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<document id="17A02B36A6831F2C15FD5484E059CF0E" ENCODING="UTF-8" ID-GBIF-Dataset="ea10c292-8516-4f2f-911e-38b9b36d434e" ModsDocID="z01614p053" checkinTime="1247241498666" checkinUser="thomas" docAuthor="Thiago N. A. Pereira &amp; Paulo H. F. Lucinda" docDate="2007" docId="077561934980E46F9CE3FAA30CEF9CB8" docLanguage="en" docName="2007_Pereira_Lucinda_gg2.xml" docOrigin="Zootaxa 1614" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:D553C255-E3DF-41BC-94F7-3979B8B4B02E" docTitle="Jupiaba elassonaktis Pereira &amp; Lucinda, 2007, new species" docType="treatment" docVersion="12" lastPageNumber="57" masterDocId="70490EF6352588A6E30920DE037A6AD7" masterDocTitle="A new species of Jupiaba Zanata, 1997 (Ostariophysi, Characiformes, Characidae) from the rio Tocantins drainage, Brazil." masterLastPageNumber="60" masterPageNumber="53" pageNumber="54" updateTime="1701314767256" updateUser="plazi">
<mods:mods id="9061F1B11DC24E413817EB03C2729687" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CD57D557A350EEA8507BB9575D26C580">
<mods:title id="FE8CF2DA462737B2C15A7EB2F87D2AC1">A new species of Jupiaba Zanata, 1997 (Ostariophysi, Characiformes, Characidae) from the rio Tocantins drainage, Brazil.</mods:title>
</mods:titleInfo>
<mods:name id="6822EF2F9B648C292555E3E79FFA9685" type="personal">
<mods:role id="142AB72A7CBD22D595F93DDA915BF58C">
<mods:roleTerm id="17BD0CE229DBB00DB43705141ECEF468">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="62D6AA55BE374BDB2F6C789B10664628">Thiago N. A. Pereira</mods:namePart>
</mods:name>
<mods:name id="CE9F6834A3A0E9AEC7CABD7F6B1016E1" type="personal">
<mods:role id="221826C2E788B797D135CFCC654B07D9">
<mods:roleTerm id="115E2E251FB8A4720BE3C4B93C5D503E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CFEB5AAAD91C1DA14054314F8ED742B5">Paulo H. F. Lucinda</mods:namePart>
</mods:name>
<mods:typeOfResource id="AA9AB0C561ABCE276C7503EEDB998527">text</mods:typeOfResource>
<mods:relatedItem id="424AE3E01CB5CA3C77FAC24EC6A5BD96" type="host">
<mods:titleInfo id="6C68F23877121BC05648E8FFCF7753EE">
<mods:title id="72BE3756FEF11519347F1C7236B22E0E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="68473DF31953295FD272F712EB57F87B">
<mods:date id="8CF3294B4D6B49ED6DA3830979D3A383">2007</mods:date>
<mods:detail id="4692C528246FF2B2B597D994219739C4" type="volume">
<mods:number id="09502B392A8A1FD8190DF52868121A4E">1614</mods:number>
</mods:detail>
<mods:extent id="A67D736BB40760A65BED7EB2BA38EDDF" unit="page">
<mods:start id="DC90146DA7E91CA3CE0D41BBEFEA63F5">53</mods:start>
<mods:end id="AF907085B775E96EAB00FE43CEF4E1A6">60</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="05638A5D187080CE23A98288FD6ED17F">
<mods:url id="3DA904810E167FE0523D52262DB330EB">http://www.zoobank.org/urn:lsid:zoobank.org:pub:D553C255-E3DF-41BC-94F7-3979B8B4B02E</mods:url>
</mods:location>
<mods:classification id="01155DA44B06997AE1E5A32AB04A4CCD">journal article</mods:classification>
<mods:identifier id="5083D2E97CAD4F218340251902605C4C" type="Plazi-Custom">z01614p053</mods:identifier>
<mods:identifier id="F02E70C31EEB951F5F59D8FF61B7CE17" type="ZooBank">D553C255-E3DF-41BC-94F7-3979B8B4B02E</mods:identifier>
</mods:mods>
<treatment id="077561934980E46F9CE3FAA30CEF9CB8" ID-DOI="http://doi.org/10.5281/zenodo.6248259" ID-GBIF-Taxon="100123738" ID-Zenodo-Dep="6248259" LSID="urn:lsid:plazi:treatment:077561934980E46F9CE3FAA30CEF9CB8" httpUri="http://treatment.plazi.org/id/077561934980E46F9CE3FAA30CEF9CB8" lastPageNumber="57" pageNumber="54">
<subSubSection id="AB4E565FC147D824E43B7915A852DD7F" type="nomenclature">
<paragraph id="5FB29BE0A58414050A20D728EBCB6835" pageNumber="54">
<taxonomicName id="AD20C214128DCC919289A032FBC16B3D" ID-CoL="6NHP9" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="elassonaktis">Jupiaba elassonaktis</taxonomicName>
,
<taxonomicNameLabel id="A03AC801884602BE5279E626D2E83555" rank="species">new species</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="29E7A2758DBA47A9F95AF51B28AFE8BD" type="description">
<paragraph id="FF202B939B9D226EDC81362909197EF0" pageNumber="54">(Figs. 1 and 2, Table 1)</paragraph>
</subSubSection>
<subSubSection id="4D77583DF2C7C505F9F6BE93631869EB" type="nomenclature">
<paragraph id="D94CD049439DFD36675B4850124D6011" pageNumber="54">
<materialsCitation id="5ABD468C4B788EA6A21C90DC29481F9E" ID-GBIF-Occurrence="923930448" collectingDate="2007-02-16" collectionCode="MCP" collectorName="NEAMB-UFT staff" country="Brazil" latitude="-12.25" location="Peixe, rio Tocantins downstream UHE Peixe-Angical" longitude="-48.383335" specimenCode="MCP 41465" stateProvince="Tocantins" typeStatus="Holotype">
<typeStatus id="2E442DC1CCF962F675C35B68813736AC">Holotype</typeStatus>
. (Fig. 1).
<specimenCode id="AE0DA73A884D3CEF6D899353D4051CD5">
<collectionCode id="C8FBC8671871DBCF061A4C03E5F92B5A">MCP</collectionCode>
41465
</specimenCode>
, 34.5 mm SL,
<location id="AC689437DA62BB9D14859B0883B940F1" LSID="urn:lsid:plazi:treatment:077561934980E46F9CE3FAA30CEF9CB8:AC689437DA62BB9D14859B0883B940F1" country="Brazil" latitude="-12.25" longitude="-48.383335" name="Peixe, rio Tocantins downstream UHE Peixe-Angical" stateProvince="Tocantins">Peixe, rio Tocantins downstream UHE Peixe-Angical</location>
, approximately
<geoCoordinate id="95944545433FDF99176F899659D01E12" direction="south" orientation="latitude" value="-12.25">12°15S</geoCoordinate>
<geoCoordinate id="1FC8C5E04A6553C067699A59EA9CC6EE" direction="west" orientation="longitude" value="-48.383335">48°23W</geoCoordinate>
,
<collectingRegion id="A43172E3CC7A525E7B89A09127BB77E4">Tocantins</collectingRegion>
,
<collectingCountry id="806986600C743ABBB72BE58E73DFF99D">Brazil</collectingCountry>
,
<date id="58BBC721B786C5D475F0D978122661B1" day="16" month="02" value="2007-02-16" year="2007">
<collectingDate id="AB145A00111026354AE8CC6F47D58CE0" day="16" month="02" value="2007-02-16" year="2007">16 Feb 2007</collectingDate>
</date>
,
<collectorName id="47ADD20D2B79FFD6A7F96CFEB594DFE6">NEAMB-UFT</collectorName>
.
</materialsCitation>
</paragraph>
<paragraph id="8468DC8C5303651C626F08E1AD1B1BDF" pageNumber="54">
<materialsCitation id="3FDD25C20F8B45E0C9CECD21884B533A" ID-GBIF-Occurrence="923930449" collectingDate="2005-03-28" collectionCode="UNT" collectorName="NEAMB-UFT staff" country="Brazil" latitude="-10.720833" location="Porto Nacional, rio Tocantins" longitude="-48.420555" specimenCode="UNT 5568" stateProvince="Tocantins" typeStatus="Paratypes">
<typeStatus id="092453A3624AFD9845C9827F46C84C49">Paratypes</typeStatus>
. All collected by
<collectorName id="0147333F247CED86B07A833FF6FBDE5F">NEAMB-UFT staff</collectorName>
. All specimens are from
<collectingCountry id="4B737618AC025009E1C5E735AF901687">Brazil</collectingCountry>
,
<collectingRegion id="20D69C3F1AFCB1F220F223361E6830C6">Tocantins</collectingRegion>
state, Rio Tocantins:
<specimenCode id="DD33E84AFF4CBF00717F98D7F1F4B2FC">
<collectionCode id="992D7A1A271A4E5BF01ED37AA2163652">UNT</collectionCode>
5568
</specimenCode>
, 8, 24.0-28.4 mm SL,
<location id="2741ED7CCF3FDC38B4ED6FBE5D67AC3C" LSID="urn:lsid:plazi:treatment:077561934980E46F9CE3FAA30CEF9CB8:2741ED7CCF3FDC38B4ED6FBE5D67AC3C" country="Brazil" latitude="-10.720833" longitude="-48.420555" name="Porto Nacional, rio Tocantins" stateProvince="Tocantins">Porto Nacional, rio Tocantins</location>
,
<geoCoordinate id="8876199790ACDD7FEFE3EA301FF52C14" direction="south" orientation="latitude" value="-10.720833">10°4315&quot;S</geoCoordinate>
<geoCoordinate id="2DC3CE31B1C99A92346AD261387A9644" direction="west" orientation="longitude" value="-48.420555">48°2514&quot;W</geoCoordinate>
,
<date id="9B26ABCBC0B97E1F29321F47C1039A06" day="28" month="03" value="2005-03-28" year="2005">
<collectingDate id="417073879BA1050D113AA1F1D009EAB8" day="28" month="03" value="2005-03-28" year="2005">28 Mar 2005</collectingDate>
</date>
.
</materialsCitation>
<materialsCitation id="A3C86BF14BDFBC77E309D3956700B1C5" ID-GBIF-Occurrence="923930454" collectingDate="2003-10-01" collectionCode="UNT" collectorName="NEAMB-UFT staff" country="Brazil" latitude="-11.05" location="Brejinho do Nazare, rio Crixas" longitude="-48.633335" specimenCode="UNT 5569" stateProvince="Tocantins" typeStatus="Paratypes">
<specimenCode id="1DDDB78E592EFAC0E31B7BEE635E8A1D">
<collectionCode id="DFCDBA4679C640B9F685ECEEF7109249">UNT</collectionCode>
5569
</specimenCode>
, 19, 20.7-29.5 mm SL,
<location id="F4C4EAA3C46E6FD822CAD0C66D64620E" LSID="urn:lsid:plazi:treatment:077561934980E46F9CE3FAA30CEF9CB8:F4C4EAA3C46E6FD822CAD0C66D64620E" latitude="-11.05" longitude="-48.633335" name="Brejinho do Nazare, rio Crixas">
Brejinho do
<normalizedToken id="694DABE0C9804287EFE8C740416AF0A2" originalValue="Nazaré">Nazare</normalizedToken>
, rio
<normalizedToken id="E6510DCE37E5B38AA1924F6DCAC8B87B" originalValue="Crixás">Crixas</normalizedToken>
</location>
, approximately
<geoCoordinate id="55D78F8BF2FC416E7CA4BAAA05FFA5AE" direction="south" orientation="latitude" value="-11.05">11°3S</geoCoordinate>
<geoCoordinate id="666A4213BE833B168DCE25F63D299BE2" direction="west" orientation="longitude" value="-48.633335">48°38W</geoCoordinate>
,
<date id="023BE79FE568825CA666B69B9626A0D8" day="01" month="10" value="2003-10-01" year="2003">
<collectingDate id="0EF7ECD8809371BC847D972FCB41F887" day="01" month="10" value="2003-10-01" year="2003">1 Oct 2003</collectingDate>
</date>
.
</materialsCitation>
<materialsCitation id="0B5058E042A1D21F5B1E71E69345BFCD" ID-GBIF-Occurrence="923930455" collectingDate="2005-07-21" collectionCode="UNT" collectorName="NEAMB-UFT staff" country="Brazil" latitude="-16.483334" location="Parana, rio Tocantins, fazenda Tracadal" longitude="-48.2" specimenCode="UNT 5573" stateProvince="Tocantins" typeStatus="Paratype">
<specimenCode id="5F331133ED9C0DB42FC465397E0DD2B0">
<collectionCode id="73C20E52D79A0CDDC209AFE14A7C0B3A">UNT</collectionCode>
5573
</specimenCode>
, 1, 35.0 mm SL,
<location id="E7EF5E4D23576A2FF27C9125FAF825DF" LSID="urn:lsid:plazi:treatment:077561934980E46F9CE3FAA30CEF9CB8:E7EF5E4D23576A2FF27C9125FAF825DF" latitude="-16.483334" longitude="-48.2" name="Parana, rio Tocantins, fazenda Tracadal">
<normalizedToken id="76DA121791C6EA8CDDBD1A7D2D4CD883" originalValue="Paranã">Parana</normalizedToken>
, rio Tocantins, fazenda
<normalizedToken id="64DC72B7118E664920BBEF8B7B06E909" originalValue="Traçadal">Tracadal</normalizedToken>
</location>
, approximately
<geoCoordinate id="88481B467183906FFB323161E746507F" direction="south" orientation="latitude" value="-16.483334">16°29S</geoCoordinate>
<geoCoordinate id="42DDD4C6DB0D490A455C12249EFEB7D5" direction="west" orientation="longitude" value="-48.2">48°12W</geoCoordinate>
,
<date id="00EF50C9F98667906DCDCDFF90AFDB32" day="21" month="07" value="2005-07-21" year="2005">
<collectingDate id="13461FEEF38A2A5BF48E0667A091A0D2" day="21" month="07" value="2005-07-21" year="2005">21 Jul 2005</collectingDate>
</date>
.
</materialsCitation>
<materialsCitation id="F8AC81339B3FB8371866BCB77E31F085" ID-GBIF-Occurrence="923930451" collectingDate="2001-10-07" collectionCode="UNT" collectorName="NEAMB-UFT staff" country="Brazil" latitude="-8.983334" location="Pedro Afonso, rio Tocantins next to the confluence of rio Sono" longitude="-48.166668" specimenCode="UNT 5575" stateProvince="Tocantins" typeStatus="Paratype">
<specimenCode id="6C2FE7187A8C6103AB596920608798B3">
<collectionCode id="CD214F9AA0498A664358EAEA2D7ECE02">UNT</collectionCode>
5575
</specimenCode>
, 1 c&amp;s, 30.9 mm SL,
<location id="B5BACC00535B759E3C2B4F0BD1EA1287" LSID="urn:lsid:plazi:treatment:077561934980E46F9CE3FAA30CEF9CB8:B5BACC00535B759E3C2B4F0BD1EA1287" latitude="-8.983334" longitude="-48.166668" name="Pedro Afonso, rio Tocantins next to the confluence of rio Sono">Pedro Afonso, rio Tocantins next to the confluence of rio Sono</location>
, aproximately
<geoCoordinate id="D581D57710D14B543F87AA15CEFE918D" direction="south" orientation="latitude" value="-8.983334">8°59S</geoCoordinate>
<geoCoordinate id="645B4B8D79E51A9A4860826F422C3CF5" direction="west" orientation="longitude" value="-48.166668">48°10W</geoCoordinate>
,
<date id="C35739771701B7E5566A87EDEC7A3920" day="07" month="10" value="2001-10-07" year="2001">
<collectingDate id="2028BF2D14AAA93046CE95B6FE536477" day="07" month="10" value="2001-10-07" year="2001">7 Oct 2001</collectingDate>
</date>
.
</materialsCitation>
<materialsCitation id="3054BB5D5F698C2487163B7AACC20781" ID-GBIF-Occurrence="923930453" collectingDate="2005-03-28" collectionCode="UNT" collectorName="NEAMB-UFT staff" country="Brazil" latitude="-11.316667" location="Ipueiras, rio Tocantins next to the confluence of rio Manuel Alves" longitude="-48.466667" specimenCode="UNT 5576" stateProvince="Tocantins" typeStatus="Paratypes">
<specimenCode id="591C132CC3A5A35D974C4B08D526FA8F">
<collectionCode id="D912B4D5F66C77BF9049272C345ACDFF">UNT</collectionCode>
5576
</specimenCode>
, 3, 19.9-32.5 mm SL,
<location id="C867B210AC38BADF2340D0E7F651C208" LSID="urn:lsid:plazi:treatment:077561934980E46F9CE3FAA30CEF9CB8:C867B210AC38BADF2340D0E7F651C208" latitude="-11.316667" longitude="-48.466667" name="Ipueiras, rio Tocantins next to the confluence of rio Manuel Alves">Ipueiras, rio Tocantins next to the confluence of rio Manuel Alves</location>
, approximately
<geoCoordinate id="04E80F5622A5D781C774EFF9A3239B2B" direction="south" orientation="latitude" value="-11.316667">11°19S</geoCoordinate>
<geoCoordinate id="5EB524F4C215DD2A0B2753C4AE94A029" direction="west" orientation="longitude" value="-48.466667">48°28W</geoCoordinate>
,
<date id="9AA9AEF4BC53B3C73CC0A9782F064E95" day="28" month="03" value="2005-03-28" year="2005">
<collectingDate id="1F77E8DDCD25C466E11E649F80EB92F5" day="28" month="03" value="2005-03-28" year="2005">28 Mar 2005</collectingDate>
</date>
.
</materialsCitation>
<materialsCitation id="6278523F81C88403AC2016F6DB9D917F" ID-GBIF-Occurrence="923930452" collectingDate="2003-10-07" collectionCode="UNT" collectorName="NEAMB-UFT staff" country="Brazil" latitude="-8.75" location="Tupirama, rio Tocantins next to the confluence with the rio Tranqueira" longitude="-48.133335" specimenCode="UNT 5577" stateProvince="Tocantins" typeStatus="Paratypes">
<specimenCode id="6930F91BAC778D69F06CB49C1CB84C61">
<collectionCode id="F48A671BE8CB372CBB5681954EDCB49F">UNT</collectionCode>
5577
</specimenCode>
, 3, 27.3-29.5 mm SL,
<location id="2ABD8E9F5627D2AD120FA5A1B0ECF28E" LSID="urn:lsid:plazi:treatment:077561934980E46F9CE3FAA30CEF9CB8:2ABD8E9F5627D2AD120FA5A1B0ECF28E" latitude="-8.75" longitude="-48.133335" name="Tupirama, rio Tocantins next to the confluence with the rio Tranqueira">Tupirama, rio Tocantins next to the confluence with the rio Tranqueira</location>
, approximately
<geoCoordinate id="D4F5549CED07013F04FBA36442BDF2D4" direction="south" orientation="latitude" value="-8.75">8°45S</geoCoordinate>
<geoCoordinate id="85A09454144BCC518E432FEB1C358508" direction="west" orientation="longitude" value="-48.133335">48°8W</geoCoordinate>
,
<date id="3C79B9DD5B736DB87091277913E998E0" day="07" month="10" value="2003-10-07" year="2003">
<collectingDate id="5D47507D7ED6F5098B0C9CB07A2366F8" day="07" month="10" value="2003-10-07" year="2003">7 Oct 2003</collectingDate>
</date>
.
</materialsCitation>
<materialsCitation id="0E4921DC1ABE370857833F0BED639226" ID-GBIF-Occurrence="923930450" collectingDate="2003-10-02" collectionCode="UNT" collectorName="NEAMB-UFT staff" country="Brazil" latitude="-11.016666" location="Brejinho de Nazare, lagoa Pedra do Santo" longitude="-48.566666" specimenCode="UNT 5578" stateProvince="Tocantins" typeStatus="Paratype">
<specimenCode id="535CC30AB28AB42587C495AE8D548E66">
<collectionCode id="7AD5AC54F0B66315F737CB8A1131DA5F">UNT</collectionCode>
5578
</specimenCode>
, 1, 26.5 mm SL,
<location id="B10A38F644DE8C7D659F09D186A1D86E" LSID="urn:lsid:plazi:treatment:077561934980E46F9CE3FAA30CEF9CB8:B10A38F644DE8C7D659F09D186A1D86E" latitude="-11.016666" longitude="-48.566666" name="Brejinho de Nazare, lagoa Pedra do Santo">
Brejinho de
<normalizedToken id="CF942B9644B524AEB0159DC0BE055457" originalValue="Nazaré">Nazare</normalizedToken>
, lagoa Pedra do Santo
</location>
, approximately
<geoCoordinate id="BD46949F74EA50767D3866C2A941027A" direction="south" orientation="latitude" value="-11.016666">11°1S</geoCoordinate>
<geoCoordinate id="F3D037655F090209918E49E031EA85A7" direction="west" orientation="longitude" value="-48.566666">48°34W</geoCoordinate>
,
<date id="0020CB22194DDAF7411CA78A4BFAC87F" day="02" month="10" value="2003-10-02" year="2003">
<collectingDate id="126EB4CBA560C03CB32E10AC49EAB76E" day="02" month="10" value="2003-10-02" year="2003">2 Oct 2003</collectingDate>
</date>
.
</materialsCitation>
<materialsCitation id="F30516B2CB1BCD63ED54B825B748A835" ID-GBIF-Occurrence="923930461" collectingDate="2004-05-08" collectionCode="UNT" collectorName="NEAMB-UFT staff" country="Brazil" latitude="-5.7166667" location="Itaguatins, rio Tocantins next to the confluence with the rio Sao Domingos" longitude="-47.5" specimenCode="UNT 5579" stateProvince="Tocantins" typeStatus="Paratype">
<specimenCode id="F0B752FC542F93B8DA9B5D843EBDFC0B">
<collectionCode id="9E0D5DF3CDC7D295C3ECAE226087C951">UNT</collectionCode>
5579
</specimenCode>
, 1, 26.1 mm SL,
<location id="72257D25351E15BFEEAA5989FE6D99D8" LSID="urn:lsid:plazi:treatment:077561934980E46F9CE3FAA30CEF9CB8:72257D25351E15BFEEAA5989FE6D99D8" latitude="-5.7166667" longitude="-47.5" name="Itaguatins, rio Tocantins next to the confluence with the rio Sao Domingos">
Itaguatins, rio Tocantins next to the confluence with the rio
<normalizedToken id="A24650573409E6D756805638021D8ABF" originalValue="São">Sao</normalizedToken>
Domingos
</location>
, approximately
<geoCoordinate id="C92B4150D2737EE2B76213B1A3065AE2" direction="south" orientation="latitude" value="-5.7166667">5°43S</geoCoordinate>
<geoCoordinate id="30FCB774C1CE24E05CC496E210A434DD" direction="west" orientation="longitude" value="-47.5">47°30W</geoCoordinate>
,
<date id="D500AC37A779A67C6B06BDE2863C41A6" day="08" month="05" value="2004-05-08" year="2004">
<collectingDate id="70017D600B1B97CB981027FB0D1450B5" day="08" month="05" value="2004-05-08" year="2004">8 May 2004</collectingDate>
</date>
.
</materialsCitation>
<materialsCitation id="2C661A21CDED472788257F10721622C8" ID-GBIF-Occurrence="923930456" collectingDate="2007-02-16" collectionCode="MCP" collectorName="NEAMB-UFT staff" country="Brazil" latitude="-12.25" location="Peixe, rio Tocantins downstream UHE Peixe-Angical" longitude="-48.383335" specimenCode="MCP 41466" stateProvince="Tocantins" typeStatus="Paratypes">
<specimenCode id="CAFDE8094FFE921143E91F5433EE2434">
<collectionCode id="557BFCBDA86708397C9C5A536E98A3CF">MCP</collectionCode>
41466
</specimenCode>
, 20, 32.1-36.0, and
</materialsCitation>
<materialsCitation id="B1B28993957309859389D2520964D369" ID-GBIF-Occurrence="923930458" collectingDate="2007-02-16" collectionCode="UNT" collectorName="NEAMB-UFT staff" country="Brazil" latitude="-12.25" location="Peixe, rio Tocantins downstream UHE Peixe-Angical" longitude="-48.383335" specimenCode="UNT 7610" stateProvince="Tocantins" typeStatus="Paratypes">
<specimenCode id="AA7A95F909C1CC4099A7AB01AC6E1EDE">
<collectionCode id="434A65AF7B50784CBC6E8C834287A6BA">UNT</collectionCode>
7610
</specimenCode>
, 132, 6 c&amp;s, 32.7-40.3 mm SL, collected with the holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="6B8C27DE5988C477D4E5B5C90C95EB6F" type="diagnosis">
<paragraph id="669ECBF54F2FC2D80DDF9B3F8B6AF622" lastPageNumber="55" pageNumber="54">
Diagnosis.
<taxonomicName id="7D8E1969355DC946D669CECFC5068C0B" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="elassonaktis">Jupiaba elassonaktis</taxonomicName>
is distinguished from
<taxonomicName id="CF83673EDC57A2E295772E3BAA543968" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="acanthogaster">J. acanthogaster</taxonomicName>
,
<taxonomicName id="55DE90670F6877B6D3BC94C3A25D4107" LSID-ZBK="urn:lsid:zoobank.org:act:FF6EBA3B-12FC-4EFD-BF0B-D24139D5AAD3" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Jupiaba atypindi Zanata 1997:111" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="atypindi">J. atypindi</taxonomicName>
,
<taxonomicName id="711001FF78A71765D0B48D287FE4BD5D" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="keithi">J. keithi</taxonomicName>
,
<taxonomicName id="9322004D49BA93F0B7EDC0FC6686803B" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="maroniensis">J. maroniensis</taxonomicName>
,
<taxonomicName id="DCEE7FA4CE815993EB78224A643E7E0F" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="meunieri">J. meunieri</taxonomicName>
,
<taxonomicName id="C3142E124EBE30B218980C91C468D1BE" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="minor">J. minor</taxonomicName>
,
<taxonomicName id="0F9C8AE7E3B0D6D80B9BDA136134A60C" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="pinnata">J. pinnata</taxonomicName>
, and
<taxonomicName id="D54C290A269D1BD077F2AA76788C919F" LSID-ZBK="urn:lsid:zoobank.org:act:9820235E-233B-4FDE-A62C-E1DE7D42BF22" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Jupiaba poekotero Zanata &amp; Lima 2005:272" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="poekotero">J. poekotero</taxonomicName>
by the medial cusps of premaxillary teeth much larger than remaining cusps (vs. cusps similar in size to each other) and by dentary teeth decreasing abruptly in size from the fifth tooth backwards (vs. decreasing gradually).
<taxonomicName id="29BB792CDD57AFCC297257BEC9703293" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="elassonaktis">Jupiaba elassonaktis</taxonomicName>
is distinguished from
<taxonomicName id="BB09B8C009AD0FA553D1D694FEC716A0" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="abramoides">J. abramoides</taxonomicName>
,
<taxonomicName id="965AE2DA00D83C2EF5567EE7AC9F3CD5" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="anteroides">J. anteroides</taxonomicName>
,
<taxonomicName id="B3FBBFCF6FD6299681B8F9E064E7BDEE" LSID-ZBK="urn:lsid:zoobank.org:act:F13828FC-18DD-426C-8F33-8DA772545C2A" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Jupiaba apenima Zanata 1997:108" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="apenima">J. apenima</taxonomicName>
,
<taxonomicName id="1E3F70B771BE13E44A35D227A899682D" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="asymmetrica">J. asymmetrica</taxonomicName>
,
<taxonomicName id="C80E13BC1D02258657DFE37DBA5A5BDD" LSID-ZBK="urn:lsid:zoobank.org:act:7536EF5A-CA7A-4B6B-999C-B46E930A2E2B" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Jupiaba pirana Zanata 1997:114" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="pirana">J. pirana</taxonomicName>
,
<taxonomicName id="682E5E3F2CAA1CEFAD1626AD978DF557" LSID-ZBK="urn:lsid:zoobank.org:act:1DD0F743-EB8E-4C5B-AFA8-D0104A4E3883" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Jupiaba poranga Zanata 1997:115" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="poranga">J. poranga</taxonomicName>
,
<taxonomicName id="EE3449D1AA81DE51233E96CE9CDB91A3" LSID-ZBK="urn:lsid:zoobank.org:act:5EBA9FA2-03E3-40EE-86FF-3091412BA350" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Jupiaba yarina Zanata 1997:119" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="yarina">J. yarina</taxonomicName>
, and
<taxonomicName id="66373101BF371E6559A32CD89B2BFAAD" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="zonata">J. zonata</taxonomicName>
by the lower number of scales on lateral line (33-39 vs. 40-66, respectively).
<taxonomicName id="602DF8F80D7693486A972E926217880D" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="elassonaktis">Jupiaba elassonaktis</taxonomicName>
is distinguished from
<taxonomicName id="2FFF24F63CEB586F5BB94DD004CB4161" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="ocellata">J. ocellata</taxonomicName>
by the first dorsal- and anal-fin rays not prolonged by an elongate filament (vs. prolonged by an elongate filament) and from
<taxonomicName id="9B53F0939A4372AB7E8055025E5D8F3F" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="mucronata">J. mucronata</taxonomicName>
by the presence of a vertically elongate humeral spot (vs. humeral spot horizontally elongate). The number of scales series above lateral line distinguishes
<taxonomicName id="CA08CDDFF11DA14135EC5E229A9B47B6" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="elassonaktis">J. elassonaktis</taxonomicName>
from
<taxonomicName id="8E64E45404EEE91ED66FD0E7C845236A" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="polylepis">J. polylepis</taxonomicName>
,
<taxonomicName id="59FD6D9A4DDD87129511A559B9C6E83B" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="potaroensis">J. potaroensis</taxonomicName>
, and
<taxonomicName id="744C0D1DDE8F362BBABEA7B3B958D689" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="zonata">J. zonata</taxonomicName>
(5-7 vs. 8-11, respectively). The number of pectoral-fin rays distinguishes
<taxonomicName id="47FC7F7F97E6F71A90CDBA99D5A12603" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="elassonaktis">J. elassonaktis</taxonomicName>
from
<taxonomicName id="DBEC3ED50FB9C5B6BBBC54A1CBC09F16" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="essequibensis">J. essequibensis</taxonomicName>
and
<taxonomicName id="CCE0F8BCCF7DE988F91BB636D3D19DE1" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="scologaster">J. scologaster</taxonomicName>
(9-11 vs. 13-16, respectively).
<taxonomicName id="5CEAF343CFE2CE7875B294C8D4E90BCB" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="elassonaktis">Jupiaba elassonaktis</taxonomicName>
is also distinguished from
<taxonomicName id="6D88BDA95664927F30D63852C57A1CA2" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="essequibensis">J. essequibensis</taxonomicName>
and
<taxonomicName id="7973C9EB99D1B57B21F148383D36DC57" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="scologaster">J. scologaster</taxonomicName>
by the incomplete series of median predorsal scales (vs. complete in later species). Futhermore,
<taxonomicName id="E325CC29D2C0980E1545C4747D62DA45" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="elassonaktis">J. elassonaktis</taxonomicName>
is also distinguished from
<taxonomicName id="2D0EFCE29D04B69374D9E81774A07F17" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="polylepis">J. polylepis</taxonomicName>
and
<taxonomicName id="F0849AFECF05CD2B72A3F2248324DA37" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="essequibensis">J.essequibensis</taxonomicName>
by the number of humeral spots.
<taxonomicName id="A85F4AAF2C1C51D848E211B60469F3D7" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="elassonaktis">Jupiaba elassonaktis</taxonomicName>
possesses two humeral spots. First humeral spot is large and vertically elongate located at fifth and sixth scales of lateral line and the second humeral spot very diffuse.
<taxonomicName id="A4532E2696DBF1AE85ACF283E351F08D" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="polylepis">Jupiaba polylepis</taxonomicName>
and
<taxonomicName id="B9F4D51EA848FE1E96ADA6ACBDC09383" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="essequibensis">J. essequibensis</taxonomicName>
exhibit only one humeral spot (surrounded by a light area and located on sixth to nineth lateral line scales in
<taxonomicName id="88726036D110F7480A22EAE65CA1EC3C" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="54" phylum="Chordata" rank="species" species="polylepis">J. polylepis</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="640D752269ACA0723A0201F3856E3CE8" type="description">
<paragraph id="3E37E0B2567B4B80565CA4716A1A6655" pageNumber="55">Description. Morphometric data summarized in Table 1. Body compressed and somewhat deep; greatest body depth at base of first dorsal-fin ray. Dorsal head profile straight or slightly concave. Profile convex to nearly straight from supraocciptal tip to base of last dorsal-fin ray, and straight towards adipose-fin origin. Anterior portion of ventral profile of head oblique. Posterior portion of ventral profile of head convex. Ventral body profile convex from pectoral-fin origin to pelvic-fin origin, and straight to slightly concave towards anal-fin origin. Body profile along anal-fin base posterodorsally slanted. Caudal peduncle nearly straight to slightly concave along dorsal and ventral margins. Snout rounded from margin of upper lip to vertical through anterior nostrils. Mouth terminal. Distal tip of maxilla anterior to vertical through center of orbit. Anterodorsal border of maxilla straight and posterodorsal border slightly concave. Anteroventral border straight and posteroventral borders convex. Maxilla slightly widened posteriorly.</paragraph>
<paragraph id="1F6A03A824CE489303E8C1FF3C7F666A" pageNumber="55">Two tooth rows on premaxilla. Outer row with three pentacuspid teeth with central cusp longer. Inner row with five teeth, gradually decreasing in length from first to third teeth; all teeth with seven cusps, except last hexacuspidate tooth. First tooth with third cusp twice or three times longer and broader than other cusps. Second, third, and fourth teeth with central cusp twice or three times longer and broader than other cusps. Fifth tooth with fourth cusp twice or three times longer and broader than other cusps. Maxilla with two, three or four teeth; first and second teeth tricuspid, tetracuspid or pentacuspid; third teeth truncate, conical or tricuspid; fourth teeth conical. Five anteriormost dentary teeth larger than remaining teeth. First tooth heptacuspid; Second to fourth teeth with six cusps, followed by one truncate medium-sized tooth, and seven tricuspid teeth; central cusp in all teeth two to three times longer and broader than other cusps. All cusp tips slightly curved posteriorly.</paragraph>
<paragraph id="07BB6766C5EAA6AC03CE10B1AA38B63E" lastPageNumber="56" pageNumber="55">Dorsal-fin rays ii, 8[20], 9 [21], 10[5] (n = 46; mean = 8.7); first unbranched ray approximately one-half length of second ray. Dorsal-fin origin located anterior to middle of SL and in same vertical through pelvic-fin origin. Adipose-fin located approximately at vertical through insertion of base of 17th -19th anal-fin rays. Anal-fin rays iii -iv, 18[7], 19[12], 20[14], 21[11], 22[3] (n = 46; mean = 19.8). Anal-fin profile concave. Anal fin origin anterior to vertical through base of last dorsal-fin ray. Pectoral-fin rays i, 9[1], 10[20], 11[25] (n = 46; mean = 10.5). Pectoral-fin tip not reaching pelvic-fin origin. Pelvic-fin rays i, 7 (n = 46). Pelvic-fin origin located anterior to vertical through dorsal-fin origin. Pelvic bone modified into strong spine. Anterior tip of two pelvic bones diverging widely and usually exserted from body wall. Dorsal procurrent rays 11 [1]. Ventral procurrent rays 9 [1]. Scales cycloid, moderately large. Lateral line complete. Scales in longitudinal series 33[4], 34[16], 35[15], 36[4], 37[1], 39[1] (n = 41; mean = 34.7). Scale rows between dorsal-fin origin and lateral line 5[4], 6[12], 7[28] (n = 44; mean = 6.5); scale rows between lateral line and pelvic fin origin 4[4], 5[38], 6[2] (n = 44; mean = 5.0). Precaudal vertebrae 15 [1], 16 [1]; caudal vertebrae 19 [2]; total vertebrae 34 [1], 35 [1]; supraneurals 4 [1], 5[1]. Gill-rakers 7+1/8 [2].</paragraph>
<paragraph id="4B1217724CFDB26D99A2A7DB45EE94BF" lastPageNumber="57" pageNumber="56">Color in alcohol. Ground color pale yelow. Head dorsum light gray. Two humeral spots separated by a light area. First humeral spot dark brown large and vertically elongate, trespassing upon fifth and sixth lateralline scales and extending over eight or nine horizontal series of scales, including lateral line; center of first humeral spot forming a circular darker spot inside. Second humeral spot very diffuse, trespassing upon ninth and tenth lateral-line scales and extending over eight to ten horizontal series of scales, including lateral line. Second humeral spot not visible in some specimens. Flanks with silvery narrow stripe extending from posterior opercular border to caudal spot. Patch of black chromatophores on caudal peduncle continued by black spot extending over middle caudal-fin rays. Yellow pigmentation on base of outer edge of caudal-fin rays. Remaining fins lacking distinctive patches of pigmentation (Fig. 1).</paragraph>
<paragraph id="BE249CC45509DFEAA4922332231C8AED" pageNumber="57">Sexual dimorphism. No apparent sexual dimorphism was observed.</paragraph>
</subSubSection>
<subSubSection id="491AB7F5F3BF0C8FA0C40B3E2949B6AB" type="etymology">
<paragraph id="B4EE373A5E50461B05F3801FE3B1A922" pageNumber="57">Etymology. The specific epithet elassonaktis, from the Greek elasson, meaning few, and aktis, meaning rays, is in allusion to the small number of anal-fin rays.</paragraph>
</subSubSection>
<subSubSection id="C8788531A53783AAB83463AAD263E5B8" type="distribution">
<paragraph id="0A36CD8E2BEAFDDD7FF77FD781D94E9E" pageNumber="57">
Distribution.
<taxonomicName id="BBC3D489C1911308BE396684EDFA0BEE" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="elassonaktis">Jupiaba elassonaktis</taxonomicName>
is known from medium and upper portions of rio Tocantins drainage, Brazil (Fig. 2).
</paragraph>
</subSubSection>
<subSubSection id="E402DE377F9FE484D1EA078BF597BA60" type="discussion">
<paragraph id="7D03429FD36C1A07E3863A2EE35EEE92" pageNumber="57">Discussion</paragraph>
<paragraph id="1A8F8BB957BE78180BE6B6D5513759E1" pageNumber="57">
We herein describe a new species in
<taxonomicName id="9E7A3A2E634436940B403E20EB221DDC" LSID-ZBK="urn:lsid:zoobank.org:act:245EB3ED-92DD-4953-8C1B-5F3E7720338F" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Jupiaba Zanata 1997:102" order="Characiformes" pageNumber="57" phylum="Chordata" rank="genus">Jupiaba</taxonomicName>
, since it conforms to the definition of Zanata (1997), who established the genus on the basis of six putative synapomorphies associated with the elongation of the pelvic spine-like bones. On the other hand, the intrageneric phylogenetic relationships of
<taxonomicName id="CF968DFC71094448055EEA52D67B1B48" LSID-ZBK="urn:lsid:zoobank.org:act:245EB3ED-92DD-4953-8C1B-5F3E7720338F" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Jupiaba Zanata 1997:102" order="Characiformes" pageNumber="57" phylum="Chordata" rank="genus">Jupiaba</taxonomicName>
species are not known. However, Zanata &amp; Lima (2005) artificially divided the genus in two distinct species-group on the basis of tooth morphology: (1) species with premaxillary and dentary teeth somewhat compressed, with cusps similar in size to each other and dentary teeth decreasing gradually in size; (2) species with more robust teeth bearing one largest median cusp and small lateral ones, and larger dentary teeth anteriorly followed by distinctly smaller teeth.
<taxonomicName id="E86CEFE3C7DBE4A0F80AE806C2BF49B4" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="elassonaktis">Jupiaba elassonaktis</taxonomicName>
belongs to the second species-group, which also contains
<taxonomicName id="729AD8A9059F5469E0A91340E8C064A8" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="mucronata">J. mucronata</taxonomicName>
,
<taxonomicName id="03ED533773A462A3C60F35D27D457B67" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="essequibensis">J. essequibensis</taxonomicName>
,
<taxonomicName id="B230768248E12A701B88E4125C2E303C" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="scologaster">J. scologaster</taxonomicName>
,
<taxonomicName id="52A6744C3F2C5B0E320D34E1F0FDB467" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="ocellata">J. ocellata</taxonomicName>
,
<taxonomicName id="DA4819BC2063BAEAACB7D20FA1DECD9A" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="polylepis">J. polylepis</taxonomicName>
,
<taxonomicName id="73A3DEC23088D2C72D5D1E0B421A7203" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="anteroides">J. anteroides</taxonomicName>
,
<taxonomicName id="1F7BE15A99D2EDF6AE706BEA5106C7F5" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="abramoides">J. abramoides</taxonomicName>
,
<taxonomicName id="F51969953FB6DCA53EFE86C20A60961D" LSID-ZBK="urn:lsid:zoobank.org:act:1DD0F743-EB8E-4C5B-AFA8-D0104A4E3883" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Jupiaba poranga Zanata 1997:115" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="poranga">J. poranga</taxonomicName>
,
<taxonomicName id="5750D3869DDD254115F8055A8D227151" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="potaroensis">J. potaroensis</taxonomicName>
,
<taxonomicName id="B336FBF5981291EA319F272C9DCC34EB" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="zonata">J. zonata</taxonomicName>
,
<taxonomicName id="9D8C8CEE602346FFF82C37B9C91D92C1" LSID-ZBK="urn:lsid:zoobank.org:act:7536EF5A-CA7A-4B6B-999C-B46E930A2E2B" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Jupiaba pirana Zanata 1997:114" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="pirana">J. pirana</taxonomicName>
,
<taxonomicName id="63E4C721D6B7CFBA8EAAB53AEC43C7AB" LSID-ZBK="urn:lsid:zoobank.org:act:5EBA9FA2-03E3-40EE-86FF-3091412BA350" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Jupiaba yarina Zanata 1997:119" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="yarina">J. yarina</taxonomicName>
,
<taxonomicName id="6674A5180C19CDE4DB05B79A677EC19E" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="asymmetrica">J. asymmetrica</taxonomicName>
, and
<taxonomicName id="CF9A753FC5F03998664B54C880384460" LSID-ZBK="urn:lsid:zoobank.org:act:F13828FC-18DD-426C-8F33-8DA772545C2A" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Jupiaba apenima Zanata 1997:108" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="apenima">J. apenima</taxonomicName>
. Nonetheless, the lack of phylogenetic information precludes any unambiguous assertions on the possible relationships of the new species, given that it is unwarranted to assume that these similarities effectively reflect common ancestry. This condition also prevents any reasonable biogeographic inferences.
</paragraph>
<paragraph id="7E1E47C22EF65E1F9424FF63B78FD9C4" pageNumber="57">
Two other
<taxonomicName id="371FCD3B159B00DDE47D1CAF1A569A8B" LSID-ZBK="urn:lsid:zoobank.org:act:245EB3ED-92DD-4953-8C1B-5F3E7720338F" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Jupiaba Zanata 1997:102" order="Characiformes" pageNumber="57" phylum="Chordata" rank="genus">Jupiaba</taxonomicName>
species inhabit the rio Tocantins drainage, namely
<taxonomicName id="74B983395E9E5F5334624EE8CF881CC8" LSID-ZBK="urn:lsid:zoobank.org:act:F13828FC-18DD-426C-8F33-8DA772545C2A" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Jupiaba apenima Zanata 1997:108" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="apenima">J. apenima Zanata</taxonomicName>
and
<taxonomicName id="665A4B9A909BF136331330E46A2E14E5" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="polylepis">
J. polylepis
<normalizedToken id="F7D4CBB49D8A1B95804F8008C3BA7636" originalValue="Günther">Guenther</normalizedToken>
</taxonomicName>
.
<taxonomicName id="F418305A4F39421B28C3F19463DB201C" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="elassonaktis">Jupiaba elassonaktis</taxonomicName>
is distinguished [[from]]
<taxonomicName id="98AD33ADFBB1E35F75C53041BAB964C8" LSID-ZBK="urn:lsid:zoobank.org:act:F13828FC-18DD-426C-8F33-8DA772545C2A" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Jupiaba apenima Zanata 1997:108" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="apenima">J. apenima</taxonomicName>
by the lower number of scales on lateral line (33- 39 vs. 55-59, respectively). The number of scales series above lateral line distinguishes
<taxonomicName id="2ED881C86696AD4A452DBF37B2B2F687" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="elassonaktis">J. elassonaktis</taxonomicName>
from
<taxonomicName id="7F77D93AF476B6905E70BF4E5E8308A4" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="polylepis">J. polylepis</taxonomicName>
(5-7 vs. 8-11, respectively). Futhermore,
<taxonomicName id="3694845C95114A4079E7D3B0C8E8B0E5" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="elassonaktis">J. elassonaktis</taxonomicName>
is also distinguished from
<taxonomicName id="8BA7086D098FF263EF54A2C12AAED691" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="polylepis">J. polylepis</taxonomicName>
by the number of humeral spots.
<taxonomicName id="C38D27936BB1AEB4408AB0660C3C3C14" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="elassonaktis">Jupiaba elassonaktis</taxonomicName>
possesses two humeral spots. First humeral spot is large and vertically elongate located at fifth and sixth scales of lateral line and the second humeral spot very diffuse.
<taxonomicName id="613992A41AF0846AF55C8E9A178F7032" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="polylepis">Jupiaba polylepis</taxonomicName>
exhibit only one humeral spot surrounded by a light area and located on sixth to nineth lateral line scales.
</paragraph>
<paragraph id="1A11E7C87491D4137DB4A15B9FA5B3CE" pageNumber="57">
<taxonomicName id="E302DAEDFBB322880EE2F2E622652146" family="Characidae" genus="Jupiaba" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="elassonaktis">Jupiaba elassonaktis</taxonomicName>
is known from upper and medium portions of the rio Tocantins drainage and probably is an additional example of endemism in this basin. The rio Tocantins drainage is an area of endemism for several Neotropical freshwater fish groups including new recently described characid species such as
<taxonomicName id="AB4AA510D64741F4B11BD54A49F6A262" LSID-ZBK="urn:lsid:zoobank.org:act:D7842DE6-AA75-43C5-8032-A77F992D1771" family="Characidae" genus="Hyphessobrycon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hyphessobrycon hamatus Bertaco &amp; Malabarba 2005:84" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="hamatus">Hyphessobrycon hamatus</taxonomicName>
,
<taxonomicName id="99F2175AC40607EBF2374E53328681D9" LSID-ZBK="urn:lsid:zoobank.org:act:E1334B2A-947E-4C19-BA1F-7745DCDAC7AA" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax elachylepis Bertaco &amp; Lucinda 2005" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="elachylepis">Astyanax elachylepis</taxonomicName>
,
<taxonomicName id="E303EF11F451FD9AF2938ABEF98EA569" LSID-ZBK="urn:lsid:zoobank.org:act:44E642B2-3E43-4F0C-AD6B-27B45C78F565" family="Characidae" genus="Moenkhausia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Moenkhausia pankilopteryx Bertaco &amp; Lucinda 2006" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="pankilopteryx">Moenkhausia pankilopteryx</taxonomicName>
, and
<taxonomicName id="8963475D6011B706C132D5CFDB83BD83" family="Characidae" genus="Moenkhausia" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="57" phylum="Chordata" rank="species" species="hysterosticta">Moenkhausia hysterosticta</taxonomicName>
(Bertaco &amp; Malabarba, 2005; Bertaco &amp; Lucinda, 2005, 2006; Lucinda et al., 2007). Especially in its upper regions, the rio Tocantins drainage appears to be a highly endemic center for the Ancistrini, as shown by the presence of three recently described species of
<taxonomicName id="A27645DE8C7DBE663C14D22D64FEF78A" LSID-ZBK="urn:lsid:zoobank.org:act:2AC0D766-F761-4436-B6F4-395BBA997694" family="Loricariidae" genus="Hemiancistrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Hemiancistrus Bleeker 1862:2" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="genus">Hemiancistrus Bleeker</taxonomicName>
(Cardoso &amp; Lucinda, 2003), and by four endemic species of
<taxonomicName id="F0615F1FDBE3408AE9B4F11F7C56FFA4" LSID-ZBK="urn:lsid:zoobank.org:act:35063369-E231-4E03-A49C-EE29CD36526F" family="Loricariidae" genus="Ancistrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ancistrus Kner 1854:272" order="Siluriformes" pageNumber="57" phylum="Chordata" rank="genus">Ancistrus Kner</taxonomicName>
(Fisch-Muller et al., 2001, 2005).
</paragraph>
</subSubSection>
</treatment>
</document>