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<document id="62B634E335EFC6BB41B8FBAA94209D2B" ID-DOI="10.11646/zootaxa.3866.1.5" ID-GBIF-Dataset="1c892016-2918-4b05-8eea-9032f9f7c6ff" ID-ISSN="1175-5326" ID-Zenodo-Dep="227564" ID-ZooBank="A3620CFB-E223-479C-8A9B-FF84F30143CF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460916005144" checkinUser="plazi" docAuthor="Nguyen, Tung T., Tran, Binh T. T. &amp; Nguyen, Anh D." docDate="2014" docId="0F1A87E0FFF1FFB2A8CAFDB2F194FA9F" docLanguage="en" docName="zt03866p121.pdf" docOrigin="Zootaxa 3866 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Polypheretima grandisetosa" docType="treatment" docVersion="10" lastPageNumber="115" masterDocId="F323FF98FFFBFFB8A85DFFF5F256FFBA" masterDocTitle="Earthworms of the ' acaecate' Pheretima group in Vietnam (Oligochaeta: Megascolecidae), with description of a new species from the Mekong delta" masterLastPageNumber="121" masterPageNumber="105" pageNumber="115" updateTime="1698361745542" updateUser="plazi">
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<mods:title id="DA73E6578A967A90AD6B6CA1BE51608B">Earthworms of the ' acaecate' Pheretima group in Vietnam (Oligochaeta: Megascolecidae), with description of a new species from the Mekong delta</mods:title>
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<treatment id="0F1A87E0FFF1FFB2A8CAFDB2F194FA9F" ID-DOI="http://doi.org/10.5281/zenodo.6144718" ID-GBIF-Taxon="127711696" ID-Zenodo-Dep="6144718" LSID="urn:lsid:plazi:treatment:0F1A87E0FFF1FFB2A8CAFDB2F194FA9F" httpUri="http://treatment.plazi.org/id/0F1A87E0FFF1FFB2A8CAFDB2F194FA9F" lastPageNumber="115" pageId="10" pageNumber="115">
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<heading id="DC44819AFFF1FFB2A8CAFDB2F0D3FDD8" bold="true" box="[151,645,583,610]" fontSize="11" level="1" pageId="10" pageNumber="115" reason="1">
<taxonomicName id="40B34D75FFF1FFB2A8CAFDB2F0D3FDD8" authority="(Thai, 1996)" baseAuthorityName="Thai" baseAuthorityYear="1996" box="[151,645,583,610]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="115" phylum="Annelida" rank="species" species="grandisetosa">
<emphasis id="B5C7EAE4FFF1FFB2A8CAFDB2F0D3FDD8" bold="true" box="[151,645,583,610]" pageId="10" pageNumber="115">
<emphasis id="B5C7EAE4FFF1FFB2A8CAFDB2F3B1FDDB" bold="true" box="[151,487,583,609]" italics="true" pageId="10" pageNumber="115">Polypheretima grandisetosa</emphasis>
(
<bibRefCitation id="E3224B07FFF1FFB2A9ABFDBDF02BFDD8" author="Thai" box="[502,637,584,610]" pageId="10" pageNumber="115" refString="Thai, T. B. (1996) Description of five new species of the acaecate earthworms of the genus Pheretima Kinberg in Vietnam and key to the species of acaecate Pheretima recorded from Indochinese area. Tap Chi Sinh Hoc Journal of Biology, 18 (1), 1 - 6. [in Vietnamese with English summary]" type="journal article" year="1996">
<collectingCountry id="FFA47666FFF1FFB2A9ABFDBDF065FDD8" box="[502,563,584,610]" name="Thailand" pageId="10" pageNumber="115">Thai</collectingCountry>
, 1996
</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="870C36F6FFF1FFB2A8CAFD9FF2BDFD39" blockId="10.[151,645,583,643]" box="[151,235,618,643]" pageId="10" pageNumber="115">
(
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)
</paragraph>
<paragraph id="870C36F6FFF1FFB2A8CAFD47F020FD72" blockId="10.[151,877,690,743]" box="[151,630,690,712]" pageId="10" pageNumber="115">
<treatmentCitationGroup id="A7A311D8FFF1FFB2A8CAFD47F020FD72" box="[151,630,690,712]" pageId="10" pageNumber="115">
<taxonomicName id="40B34D75FFF1FFB2A8CAFD47F04EFD72" authority="Thai, 1996: 3" authorityName="Thai" authorityPageNumber="3" authorityYear="1996" box="[151,536,690,712]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="115" phylum="Annelida" rank="species" species="grandisetosa">
<emphasis id="B5C7EAE4FFF1FFB2A8CAFD47F3DFFD72" box="[151,393,690,712]" italics="true" pageId="10" pageNumber="115">Pheretima grandisetosa</emphasis>
<treatmentCitation id="061210E7FFF1FFB2A9D2FD47F04EFD72" author="Thai" box="[399,536,690,712]" page="3" pageId="10" pageNumber="115" year="1996">
<bibRefCitation id="E3224B07FFF1FFB2A9D2FD47F056FD72" author="Thai" box="[399,512,690,712]" pageId="10" pageNumber="115" refString="Thai, T. B. (1996) Description of five new species of the acaecate earthworms of the genus Pheretima Kinberg in Vietnam and key to the species of acaecate Pheretima recorded from Indochinese area. Tap Chi Sinh Hoc Journal of Biology, 18 (1), 1 - 6. [in Vietnamese with English summary]" type="journal article" year="1996">
<collectingCountry id="FFA47666FFF1FFB2A9D2FD47F396FD72" box="[399,448,690,712]" name="Thailand" pageId="10" pageNumber="115">Thai</collectingCountry>
, 1996
</bibRefCitation>
: 3
</treatmentCitation>
</taxonomicName>
,
<figureCitation id="1F882A73FFF1FFB2AA7EFD47F036FD72" box="[547,608,690,712]" captionStart="FIGURE 2" captionStartId="5.[151,250,1690,1712]" captionTargetBox="[151,1436,241,1658]" captionTargetId="figure@5.[151,1436,195,1668]" captionTargetPageId="5" captionText="FIGURE 2. Distribution of species of the genus Planapheretima in Vietnam." httpUri="https://zenodo.org/record/227566/files/figure.png" pageId="10" pageNumber="115">Fig. 2</figureCitation>
B.
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</paragraph>
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<treatmentCitationGroup id="A7A311D8FFF1FFB2A8CAFD24F13BFD5D" box="[151,877,721,743]" pageId="10" pageNumber="115">
<taxonomicName id="40B34D75FFF1FFB2A8CAFD24F3E2FD5D" box="[151,436,721,743]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="115" phylum="Annelida" rank="species" species="grandisetosa">
<emphasis id="B5C7EAE4FFF1FFB2A8CAFD24F3E2FD5D" box="[151,436,721,743]" italics="true" pageId="10" pageNumber="115">Polypheretima grandisetosa</emphasis>
</taxonomicName>
<treatmentCitation id="061210E7FFF1FFB2A990FD24F0DAFD5D" author="Nakamura" box="[461,652,721,743]" page="7" pageId="10" pageNumber="115" year="1999">
<bibRefCitation id="E3224B07FFF1FFB2A990FD24F025FD5D" author="Nakamura" box="[461,627,721,743]" pageId="10" pageNumber="115" refString="Nakamura, Y. (1999) Checklist of earthworms of Pheretima genus group (Megascolecidae: Oligochaeta) of the world. Edaphologia, 64, 1 - 78." type="journal article" year="1999">Nakamura 1999</bibRefCitation>
: 7
</treatmentCitation>
;
<treatmentCitation id="061210E7FFF1FFB2AACAFD24F13EFD5D" author="Blakemore" box="[663,872,721,743]" page="45" pageId="10" pageNumber="115" year="2007">
<bibRefCitation id="E3224B07FFF1FFB2AACAFD24F114FD5D" author="Blakemore" box="[663,834,721,743]" pageId="10" pageNumber="115" refString="Blakemore, R. J. (2007) Updated checklist of Pheretimoids (Oligochaeta: Megascolecidae: Pheretima auct.) taxa. Available from: www. annelida. net / earthworm / Pheretimoids. pdf (accessed 26 August 2014)" type="journal article" year="2007">Blakemore 2007</bibRefCitation>
: 45
</treatmentCitation>
.
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</paragraph>
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<subSubSection id="CFA9657DFFF1FFB2A8CAFCE1F390FC22" pageId="10" pageNumber="115" type="materials_examined">
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<emphasis id="B5C7EAE4FFF1FFB2A8CAFCE1F313FC97" bold="true" box="[151,325,788,813]" pageId="10" pageNumber="115">
<typeStatus id="58088854FFF1FFB2A8CAFCE1F284FC97" box="[151,210,788,813]" pageId="10" pageNumber="115">Type</typeStatus>
material.
</emphasis>
<typeStatus id="58088854FFF1FFB2A911FCE1F3EFFC97" box="[332,441,788,813]" pageId="10" pageNumber="115" type="holotype">Holotype</typeStatus>
: 1A (SORC-HV.223) Ba Ria-Vung
<collectingCountry id="FFA47666FFF1FFB2AB17FCE0F122FC96" box="[842,884,789,812]" name="China" pageId="10" pageNumber="115">Tau</collectingCountry>
province, Conlon
<collectingCountry id="FFA47666FFF1FFB2AC18FCE1F6DAFC97" box="[1093,1164,788,813]" name="Iceland" pageId="10" pageNumber="115">Island</collectingCountry>
, slope near Datdoc Bay,
<quantity id="404B9B13FFF1FFB2A8CAFCCDF29EFCEA" box="[151,200,824,848]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="115" unit="m" value="10.0">10m</quantity>
asl,
<date id="F30D1036FFF1FFB2A8A0FCC2F3C3FCEA" box="[253,405,823,848]" pageId="10" pageNumber="115" value="1989-06-25">25 June 1989</date>
, coll.
<collectingCountry id="FFA47666FFF1FFB2A985FCC2F05DFCEA" box="[472,523,823,848]" name="Thailand" pageId="10" pageNumber="115">Thai</collectingCountry>
Tran Bai.
<typeStatus id="58088854FFF1FFB2AADFFCCDF0BFFCEA" box="[642,745,824,848]" pageId="10" pageNumber="115" type="paratype">Paratype</typeStatus>
: 1A, same data as
<typeStatus id="58088854FFF1FFB2ABE5FCC2F64CFCEA" box="[952,1050,823,848]" pageId="10" pageNumber="115" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="E3224B07FFF1FFB2AC75FCC2F6F6FCEA" author="Thai" box="[1064,1184,823,848]" pageId="10" pageNumber="115" refString="Thai, T. B. (1996) Description of five new species of the acaecate earthworms of the genus Pheretima Kinberg in Vietnam and key to the species of acaecate Pheretima recorded from Indochinese area. Tap Chi Sinh Hoc Journal of Biology, 18 (1), 1 - 6. [in Vietnamese with English summary]" type="journal article" year="1996">
<collectingCountry id="FFA47666FFF1FFB2AC75FCC2F60BFCEA" box="[1064,1117,823,848]" name="Thailand" pageId="10" pageNumber="115">Thai</collectingCountry>
1996
</bibRefCitation>
).
</paragraph>
<paragraph id="870C36F6FFF1FFB2A89AFCA9F390FC22" blockId="10.[151,1437,788,1317]" pageId="10" pageNumber="115">
<emphasis id="B5C7EAE4FFF1FFB2A89AFCA9F3CAFCCF" bold="true" box="[199,412,860,885]" pageId="10" pageNumber="115">Further material.</emphasis>
5A (
<collectionCode id="E1A2AE33FFF1FFB2A987FCA9F07FFCCE" box="[474,553,860,884]" pageId="10" pageNumber="115">SORC</collectionCode>
), Ba Ria-Vung
<collectingCountry id="FFA47666FFF1FFB2AA83FCA8F15EFCCE" box="[734,776,861,884]" name="China" pageId="10" pageNumber="115">Tau</collectingCountry>
province, Baycanh
<collectingCountry id="FFA47666FFF1FFB2ABADFCA9F661FCCF" box="[1008,1079,860,885]" name="Iceland" pageId="10" pageNumber="115">Island</collectingCountry>
,
<quantity id="404B9B13FFF1FFB2AC1BFCA9F6D3FCCE" box="[1094,1157,860,884]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="10" pageNumber="115" unit="m" value="210.0">210m</quantity>
asl,
<date id="F30D1036FFF1FFB2ACE2FCA9F70BFCCE" box="[1215,1373,860,884]" pageId="10" pageNumber="115" value="1989-06-20">20 June 1989</date>
, coll.
<collectingCountry id="FFA47666FFF1FFB2A8CAFC8AF29DFC22" box="[151,203,895,920]" name="Thailand" pageId="10" pageNumber="115">Thai</collectingCountry>
Tran Bai (
<bibRefCitation id="E3224B07FFF1FFB2A91FFC8AF3EFFC22" author="Thai" box="[322,441,895,920]" pageId="10" pageNumber="115" refString="Thai, T. B. (1996) Description of five new species of the acaecate earthworms of the genus Pheretima Kinberg in Vietnam and key to the species of acaecate Pheretima recorded from Indochinese area. Tap Chi Sinh Hoc Journal of Biology, 18 (1), 1 - 6. [in Vietnamese with English summary]" type="journal article" year="1996">
<collectingCountry id="FFA47666FFF1FFB2A91FFC8AF321FC22" box="[322,375,895,920]" name="Thailand" pageId="10" pageNumber="115">Thai</collectingCountry>
1996
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CFA9657DFFF1FFB2A89AFC51F6D8FB02" pageId="10" pageNumber="115" type="description">
<paragraph id="870C36F6FFF1FFB2A89AFC51F723FB2E" blockId="10.[151,1437,788,1317]" pageId="10" pageNumber="115">
<emphasis id="B5C7EAE4FFF1FFB2A89AFC51F30FFC07" bold="true" box="[199,345,932,957]" pageId="10" pageNumber="115">Description.</emphasis>
Body cylindrical, small size, unpigmented. Prostomium prolobous. First dorsal pore in 11/12. Setae all fine and regularly distributed on pre- and postclitellar segments, 60(16) in V,
<quantity id="404B9B13FFF1FFB2AC00FC32F6C0FC5A" box="[1117,1174,967,992]" metricMagnitude="0" metricUnit="m" metricValue="1.3716" pageId="10" pageNumber="115" unit="in" value="54.0">54 in</quantity>
VI,
<quantity id="404B9B13FFF1FFB2AC95FC32F757FC5A" box="[1224,1281,967,992]" metricMagnitude="0" metricUnit="m" metricValue="1.27" pageId="10" pageNumber="115" unit="in" value="50.0">50 in</quantity>
VIII, 40+
<quantity id="404B9B13FFF1FFB2AD2FFC3DF7CAFC5A" box="[1394,1436,967,992]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="10" pageNumber="115" unit="in" value="4.0">4 in</quantity>
XVIII,
<quantity id="404B9B13FFF1FFB2A8B0FC19F37DFBBF" box="[237,299,1004,1029]" metricMagnitude="0" metricUnit="m" metricValue="1.1176" pageId="10" pageNumber="115" unit="in" value="44.0">44 in</quantity>
XXV, aa=1.52ab, zz=zy; segment VI with 46 enlarged setae on the dorsal side.
<collectingCountry id="FFA47666FFF1FFB2ACAEFC19F77BFBBF" box="[1267,1325,1004,1029]" name="Malta" pageId="10" pageNumber="115">Male</collectingCountry>
pores on transversely oval porophores. No genital markings in both spermathecal and male regions. Spermathecal pores 2 pairs, ventral in furrows 5/6/7, about 0.25 body circumference apart. Metandric. Seminal vesicles in XII. Ovaries small in XIII. Last hearts in XIII. Intestine beginning at XVII, without caeca. Typhlosole simple, lamelliform, T=0.15.
<collectingCountry id="FFA47666FFF1FFB2A8AAFB89F363FB2E" box="[247,309,1148,1172]" name="Spain" pageId="10" pageNumber="115">Septa</collectingCountry>
<date id="F30D1036FFF1FFB2A961FB89F3D5FB2E" box="[316,387,1148,1172]" pageId="10" pageNumber="115" value="2010-09-08">8/9/10</date>
absent, 5/6/7/8 thickened. Pharyngeal micronephridia in 7/8. (Data from
<bibRefCitation id="E3224B07FFF1FFB2ACE7FB89F766FB2F" author="Thai" box="[1210,1328,1148,1173]" pageId="10" pageNumber="115" refString="Thai, T. B. (1996) Description of five new species of the acaecate earthworms of the genus Pheretima Kinberg in Vietnam and key to the species of acaecate Pheretima recorded from Indochinese area. Tap Chi Sinh Hoc Journal of Biology, 18 (1), 1 - 6. [in Vietnamese with English summary]" type="journal article" year="1996">
<collectingCountry id="FFA47666FFF1FFB2ACE7FB89F6BBFB2F" box="[1210,1261,1148,1173]" name="Thailand" pageId="10" pageNumber="115">Thai</collectingCountry>
1996
</bibRefCitation>
: 34.)
</paragraph>
<paragraph id="870C36F6FFF1FFB2A89AFB6AF6D8FB02" blockId="10.[151,1437,788,1317]" box="[199,1166,1183,1208]" pageId="10" pageNumber="115">
<emphasis id="B5C7EAE4FFF1FFB2A89AFB6AF388FB02" bold="true" box="[199,478,1183,1208]" pageId="10" pageNumber="115">
Records from
<collectingCountry id="FFA47666FFF1FFB2A92CFB6AF38FFB02" box="[369,473,1183,1208]" name="Vietnam" pageId="10" pageNumber="115">Vietnam</collectingCountry>
.
</emphasis>
Ba Ria-Vung
<collectingCountry id="FFA47666FFF1FFB2AA22FB55F0FFFB0D" box="[639,681,1184,1207]" name="China" pageId="10" pageNumber="115">Tau</collectingCountry>
Province (Conlon
<collectingCountry id="FFA47666FFF1FFB2AB20FB6AF193FB02" box="[893,965,1183,1208]" name="Iceland" pageId="10" pageNumber="115">Island</collectingCountry>
; Baycanh
<collectingCountry id="FFA47666FFF1FFB2AC64FB6AF6D4FB02" box="[1081,1154,1183,1208]" name="Iceland" pageId="10" pageNumber="115">Island</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="CFA9657DFFF1FFB2A89AFB31F0CBFB67" box="[199,669,1220,1245]" pageId="10" pageNumber="115" type="distribution">
<paragraph id="870C36F6FFF1FFB2A89AFB31F0CBFB67" blockId="10.[151,1437,788,1317]" box="[199,669,1220,1245]" pageId="10" pageNumber="115">
<emphasis id="B5C7EAE4FFF1FFB2A89AFB31F336FB67" bold="true" box="[199,352,1220,1245]" pageId="10" pageNumber="115">Distribution.</emphasis>
Only known from
<collectingCountry id="FFA47666FFF1FFB2AA6AFB31F0CCFB67" box="[567,666,1220,1245]" name="Vietnam" pageId="10" pageNumber="115">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="CFA9657DFFF1FFB2A89AFB12F194FA9F" pageId="10" pageNumber="115" type="discussion">
<paragraph id="870C36F6FFF1FFB2A89AFB12F194FA9F" blockId="10.[151,1437,788,1317]" pageId="10" pageNumber="115">
<emphasis id="B5C7EAE4FFF1FFB2A89AFB12F36DFABA" bold="true" box="[199,315,1255,1280]" pageId="10" pageNumber="115">Remarks.</emphasis>
The species is relatively close to
<taxonomicName id="40B34D75FFF1FFB2AA9CFB1DF63DFABA" authority="Thai, 1996" authorityName="Thai" authorityYear="1996" box="[705,1131,1255,1280]" class="Clitellata" family="Megascolecidae" genus="Polypheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="115" phylum="Annelida" rank="species" species="colonensis">
<emphasis id="B5C7EAE4FFF1FFB2AA9CFB1DF1B5FABA" box="[705,995,1256,1280]" italics="true" pageId="10" pageNumber="115">Polypheretima colonensis</emphasis>
<bibRefCitation id="E3224B07FFF1FFB2ABB0FB12F63DFABA" author="Thai" box="[1005,1131,1255,1280]" pageId="10" pageNumber="115" refString="Thai, T. B. (1996) Description of five new species of the acaecate earthworms of the genus Pheretima Kinberg in Vietnam and key to the species of acaecate Pheretima recorded from Indochinese area. Tap Chi Sinh Hoc Journal of Biology, 18 (1), 1 - 6. [in Vietnamese with English summary]" type="journal article" year="1996">
<collectingCountry id="FFA47666FFF1FFB2ABB0FB12F675FABA" box="[1005,1059,1255,1280]" name="Thailand" pageId="10" pageNumber="115">Thai</collectingCountry>
, 1996
</bibRefCitation>
</taxonomicName>
, but differs in smaller size and the presence of big setae in VI.
<typeStatus id="58088854FFF1FFB2AA74FAF8F03AFA9F" box="[553,620,1293,1317]" pageId="10" pageNumber="115">Types</typeStatus>
were rechecked for this study.
</paragraph>
</subSubSection>
</treatment>
</document>