCode>
</ext-link>
[tef- 1 α].
</materialsCitation>
</paragraph>
<caption id="9577C76715DC20EA3023F6D583CD4DFB" ID-DOI="10.3897/mycokeys.112.141766.figure7" ID-arpha="91AAA3CD-A734-59C9-9221-433EDE294D51" httpUri="https://binary.pensoft.net/fig/1221199" startId="F7">
<paragraph id="C26238D6D9D507718EE85D9C202469B1">
<label id="5E98D812E400F8937C859B9597BF1C22">Figure 7.</label>
</paragraph>
<paragraph id="B876520F4FFE0E6BE93CDB277DFD7C5A">
<emphasis id="BC10DC0A9B81E3C7B662993E0CC5EFEF" bold="true">A – C</emphasis>
basidiomata of 
<taxonomicName id="15CC7B1AB8555556617F486FE8B9E2AD" authorityName="Haqnawaz, Niazi &amp; Khalid" authorityYear="2025" class="Agaricomycetes" family="Psathyrellaceae" genus="Candolleomyces" higherTaxonomySource="CoL,GBIF" kingdom="Fungi" order="Agaricales" phylum="Basidiomycota" rank="species" species="virgatus" status="sp. nov.">
<emphasis id="F61D32446B98BE941B474F8B7626C0AA" italics="true">Candolleomyces virgatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DF7BBFCE37E8D76F19F875E42157F9DE" rank="species">sp. nov.</taxonomicNameLabel>
(Holotype LAH 38259). Scale bars: 10 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0E5OAI" type="diagnosis">
<paragraph id="E7F9B49F98C57E6B4C44CBC315F5CA96">
<heading id="19A1AB29C9DFC55FB9027CF5B3EF4308" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="FD7321147B9660C612FF4BFFFCD7AF0D">
This species is different from 
<taxonomicName id="5EF067C2290AC796EB98607F017C5EEB" class="Agaricomycetes" family="Psathyrellaceae" genus="Candolleomyces" higherTaxonomySource="CoL,GBIF" kingdom="Fungi" order="Agaricales" phylum="Basidiomycota" rank="species" species="campanulatus">
<emphasis id="78BF0C682830FF843A555631F170E6B0" italics="true">Candolleomyces campanulatus</emphasis>
</taxonomicName>
by its parabolic to plane, distinct umbo, virgate surface of pileus, crinkled, 1–7 tiers of forking lamellae, marginate base, longitudinally striation on surface of stipe, lageniform with moniliform at top, moniliform, cylindrical, tibiiform, conical cheilocystidia.
</paragraph>
<caption id="D11F490F780FA47744BF75124E67E727" ID-DOI="10.3897/mycokeys.112.141766.figure8" ID-arpha="1018B4EC-4C18-5F04-8781-558C65F5ECE9" httpUri="https://binary.pensoft.net/fig/1221200" startId="F8">
<paragraph id="51B81ACD1CED61BF5138C41715C5AB03">
<label id="2463794AFC6117A57D26D402B3F1CEF6">Figure 8.</label>
</paragraph>
<paragraph id="4020E3DB23A055AD2A8F06D4921E79F3">
<emphasis id="AE3BE481AD18AE559FAA0F2DE15BB228" bold="true">A – D</emphasis>
microscopic structures of 
<taxonomicName id="8E060E23438D49F308C4711C4D235EEE" authorityName="Haqnawaz, Niazi &amp; Khalid" authorityYear="2025" class="Agaricomycetes" family="Psathyrellaceae" genus="Candolleomyces" higherTaxonomySource="CoL,GBIF" kingdom="Fungi" order="Agaricales" phylum="Basidiomycota" rank="species" species="virgatus">
<emphasis id="48AF965FF392C41E55AD6D43C12B7485" italics="true">Candolleomyces virgatus</emphasis>
</taxonomicName>
(Holotype. LAH 38259) 
<emphasis id="46C84A42326C1A47B799EB35BE4538C7" bold="true">A</emphasis>
basidia 
<emphasis id="E452C2FC79B72C153501B12F68469573" bold="true">B</emphasis>
basidiospores 
<emphasis id="8E81C106433D456E0464EC7D6A7B2B97" bold="true">C</emphasis>
cheilocystidia 
<emphasis id="B6ACD0630D5741240F6BEC26FE0F5BB4" bold="true">D</emphasis>
pileipellis. Scale bars: 10 µm (
<emphasis id="28531CA5A56BF897BB9A3B0C939396D4" bold="true">A – C</emphasis>
); (
<emphasis id="45A15FEA404527C99A41E8BFFB18F1FC" bold="true">D</emphasis>
) 20 µm (
<emphasis id="D90C0C4132E2AEDFC2C22B0EF97650E3" bold="true">D</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EXQAI" type="description">
<paragraph id="9A1F1D31949F00D06DBD81D70915B39D">
<heading id="9FA1B85928C9544F9A540F9A79360001" reason="title">Description.</heading>
</paragraph>
<paragraph id="0783EA42E423ED836121A153F9872E5F">
Basidiomata medium-sized. 
<emphasis id="1F4131277C0C2D50AFCE2269975C45E4" bold="true" italics="true">Pileus</emphasis>
<quantity id="7E04FB9BD3DECE56DE4EA58DEA813538" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="3.0" metricValueMin="1.7" unit="mm" value="23.5" valueMax="30.0" valueMin="17.0">17–30 mm</quantity>
diameter, parabolic at young stage, campanulate to plane, umbonatus, straight shape and regular cap of margins, virgate, light brownish gray (7.5 YR 7 / 1), and dull reddish brown (5 YR 5 / 4) center. 
<emphasis id="92C570659252111C2292A3C4066869F1" bold="true" italics="true">Lamellae</emphasis>
adnate, broad, crinkled, 1–7 tiers of lamellae, branched or forking; some are not branched, grayish red (10 R 6 / 2). 
<emphasis id="A4433005CE8B1CF79DC6A4B67071F813" bold="true" italics="true">Stipe</emphasis>
40–80 × 
<quantity id="C8F6E51A430C1137149926C285E27D67" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" unit="mm" value="2.5" valueMax="4.0" valueMin="1.0">1–4 mm</quantity>
, white gray (N 8 / 0), slightly flexuous, hollow, tomentose, and broad at base, marginate, longitudinally striation. Annulus absent.
</paragraph>
<paragraph id="71CC4AF387ADB65CDDB353ABD4E046D8">
<emphasis id="631E59F12872FDC09C6EC95EA9505B1D" bold="true" italics="true">Basidiospores</emphasis>
[150 / 2 / 3], (6 –) 7–9 (– 10) × (4 –) 4.5–5 (– 6) µm, Q = 1.2–1.8, Qav = 1.61, ellipsoid to broadly ellipsoid, thick-walled, smooth, guttulate, germ pore present, purplish to brownish with greenish to yellow guttule in 5 % KOH and water. 
<emphasis id="204EF277865A812005AC41AE916BEE9C" bold="true" italics="true">Basidia</emphasis>
(12 –) 13–29 (– 30) × (6 –) 7–9 (– 10) µm, clavate, broadly clavate, hyaline to light yellowish in 5 % KOH, thick-walled, smooth, with 2–4 sterigmata. 
<emphasis id="2A50D2EF56E8CDC67C4A00435BE26D35" bold="true" italics="true">Cheilocystidia</emphasis>
(26 –) 27–46 (– 47) × (8 –) 9–13 (– 14) µm, utriform, lageniform with moniliform at top, moniliform, cylindrical, tibiiform, conical, hyaline to light yellowish in 5 % KOH. 
<emphasis id="364AE38E6BB3694B5972BE31A5E1CB15" bold="true" italics="true">Pleurocystidia</emphasis>
and 
<emphasis id="0C1038E8CD52A6A6449AE89A3BAD2089" bold="true" italics="true">Caulocystidia</emphasis>
absent. 
<emphasis id="A9145B26AE030681F1B9FD919B969B75" bold="true" italics="true">Pileipellis</emphasis>
an irregular epithelium; pileipellis cells 30–60 × 20–42 µm diameter, globose to subglobose and ellipsoid to broadly ellipsoid, clavate, hyaline to light yellowish in 5 % KOH. 
<emphasis id="573D9AFADF0EE64ECE6DBB9966BBB196" bold="true" italics="true">Stipitipellis</emphasis>
a cutis, hyphae subregular, branched 6–15 µm in diameter, thin-walled, septate, hyaline to light yellowish in 5 % KOH. Clamp connection present in all tissues.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EISAI" type="ecology">
<paragraph id="0741C16F2AB9B391B6D217969DABC25F">
<heading id="6EC6EAF78A4036A0F9E47B8DF97FBF21" reason="title">Ecology and habitat.</heading>
</paragraph>
<paragraph id="4E0EA079E20DEB5D8CE684088ABFB4B9">Terrestrial, gregarious, scattered, on loamy soil rich in organic matter.</paragraph>
</subSubSection>
<subSubSection id="SECID0ENSAI" type="material">
<paragraph id="E90AF969A6B2F0779DFFF52CC3F9B30E">
<heading id="9D67698382D5149F7E9EFA0AB3BD9D3B" reason="title">Additional material examined.</heading>
</paragraph>
<paragraph id="E8FA3AD3D78E0E534AA313F738C8DFCD">
<materialsCitation id="16FEBD66777A697CA01CBAA626BFFA9E" ID-GBIF-Occurrence="5018575301" collectedFrom="on loamy soil" collectingDate="2020-09-02" collectionCode="ITS, LSU" collectorName="Muhammad Haqnawaz" country="Pakistan" elevation="132" latitude="30.463057" location="Noor Shah Thal" longLatPrecision="20" longitude="71.03472" municipality="Kot Addu" specimenCode="LAH 38260, PQ 329550, PQ 369431" specimenCount="1" stateProvince="Punjab Province" typeStatus="paratype">
<collectingCountry id="E5BEC2F42E478FD9C743E55B1871C71E" name="Pakistan">Pakistan</collectingCountry>
• 
<collectingRegion id="EDD0DDB03872873FE5798D0F04F61E90" country="Pakistan" name="Punjab">Punjab Province</collectingRegion>
: 
<collectingMunicipality id="D5A7ABE81FFA6E9A1066BD9C5211CF7A">Kot Addu</collectingMunicipality>
, 
<location id="CE9E618074CE9238F3F6CD01D033D11F" LSID="urn:lsid:plazi:treatment:25AC64DF00CF5404A087DD1248A654FD:CE9E618074CE9238F3F6CD01D033D11F" country="Pakistan" latitude="30.463057" longLatPrecision="20" longitude="71.03472" municipality="Kot Addu" name="Noor Shah Thal" stateProvince="Punjab Province">Noor Shah Thal</location>
, (
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EWSAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[71.034722,30.463056]}">
<geoCoordinate id="4696D9DD3977FD886932B94CD27010D8" degrees="30" direction="north" minutes="27" orientation="latitude" precision="15" seconds="47" value="30.463057">30°27'47&quot;N</geoCoordinate>
, 
<geoCoordinate id="876908405DD214D19E122EF2C52A2603" degrees="71" direction="east" minutes="02" orientation="longitude" precision="15" seconds="05" value="71.03472">71°02'05&quot;E</geoCoordinate>
</named-content>
, 
<elevation id="84AB9BF9A1AE18BE3B844DED90F907F5" metricMagnitude="2" metricUnit="m" metricValue="1.3199999999999998" unit="m" value="132.0">
<quantity id="D0B5CFE878FDE4292848FBE37FB9EE42" metricMagnitude="2" metricUnit="m" metricValue="1.3199999999999998" unit="m" value="132.0">132 m</quantity>
a. s. l.
</elevation>
), 
<collectedFrom id="51F60911CA433B48B6D99C9DA9CF5826">on loamy soil</collectedFrom>
, 
<date id="BFF79E8C2DA0DF4AA3F3853289036C46" value="2020-09-02">
<collectingDate id="BA175D357700C1921359518EF2F421AE" value="2020-09-02">02 September 2020</collectingDate>
</date>
, 
<collectorName id="4A0642A4F94268AD4F782B22165DC193">Muhammad Haqnawaz</collectorName>
, KA – 59 (
<emphasis id="7FA7763886F0E443A076D692845BE1C4" bold="true" italics="true">
<typeStatus id="88B657303EA410F0B58FC1D496234AFB" type="paratype">Paratype</typeStatus>
</emphasis>
; 
<specimenCode id="BB9EB5BE4649F7C9DC5AD04C250DC81A" collectionCode="LAH" country="Pakistan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14637" lsid="urn:lsid:biocol.org:col:14637" name="University of the Punjab" type="Herbarium">LAH 38260</specimenCode>
) GenBank 
<ext-link id="F7F28F469A07674B10E0D711D0ED9EE6" ext-link-type="gen" xlink_href="PQ329544" xlink_type="simple">PQ 329544</ext-link>
[
<collectionCode id="A68B72744606068C75E01887C7393559">ITS</collectionCode>
], 
<ext-link id="16A9CD535D308C231CE3AC6270F1E7A6" ext-link-type="gen" xlink_href="PQ329550" xlink_type="simple">
<specimenCode id="9213021115E99A355ED1A5174EF23EFE" collectionCode="ITS">PQ 329550</specimenCode>
</ext-link>
[
<collectionCode id="B14A83BC9170AC1A8686D01464D5BE6B" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15590" lsid="urn:lsid:biocol.org:col:15590" name="Louisiana State University - Herbarium" type="Herbarium">LSU</collectionCode>
], 
<ext-link id="2671E285387340D6CD62B175A47F2677" ext-link-type="gen" xlink_href="PQ369431" xlink_type="simple">
<specimenCode id="758B836564D4ED64A2CDCCB2BF8C0994" collectionCode="LSU" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15590" lsid="urn:lsid:biocol.org:col:15590" name="Louisiana State University - Herbarium" type="Herbarium">PQ 369431</specimenCode>
</ext-link>
[tef- 1 α]
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>