158 lines
18 KiB
XML
158 lines
18 KiB
XML
<document id="DD44169F4DF4965CBA37CF094A0C7957" ID-DOI="10.5281/zenodo.5358903" ID-ISSN="2345-7600" ID-Zenodo-Dep="5358903" ID-ZooBank="B04A2DD9-91C2-48D1-9EB2-965D3FFBF879" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630225750785" checkinUser="valdenar" docAuthor="Goulding, Tricia C., Khalil, Munawar, Tan, Shau Hwai & Dayrat, Benoît" docDate="2018" docId="0702712AFFFBDA5BFF69DE924B6D1C19" docLanguage="en" docName="66rbz337-349.pdf" docOrigin="Raffles Bulletin of Zoology 66" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Alionchis Goulding & Dayrat 2018, new genus" docType="treatment" docUuid="FAEB68BF-D52F-48DC-804B-427071F85854" docUuidSource="ZooBank" docVersion="4" lastPageNumber="341" masterDocId="FB3B0952FFFFDA5FFFB3DC2A4F521848" masterDocTitle="A new genus and a new species of onchidiid slugs from eastern Indonesia (Gastropoda: Euthyneura: Onchidiidae)" masterLastPageNumber="349" masterPageNumber="337" pageNumber="341" updateTime="1699191983827" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="8C79C4649A962D9E06A4634576AECAF2">
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<mods:title id="45E335633D211AF4CFF6144366CF377B">A new genus and a new species of onchidiid slugs from eastern Indonesia (Gastropoda: Euthyneura: Onchidiidae)</mods:title>
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<mods:name id="95A7B6FEE3D08DBA0450F8F991F95931" type="personal">
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<mods:roleTerm id="1E32ECCA15F00128CFB5B67D03491BAC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0A0456359F1F655B2BB14D08C0703FD2">Goulding, Tricia C.</mods:namePart>
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<mods:roleTerm id="9F8FB2967D881C21F31CBA14FF5ECFC7">Author</mods:roleTerm>
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<mods:namePart id="662911AEACB0A3AC664E46ADDAE23C5B">Khalil, Munawar</mods:namePart>
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</mods:name>
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<mods:name id="B93D8FCA1BD92274148C28AEA7AA0ABD" type="personal">
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<mods:roleTerm id="3046AED55A638ECF0BB2A5373C14DBE4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DB950B873F117F4E07BC973F84C82D94">Tan, Shau Hwai</mods:namePart>
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</mods:name>
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<mods:name id="0F1DDE2919A05B1F965904273F9A0893" type="personal">
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<mods:roleTerm id="8AF0C1300E6D19713AFC29E3F5862EE9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C952C13F79ED18971964A08AE76A7C43">Dayrat, Benoît</mods:namePart>
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<mods:typeOfResource id="EB142937CE94B507AB02DAF2CE124DD2">text</mods:typeOfResource>
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<mods:title id="BAEA2B4959AF18546683D2646F9E6114">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="6C63CF24B0404352F2EB8774A747B6B1">
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<mods:date id="EAE9246625B62FFDCD1CA0093E716115">2018</mods:date>
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<mods:number id="9C27DF422EC8AA4AF1817E0573B7C53D">2018-05-21</mods:number>
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<mods:number id="6D56BF859A50E91D3CB2FDC9AA11819D">66</mods:number>
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<mods:start id="51C8331E85F9E0B61E51BA5716691C20">337</mods:start>
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<mods:classification id="5CBB1E6A5FE6FF8D6EDEF5500962BDF8">journal article</mods:classification>
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<mods:identifier id="B88839C10AC1BA523C853E08B52A4AF3" type="DOI">10.5281/zenodo.5358903</mods:identifier>
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<treatment id="0702712AFFFBDA5BFF69DE924B6D1C19" LSID="urn:lsid:zoobank.org:act:FAEB68BF-D52F-48DC-804B-427071F85854" httpUri="http://treatment.plazi.org/id/0702712AFFFBDA5BFF69DE924B6D1C19" lastPageNumber="341" pageId="4" pageNumber="341">
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<subSubSection id="C7B193B7FFFBDA5BFF69DE924DE71A98" box="[218,693,696,720]" pageId="4" pageNumber="341" type="nomenclature">
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<paragraph id="8F14C03CFFFBDA5BFF69DE924DE71A98" blockId="4.[218,693,696,720]" box="[218,693,696,720]" pageId="4" pageNumber="341">
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<heading id="D45C7750FFFBDA5BFF69DE924DE71A98" bold="true" box="[218,693,696,720]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="341" reason="2">
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<emphasis id="BDDF1C2EFFFBDA5BFF69DE924DE71A98" bold="true" box="[218,693,696,720]" pageId="4" pageNumber="341">
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<taxonomicName id="48ABBBBFFFFBDA5BFF69DE924D621A98" ID-CoL="7NNKD" authority="Goulding & Dayrat" authorityName="Goulding & Dayrat" authorityYear="2018" box="[218,560,696,720]" class="Gastropoda" family="Onchidiidae" genus="Alionchis" kingdom="Animalia" order="Systellommatophora" pageId="4" pageNumber="341" phylum="Mollusca" rank="genus" status="gen. nov.">
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<emphasis id="BDDF1C2EFFFBDA5BFF69DE924E111A98" bold="true" box="[218,323,696,720]" italics="true" pageId="4" pageNumber="341">Alionchis</emphasis>
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Goulding & Dayrat
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</taxonomicName>
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,
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<taxonomicNameLabel id="A6ECA155FFFBDA5BFD8EDE924DE71A98" box="[573,693,696,720]" pageId="4" pageNumber="341" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C7B193B7FFFBDA5BFF3BDED24D8E1B58" box="[136,732,760,784]" pageId="4" pageNumber="341" type="type_taxon">
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<paragraph id="8F14C03CFFFBDA5BFF3BDED24D8E1B58" blockId="4.[136,732,760,784]" box="[136,732,760,784]" pageId="4" pageNumber="341">
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||
<emphasis id="BDDF1C2EFFFBDA5BFF3BDED24D8E1B58" bold="true" box="[136,732,760,784]" pageId="4" pageNumber="341">
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<typeStatus id="50107E9EFFFBDA5BFF3BDED24F901B58" box="[136,194,760,784]" pageId="4" pageNumber="341">Type</typeStatus>
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species:
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<taxonomicName id="48ABBBBFFFFBDA5BFE9EDED24D4A1B58" ID-CoL="BSCB" authorityName="Goulding & Dayrat" authorityYear="2018" box="[301,536,760,784]" class="Gastropoda" family="Onchidiidae" genus="Alionchis" kingdom="Animalia" order="Systellommatophora" pageId="4" pageNumber="341" phylum="Mollusca" rank="species" species="jailoloensis">
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<emphasis id="BDDF1C2EFFFBDA5BFE9EDED24D4A1B58" bold="true" box="[301,536,760,784]" italics="true" pageId="4" pageNumber="341">Alionchis jailoloensis</emphasis>
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</taxonomicName>
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, designated here
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C7B193B7FFFBDA5BFF3BDF124C5A1C98" pageId="4" pageNumber="341" type="etymology">
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<paragraph id="8F14C03CFFFBDA5BFF3BDF124C5A1C98" blockId="4.[134,776,824,1233]" pageId="4" pageNumber="341">
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<emphasis id="BDDF1C2EFFFBDA5BFF3BDF124E581B18" bold="true" box="[136,266,824,848]" pageId="4" pageNumber="341">Etymology.</emphasis>
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The genus
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<taxonomicName id="48ABBBBFFFFBDA5BFE3BDF124EBF1B18" authorityName="Goulding & Dayrat" authorityYear="2018" box="[392,493,824,848]" class="Gastropoda" family="Onchidiidae" genus="Alionchis" kingdom="Animalia" order="Systellommatophora" pageId="4" pageNumber="341" phylum="Mollusca" rank="genus">
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<emphasis id="BDDF1C2EFFFBDA5BFE3BDF124EBF1B18" box="[392,493,824,848]" italics="true" pageId="4" pageNumber="341">Alionchis</emphasis>
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</taxonomicName>
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is a combination of Ali, a collector and assistant to Alfred Russel Wallace during his travels across the Malay Archipelago, and
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<taxonomicName id="48ABBBBFFFFBDA5BFDF5DF524DC01BD8" authorityName="Ferussac" authorityYear="1822" box="[582,658,888,912]" class="Gastropoda" family="Onchidiidae" genus="Onchis" kingdom="Animalia" order="Systellommatophora" pageId="4" pageNumber="341" phylum="Mollusca" rank="genus">
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<emphasis id="BDDF1C2EFFFBDA5BFDF5DF524DC01BD8" box="[582,658,888,912]" italics="true" pageId="4" pageNumber="341">Onchis</emphasis>
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</taxonomicName>
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, one of the names used to refer to onchidiid slugs. Ali began working for Wallace in Borneo, first serving as cook, and later collecting and preparing specimens (
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<bibRefCitation id="EB3ABDCDFFFBDA5BFE14DFF24DAE1BB8" author="Van Wyhe J & Drawhorn GM" box="[423,764,984,1008]" pageId="4" pageNumber="341" pagination="3 - 31" refId="ref9315" refString="Van Wyhe J & Drawhorn GM (2015) " I am Ali Wallace ": The Malay assistant of Alfred Russel Wallace. Journal of the Malaysian Branch of the Royal Asiatic Society, 88 (1): 3 - 31." type="journal article" year="2015">Van Wyhe & Drawhorn, 2015</bibRefCitation>
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). Ali’s collections of birds and natural history observations greatly contributed to Wallace’s knowledge of the fauna in the region. This genus is named to honor Ali’s contribution to studies of biodiversity in the Malay Archipelago. Ali actually got married on the island of
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<collectingCountry id="F7BC80ACFFFBDA5BFDB3D8534D011CD8" box="[512,595,1145,1168]" name="Indonesia" pageId="4" pageNumber="341">Ternate</collectingCountry>
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, near Halmahera, and settled there, which, given the geographic distribution of
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<taxonomicName id="48ABBBBFFFFBDA5BFF14D8924E5E1C98" authorityName="Goulding & Dayrat" authorityYear="2018" box="[167,268,1208,1232]" class="Gastropoda" family="Onchidiidae" genus="Alionchis" kingdom="Animalia" order="Systellommatophora" pageId="4" pageNumber="341" phylum="Mollusca" rank="genus">
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<emphasis id="BDDF1C2EFFFBDA5BFF14D8924E5E1C98" box="[167,268,1208,1232]" italics="true" pageId="4" pageNumber="341">Alionchis</emphasis>
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</taxonomicName>
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, makes the new genus name especially fitting.
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</paragraph>
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</subSubSection>
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<subSubSection id="C7B193B7FFFBDA5BFF3BD8D24D2C1EB8" pageId="4" pageNumber="341" type="diagnosis">
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<paragraph id="8F14C03CFFFBDA5BFF3BD8D24ED71E78" blockId="4.[135,776,1272,1585]" pageId="4" pageNumber="341">
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<emphasis id="BDDF1C2EFFFBDA5BFF3BD8D24FAF1D58" bold="true" box="[136,253,1272,1296]" pageId="4" pageNumber="341">Diagnosis.</emphasis>
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Body slightly flattened. Notum soft (gelatinous). No dorsal gills. Dorsal eyes present on notum. Retractable, central papilla (with four dorsal eyes) present. Foot wide, approximately two thirds of total width. Eye tentacles thick and long. Eyes at tip of ocular tentacles. Pneumostome median, at posterior margin of hyponotum. Female opening adjacent to anus. Male opening below right ocular tentacle. Intestinal loops of
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<typeStatus id="50107E9EFFFBDA5BFEFFD9F34E281DB9" box="[332,378,1497,1521]" pageId="4" pageNumber="341">type</typeStatus>
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II,
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<typeStatus id="50107E9EFFFBDA5BFE2ED9F34E991DB9" box="[413,459,1497,1521]" pageId="4" pageNumber="341">type</typeStatus>
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III, and intermediate between
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<typeStatus id="50107E9EFFFBDA5BFF34D9D34F901E59" box="[135,194,1529,1553]" pageId="4" pageNumber="341">types</typeStatus>
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II and III. No rectal gland. No penial hooks. No accessory penial gland.
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</paragraph>
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<paragraph id="8F14C03CFFFBDA5BFF3BDA724D2C1EB8" blockId="4.[135,776,1624,1776]" pageId="4" pageNumber="341">
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<emphasis id="BDDF1C2EFFFBDA5BFF3BDA724E3F1E38" bold="true" box="[136,365,1624,1648]" pageId="4" pageNumber="341">Diagnostic features.</emphasis>
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Externally,
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<taxonomicName id="48ABBBBFFFFBDA5BFE46DA724D081E38" authorityName="Goulding & Dayrat" authorityYear="2018" box="[501,602,1624,1648]" class="Gastropoda" family="Onchidiidae" genus="Alionchis" kingdom="Animalia" order="Systellommatophora" pageId="4" pageNumber="341" phylum="Mollusca" rank="genus">
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<emphasis id="BDDF1C2EFFFBDA5BFE46DA724D081E38" box="[501,602,1624,1648]" italics="true" pageId="4" pageNumber="341">Alionchis</emphasis>
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</taxonomicName>
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can be reliably distinguished from other onchidiid genera by the unusually thick eye tentacles and the pneumostome at the very posterior end of the hyponotum. The squishy, gelatinous texture of the notum is also distinctive (in live animals).
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</paragraph>
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</subSubSection>
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<subSubSection id="C7B193B7FFFBDA5BFF3BDB324B6D1C19" pageId="4" pageNumber="341" type="discussion">
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<paragraph id="8F14C03CFFFBDA5BFF3BDB324B6D1C19" blockId="4.[136,776,1816,2064]" lastBlockId="4.[835,1475,569,1105]" pageId="4" pageNumber="341">
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<emphasis id="BDDF1C2EFFFBDA5BFF3BDB324FAE1F78" bold="true" box="[136,252,1816,1840]" pageId="4" pageNumber="341">Remarks.</emphasis>
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A new generic name is needed for the new species described here because it does not belong to any of the known genera and no existing generic name could apply to it (for a recent review of the application of all onchidiid genus-group names, see Dayrat et al., 2017). The position of the pneumostome in
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<taxonomicName id="48ABBBBFFFFBDA5BFE55DB924D191F98" authorityName="Goulding & Dayrat" authorityYear="2018" box="[486,587,1976,2000]" class="Gastropoda" family="Onchidiidae" genus="Alionchis" kingdom="Animalia" order="Systellommatophora" pageId="4" pageNumber="341" phylum="Mollusca" rank="genus">
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<emphasis id="BDDF1C2EFFFBDA5BFE55DB924D191F98" box="[486,587,1976,2000]" italics="true" pageId="4" pageNumber="341">Alionchis</emphasis>
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</taxonomicName>
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is unique. It is at the posterior end of the hyponotum, close to the hyponotum edge (but still on the hyponotum surface), which cannot be confused with any other onchidiid genus. In
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<taxonomicName id="48ABBBBFFFFBDA5BFAEDDE134C931A39" authority="Plate, 1893" authorityName="Plate" authorityYear="1893" class="Gastropoda" family="Onchidiidae" genus="Peronina" kingdom="Animalia" order="Systellommatophora" pageId="4" pageNumber="341" phylum="Mollusca" rank="genus">
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<emphasis id="BDDF1C2EFFFBDA5BFAEDDE134A901A18" box="[1374,1474,569,592]" italics="true" pageId="4" pageNumber="341">Peronina</emphasis>
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<bibRefCitation id="EB3ABDCDFFFBDA5BFCF0DE734C931A39" author="Plate L von" box="[835,961,601,625]" pageId="4" pageNumber="341" pagination="93 - 234" refId="ref9005" refString="Plate L von (1893) Studien uber opisthopneumone Lungenschnecken, II, Die Onciidien. Zoologische Jahrbucher, Anatomie, 7: 93 - 234, pl. 7 - 12." type="journal article" year="1893">Plate, 1893</bibRefCitation>
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</taxonomicName>
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, the pneumostome also is at the posterior end of the hyponotum, but it is not on the hyponotum surface (Goulding et al., in press-b). Instead, it is right at the margin between the ventral hyponotum and the dorsal notum. In other onchidiids, the pneumostome is on the hyponotum, approximately equally distant between the foot and the hyponotum edge. The eye tentacles of
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<taxonomicName id="48ABBBBFFFFBDA5BFB48DF334A331B79" authorityName="Goulding & Dayrat" authorityYear="2018" box="[1275,1377,793,817]" class="Gastropoda" family="Onchidiidae" genus="Alionchis" kingdom="Animalia" order="Systellommatophora" pageId="4" pageNumber="341" phylum="Mollusca" rank="genus">
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<emphasis id="BDDF1C2EFFFBDA5BFB48DF334A331B79" box="[1275,1377,793,817]" italics="true" pageId="4" pageNumber="341">Alionchis</emphasis>
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</taxonomicName>
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are also unique. They are much thicker than in all other onchidiid genera. Only one species of
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<taxonomicName id="48ABBBBFFFFBDA5BFB38DF734BA01B39" authorityName="Goulding & Dayrat" authorityYear="2018" box="[1163,1266,857,881]" class="Gastropoda" family="Onchidiidae" genus="Alionchis" kingdom="Animalia" order="Systellommatophora" pageId="4" pageNumber="341" phylum="Mollusca" rank="genus">
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<emphasis id="BDDF1C2EFFFBDA5BFB38DF734BA01B39" box="[1163,1266,857,881]" italics="true" pageId="4" pageNumber="341">Alionchis</emphasis>
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</taxonomicName>
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is described here, but the position of the pneumostome and the unique eye tentacles will likely be shared by other species in this genus because these traits usually do not vary within a genus. For instance, all three species of
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<taxonomicName id="48ABBBBFFFFBDA5BFB26DFF34A5D1BB9" authorityName="Buchannan" authorityYear="1800" box="[1173,1295,985,1009]" class="Gastropoda" family="Onchidiidae" genus="Onchidium" kingdom="Animalia" order="Systellommatophora" pageId="4" pageNumber="341" phylum="Mollusca" rank="genus">
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<emphasis id="BDDF1C2EFFFBDA5BFB26DFF34A5D1BB9" box="[1173,1295,985,1009]" italics="true" pageId="4" pageNumber="341">Onchidium</emphasis>
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</taxonomicName>
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share long and thin eye tentacles (
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<bibRefCitation id="EB3ABDCDFFFBDA5BFBBEDFD34B8B1C59" author="Dayrat B & Goulding TC & Apte D & Bhave V & Comendador J & Quang NX & Tan SK & Tan SH" box="[1037,1241,1017,1041]" pageId="4" pageNumber="341" pagination="1 - 40" refId="ref8434" refString="Dayrat B, Goulding TC, Apte D, Bhave V, Comendador J, Quang NX, Tan SK & Tan SH (2016) Integrative taxonomy of the genus Onchidium Buchannan, 1800 (Mollusca: Gastropoda: Pulmonata: Onchidiidae). ZooKeys, 636: 1 - 40." type="journal article" year="2016">Dayrat et al., 2016</bibRefCitation>
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) and all three species of
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<taxonomicName id="48ABBBBFFFFBDA5BFCD7D8334C9B1C78" authorityName="Plate" authorityYear="1893" box="[868,969,1049,1072]" class="Gastropoda" family="Onchidiidae" genus="Peronina" kingdom="Animalia" order="Systellommatophora" pageId="4" pageNumber="341" phylum="Mollusca" rank="genus">
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<emphasis id="BDDF1C2EFFFBDA5BFCD7D8334C9B1C78" box="[868,969,1049,1072]" italics="true" pageId="4" pageNumber="341">Peronina</emphasis>
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</taxonomicName>
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(Goulding et al., in press-b) share the same pneumostome position.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |