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<document id="E8F55F5188E93AE4BC39D78BDC6FB068" ID-DOI="10.5281/zenodo.206562" ID-GBIF-Dataset="031e9046-3e51-49d1-9fa6-b712840412bc" ID-ISSN="1175-5326" ID-Zenodo-Dep="206562" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460261978985" checkinUser="plazi" docAuthor="Scarponi, Daniele, Bella, Giano Della &amp; Ceregato, Alessandro" docDate="2011" docId="0391940F1C7A4409FF14FB3C5E5DFE17" docLanguage="en" docName="zt02796p055.pdf" docOrigin="Zootaxa 2796" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Haedropleura parva Scarponi, Bella &amp; Ceregato, 2011, n. sp." docType="treatment" docVersion="7" lastPageNumber="47" masterDocId="FFA8EC771C734403FF83FFFB5B47FFE3" masterDocTitle="The genus Haedropleura (Neogastropoda, Toxoglossa = Conoidea) in the Plio Quaternary of the Mediterranean basin" masterLastPageNumber="55" masterPageNumber="37" pageNumber="46" updateTime="1698244664594" updateUser="plazi">
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<mods:title id="01EAF47156D46EB95DC3162C1FF4484D">The genus Haedropleura (Neogastropoda, Toxoglossa = Conoidea) in the Plio Quaternary of the Mediterranean basin</mods:title>
</mods:titleInfo>
<mods:name id="56E181AE59CD2D2F1FC73B9A6F5ECF99" type="personal">
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<mods:namePart id="8C298CA96D9E4CA9B1F3758AA5303CE3">Scarponi, Daniele</mods:namePart>
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<mods:namePart id="72500746A528B93705E4D8CE27152871">Bella, Giano Della</mods:namePart>
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<mods:namePart id="3B5E2F4EF8131F417E7F212E23261D1A">Ceregato, Alessandro</mods:namePart>
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<mods:title id="A53048CA334F34720B9866A1D3DBA485">Zootaxa</mods:title>
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<mods:part id="43DB4262EBD8F44849DDFC581911A117">
<mods:date id="5DB5C25347E46E258755333CB5EF81E2">2011</mods:date>
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<mods:number id="B8E1971BD30144E627E1514DB229E39C">2796</mods:number>
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<treatment id="0391940F1C7A4409FF14FB3C5E5DFE17" ID-DOI="http://doi.org/10.5281/zenodo.6184261" ID-GBIF-Taxon="119393532" ID-Zenodo-Dep="6184261" LSID="urn:lsid:plazi:treatment:0391940F1C7A4409FF14FB3C5E5DFE17" httpUri="http://treatment.plazi.org/id/0391940F1C7A4409FF14FB3C5E5DFE17" lastPageId="10" lastPageNumber="47" pageId="9" pageNumber="46">
<subSubSection id="C32276921C7A440AFF14FB3C5A28FAE1" pageId="9" pageNumber="46" type="nomenclature">
<paragraph id="8B8725191C7A440AFF14FB3C5A92FB02" blockId="9.[151,469,1223,1282]" box="[151,469,1223,1249]" pageId="9" pageNumber="46">
<heading id="D0CF92751C7A440AFF14FB3C5A92FB02" bold="true" box="[151,469,1223,1249]" fontSize="11" level="1" pageId="9" pageNumber="46" reason="1">
<emphasis id="B94CF90B1C7A440AFF14FB3C5A92FB02" bold="true" box="[151,469,1223,1249]" pageId="9" pageNumber="46">
<taxonomicName id="4C385E9A1C7A440AFF14FB3C5ACCFB02" box="[151,395,1223,1249]" class="Gastropoda" family="Horaiclavidae" genus="Haedropleura" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="46" phylum="Mollusca" rank="species" species="parva" status="sp. nov.">
<emphasis id="B94CF90B1C7A440AFF14FB3C5ACCFB02" bold="true" box="[151,395,1223,1249]" italics="true" pageId="9" pageNumber="46">Haedropleura parva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F44701C7A440AFE10FB3C5A92FB02" box="[403,469,1223,1249]" pageId="9" pageNumber="46" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8725191C7A440AFF14FB115A28FAE1" blockId="9.[151,469,1223,1282]" box="[151,367,1258,1282]" pageId="9" pageNumber="46">Figs. 2836, 6769</paragraph>
</subSubSection>
<subSubSection id="C32276921C7A440AFF14FAC85B82F9C1" pageId="9" pageNumber="46" type="materials_examined">
<paragraph id="8B8725191C7A440AFF14FAC85849FA8D" blockId="9.[151,1436,1330,2002]" pageId="9" pageNumber="46">
<emphasis id="B94CF90B1C7A440AFF14FAC85A02FAA8" bold="true" box="[151,325,1330,1355]" pageId="9" pageNumber="46">
<typeStatus id="54839BBB1C7A440AFF14FAC85B95FAA8" box="[151,210,1331,1355]" pageId="9" pageNumber="46">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54839BBB1C7A440AFECFFAC95AF3FAA9" box="[332,436,1330,1354]" pageId="9" pageNumber="46" type="holotype">Holotype</typeStatus>
and 14
<typeStatus id="54839BBB1C7A440AFD8EFAC8593FFAA8" box="[525,632,1331,1355]" pageId="9" pageNumber="46" type="paratype">paratypes</typeStatus>
(Appendix 1), in Della Bella Collection,
<collectionCode id="ED29BDDC1C7A440AFBCAFAC95FDAFAA9" box="[1097,1181,1330,1354]" pageId="9" pageNumber="46">MGGC</collectionCode>
(Bologna,
<collectingCountry id="F32F65891C7A440AFA99FAC95E15FAA9" box="[1306,1362,1330,1354]" name="Italy" pageId="9" pageNumber="46">Italy</collectingCountry>
); catalogue numbers 23457 (
<typeStatus id="54839BBB1C7A440AFE1BFAAD5ABAFA8D" box="[408,509,1366,1390]" pageId="9" pageNumber="46" type="holotype">holotype</typeStatus>
) and 23458 (
<typeStatus id="54839BBB1C7A440AFD13FAAD5845FA8D" box="[656,770,1366,1390]" pageId="9" pageNumber="46" type="paratype">paratypes</typeStatus>
).
</paragraph>
<paragraph id="8B8725191C7A440AFF44FA805AB6FA55" blockId="9.[151,1436,1330,2002]" pageId="9" pageNumber="46">
<emphasis id="B94CF90B1C7A440AFF44FA805A23FA70" bold="true" box="[199,356,1402,1427]" pageId="9" pageNumber="46">
<typeStatus id="54839BBB1C7A440AFF44FA805A45FA70" box="[199,258,1403,1427]" pageId="9" pageNumber="46">Type</typeStatus>
locality.
</emphasis>
Poggio alla Staffa (Colle Val dElsa, Siena),
<collectingCountry id="F32F65891C7A440AFCD8FA8158C8FA71" box="[859,911,1402,1426]" name="Italy" pageId="9" pageNumber="46">Italy</collectingCountry>
, (information on the
<typeStatus id="54839BBB1C7A440AFBF9FA805FEDFA70" box="[1146,1194,1403,1427]" pageId="9" pageNumber="46">type</typeStatus>
locality in
<bibRefCitation id="EFA958E81C7A440AFAA4FA815B94FA55" author="Bogi" pageId="9" pageNumber="46" refString="Bogi, C., Cauli, L. &amp; Pagli, F. (2002) Le Architectonicidae Gray J. E., 1840 del Pliocene toscano. Bollettino Malacologico, 38, 31 - 40." type="journal article" year="2002">
Bogi
<emphasis id="B94CF90B1C7A440AFAE7FA875EDBFA70" box="[1380,1436,1403,1427]" italics="true" pageId="9" pageNumber="46">et al.</emphasis>
2002
</bibRefCitation>
;
<bibRefCitation id="EFA958E81C7A440AFF5EFA655AA1FA55" author="Pedriali" box="[221,486,1438,1462]" pageId="9" pageNumber="46" refString="Pedriali, L. &amp; Robba, E. (2005) A revision of the Pliocene naticids of northern and central Italy. The subfamily Naticinae. Rivista Italiana di Paleontologia e Stratigrafia, 111, 371 - 429." type="journal article" year="2005">Pedriali &amp; Robba 2005</bibRefCitation>
).
</paragraph>
<paragraph id="8B8725191C7A440AFF44FA395B82F9C1" blockId="9.[151,1436,1330,2002]" pageId="9" pageNumber="46">
<emphasis id="B94CF90B1C7A440AFF44FA395959FA38" bold="true" box="[199,542,1474,1499]" pageId="9" pageNumber="46">Material examined. Pliocene</emphasis>
<emphasis id="B94CF90B1C7A440AFDA8FA3859D4FA38" box="[555,659,1475,1499]" italics="true" pageId="9" pageNumber="46">Zanclean</emphasis>
: Villa Filicaia (Florence),
<geoCoordinate id="EE0C43DE1C7A440AFC41FA395F09FA39" box="[962,1102,1474,1498]" direction="north" orientation="latitude" pageId="9" pageNumber="46" precision="15" value="43.540554">43°3226”N</geoCoordinate>
,
<geoCoordinate id="EE0C43DE1C7A440AFBD8FA395FA4FA39" box="[1115,1251,1474,1498]" direction="east" orientation="longitude" pageId="9" pageNumber="46" precision="15" value="10.927222">10°5538”E</geoCoordinate>
, 1 sh.;
<emphasis id="B94CF90B1C7A440AFAC6FA3F5B81FA1D" italics="true" pageId="9" pageNumber="46">Piacenzian</emphasis>
: La Speranza (Siena),
<geoCoordinate id="EE0C43DE1C7A440AFE4EFA1D591FFA1D" box="[461,600,1510,1534]" direction="north" orientation="latitude" pageId="9" pageNumber="46" precision="15" value="43.44139">43°2629”N</geoCoordinate>
,
<geoCoordinate id="EE0C43DE1C7A440AFDE5FA1D59AAFA1D" box="[614,749,1510,1534]" direction="east" orientation="longitude" pageId="9" pageNumber="46" precision="15" value="11.100278">11°0601”E</geoCoordinate>
, 5 sh.; Poggio alla Staffa (Siena),
<geoCoordinate id="EE0C43DE1C7A440AFBF4FA1D5E44FA1D" box="[1143,1283,1510,1534]" direction="north" orientation="latitude" pageId="9" pageNumber="46" precision="15" value="43.444447">43°2640”N</geoCoordinate>
,
<geoCoordinate id="EE0C43DE1C7A440AFA93FA1D5ED0FA1D" box="[1296,1431,1510,1534]" direction="east" orientation="longitude" pageId="9" pageNumber="46" precision="15" value="11.0925">11°0533”E</geoCoordinate>
, 9sh.
</paragraph>
</subSubSection>
<subSubSection id="C32276921C7A4409FF44F9D55823FF14" lastPageId="10" lastPageNumber="47" pageId="9" pageNumber="46" type="description">
<paragraph id="8B8725191C7A4409FF44F9D55823FF14" blockId="9.[151,1436,1330,2002]" lastBlockId="10.[151,1437,151,500]" lastPageId="10" lastPageNumber="47" pageId="9" pageNumber="46">
<emphasis id="B94CF90B1C7A440AFF44F9D55A1EF9A4" bold="true" box="[199,345,1582,1607]" pageId="9" pageNumber="46">Description.</emphasis>
Shell very small (max length
<quantity id="4CC088FC1C7A440AFD2BF9D559B8F9A5" box="[680,767,1582,1606]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="9" pageNumber="46" unit="mm" value="4.7">4.7 mm</quantity>
, max diameter 2.0 mm, see Appendix 1), ovate, solid, whitish or buff-coloured with several paler lines faintly visible on last whorl (
<figureCitation id="1303399C1C7A440AFC46F9A95F65F989" box="[965,1058,1618,1642]" captionStart="FIGURES 19 27" captionStartId="8.[151,264,1649,1672]" captionTargetBox="[151,1436,195,1628]" captionTargetId="figure@8.[151,1436,195,1628]" captionTargetPageId="8" captionText="FIGURES 19 27. Haedropleura formosa n. sp. 19 21, # 54, Maiola (Bologna), Gelasian (Pleistocene); 22 24, # 46 holotype, 25 27, # 51 paratype, Codrignano (Bologna), Calabrian (Pleistocene). FIGURES 28 36. H. parva n. sp. 28 30, # 72, La Speranza (Siena), Piacenzian (Pliocene); 31 33, # 63 paratype, 34 36, # 62 holotype, Poggio alla Staffa (Siena), Zanclean (Pliocene). Scale bars 1 mm; # = shell number in Appendix 1." httpUri="https://zenodo.org/record/206564/files/figure.png" pageId="9" pageNumber="46">Figs. 22</figureCitation>
, 28). Protoconch entirely smooth, paucispiral (of 1.31.4 whorls; average diameter
<quantity id="4CC088FC1C7A440AFD38F98D5865F96D" box="[699,802,1654,1678]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="9" pageNumber="46" unit="mm" value="0.58">0.58 mm</quantity>
, SD=
<quantity id="4CC088FC1C7A440AFCDDF98D5881F96D" box="[862,966,1654,1678]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="46" unit="mm" value="0.02">0.02 mm</quantity>
; see Appendix 1). Protoconchteleoconch transition marked well by relatively strong fold or, in most specimens, by narrow sigmoid plica. Teleoconch glossy if well-preserved, of max. ~4.0 whorls, with cyrtoconic spire. Whorls sides almost flat in large specimens, whereas young specimens tend to have moderately convex whorls with periphery below mid-whorl, sutural ramp slightly concave. Suture impressed, undulated. Aperture oblong with evident parietal callus (see Appendix 1). Columellar lip straight, thin. Outer lip thin, backed by wide, round varix, visible only in specimens with more than three teleoconch whorls; varix spreading forwards, corresponding to shallow but broad subsutural sinus, then curving rapidly downwards, extending almost straight to ill-defined anterior siphonal canal; canal wide, without notch. Last whorl about 2/3 of shell length. Axial sculpture consists of 79 prominent, straight axial ribs and well-marked grow lines (see Appendix 1). Axial ribs on spire whorls pinched adapically, extending from suture to suture; less numerous on last whorl in most specimens (mode=8, Appendix 1), fading rapidly towards neck. Spiral sculpture consists of dense, faint, close-set threads, with very shallow incised interspaces, evident only in well-preserved specimens.
<emphasis id="B94CF90B1C794409FF44FF1B5A09FF1B" bold="true" box="[199,334,224,248]" pageId="10" pageNumber="47">Etymology.</emphasis>
This species is named for its small dimensions.
</paragraph>
</subSubSection>
<subSubSection id="C32276921C794409FF44FEF85E34FEFF" box="[199,1395,259,285]" pageId="10" pageNumber="47" type="distribution">
<paragraph id="8B8725191C794409FF44FEF85E34FEFF" blockId="10.[151,1437,151,500]" box="[199,1395,259,285]" pageId="10" pageNumber="47">
<emphasis id="B94CF90B1C794409FF44FEF85A27FEFF" bold="true" box="[199,352,259,284]" pageId="10" pageNumber="47">Distribution.</emphasis>
<taxonomicName id="4C385E9A1C794409FEE4FEFF590AFEFF" box="[359,589,260,284]" class="Gastropoda" family="Horaiclavidae" genus="Haedropleura" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="47" phylum="Mollusca" rank="species" species="parva" status="sp. nov.">
<emphasis id="B94CF90B1C794409FEE4FEFF590AFEFF" box="[359,589,260,284]" italics="true" pageId="10" pageNumber="47">Haedropleura parva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F44701C794409FDD7FEFF59D4FEFF" box="[596,659,260,284]" pageId="10" pageNumber="47" rank="species">
<emphasis id="B94CF90B1C794409FDD7FEFF59D4FEFF" bold="true" box="[596,659,260,284]" pageId="10" pageNumber="47">n. sp.</emphasis>
</taxonomicNameLabel>
is known from several Pliocene outcrops in Tuscany (see above).
</paragraph>
</subSubSection>
<subSubSection id="C32276921C794409FF44FED35E5DFE17" pageId="10" pageNumber="47" type="discussion">
<paragraph id="8B8725191C794409FF44FED35E5DFE17" blockId="10.[151,1437,151,500]" pageId="10" pageNumber="47">
<emphasis id="B94CF90B1C794409FF44FED35A7CFEA3" bold="true" box="[199,315,296,320]" pageId="10" pageNumber="47">Remarks.</emphasis>
<taxonomicName id="4C385E9A1C794409FEC5FED35977FEA3" box="[326,560,296,320]" class="Gastropoda" family="Horaiclavidae" genus="Haedropleura" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="47" phylum="Mollusca" rank="species" species="parva">
<emphasis id="B94CF90B1C794409FEC5FED35977FEA3" box="[326,560,296,320]" italics="true" pageId="10" pageNumber="47">Haedropleura parva</emphasis>
</taxonomicName>
has all the principal characters of the genus
<taxonomicName id="4C385E9A1C794409FBC8FED35FAFFEA3" box="[1099,1256,296,320]" class="Gastropoda" family="Horaiclavidae" genus="Haedropleura" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B94CF90B1C794409FBC8FED35FAFFEA3" box="[1099,1256,296,320]" italics="true" pageId="10" pageNumber="47">Haedropleura</emphasis>
</taxonomicName>
. Its very small dimensions coupled with adult features (that is, parietal callus and variced outer lip) make the species distinctive. The new species has a certain teleoconch affinity with
<taxonomicName id="4C385E9A1C794409FD75FE8A58F7FE6B" box="[758,944,368,392]" class="Gastropoda" family="Horaiclavidae" genus="Haedropleura" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="47" phylum="Mollusca" rank="species" species="septangularis">
<emphasis id="B94CF90B1C794409FD75FE8A58F7FE6B" box="[758,944,368,392]" italics="true" pageId="10" pageNumber="47">H. septangularis</emphasis>
</taxonomicName>
. However, it is easily differentiated from
<taxonomicName id="4C385E9A1C794409FA02FE8A5A77FE4F" class="Gastropoda" family="Horaiclavidae" genus="Haedropleura" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="47" phylum="Mollusca" rank="species" species="septangularis">
<emphasis id="B94CF90B1C794409FA02FE8A5A77FE4F" italics="true" pageId="10" pageNumber="47">H. septangularis</emphasis>
</taxonomicName>
by its smaller teleoconch (average height
<quantity id="4CC088FC1C794409FC9CFE6F583DFE4F" box="[799,890,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="10" pageNumber="47" unit="mm" value="4.1">4.1 mm</quantity>
, SD=
<quantity id="4CC088FC1C794409FC3FFE6F5F51FE4F" box="[956,1046,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="47" unit="mm" value="0.6">0.6 mm</quantity>
vs.
<quantity id="4CC088FC1C794409FBC9FE6F5FE2FE4F" box="[1098,1189,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="10" pageNumber="47" unit="mm" value="9.3">9.3 mm</quantity>
, SD=
<quantity id="4CC088FC1C794409FB64FE6F5E06FE4F" box="[1255,1345,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="47" unit="mm" value="2.4">2.4 mm</quantity>
respectively) and by its different protoconch development (paucispiral vs. multispiral). The combination of adult features and small size make
<taxonomicName id="4C385E9A1C794409FE03FE265AA4FE17" box="[384,483,477,500]" class="Gastropoda" family="Horaiclavidae" genus="Haedropleura" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="47" phylum="Mollusca" rank="species" species="parva">
<emphasis id="B94CF90B1C794409FE03FE265AA4FE17" box="[384,483,477,500]" italics="true" pageId="10" pageNumber="47">H. parva</emphasis>
</taxonomicName>
easily distinguishable also from all congeneric species (see Appendix 1).
</paragraph>
</subSubSection>
</treatment>
</document>