302 lines
40 KiB
XML
302 lines
40 KiB
XML
<document id="22229447617EEE214974FE14A06976D7" ID-CLB-Dataset="36856" ID-DOI="10.11646/zootaxa.3795.1.9" ID-GBIF-Dataset="d113b7c3-8cf5-4814-be3a-c47535935da1" ID-ISSN="1175-5326" ID-Zenodo-Dep="286047" ID-ZooBank="03AFA16A-9DED-4338-A2CF-B0A0515AB7CD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460417749826" checkinUser="plazi" docAuthor="López-García, Margarita M. & Méndez-Rojas, Diana M." docDate="2014" docId="07363D27FF8BFFDAFF36FC29990CFC5A" docLanguage="en" docName="zt03795p090.pdf" docOrigin="Zootaxa 3795 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Linoderus navarretei Lopez-Garcia & Mendez-Rojas, new species" docType="treatment" docVersion="7" lastPageNumber="89" masterDocId="FB0F455FFF8AFFDEFFA1FF8D9A75FFFE" masterDocTitle="A new species of the genus Linoderus Sharp, 1885 (Coleoptera, Staphylinidae, Philonthina) from the Colombian Andes" masterLastPageNumber="90" masterPageNumber="85" pageNumber="86" updateTime="1698329171632" updateUser="ExternalLinkService">
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<mods:titleInfo id="D34B36E5D2A615D798257260A49DB521">
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<mods:title id="3DC9F8D35C16BA643971E114456B0019">A new species of the genus Linoderus Sharp, 1885 (Coleoptera, Staphylinidae, Philonthina) from the Colombian Andes</mods:title>
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<mods:name id="C937C1A417AD7A9D89E791882C6432B3" type="personal">
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<mods:namePart id="4B676208D65DA534F1E0EBC31D43D869">López-García, Margarita M.</mods:namePart>
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<mods:name id="313CC31F1AE16FDD536FD2712435EACD" type="personal">
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<mods:roleTerm id="8ECC60F5942388B41B3A111A4A17DD5D">Author</mods:roleTerm>
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<mods:namePart id="BCC072BB27AA19F420E36BCA226E2E88">Méndez-Rojas, Diana M.</mods:namePart>
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<mods:typeOfResource id="85659A0B41716E327B6FEAC562C9D9E9">text</mods:typeOfResource>
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<mods:title id="EAAE5640071729DBFC75EA2F73A113A7">Zootaxa</mods:title>
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<mods:part id="5ABFB78FD0F2AC6C3C30406FAF35F6AD">
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<mods:date id="E1C825C29E9644044C00FD6190CDB291">2014</mods:date>
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<mods:number id="2CE10BD78922606F58ACCBDD94D7343B">3795</mods:number>
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<mods:classification id="CCF478DA2733D3013A4FFC1E4E52E598">journal article</mods:classification>
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<mods:identifier id="322450A620677E1FFAC79DE243E5ED8F" type="DOI">10.11646/zootaxa.3795.1.9</mods:identifier>
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<treatment id="07363D27FF8BFFDAFF36FC29990CFC5A" ID-DOI="http://doi.org/10.5281/zenodo.6133669" ID-GBIF-Taxon="127667816" ID-Zenodo-Dep="6133669" LSID="urn:lsid:plazi:treatment:07363D27FF8BFFDAFF36FC29990CFC5A" httpUri="http://treatment.plazi.org/id/07363D27FF8BFFDAFF36FC29990CFC5A" lastPageId="4" lastPageNumber="89" pageId="1" pageNumber="86">
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<subSubSection id="C785DFBAFF8BFFDFFF36FC2999B6FC40" box="[151,963,932,958]" pageId="1" pageNumber="86" type="nomenclature">
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<paragraph id="8F208C31FF8BFFDFFF36FC2999B6FC40" blockId="1.[151,963,932,958]" box="[151,963,932,958]" pageId="1" pageNumber="86">
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<heading id="D4683B5DFF8BFFDFFF36FC2999B6FC40" bold="true" box="[151,963,932,958]" fontSize="11" level="1" pageId="1" pageNumber="86" reason="1">
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<emphasis id="BDEB5023FF8BFFDFFF36FC2999B6FC40" bold="true" box="[151,963,932,958]" pageId="1" pageNumber="86">
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<taxonomicName id="489FF7B2FF8BFFDFFF36FC299953FC40" ID-CoL="3V3RZ" authority="Lopez-Garcia & Mendez-Rojas" authorityName="Lopez-Garcia & Mendez-Rojas" box="[151,806,932,958]" class="Insecta" family="Staphylinidae" genus="Linoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="86" phylum="Arthropoda" rank="species" species="navarretei" status="sp. nov.">
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<emphasis id="BDEB5023FF8BFFDFFF36FC299BE3FC40" bold="true" box="[151,406,932,958]" italics="true" pageId="1" pageNumber="86">Linoderus navarretei</emphasis>
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López-García & Méndez-Rojas
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</taxonomicName>
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,
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<taxonomicNameLabel id="A6D8ED58FF8BFFDFFC92FC2999B6FC40" box="[819,963,932,958]" pageId="1" pageNumber="86" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C785DFBAFF8BFFDFFF36FC679F79FB48" pageId="1" pageNumber="86" type="diagnosis">
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<paragraph id="8F208C31FF8BFFDFFF36FC679F79FB48" blockId="1.[151,1437,1002,2035]" pageId="1" pageNumber="86">
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<emphasis id="BDEB5023FF8BFFDFFF36FC679B64FBFD" bold="true" box="[151,273,1002,1027]" pageId="1" pageNumber="86">Diagnosis.</emphasis>
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<taxonomicName id="489FF7B2FF8BFFDFFEBBFC669872FBFC" box="[282,519,1003,1027]" class="Insecta" family="Staphylinidae" genus="Linoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="86" phylum="Arthropoda" rank="species" species="navarretei" status="sp. nov.">
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<emphasis id="BDEB5023FF8BFFDFFEBBFC669872FBFC" box="[282,519,1003,1027]" italics="true" pageId="1" pageNumber="86">Linoderus navarretei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A6D8ED58FF8BFFDFFDB0FC679819FBFD" box="[529,620,1002,1027]" pageId="1" pageNumber="86" rank="species">
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<emphasis id="BDEB5023FF8BFFDFFDB0FC679819FBFD" bold="true" box="[529,620,1002,1027]" pageId="1" pageNumber="86">sp. nov.</emphasis>
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</taxonomicNameLabel>
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may be easily distinguished from
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<taxonomicName id="489FF7B2FF8BFFDFFC5BFC669EFCFBFD" box="[1018,1161,1003,1027]" class="Insecta" family="Staphylinidae" genus="Linoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="86" phylum="Arthropoda" rank="species" species="gracilipes">
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<emphasis id="BDEB5023FF8BFFDFFC5BFC669EFCFBFD" box="[1018,1161,1003,1027]" italics="true" pageId="1" pageNumber="86">L. gracilipes</emphasis>
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</taxonomicName>
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by its color pattern and other characters. The new species has legs and body black, except by abdomen apex golden orange and apex of antenna pale yellow.
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<taxonomicName id="489FF7B2FF8BFFDFFE2AFBBE986EFBB5" box="[395,539,1075,1099]" class="Insecta" family="Staphylinidae" genus="Linoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="86" phylum="Arthropoda" rank="species" species="gracilipes">
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<emphasis id="BDEB5023FF8BFFDFFE2AFBBE986EFBB5" box="[395,539,1075,1099]" italics="true" pageId="1" pageNumber="86">L. gracilipes</emphasis>
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</taxonomicName>
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has head and elytra teal dark; pronotum, legs and abdomen until apex of VII orange red, apex of abdomen black, antenna black with the three basal and the two apical segments yellow. Styli of
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<taxonomicName id="489FF7B2FF8BFFDFFF36FBF69B52FB6D" box="[151,295,1147,1171]" class="Insecta" family="Staphylinidae" genus="Linoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="86" phylum="Arthropoda" rank="species" species="gracilipes">
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<emphasis id="BDEB5023FF8BFFDFFF36FBF69B52FB6D" box="[151,295,1147,1171]" italics="true" pageId="1" pageNumber="86">L. gracilipes</emphasis>
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</taxonomicName>
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are more dilated apically than those of
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<taxonomicName id="489FF7B2FF8BFFDFFD56FBF699FEFB6C" box="[759,907,1147,1170]" class="Insecta" family="Staphylinidae" genus="Linoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="86" phylum="Arthropoda" rank="species" species="navarretei" status="sp. nov.">
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<emphasis id="BDEB5023FF8BFFDFFD56FBF699FEFB6C" box="[759,907,1147,1170]" italics="true" pageId="1" pageNumber="86">L. navarretei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A6D8ED58FF8BFFDFFC35FBF69998FB6C" box="[916,1005,1147,1170]" pageId="1" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
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which are thinner; apical segment of maxillary palp twice longer than penultimate in
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<taxonomicName id="489FF7B2FF8BFFDFFD13FB12994AFB48" box="[690,831,1182,1206]" class="Insecta" family="Staphylinidae" genus="Linoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="86" phylum="Arthropoda" rank="species" species="gracilipes">
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<emphasis id="BDEB5023FF8BFFDFFD13FB12994AFB48" box="[690,831,1182,1206]" italics="true" pageId="1" pageNumber="86">L. gracilipes</emphasis>
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</taxonomicName>
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, while 1.3 times in
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<taxonomicName id="489FF7B2FF8BFFDFFBBCFB129EDBFB48" box="[1053,1198,1183,1206]" class="Insecta" family="Staphylinidae" genus="Linoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="86" phylum="Arthropoda" rank="species" species="navarretei" status="sp. nov.">
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<emphasis id="BDEB5023FF8BFFDFFBBCFB129EDBFB48" box="[1053,1198,1183,1206]" italics="true" pageId="1" pageNumber="86">L. navarretei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A6D8ED58FF8BFFDFFB14FB129F79FB48" box="[1205,1292,1183,1206]" pageId="1" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C785DFBAFF8BFFDDFF66FB4F9E7DF8D2" lastPageId="3" lastPageNumber="88" pageId="1" pageNumber="86" type="description">
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<paragraph id="8F208C31FF8BFFDFFF66FB4F9829FAB9" blockId="1.[151,1437,1002,2035]" pageId="1" pageNumber="86">
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<emphasis id="BDEB5023FF8BFFDFFF66FB4F9B2CFB25" bold="true" box="[199,345,1218,1243]" pageId="1" pageNumber="86">Description.</emphasis>
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Habitus as in
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<figureCitation id="17A490B4FF8BFFDFFDAEFB4F982DFB25" box="[527,600,1218,1243]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1932,1955]" captionTargetBox="[166,1424,365,1931]" captionTargetId="figure@2.[151,1436,337,1940]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Linoderus navarretei sp. nov. HOLOTYPE (1), PARATYPES (2 – 9). 1, Habitus; 2, head, 3, male sternum VIII; 4, male sternum IX; 5, aedeagus in dorsal view; 6, aedeagus in ventral view; 7, aedeagus in lateral view; 8, female tergum X; 9, gonocoxite of female genital segment. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/286048/files/figure.png" pageId="1" pageNumber="86">Fig. 1</figureCitation>
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. Body length 7.50–10.0 mm (until apex elytra
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<quantity id="486721D4FF8BFFDFFB35FB4E9F34FB25" box="[1172,1345,1219,1243]" metricMagnitude="-3" metricUnit="m" metricValue="3.995" metricValueMax="4.29" metricValueMin="3.7" pageId="1" pageNumber="86" unit="mm" value="3.995" valueMax="4.29" valueMin="3.7">3.70–4.29 mm</quantity>
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). Head, pronotum and elytra shiny black. Antennomeres 1–3 black, 4–6 dark brown, 7 light brown and 8–11 pale yellow. Abdomen shiny black except posterior 1/4 of VII segment, VIII and XI golden-yellow. Basal half of styli goldenyellow and distal half black. Legs black.
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</paragraph>
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<paragraph id="8F208C31FF8BFFDFFF66FADF99A7FA05" blockId="1.[151,1437,1002,2035]" pageId="1" pageNumber="86">Head slightly longer than wide (HL/HW:1.15–1.20), narrowed behind eyes until neck constriction and slightly wider than pronotum (HW/PW:1.23). Eyes distinctly shorter than temples seen from above (EL/TL:0.43–0.5). Epicranium with four interocular punctures bearing long and black setas, shaping a transverse line between the eyes; postocular region with a few similar punctures dispersed. Postocular ridge rudimentary, reaching the postgenal ridge but not extending beyond it; postmandibular ridge absent.</paragraph>
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<paragraph id="8F208C31FF8BFFDFFF66F98B99EDF878" blockId="1.[151,1437,1002,2035]" pageId="1" pageNumber="86">
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Antennal insertions separated from anterior margin of frontoclypeus and from eyes by a subequal distance; antennae slender, a bit longer than head and pronotum combined, the three basal segments elongated, segments 1 and 3 longer than 2, segments 4–7 cylindrical, the fourth shorter, segments 8–10 with apex slightly wider than base,, segment 11 dorsally emarginated; segments 1–3 glossy with a few long black setae, 4–11 pubescent and with a few white setae decreasing in length towards apex. Labrum deeply emarginated and bilobulated with two long black setae and six more of moderate length (
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<figureCitation id="17A490B4FF8BFFDFFD08F9379884F92D" box="[681,753,1722,1747]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1932,1955]" captionTargetBox="[166,1424,365,1931]" captionTargetId="figure@2.[151,1436,337,1940]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Linoderus navarretei sp. nov. HOLOTYPE (1), PARATYPES (2 – 9). 1, Habitus; 2, head, 3, male sternum VIII; 4, male sternum IX; 5, aedeagus in dorsal view; 6, aedeagus in ventral view; 7, aedeagus in lateral view; 8, female tergum X; 9, gonocoxite of female genital segment. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/286048/files/figure.png" pageId="1" pageNumber="86">Fig. 2</figureCitation>
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). Frontoclypeus with anterior edge truncate, with a median longitudinal fovea deeply (some males,
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<figureCitation id="17A490B4FF8BFFDFFDFFF95398D4F909" box="[606,673,1758,1783]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1932,1955]" captionTargetBox="[166,1424,365,1931]" captionTargetId="figure@2.[151,1436,337,1940]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Linoderus navarretei sp. nov. HOLOTYPE (1), PARATYPES (2 – 9). 1, Habitus; 2, head, 3, male sternum VIII; 4, male sternum IX; 5, aedeagus in dorsal view; 6, aedeagus in ventral view; 7, aedeagus in lateral view; 8, female tergum X; 9, gonocoxite of female genital segment. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/286048/files/figure.png" pageId="1" pageNumber="86">Fig. 2</figureCitation>
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) to slightly impressed. Maxilar palpi elongated, segment 1 minute, segment 2 more than thrice as long as maximum width and longer than segment 3, last segment subcylindrical, 1.2–1.3 longer than preceding segment; gular sutures fused on its lower half; mentum rectangular and shorter than submentum; labial palpi slender, segment 2 distinctly longer than 1 and 3. Neck no more than 0.4 times as wide as head at widest point (NW/HW:0.33–0.40); transverse carina present.
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</paragraph>
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<paragraph id="8F208C31FF8BFFDCFF66F81E9E49FEC1" blockId="1.[151,1437,1002,2035]" lastBlockId="2.[151,1436,151,320]" lastPageId="2" lastPageNumber="87" pageId="1" pageNumber="86">Pronotum longer than wide (PL/PW:1.30–1.39), widened apically; prosternal carina evident; superior line of hypomeron not completely visible in dorsal view; lateral puncture of pronotum bearing long macroseta separated from superior line by a distance three times as large as diameter of the puncture; dorsal surface of pronotal disk with two medial rows of six to seven punctures each. Legs slender, claws small, hind leg longer than abdomen, anterior femora with two short spines each, tibiae with spines and long hairs, tarsi elongated- nearly as long as tibiae- and with long whitish yellow hairs more abundant on first tarsomere, which is longer than tarsomeres 2 and 3 combined, hind tarsus with first segment longer than the last one (S1/S5=1.57–2.14), S1 a little longer in males (S1=0.70–0.73) than in females (S1=0.50–0.52); front tarsi undilated in both sexes.</paragraph>
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<caption id="DBE0DCB9FF88FFDCFF36F8019884F81E" httpUri="https://zenodo.org/record/286048/files/figure.png" pageId="2" pageNumber="87" targetBox="[166,1424,365,1931]" targetPageId="2">
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<paragraph id="8F208C31FF88FFDCFF36F8019884F81E" blockId="2.[151,1436,1932,2016]" pageId="2" pageNumber="87">
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<emphasis id="BDEB5023FF88FFDCFF36F8019B4CF85C" bold="true" box="[151,313,1932,1955]" pageId="2" pageNumber="87">FIGURES 1–9.</emphasis>
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<taxonomicName id="489FF7B2FF88FFDCFE9EF8009860F85D" box="[319,533,1933,1955]" class="Insecta" family="Staphylinidae" genus="Linoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="87" phylum="Arthropoda" rank="species" species="navarretei" status="sp. nov.">
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<emphasis id="BDEB5023FF88FFDCFE9EF8009860F85D" box="[319,533,1933,1955]" italics="true" pageId="2" pageNumber="87">Linoderus navarretei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A6D8ED58FF88FFDCFDBBF800981FF85C" box="[538,618,1933,1954]" pageId="2" pageNumber="87" rank="species">
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<emphasis id="BDEB5023FF88FFDCFDBBF800981FF85C" bold="true" box="[538,618,1933,1954]" pageId="2" pageNumber="87">sp. nov.</emphasis>
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</taxonomicNameLabel>
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HOLOTYPE (1), PARATYPES (2–9). 1, Habitus; 2, head, 3, male sternum VIII; 4, male sternum IX; 5, aedeagus in dorsal view; 6, aedeagus in ventral view; 7, aedeagus in lateral view; 8, female tergum X; 9, gonocoxite of female genital segment. Scale bars= 0.2 mm.
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</paragraph>
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</caption>
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<caption id="DBE0DCB9FF89FFDDFF36FA8C9EA6FAC8" httpUri="https://zenodo.org/record/286049/files/figure.png" pageId="3" pageNumber="88" targetBox="[151,1429,241,1254]" targetPageId="3">
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<paragraph id="8F208C31FF89FFDDFF36FA8C9EA6FAC8" blockId="3.[151,1436,1281,1334]" pageId="3" pageNumber="88">
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<emphasis id="BDEB5023FF89FFDDFF36FA8C9B22FAE9" bold="true" box="[151,343,1281,1304]" pageId="3" pageNumber="88">FIGURES 10–13.</emphasis>
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Habitat of
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<taxonomicName id="489FF7B2FF89FFDDFE73FA8F98DEFAE6" box="[466,683,1282,1304]" class="Insecta" family="Staphylinidae" genus="Linoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="88" phylum="Arthropoda" rank="species" species="navarretei" status="sp. nov.">
|
||
<emphasis id="BDEB5023FF89FFDDFE73FA8F98DEFAE6" box="[466,683,1282,1304]" italics="true" pageId="3" pageNumber="88">Linoderus navarretei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A6D8ED58FF89FFDDFD15FA8F9972FAE9" box="[692,775,1282,1303]" pageId="3" pageNumber="88" rank="species">
|
||
<emphasis id="BDEB5023FF89FFDDFD15FA8F9972FAE9" bold="true" box="[692,775,1282,1303]" pageId="3" pageNumber="88">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
10–11 under leaves (Photos by MMLG); 12,
|
||
<emphasis id="BDEB5023FF89FFDDFB4CFA8E9BFEFAC8" italics="true" pageId="3" pageNumber="88">Parque Regional Natural Barbas-Bremen</emphasis>
|
||
: forest, view from above; 13, riparian vegetation (Photos by O. H. Marín-Gómez).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8F208C31FF89FFDDFF66FAEE9B56FA16" blockId="3.[151,1437,1379,2016]" pageId="3" pageNumber="88">Elytra wider (EtW/PW=1.44–1.55) and longer than pronotum (EtL/PL=1.17–1.25), punctuation coarse and denser than that of pronotum, with white microsetae and black macrosetae, transversal distance between punctures shorter near the base of elytra; scutellum with margins glabrous and medial area with deep umbilicated punctuation.</paragraph>
|
||
<paragraph id="8F208C31FF89FFDDFF66FA7E9F6DF9D1" blockId="3.[151,1437,1379,2016]" pageId="3" pageNumber="88">Abdomen with few long and black macrosetae near hind margin of each segment, III to V with medial transverse carina. Styli slightly dilated towards the apex and with numerous long and thick black setae.</paragraph>
|
||
<paragraph id="8F208C31FF89FFDDFF66F9B698B6F93E" blockId="3.[151,1437,1379,2016]" pageId="3" pageNumber="88">
|
||
Male: hind margin of sternum VIII medially emarginated and with disperse black setae (
|
||
<figureCitation id="17A490B4FF89FFDDFB1CF9B69F77F9AA" box="[1213,1282,1595,1620]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1932,1955]" captionTargetBox="[166,1424,365,1931]" captionTargetId="figure@2.[151,1436,337,1940]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Linoderus navarretei sp. nov. HOLOTYPE (1), PARATYPES (2 – 9). 1, Habitus; 2, head, 3, male sternum VIII; 4, male sternum IX; 5, aedeagus in dorsal view; 6, aedeagus in ventral view; 7, aedeagus in lateral view; 8, female tergum X; 9, gonocoxite of female genital segment. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/286048/files/figure.png" pageId="3" pageNumber="88">Fig. 3</figureCitation>
|
||
); sternum IX acutely emarginated apically and with inwardly directed short setae on the apical half with proximal portion symmetric (
|
||
<figureCitation id="17A490B4FF89FFDDFEBDF90E9B14F962" box="[284,353,1667,1692]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1932,1955]" captionTargetBox="[166,1424,365,1931]" captionTargetId="figure@2.[151,1436,337,1940]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Linoderus navarretei sp. nov. HOLOTYPE (1), PARATYPES (2 – 9). 1, Habitus; 2, head, 3, male sternum VIII; 4, male sternum IX; 5, aedeagus in dorsal view; 6, aedeagus in ventral view; 7, aedeagus in lateral view; 8, female tergum X; 9, gonocoxite of female genital segment. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/286048/files/figure.png" pageId="3" pageNumber="88">Fig. 4</figureCitation>
|
||
).
|
||
<emphasis id="BDEB5023FF89FFDDFED4F90E9B97F965" box="[373,482,1667,1691]" italics="true" pageId="3" pageNumber="88">Aedeagus</emphasis>
|
||
with parameres long and bilobulated, without pegsetae and shorter than median lobe, internal sac with sclerotized structures (
|
||
<figureCitation id="17A490B4FF89FFDDFDF2F92A98C3F93E" box="[595,694,1703,1728]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1932,1955]" captionTargetBox="[166,1424,365,1931]" captionTargetId="figure@2.[151,1436,337,1940]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Linoderus navarretei sp. nov. HOLOTYPE (1), PARATYPES (2 – 9). 1, Habitus; 2, head, 3, male sternum VIII; 4, male sternum IX; 5, aedeagus in dorsal view; 6, aedeagus in ventral view; 7, aedeagus in lateral view; 8, female tergum X; 9, gonocoxite of female genital segment. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/286048/files/figure.png" pageId="3" pageNumber="88">Fig. 5–7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8F208C31FF89FFDDFF66F9469E7DF8D2" blockId="3.[151,1437,1379,2016]" pageId="3" pageNumber="88">
|
||
Female: with no secondary sexual structures. Genital segment with styli of tergum IX similar to those of male; tergum X with six long thick setae at the apical margin and with short and pale setae on the median surface (
|
||
<figureCitation id="17A490B4FF89FFDDFAE8F9629FF9F8F9" box="[1353,1420,1775,1800]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1932,1955]" captionTargetBox="[166,1424,365,1931]" captionTargetId="figure@2.[151,1436,337,1940]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Linoderus navarretei sp. nov. HOLOTYPE (1), PARATYPES (2 – 9). 1, Habitus; 2, head, 3, male sternum VIII; 4, male sternum IX; 5, aedeagus in dorsal view; 6, aedeagus in ventral view; 7, aedeagus in lateral view; 8, female tergum X; 9, gonocoxite of female genital segment. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/286048/files/figure.png" pageId="3" pageNumber="88">Fig. 8</figureCitation>
|
||
); second gonocoxites laterally flattened and with a long seta at the apex (
|
||
<figureCitation id="17A490B4FF89FFDDFC14F89E998FF8D2" box="[949,1018,1811,1836]" captionStart="FIGURES 1 – 9" captionStartId="2.[151,264,1932,1955]" captionTargetBox="[166,1424,365,1931]" captionTargetId="figure@2.[151,1436,337,1940]" captionTargetPageId="2" captionText="FIGURES 1 – 9. Linoderus navarretei sp. nov. HOLOTYPE (1), PARATYPES (2 – 9). 1, Habitus; 2, head, 3, male sternum VIII; 4, male sternum IX; 5, aedeagus in dorsal view; 6, aedeagus in ventral view; 7, aedeagus in lateral view; 8, female tergum X; 9, gonocoxite of female genital segment. Scale bars = 0.2 mm." httpUri="https://zenodo.org/record/286048/files/figure.png" pageId="3" pageNumber="88">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C785DFBAFF89FFDDFF66F8BA9970F88A" pageId="3" pageNumber="88" type="etymology">
|
||
<paragraph id="8F208C31FF89FFDDFF66F8BA9970F88A" blockId="3.[151,1437,1379,2016]" pageId="3" pageNumber="88">
|
||
<emphasis id="BDEB5023FF89FFDDFF66F8BA9B3BF8AE" bold="true" box="[199,334,1847,1872]" pageId="3" pageNumber="88">Etymology.</emphasis>
|
||
The name is in honor of José Luís Navarrete Heredia, who has been the main contributor to the advancement of
|
||
<taxonomicName id="489FF7B2FF89FFDDFEF1F8D69B9FF88A" box="[336,490,1883,1908]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="88" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
|
||
knowledge in
|
||
<collectingCountry id="F788CCA1FF89FFDDFD2EF8D69974F88A" box="[655,769,1883,1908]" name="Colombia" pageId="3" pageNumber="88">Colombia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C785DFBAFF89FFDAFF66F8F29EFFFE9A" lastPageId="4" lastPageNumber="89" pageId="3" pageNumber="88" type="materials_examined">
|
||
<paragraph id="8F208C31FF89FFDAFF66F8F29EFFFE9A" blockId="3.[151,1437,1379,2016]" lastBlockId="4.[151,1437,151,933]" lastPageId="4" lastPageNumber="89" pageId="3" pageNumber="88">
|
||
<emphasis id="BDEB5023FF89FFDDFF66F8F29B9DF866" bold="true" box="[199,488,1919,1944]" pageId="3" pageNumber="88">
|
||
<typeStatus id="50243293FF89FFDDFF66F8F29B77F866" box="[199,258,1919,1944]" pageId="3" pageNumber="88">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="50243293FF89FFDDFEDAF8F29B9DF866" box="[379,488,1919,1944]" pageId="3" pageNumber="88" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
♂, with labels:
|
||
<collectingCountry id="F788CCA1FF89FFDDFD38F8F2997FF866" box="[665,778,1919,1944]" name="Colombia" pageId="3" pageNumber="88">Colombia</collectingCountry>
|
||
, Quindío Dept., Filandia, Cruces, Parque Regional Natural Barbas-Bremen,
|
||
<geoCoordinate id="EAABEAF6FF89FFDDFEF8F82E9B92F845" box="[345,487,1955,1980]" direction="north" orientation="latitude" pageId="3" pageNumber="88" precision="1" value="4.7017775">4°42’6.4” N</geoCoordinate>
|
||
<geoCoordinate id="EAABEAF6FF89FFDDFE52F82E98D1F845" box="[499,676,1955,1980]" direction="west" orientation="longitude" pageId="3" pageNumber="88" precision="1" value="-75.637856">75°38’16.3” W</geoCoordinate>
|
||
,
|
||
<quantity id="486721D4FF89FFDDFD15F829997AF845" box="[692,783,1956,1980]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="88" unit="m" value="2000.0">2000 m</quantity>
|
||
,
|
||
<date id="FB21AAF1FF89FFDDFC81F82E99F9F842" box="[800,908,1955,1980]" pageId="3" pageNumber="88" value="2013-01-22">22.i.2013</date>
|
||
, López-García M. M. col., manual on forest vegetation,
|
||
<typeStatus id="50243293FF89FFDDFEBBF84A9BF7F81E" box="[282,386,1991,2016]" pageId="3" pageNumber="88" type="holotype">Holotype</typeStatus>
|
||
<taxonomicName id="489FF7B2FF89FFDDFE28F84A9E54F821" authority="Lopez-Garcia & Mendez-Rojas 2014" authorityName="Lopez-Garcia & Mendez-Rojas" authorityYear="2014" box="[393,1057,1991,2016]" class="Insecta" family="Staphylinidae" genus="Linoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="88" phylum="Arthropoda" rank="species" species="navarretei">
|
||
<emphasis id="BDEB5023FF89FFDDFE28F84A9803F821" box="[393,630,1991,2015]" italics="true" pageId="3" pageNumber="88">Linoderus navarretei</emphasis>
|
||
López-García & Méndez-Rojas 2014
|
||
</taxonomicName>
|
||
, (ICN-MHN).
|
||
<emphasis id="BDEB5023FF89FFDDFB69F84A9F32F81E" bold="true" box="[1224,1351,1991,2016]" pageId="3" pageNumber="88">
|
||
<typeStatus id="50243293FF89FFDDFB69F84A9F37F81E" box="[1224,1346,1991,2016]" pageId="3" pageNumber="88" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Eleven
|
||
<typeStatus id="50243293FF8EFFDAFF36FF159B77FF4E" box="[151,258,152,176]" pageId="4" pageNumber="89" type="paratype">paratypes</typeStatus>
|
||
with the same data, ♂ ♀ (ICN-MHN),
|
||
<specimenCount id="999947B8FF8EFFDAFD1EFF15989FFF51" box="[703,746,152,176]" pageId="4" pageNumber="89" type="generic">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="E98E14F4FF8EFFDAFD58FF159931FF4E" box="[761,836,152,176]" pageId="4" pageNumber="89">CIUQ</collectionCode>
|
||
). Two
|
||
<typeStatus id="50243293FF8EFFDAFC31FF15998EFF4E" box="[912,1019,152,176]" pageId="4" pageNumber="89" type="paratype">paratypes</typeStatus>
|
||
with the labels:
|
||
<collectingCountry id="F788CCA1FF8EFFDAFB14FF1A9F53FF4E" box="[1205,1318,151,176]" name="Colombia" pageId="4" pageNumber="89">Colombia</collectingCountry>
|
||
, Risaralda Dept., La Celia, Parque Regional Natural Verdum,
|
||
<geoCoordinate id="EAABEAF6FF8EFFDAFD76FF31991BFF2A" box="[727,878,188,213]" direction="north" orientation="latitude" pageId="4" pageNumber="89" precision="1" value="4.437028">4°26’13.3” N</geoCoordinate>
|
||
<geoCoordinate id="EAABEAF6FF8EFFDAFCD7FF319E7BFF2A" box="[886,1038,188,213]" direction="west" orientation="longitude" pageId="4" pageNumber="89" precision="15" value="-75.624725">75°37’29” W</geoCoordinate>
|
||
,
|
||
<quantity id="486721D4FF8EFFDAFBB8FF319E04FF2A" box="[1049,1137,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="89" unit="m" value="1800.0">1800 m</quantity>
|
||
,
|
||
<date id="FB21AAF1FF8EFFDAFBDCFF319E9CFF2B" box="[1149,1257,188,213]" pageId="4" pageNumber="89" value="2013-01-22">22.i.2013</date>
|
||
, Delgado-Santa L. col., manual on forest soil,
|
||
<typeStatus id="50243293FF8EFFDAFE53FF6D9821FF06" box="[498,596,224,248]" pageId="4" pageNumber="89" type="paratype">Paratype</typeStatus>
|
||
<taxonomicName id="489FF7B2FF8EFFDAFDFCFF6D9E89FF06" authority="Lopez-Garcia & Mendez-Rojas 2014" authorityName="Lopez-Garcia & Mendez-Rojas" authorityYear="2014" box="[605,1276,223,248]" class="Insecta" family="Staphylinidae" genus="Linoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="89" phylum="Arthropoda" rank="species" species="navarretei">
|
||
<emphasis id="BDEB5023FF8EFFDAFDFCFF6D993EFF09" box="[605,843,224,248]" italics="true" pageId="4" pageNumber="89">Linoderus navarretei</emphasis>
|
||
López-García & Méndez-Rojas 2014
|
||
</taxonomicName>
|
||
, 2 ♀ (
|
||
<collectionCode id="E98E14F4FF8EFFDAFAE7FF6D9FE4FF06" box="[1350,1425,224,248]" pageId="4" pageNumber="89">CIUQ</collectionCode>
|
||
). Three
|
||
<typeStatus id="50243293FF8EFFDAFF41FE889B3EFEE3" box="[224,331,261,285]" pageId="4" pageNumber="89" type="paratype">paratypes</typeStatus>
|
||
with the labels:
|
||
<collectingCountry id="F788CCA1FF8EFFDAFDA9FE89980CFEE3" box="[520,633,260,285]" name="Colombia" pageId="4" pageNumber="89">Colombia</collectingCountry>
|
||
, Quindío Dept., Filandia, Reserva Forestal y de Investigación Bremen- La Popa,
|
||
<geoCoordinate id="EAABEAF6FF8EFFDAFEA4FEAA9BEBFEC1" box="[261,414,295,320]" direction="north" orientation="latitude" pageId="4" pageNumber="89" precision="1" value="4.671222">4°40’16.4” N</geoCoordinate>
|
||
<geoCoordinate id="EAABEAF6FF8EFFDAFE06FEAA9820FEC1" box="[423,597,295,320]" direction="west" orientation="longitude" pageId="4" pageNumber="89" precision="1" value="-75.61591">75°36’57.3” W</geoCoordinate>
|
||
,
|
||
<quantity id="486721D4FF8EFFDAFDC0FEA598CFFEC1" box="[609,698,296,320]" metricMagnitude="3" metricUnit="m" metricValue="2.04" pageId="4" pageNumber="89" unit="m" value="2040.0">2040 m</quantity>
|
||
,
|
||
<date id="FB21AAF1FF8EFFDAFD66FEAA993CFEBE" box="[711,841,295,320]" pageId="4" pageNumber="89" value="2013-07-19">19.vii.2013</date>
|
||
, Soto-Garzón C. col., manual on leaves of
|
||
<taxonomicName id="489FF7B2FF8EFFDAFA9BFEA59FECFEBE" box="[1338,1433,296,320]" class="Liliopsida" family="Araceae" kingdom="Plantae" order="Alismatales" pageId="4" pageNumber="89" phylum="Tracheophyta" rank="family">Araceae</taxonomicName>
|
||
,
|
||
<typeStatus id="50243293FF8EFFDAFF36FEC09A8CFE9B" box="[151,249,333,357]" pageId="4" pageNumber="89" type="paratype">Paratype</typeStatus>
|
||
<taxonomicName id="489FF7B2FF8EFFDAFEA1FEC199E3FE9A" authority="Lopez-Garcia & Mendez-Rojas 2014" authorityName="Lopez-Garcia & Mendez-Rojas" authorityYear="2014" box="[256,918,332,357]" class="Insecta" family="Staphylinidae" genus="Linoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="89" phylum="Arthropoda" rank="species" species="navarretei">
|
||
<emphasis id="BDEB5023FF8EFFDAFEA1FEC19B99FE9A" box="[256,492,332,356]" italics="true" pageId="4" pageNumber="89">Linoderus navarretei</emphasis>
|
||
López-García & Méndez-Rojas 2014
|
||
</taxonomicName>
|
||
,
|
||
<specimenCount id="999947B8FF8EFFDAFC00FEC199B1FE9A" box="[929,964,332,356]" pageId="4" pageNumber="89" type="generic">1♂</specimenCount>
|
||
2♀ (ICN-MHN).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C785DFBAFF8EFFDAFF66FEE29B2FFD56" pageId="4" pageNumber="89" type="distribution">
|
||
<paragraph id="8F208C31FF8EFFDAFF66FEE29941FE53" blockId="4.[151,1437,151,933]" pageId="4" pageNumber="89">
|
||
<emphasis id="BDEB5023FF8EFFDAFF66FEE29B15FE76" bold="true" box="[199,352,367,392]" pageId="4" pageNumber="89">Distribution.</emphasis>
|
||
The new species of
|
||
<taxonomicName id="489FF7B2FF8EFFDAFDC3FEFD98A1FE76" box="[610,724,368,392]" class="Insecta" family="Staphylinidae" genus="Linoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="89" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BDEB5023FF8EFFDAFDC3FEFD98A1FE76" box="[610,724,368,392]" italics="true" pageId="4" pageNumber="89">Linoderus</emphasis>
|
||
</taxonomicName>
|
||
is known only from Central and Western Cordillera of the Colombian Andes (Departments of Quindío and Risaralda).
|
||
</paragraph>
|
||
<paragraph id="8F208C31FF8EFFDAFF66FE3A9B2FFD56" blockId="4.[151,1437,151,933]" pageId="4" pageNumber="89">
|
||
<emphasis id="BDEB5023FF8EFFDAFF66FE3A9B91FE2E" bold="true" box="[199,484,439,464]" pageId="4" pageNumber="89">Habitat and bionomics.</emphasis>
|
||
The species was found in riparian areas of montane forest. Most individuals were collected directly from vegetation, during the day. Thirteen individuals were observed on the lower leaf surface that they apparently use for perching behavior (
|
||
<figureCitation id="17A490B4FF8EFFDAFDD6FE729880FDE6" box="[631,757,511,536]" captionStart="FIGURES 10 – 13" captionStartId="3.[151,264,1281,1304]" captionTargetBox="[151,1429,241,1254]" captionTargetId="figure@3.[151,1436,237,1260]" captionTargetPageId="3" captionText="FIGURES 10 – 13. Habitat of Linoderus navarretei sp. nov. 10 – 11 under leaves (Photos by MMLG); 12, Parque Regional Natural Barbas-Bremen: forest, view from above; 13, riparian vegetation (Photos by O. H. Marín-Gómez)." httpUri="https://zenodo.org/record/286049/files/figure.png" pageId="4" pageNumber="89">Fig. 10–11</figureCitation>
|
||
). More than two individuals were often found in a same leaf; even six of them were together in two occasions. Only two individuals, a male and a female, were observed on the upper leaf. However, they were covered by a piece of fallen leaf. In addition, a male was observed moving actively on inflorescences of
|
||
<taxonomicName id="489FF7B2FF8EFFDAFE25FDE19909FD7B" authority="Abalo & G. Morales. The" authorityName="Abalo & G. Morales. The" box="[388,892,620,645]" class="Liliopsida" family="Heliconiaceae" genus="Heliconia" kingdom="Animalia" order="Zingiberales" pageId="4" pageNumber="89" phylum="Tracheophyta" rank="species" species="venusta">
|
||
<emphasis id="BDEB5023FF8EFFDAFE25FDE19825FD7A" box="[388,592,620,644]" italics="true" pageId="4" pageNumber="89">Heliconia venusta</emphasis>
|
||
Abalo & G. Morales. The
|
||
</taxonomicName>
|
||
only two females of Risaralda were found on a highly moist soil.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C785DFBAFF8EFFDAFF66FD39990CFC5A" pageId="4" pageNumber="89" type="discussion">
|
||
<paragraph id="8F208C31FF8EFFDAFF66FD39990CFC5A" blockId="4.[151,1437,151,933]" pageId="4" pageNumber="89">
|
||
<emphasis id="BDEB5023FF8EFFDAFF66FD399B4EFD33" bold="true" box="[199,315,692,717]" pageId="4" pageNumber="89">Remarks.</emphasis>
|
||
The color pattern of
|
||
<taxonomicName id="489FF7B2FF8EFFDAFD89FD399966FD32" box="[552,787,692,716]" class="Insecta" family="Staphylinidae" genus="Linoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="89" phylum="Arthropoda" rank="species" species="navarretei">
|
||
<emphasis id="BDEB5023FF8EFFDAFD89FD399966FD32" box="[552,787,692,716]" italics="true" pageId="4" pageNumber="89">Linoderus navarretei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="BDEB5023FF8EFFDAFCBAFD399918FD33" bold="true" box="[795,877,692,717]" pageId="4" pageNumber="89">sp nov.</emphasis>
|
||
is widely similar to that of
|
||
<taxonomicName id="489FF7B2FF8EFFDAFB03FD399E86FD32" box="[1186,1267,692,716]" class="Insecta" family="Staphylinidae" genus="Nordus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="89" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BDEB5023FF8EFFDAFB03FD399E86FD32" box="[1186,1267,692,716]" italics="true" pageId="4" pageNumber="89">Nordus</emphasis>
|
||
</taxonomicName>
|
||
cf.
|
||
<emphasis id="BDEB5023FF8EFFDAFABFFD399FE9FD32" box="[1310,1436,692,716]" italics="true" pageId="4" pageNumber="89">maculiceps</emphasis>
|
||
(Xanthophygina), which was also found on riparian vegetation in
|
||
<emphasis id="BDEB5023FF8EFFDAFC27FD549F28FD0E" box="[902,1373,728,752]" italics="true" pageId="4" pageNumber="89">Parque Regional Natural Barbas-Bremen</emphasis>
|
||
(
|
||
<figureCitation id="17A490B4FF8EFFDAFACFFD5A9A94FCEA" captionStart="FIGURES 10 – 13" captionStartId="3.[151,264,1281,1304]" captionTargetBox="[151,1429,241,1254]" captionTargetId="figure@3.[151,1436,237,1260]" captionTargetPageId="3" captionText="FIGURES 10 – 13. Habitat of Linoderus navarretei sp. nov. 10 – 11 under leaves (Photos by MMLG); 12, Parque Regional Natural Barbas-Bremen: forest, view from above; 13, riparian vegetation (Photos by O. H. Marín-Gómez)." httpUri="https://zenodo.org/record/286049/files/figure.png" pageId="4" pageNumber="89">Fig. 12–13</figureCitation>
|
||
). In
|
||
<taxonomicName id="489FF7B2FF8EFFDAFEB0FD719B16FCEA" box="[273,355,764,788]" class="Insecta" family="Staphylinidae" genus="Nordus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="89" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BDEB5023FF8EFFDAFEB0FD719B16FCEA" box="[273,355,764,788]" italics="true" pageId="4" pageNumber="89">Nordus</emphasis>
|
||
</taxonomicName>
|
||
, it has been thought that tail color (yellow) can act with abdomen waving and chemical secretion as a defense mechanism by combination of visual and chemical signals (
|
||
<bibRefCitation id="EB0EF1C0FF8EFFDAFC6DFC929EC1FCC6" author="Chatzimanolis" box="[972,1204,799,824]" pageId="4" pageNumber="89" refString="Chatzimanolis, S. (2003) Natural history and behavior of Nordus fungicola (Sharp) (Coleoptera: Staphylinidae). Annals of The Entomological Society of America, 96 (3), 225 - 230. http: // dx. doi. org / 10.1603 / 0013 - 8746 (2003) 096 [0225: nhabon] 2.0. co; 2" type="journal article" year="2003">Chatzimanolis 2003</bibRefCitation>
|
||
). It convergences do not reflect a closer phyletic affinity and rather it can be a product of sharing the same habitat or mimicry, but it is necessary to carry out further studies that allow understand which species is model and which is mimic. Similar color patterns have been observed in some species of
|
||
<taxonomicName id="489FF7B2FF8EFFDAFD51FC019907FC5A" box="[752,882,908,932]" class="Insecta" family="Staphylinidae" genus="Belonuchus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="89" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BDEB5023FF8EFFDAFD51FC019907FC5A" box="[752,882,908,932]" italics="true" pageId="4" pageNumber="89">Belonuchus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |