329 lines
36 KiB
XML
329 lines
36 KiB
XML
<document id="AC57C86B6F6FDF84C0D30B986DC634B1" ID-DOI="10.5281/zenodo.5169251" ID-ISSN="1942-1354" ID-Zenodo-Dep="5169251" ID-ZooBank="CC0A1BA6-BA06-402C-BBAF-D794DE86C22A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628275262044" checkinUser="felipe" docAuthor="Moskalenko, Sergey" docDate="2017" docId="039187F0FFE60571FF05FEADFDAD02BD" docLanguage="en" docName="InsectaMundi.0580.1-10.pdf" docOrigin="Insecta Mundi 2017 (580)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Eupatorus gracillicornis subsp. prandii Moskalenko 2017, subsp. nov." docType="treatment" docVersion="3" lastPageNumber="3" masterDocId="FFA8FF88FFE50575FFC5FFAFFFED0307" masterDocTitle="A new subspecies of Eupatorus gracillicornis Arrow, 1908 from Southern Vietnam (Coleoptera, Scarabaeidae, Dynastinae)" masterLastPageNumber="10" masterPageNumber="1" pageNumber="2" updateTime="1699039078968" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="512E19D368206F1E2C314D63DFCB270B">
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<mods:title id="1C1A5B0444F45CE84AB00E75FF43821E">A new subspecies of Eupatorus gracillicornis Arrow, 1908 from Southern Vietnam (Coleoptera, Scarabaeidae, Dynastinae)</mods:title>
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<mods:name id="44442CB99423BBDB47E7FDDF9735525E" type="personal">
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<mods:roleTerm id="B7EDE9ADB53F7B568494A1221DA97702">Author</mods:roleTerm>
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<mods:namePart id="9C7106FD5D6B266B77C0937242E9124B">Moskalenko, Sergey</mods:namePart>
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<mods:typeOfResource id="89C972176A925E68C77A509666879E17">text</mods:typeOfResource>
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<mods:title id="77409E2D486669E7ECF0F8650F4FC0B5">Insecta Mundi</mods:title>
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<mods:part id="90373D20B5D0B509520B4B45288F4857">
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<mods:date id="5411A957C2C1E2D560B1F736F9E607F0">2017</mods:date>
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<mods:number id="7C405AFCC51E987D3DCA1E00A2044101">2017-10-27</mods:number>
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<mods:detail id="9C269027D3CCADB6A191BF14ED3BAB4C" type="volume">
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<mods:number id="FD23EC6F0C51183BA521ADDB14302934">2017</mods:number>
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<mods:detail id="E06685D270B57CD921533988A92F0496" type="issue">
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<mods:number id="06417FE11EC1811F9C95F50333070411">580</mods:number>
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<mods:classification id="E8F912619854185E9ABFA762043C6AF3">journal article</mods:classification>
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<mods:identifier id="7A9B56816BABA8FC819A4E1931150A92" type="DOI">10.5281/zenodo.5169251</mods:identifier>
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<mods:identifier id="4A8B02A07A4C26126D4C5BC0618678AF" type="ISSN">1942-1354</mods:identifier>
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<treatment id="039187F0FFE60571FF05FEADFDAD02BD" LSID="urn:lsid:plazi:treatment:039187F0FFE60571FF05FEADFDAD02BD" httpUri="http://treatment.plazi.org/id/039187F0FFE60571FF05FEADFDAD02BD" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
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<subSubSection id="C322656DFFE60576FF05FEADFCCA021D" box="[192,807,257,282]" pageId="3" pageNumber="2" type="nomenclature">
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<paragraph id="8B8736E6FFE60576FF05FEADFCCA021D" blockId="3.[192,807,257,314]" box="[192,807,257,282]" pageId="3" pageNumber="2">
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<heading id="D0CF818AFFE60576FF05FEADFCCA021D" bold="true" box="[192,807,257,282]" fontSize="10" level="2" pageId="3" pageNumber="2" reason="3">
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<emphasis id="B94CEAF4FFE60576FF05FEADFCCA021D" bold="true" box="[192,807,257,282]" pageId="3" pageNumber="2">
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<taxonomicName id="4C384D65FFE60576FF05FEADFD6B021D" authority="Moskalenko, 2017" authorityName="Moskalenko" authorityYear="2017" box="[192,646,257,282]" class="Insecta" family="Dynastidae" genus="Eupatorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="subSpecies" species="gracillicornis" status="subsp. nov." subSpecies="prandii">
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<emphasis id="B94CEAF4FFE60576FF05FEADFD6B021D" bold="true" box="[192,646,257,282]" italics="true" pageId="3" pageNumber="2">Eupatorus gracillicornis prandii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27F578FFFE60576FD4BFEADFCCA021D" box="[654,807,258,282]" pageId="3" pageNumber="2" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C322656DFFE60576FF05FE8DFAB5027D" pageId="3" pageNumber="2" type="description">
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<paragraph id="8B8736E6FFE60576FF05FE8DFD80023D" blockId="3.[192,807,257,314]" box="[192,621,290,314]" pageId="3" pageNumber="2">
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(
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<figureCitation id="13032A63FFE60576FF0CFE8DFEC3023D" box="[201,302,290,314]" captionStart="Figures 1–32" captionStartId="5.[192,288,1592,1614]" captionTargetBox="[200,1434,207,1560]" captionTargetId="figure-156@5.[192,1440,192,1568]" captionTargetPageId="5" captionText="Figures 1–32. Eupatorus gracillicornis ssp. males in dorsal view (Scale bar – 2 cm). 1–8) Eupatorus gracillicornis prandii ssp. nov. from S. Vietnam, Lam Dong Prov. 1) Holotype. 2–8) Paratypes. 9–13) Eupatorus g. gracillicornis from N. Vietnam, Sa Pa, Lao Cai Prov. 14) Same from N. Vietnam, Pioac Mt. Cao Bang Prov. 15) Same from W. Malaysia, Perak. 16) Eupatorus g. davidgohi from W. Malaysia, Genting Highlands 17) Eupatorus g. gracillicornis from N. Thailand, Wiang Pa Pao, Chang Rai 18, 19, 21) Same from NW. Thailand, Ban Mae, Umphang 20) Same from N. Thailand, Mae Wang, Chang Mai 22) Same from N. Thailand, Doi Saket 23–24) Eupatorus g. edai paratypes from W. Thailand, W. Tak Prov. 25–26) Eupatorus g. gracillicornis from S. China, S. Yunnan, Liyngshan Mt. 27–30) Same from NE. Laos, Mt. Phan 31–32) Eupatorus g. kimioi from SW. Thailand, Kanchanaburi Prov." figureDoi="http://doi.org/10.5281/zenodo.5169253" httpUri="https://zenodo.org/record/5169253/files/figure.png" pageId="3" pageNumber="2">Fig. 1–8</figureCitation>
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,
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<figureCitation id="13032A63FFE60576FEF9FE8DFE64023D" box="[316,393,290,314]" captionStart="Figures 33–66" captionStartId="6.[192,287,1702,1724]" captionTargetBox="[280,1356,195,1679]" captionTargetId="figure-17@6.[275,1367,192,1682]" captionTargetPageId="6" captionText="Figures 33–66. Eupatorus gracillicornis ssp. males in lateral view (Scale bar – 2 cm). 33–42) Eupatorus g. prandii ssp. nov. from S. Vietnam, Lam Dong. 33) Holotype. 34–42) Paratypes. 43–46) Eupatorus g. gracillicornis from NE. Laos, Mt. Phan 47, 48) Same, from S. China, S. Yunnan, Liyngshan Mt. 49) Same, from N. Thailand, Doi Saket 50–52) Same, from NW. Thailand, Ban Mae, Umphang 53) Same, from N. Thailand, Mae Wang, Chang Mai 54) Same, from N. Thailand, Wiang Pa Pao, Chang Rai 55) Same, from N. Vietnam. Pioac Mt. Cao Bang Prov. 56, 57) Eupatorus g. kimioi from SW. Thailand, Kanchanaburi Prov. 58, 59) Eupatorus g. edai paratypes from W. Thailand, W. Tak Prov. 60) Eupatorus g. davidgohi from W. Malaysia, Genting Highlands 61) Eupatorus g. gracillicornis from W. Malaysia, Perak 62–66) Eupatorus g. gracillicornis from N. Vietnam, Sa Pa, Lao Cai Prov." figureDoi="http://doi.org/10.5281/zenodo.5169255" httpUri="https://zenodo.org/record/5169255/files/figure.png" pageId="3" pageNumber="2">33–42</figureCitation>
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,
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<figureCitation id="13032A63FFE60576FE51FE8DFE0C023D" box="[404,481,290,314]" captionStart="Figures 67–73" captionStartId="7.[192,289,592,614]" captionTargetBox="[207,1432,197,568]" captionTargetId="figure-163@7.[192,1440,192,577]" captionTargetPageId="7" captionText="Figures 67–73. Eupatorus gracillicornis ssp. females in dorsal view (Scale bar – 2cm). 67, 68) Eupatorus gracillicornis prandii ssp. nov. from S. Vietnam, Lam Dong Prov. 67) Allotype. 68) Paratype. 69) Eupatorus g. gracillicornis from N. Vietnam, Pioac Mt. Cao Bang Prov. 70) Same from N. Vietnam, Sa Pa, Lao Cai Prov. 71) Same from NE. Laos, Mt. Phan 72) Same from N. Thailand, Doi Saket 73) Eupatorus g. davidgohi from W. Malaysia, Genting Highlands." figureDoi="http://doi.org/10.5281/zenodo.5169257" httpUri="https://zenodo.org/record/5169257/files/figure.png" pageId="3" pageNumber="2">67, 68</figureCitation>
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,
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<figureCitation id="13032A63FFE60576FE28FE8DFDE0023D" box="[493,525,290,314]" captionStart="Figures 74–79" captionStartId="7.[192,288,1788,1810]" captionTargetBox="[205,1427,858,1768]" captionTargetId="figure-165@7.[192,1440,858,1769]" captionTargetPageId="7" captionText="Figures 74–79. Eupatorus gracillicornis ssp. major males head and pronotum in dorsal view. 74) Eupatorus gracillicornis prandii ssp. nov. holotype from S. Vietnam, Lam Dong Prov. 75) Eupatorus g. gracillicornis from N. Thailand, Wiang Pa Pao 76) Same from N. Vietnam, Sa Pa, Lao Cai Prov 77) Eupatorus g. davidgohi from W. Malaysia, Genting Highlands 78) Eupatorus g. edai paratype from W. Thailand, W. Tak Prov. 79) Eupatorus g. kimioi from SW. Thailand, Kanchanaburi Prov." figureDoi="http://doi.org/10.5281/zenodo.5169259" httpUri="https://zenodo.org/record/5169259/files/figure.png" pageId="3" pageNumber="2">74</figureCitation>
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,
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<figureCitation id="13032A63FFE60576FDDFFE8DFDD7023D" box="[538,570,290,314]" captionStart="Figures 80–84" captionStartId="8.[192,290,1004,1026]" captionTargetBox="[246,1403,196,986]" captionTargetId="figure-93@8.[237,1403,192,986]" captionTargetPageId="8" captionText="Figures 80–84. Male genitalia of Eupatorus gracillicornis ssp. in caudal and lateral view. 80) Eupatorus gracillicornis prandii ssp. nov. from S.Vietnam, Lam Dong Prov. 81) Eupatorus g. gracillicornis from N. Thailand, Chang Rai 82) Eupatorus g. kimioi from SW.Thailand, Kanchanaburi Prov. 83) Eupatorus g. edai from W. Thailand, W. Tak Prov. 84) Eupatorus g. davidgohi holotype from W. Malaysia, Fraser’s Hill; this illustration is derived from a photograph of the holotype presented in Yamaya (2013)." figureDoi="http://doi.org/10.5281/zenodo.5169261" httpUri="https://zenodo.org/record/5169261/files/figure.png" pageId="3" pageNumber="2">80</figureCitation>
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,
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<figureCitation id="13032A63FFE60576FD83FE8DFD89023D" box="[582,612,290,314]" captionStart="Figure 88" captionStartId="11.[192,276,1711,1733]" captionTargetBox="[211,1369,209,1687]" captionTargetId="figure-30@11.[192,1376,192,1694]" captionTargetPageId="11" captionText="Figure 88. Eupatorus gracillicornis prandii ssp. nov. holotype and allotype close-up view (Scale bar – 1 cm)." figureDoi="http://doi.org/10.5281/zenodo.5169269" httpUri="https://zenodo.org/record/5169269/files/figure.png" pageId="3" pageNumber="2">88</figureCitation>
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)
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</paragraph>
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<paragraph id="8B8736E6FFE60576FF05FECEFAB5027D" blockId="3.[192,1368,353,378]" box="[192,1368,353,378]" pageId="3" pageNumber="2">
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<emphasis id="B94CEAF4FFE60576FF05FECEFE86027E" bold="true" box="[192,363,353,377]" pageId="3" pageNumber="2">Description.</emphasis>
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Mostly black, elytra straw-colored except for black sutural and external margins.
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</paragraph>
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</subSubSection>
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<subSubSection id="C322656DFFE60576FF05FE0DFD2C01FD" pageId="3" pageNumber="2" type="materials_examined">
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<paragraph id="8B8736E6FFE60576FF05FE0DFDC0015D" blockId="3.[192,1441,418,602]" pageId="3" pageNumber="2">
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<emphasis id="B94CEAF4FFE60576FF05FE0DFED302BD" bold="true" box="[192,318,418,442]" pageId="3" pageNumber="2">
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<typeStatus id="54838844FFE60576FF05FE0DFED302BD" box="[192,318,418,442]" pageId="3" pageNumber="2" type="holotype">Holotype</typeStatus>
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</emphasis>
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. Length
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<quantity id="4CC09B03FFE60576FE6AFE0DFDF002BE" box="[431,541,418,442]" metricMagnitude="-2" metricUnit="m" metricValue="6.279999999999999" pageId="3" pageNumber="2" unit="mm" value="62.8">62.8 mm</quantity>
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, width
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<quantity id="4CC09B03FFE60576FDBEFE0DFD0702BE" box="[635,746,418,442]" metricMagnitude="-2" metricUnit="m" metricValue="3.4799999999999995" pageId="3" pageNumber="2" unit="mm" value="34.8">34.8 mm</quantity>
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. Clypeus with two small teeth. Head with long cephalic horn that may be strongly curved backwards or more or less straight. Pronotum sparsely punctate, mainly on the sides, shiny, with two anterior pronotal horns and two posterior horns, pair of posterior horns ~33% shorter than anterior pair. Elytra elongate, oval, 1.17 times as long as wide, widest behind the middle part. Elytra shiny, sparsely punctate near the sutural stripe. The sutural stripe is thick, expanded near the scutellum.
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</paragraph>
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<paragraph id="8B8736E6FFE60576FF05FD2EFD2C01FD" blockId="3.[192,1440,641,762]" pageId="3" pageNumber="2">
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<emphasis id="B94CEAF4FFE60576FF05FD2EFED9019E" bold="true" box="[192,308,641,665]" pageId="3" pageNumber="2">
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<typeStatus id="54838844FFE60576FF05FD2EFED9019E" box="[192,308,641,665]" pageId="3" pageNumber="2" type="allotype">Allotype</typeStatus>
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</emphasis>
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. Length
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<quantity id="4CC09B03FFE60576FE60FD2EFDFE019E" box="[421,531,641,665]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" pageId="3" pageNumber="2" unit="mm" value="62.5">62.5 mm</quantity>
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, width
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<quantity id="4CC09B03FFE60576FDB5FD2DFD0D019E" box="[624,736,642,666]" metricMagnitude="-2" metricUnit="m" metricValue="3.19" pageId="3" pageNumber="2" unit="mm" value="31.9">31.9 mm</quantity>
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. Clypeus with two small teeth, pronotum simply convex, strongly punctate, and closely rugose at sides. Pronotum with oval dimple at center of basal part. Elytra elongate, oval, 1.17 times as long as wide, widest behind the middle part, strongly punctate on the disc, covered with extremely short pubescence.
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</paragraph>
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</subSubSection>
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<subSubSection id="C322656DFFE60576FF05FC8EFC9F06FD" pageId="3" pageNumber="2" type="diagnosis">
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<paragraph id="8B8736E6FFE60576FF05FC8EFEA5003E" blockId="3.[192,328,801,825]" box="[192,328,801,825]" pageId="3" pageNumber="2">
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<emphasis id="B94CEAF4FFE60576FF05FC8EFEA5003E" bold="true" box="[192,328,801,825]" pageId="3" pageNumber="2">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="8B8736E6FFE60576FF05FCCDFAF0007D" blockId="3.[192,1309,866,890]" box="[192,1309,866,890]" pageId="3" pageNumber="2">
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The new subspecies is most similar to
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<emphasis id="B94CEAF4FFE60576FD5DFCCDFC97007D" box="[664,890,866,890]" italics="true" pageId="3" pageNumber="2">
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E. g.
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<taxonomicName id="4C384D65FFE60576FD12FCCDFC97007D" authorityName="Arrow" authorityYear="1908" box="[727,890,866,890]" class="Insecta" family="Dynastidae" genus="Eupatorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="gracillicornis">gracillicornis</taxonomicName>
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</emphasis>
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, but differs in the following ways:
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</paragraph>
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<paragraph id="8B8736E6FFE60576FF05FC0DFE5D073D" blockId="3.[192,1440,930,1338]" pageId="3" pageNumber="2">
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<emphasis id="B94CEAF4FFE60576FF05FC0DFEF500BD" bold="true" box="[192,280,930,954]" pageId="3" pageNumber="2">Males.</emphasis>
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The posterior pronotal horns are distinctly shorter than the anterior ones (all pronotal horns of
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<emphasis id="B94CEAF4FFE60576FF18FC6DFE5000DD" box="[221,445,962,986]" italics="true" pageId="3" pageNumber="2">
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E. g.
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<taxonomicName id="4C384D65FFE60576FEDFFC6DFE5000DD" authorityName="Arrow" authorityYear="1908" box="[282,445,962,986]" class="Insecta" family="Dynastidae" genus="Eupatorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="gracillicornis">gracillicornis</taxonomicName>
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</emphasis>
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are roughly equal in length). In medium-sized males, the anterior pronotal horns are directed more forward and pointed inwards at the apex (anterior pronotal horns of medium-sized
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<emphasis id="B94CEAF4FFE60576FF05FBADFE4F071D" box="[192,418,1026,1050]" italics="true" pageId="3" pageNumber="2">
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E. g.
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<taxonomicName id="4C384D65FFE60576FF3AFBADFE4F071D" authorityName="Arrow" authorityYear="1908" box="[255,418,1026,1050]" class="Insecta" family="Dynastidae" genus="Eupatorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="gracillicornis">gracillicornis</taxonomicName>
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</emphasis>
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males are directed more outward). The sutural stripe is distinctly wider, expanded near the scutellum.
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</paragraph>
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<paragraph id="8B8736E6FFE60576FF35FBEDFB6907BE" blockId="3.[192,1440,930,1338]" pageId="3" pageNumber="2">
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Genitalia differ from those of
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<emphasis id="B94CEAF4FFE60576FDA4FBEDFCA8075D" box="[609,837,1090,1114]" italics="true" pageId="3" pageNumber="2">
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E. g.
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<taxonomicName id="4C384D65FFE60576FD67FBEDFCA8075D" authorityName="Arrow" authorityYear="1908" box="[674,837,1090,1114]" class="Insecta" family="Dynastidae" genus="Eupatorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="gracillicornis">gracillicornis</taxonomicName>
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</emphasis>
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by having parameres that are wider at the apex and are smoothly curved in the central part of the outer margin, (when viewed dorsally;
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<figureCitation id="13032A63FFE60576FAC6FBCDFAB1077D" box="[1283,1372,1122,1146]" captionStart="Figures 80–84" captionStartId="8.[192,290,1004,1026]" captionTargetBox="[246,1403,196,986]" captionTargetId="figure-93@8.[237,1403,192,986]" captionTargetPageId="8" captionText="Figures 80–84. Male genitalia of Eupatorus gracillicornis ssp. in caudal and lateral view. 80) Eupatorus gracillicornis prandii ssp. nov. from S.Vietnam, Lam Dong Prov. 81) Eupatorus g. gracillicornis from N. Thailand, Chang Rai 82) Eupatorus g. kimioi from SW.Thailand, Kanchanaburi Prov. 83) Eupatorus g. edai from W. Thailand, W. Tak Prov. 84) Eupatorus g. davidgohi holotype from W. Malaysia, Fraser’s Hill; this illustration is derived from a photograph of the holotype presented in Yamaya (2013)." figureDoi="http://doi.org/10.5281/zenodo.5169261" httpUri="https://zenodo.org/record/5169261/files/figure.png" pageId="3" pageNumber="2">Fig. 80</figureCitation>
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). The parameres of
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<emphasis id="B94CEAF4FFE60576FEAFFB2DFDA3079D" box="[362,590,1154,1178]" italics="true" pageId="3" pageNumber="2">
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E. g.
|
||
<taxonomicName id="4C384D65FFE60576FE6EFB2DFDA3079D" authorityName="Arrow" authorityYear="1908" box="[427,590,1154,1178]" class="Insecta" family="Dynastidae" genus="Eupatorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="gracillicornis">gracillicornis</taxonomicName>
|
||
</emphasis>
|
||
are distinctly protruded (
|
||
<figureCitation id="13032A63FFE60576FC4AFB2DFC0A079D" box="[911,999,1154,1178]" captionStart="Figures 80–84" captionStartId="8.[192,290,1004,1026]" captionTargetBox="[246,1403,196,986]" captionTargetId="figure-93@8.[237,1403,192,986]" captionTargetPageId="8" captionText="Figures 80–84. Male genitalia of Eupatorus gracillicornis ssp. in caudal and lateral view. 80) Eupatorus gracillicornis prandii ssp. nov. from S.Vietnam, Lam Dong Prov. 81) Eupatorus g. gracillicornis from N. Thailand, Chang Rai 82) Eupatorus g. kimioi from SW.Thailand, Kanchanaburi Prov. 83) Eupatorus g. edai from W. Thailand, W. Tak Prov. 84) Eupatorus g. davidgohi holotype from W. Malaysia, Fraser’s Hill; this illustration is derived from a photograph of the holotype presented in Yamaya (2013)." figureDoi="http://doi.org/10.5281/zenodo.5169261" httpUri="https://zenodo.org/record/5169261/files/figure.png" pageId="3" pageNumber="2">Fig. 81</figureCitation>
|
||
). When viewed laterally, the dorsal margin of the phallobase is much more convex than that of
|
||
<emphasis id="B94CEAF4FFE60576FC5EFB0DFB9307BD" box="[923,1150,1186,1210]" italics="true" pageId="3" pageNumber="2">
|
||
E. g.
|
||
<taxonomicName id="4C384D65FFE60576FC1EFB0DFB9307BD" authorityName="Arrow" authorityYear="1908" box="[987,1150,1186,1210]" class="Insecta" family="Dynastidae" genus="Eupatorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="gracillicornis">gracillicornis</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8736E6FFE60576FF35FB6DFE9C063D" blockId="3.[192,1440,930,1338]" pageId="3" pageNumber="2">
|
||
Males are also distinguishable from those of the Malaysian subspecies
|
||
<emphasis id="B94CEAF4FFE60576FB96FB6DFAE007DD" box="[1107,1293,1218,1242]" italics="true" pageId="3" pageNumber="2">E. g. davidgohi</emphasis>
|
||
by the relatively long posterior pronotal horns (they are 2/3 as long as the anterior pronotal horns, compared to 1/3 as long in
|
||
<emphasis id="B94CEAF4FFE60576FEACFAADFDCF061D" box="[361,546,1282,1306]" italics="true" pageId="3" pageNumber="2">E. g. davidgohi</emphasis>
|
||
), the more punctate pronotum, the slightly shorter tarsus, and the wider sutural stripe.
|
||
</paragraph>
|
||
<paragraph id="8B8736E6FFE60576FF05FACDFB3306BD" blockId="3.[192,1440,1378,1530]" pageId="3" pageNumber="2">
|
||
<emphasis id="B94CEAF4FFE60576FF05FACDFED7067D" bold="true" box="[192,314,1378,1402]" pageId="3" pageNumber="2">Females.</emphasis>
|
||
The elytra are more strongly punctate on the disc, less shiny, and covered with a very short brown pubescence that gives them a velvety appearance. Some individuals have an oval dimple in the middle of the basal part of pronotum, that is not present in nominotypical subspecies.
|
||
</paragraph>
|
||
<paragraph id="8B8736E6FFE60576FF35FA6DFC9F06FD" blockId="3.[192,1440,1378,1530]" pageId="3" pageNumber="2">
|
||
Females are also easily distinguishable from those of
|
||
<emphasis id="B94CEAF4FFE60576FC51FA6DFBBE06DD" box="[916,1107,1474,1498]" italics="true" pageId="3" pageNumber="2">E. g. davidgohi</emphasis>
|
||
, which have less punctate elytra without pubescence, and have more slender tarsi.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C322656DFFE60576FF05F98DFE93055D" pageId="3" pageNumber="2" type="description">
|
||
<paragraph id="8B8736E6FFE60576FF05F98DFE93055D" blockId="3.[192,1440,1569,1626]" pageId="3" pageNumber="2">
|
||
<emphasis id="B94CEAF4FFE60576FF05F98DFE7D053D" bold="true" box="[192,400,1570,1594]" pageId="3" pageNumber="2">Measurements.</emphasis>
|
||
Males (n = 248), length: 48.1–73.9, width: 26.3–39.2. Females (n = 92), length: 46–62.5, width: 24.7–33.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C322656DFFE60571FF05F92DFCE2021D" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2" type="materials_examined">
|
||
<paragraph id="8B8736E6FFE60576FF05F92DFBBA043D" blockId="3.[192,1440,1665,1850]" pageId="3" pageNumber="2">
|
||
<emphasis id="B94CEAF4FFE60576FF05F92DFE65059E" bold="true" box="[192,392,1665,1690]" pageId="3" pageNumber="2">Type material.</emphasis>
|
||
<materialsCitation id="3B503CBBFFE60576FE4BF92DFBA0059E" box="[398,1101,1665,1690]" collectingDate="2015-11" collectorName="Da Lat & Lam Dong Prov. S." country="Vietnam" location="Lam Dong Prov." pageId="3" pageNumber="2" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<typeStatus id="54838844FFE60576FE4BF92DFDEC059D" box="[398,513,1666,1690]" pageId="3" pageNumber="2" type="holotype">Holotype</typeStatus>
|
||
.
|
||
<specimenCount id="9D3EFD6FFFE60576FDCCF92DFDF0059E" box="[521,541,1666,1689]" pageId="3" pageNumber="2" type="male">♂</specimenCount>
|
||
,
|
||
<collectorName id="26CD5330FFE60576FDEFF92DFD6C059E" box="[554,641,1666,1689]" pageId="3" pageNumber="2">Da Lat</collectorName>
|
||
,
|
||
<collectorName id="26CD5330FFE60576FD4EF92DFC83059D" box="[651,878,1666,1690]" pageId="3" pageNumber="2">
|
||
<location id="8EE7603DFFE60576FD4EF92DFCBD059E" LSID="urn:lsid:plazi:treatment:039187F0FFE60571FF05FEADFDAD02BD:8EE7603DFFE60576FD4EF92DFCBD059E" box="[651,848,1666,1690]" country="Vietnam" name="Lam Dong Prov." pageId="3" pageNumber="2">Lam Dong Prov.</location>
|
||
S.
|
||
</collectorName>
|
||
<collectingCountry id="F32F7676FFE60576FCB6F92DFC0C059D" box="[883,993,1666,1690]" name="Vietnam" pageId="3" pageNumber="2">Vietnam</collectingCountry>
|
||
.
|
||
<date id="FF861026FFE60576FC2EF92EFBA0059E" box="[1003,1101,1665,1689]" pageId="3" pageNumber="2" value="2015-11">
|
||
<collectingDate id="EFC2E9CEFFE60576FC2EF92EFBA0059E" box="[1003,1101,1665,1689]" pageId="3" pageNumber="2" value="2015-11">XI.2015</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B503CBBFFE60576FB9DF92DFCC105BD" collectingDate="2014-10" collectorName="Bidoup Nui Ba N. P." country="Vietnam" location="Lam Dong Prov." pageId="3" pageNumber="2" specimenCount="1" specimenCount-female="1" stateProvince="Lam Dong" typeStatus="allotype">
|
||
<typeStatus id="54838844FFE60576FB9DF92DFB2E059D" box="[1112,1219,1666,1690]" pageId="3" pageNumber="2" type="allotype">Allotype</typeStatus>
|
||
.
|
||
<specimenCount id="9D3EFD6FFFE60576FB09F92DFB0D059E" box="[1228,1248,1666,1689]" pageId="3" pageNumber="2" type="female">♀</specimenCount>
|
||
,
|
||
<collectorName id="26CD5330FFE60576FB29F92DFF1B05BE" pageId="3" pageNumber="2">Bidoup Nui Ba N.P.</collectorName>
|
||
1550 alt.
|
||
<collectingRegion id="49FCF804FFE60576FEAAF90DFDDA05BE" box="[367,567,1698,1722]" country="Vietnam" name="Lam Dong" pageId="3" pageNumber="2">Lam Dong Prov.</collectingRegion>
|
||
S.
|
||
<collectingCountry id="F32F7676FFE60576FD99F90DFD2705BD" box="[604,714,1698,1722]" name="Vietnam" pageId="3" pageNumber="2">Vietnam</collectingCountry>
|
||
.
|
||
<date id="FF861026FFE60576FD11F90DFCC105BD" box="[724,812,1698,1722]" pageId="3" pageNumber="2" value="2014-10">
|
||
<collectingDate id="EFC2E9CEFFE60576FD11F90DFCC105BD" box="[724,812,1698,1722]" pageId="3" pageNumber="2" value="2014-10">X.2014</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B503CBBFFE60576FCFDF90DFEC105DD" collectingDate="2015-11" collectorName="Da Lat & Lam Dong Prov. S." country="Vietnam" location="Lam Dong Prov." pageId="3" pageNumber="2" specimenCount="291" specimenCount-female="81" specimenCount-male="210" typeStatus="paratype">
|
||
<typeStatus id="54838844FFE60576FCFDF90DFC5405BD" box="[824,953,1698,1722]" pageId="3" pageNumber="2" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D3EFD6FFFE60576FC06F90DFBFA05BE" box="[963,1047,1698,1722]" pageId="3" pageNumber="2" type="male">210♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3EFD6FFFE60576FBE3F90DFB8605BE" box="[1062,1131,1698,1722]" pageId="3" pageNumber="2" type="female">81♀♀</specimenCount>
|
||
, the same data as for the holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B503CBBFFE60576FEFDF96DFC2305DD" box="[312,974,1730,1754]" collectingDate="2015-11" collectorName="Da Lat & Lam Dong Prov. S." country="Vietnam" location="Lam Dong Prov." pageId="3" pageNumber="2" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
|
||
<specimenCount id="9D3EFD6FFFE60576FEFDF96DFE8205DE" box="[312,367,1730,1754]" pageId="3" pageNumber="2" type="male">3♂♂</specimenCount>
|
||
, the same data as for the holotype but XII.20016
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B503CBBFFE60576FC1EF96DFEC805FD" collectingDate="2014-10" collectorName="Bidoup Nui Ba N. P." country="Vietnam" location="Lam Dong Prov." pageId="3" pageNumber="2" specimenCount="37" specimenCount-female="8" specimenCount-male="29" stateProvince="Lam Dong" typeStatus="paratype">
|
||
<specimenCount id="9D3EFD6FFFE60576FC1EF96DFBCD05DE" box="[987,1056,1730,1754]" pageId="3" pageNumber="2" type="male">29♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3EFD6FFFE60576FBEAF96DFB8B05DE" box="[1071,1126,1730,1754]" pageId="3" pageNumber="2" type="female">8♀♀</specimenCount>
|
||
, the same data as for the allotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B503CBBFFE60576FEF4F94DFC7305FD" box="[305,926,1762,1786]" collectingDate="2014-06" collectionCode="VI" collectorName="Bidoup Nui Ba N. P." country="Vietnam" location="Lam Dong Prov." pageId="3" pageNumber="2" specimenCount="2" specimenCount-male="2" stateProvince="Lam Dong" typeStatus="paratype">
|
||
<specimenCount id="9D3EFD6FFFE60576FEF4F94DFE8505FE" box="[305,360,1762,1786]" pageId="3" pageNumber="2" type="male">2♂♂</specimenCount>
|
||
the same data as for the allotype but
|
||
<date id="FF861026FFE60576FCFEF94DFC7305FD" box="[827,926,1762,1786]" pageId="3" pageNumber="2" value="2014-06">
|
||
<collectingDate id="EFC2E9CEFFE60576FCFEF94DFC7305FD" box="[827,926,1762,1786]" pageId="3" pageNumber="2" value="2014-06">
|
||
<collectionCode id="ED29AE23FFE60576FCFEF94DFCB105FD" box="[827,860,1762,1786]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="2">VI</collectionCode>
|
||
.2014
|
||
</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B503CBBFFE60576FC6FF94DFEF5041D" collectingDate="2013-10" collectorName="Da Tro" country="Vietnam" location="Binh Thuan Prov." pageId="3" pageNumber="2" specimenCount="1" specimenCount-male="1" stateProvince="Binh Thuan" typeStatus="paratype">
|
||
<specimenCount id="9D3EFD6FFFE60576FC6FF94DFC5305FE" box="[938,958,1762,1785]" pageId="3" pageNumber="2" type="male">♂</specimenCount>
|
||
<collectorName id="26CD5330FFE60576FC03F94DFBF205FE" box="[966,1055,1762,1785]" pageId="3" pageNumber="2">Da Tro</collectorName>
|
||
,
|
||
<collectingRegion id="49FCF804FFE60576FBEFF94DFAE505FE" box="[1066,1288,1762,1786]" country="Vietnam" name="Binh Thuan" pageId="3" pageNumber="2">Binh Thuan Prov.</collectingRegion>
|
||
S.
|
||
<collectingCountry id="F32F7676FFE60576FAEBF94DFA7105FD" box="[1326,1436,1762,1786]" name="Vietnam" pageId="3" pageNumber="2">Vietnam</collectingCountry>
|
||
.
|
||
<date id="FF861026FFE60576FF05F8ADFEF5041D" box="[192,280,1794,1818]" pageId="3" pageNumber="2" value="2013-10">
|
||
<collectingDate id="EFC2E9CEFFE60576FF05F8ADFEF5041D" box="[192,280,1794,1818]" pageId="3" pageNumber="2" value="2013-10">X.2013</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B503CBBFFE60576FEE1F8ADFC08041D" box="[292,997,1794,1818]" collectingDate="2014-10" collectorName="Ton" country="Vietnam" location="Lam Dong Prov." pageId="3" pageNumber="2" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong" typeStatus="paratype">
|
||
<specimenCount id="9D3EFD6FFFE60576FEE1F8ADFED5041E" box="[292,312,1794,1817]" pageId="3" pageNumber="2" type="male">♂</specimenCount>
|
||
Dạ
|
||
<collectorName id="26CD5330FFE60576FEADF8ADFE77041E" box="[360,410,1794,1817]" pageId="3" pageNumber="2">Ton</collectorName>
|
||
, Bao Lam,
|
||
<collectingRegion id="49FCF804FFE60576FDE1F8ADFD00041E" box="[548,749,1794,1818]" country="Vietnam" name="Lam Dong" pageId="3" pageNumber="2">Lam Dong Prov.</collectingRegion>
|
||
S.
|
||
<collectingCountry id="F32F7676FFE60576FCD1F8ADFC6F041D" box="[788,898,1794,1818]" name="Vietnam" pageId="3" pageNumber="2">Vietnam</collectingCountry>
|
||
.
|
||
<date id="FF861026FFE60576FC48F8ADFC08041D" box="[909,997,1794,1818]" pageId="3" pageNumber="2" value="2014-10">
|
||
<collectingDate id="EFC2E9CEFFE60576FC48F8ADFC08041D" box="[909,997,1794,1818]" pageId="3" pageNumber="2" value="2014-10">X.2014</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B503CBBFFE60576FC34F8ADFE14043D" collectingDate="2014-10" collectorName="Loc Tan" country="Vietnam" location="Lam Dong Prov." pageId="3" pageNumber="2" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Lam Dong" typeStatus="paratype">
|
||
<specimenCount id="9D3EFD6FFFE60576FC34F8ADFBE8041E" box="[1009,1029,1794,1817]" pageId="3" pageNumber="2" type="male">♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3EFD6FFFE60576FBD1F8ADFBC5041E" box="[1044,1064,1794,1817]" pageId="3" pageNumber="2" type="female">♀</specimenCount>
|
||
<collectorName id="26CD5330FFE60576FBEAF8ADFB7B041E" box="[1071,1174,1794,1817]" pageId="3" pageNumber="2">Loc Tan</collectorName>
|
||
, Bao Lam,
|
||
<collectingRegion id="49FCF804FFE60576FAE5F8ADFEEC043E" country="Vietnam" name="Lam Dong" pageId="3" pageNumber="2">Lam Dong Prov.</collectingRegion>
|
||
S.
|
||
<collectingCountry id="F32F7676FFE60576FEEDF88DFE7B043D" box="[296,406,1826,1850]" name="Vietnam" pageId="3" pageNumber="2">Vietnam</collectingCountry>
|
||
.
|
||
<date id="FF861026FFE60576FE64F88DFE14043D" box="[417,505,1826,1850]" pageId="3" pageNumber="2" value="2014-10">
|
||
<collectingDate id="EFC2E9CEFFE60576FE64F88DFE14043D" box="[417,505,1826,1850]" pageId="3" pageNumber="2" value="2014-10">X.2014</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B503CBBFFE60576FDC0F88DFBBE043D" box="[517,1107,1826,1850]" collectingDate="2014-10" collectorName="Bao Loc" country="Vietnam" location="Lam Dong Prov." pageId="3" pageNumber="2" specimenCount="1" specimenCount-female="1" stateProvince="Lam Dong" typeStatus="paratype">
|
||
<specimenCount id="9D3EFD6FFFE60576FDC0F88DFDF4043E" box="[517,537,1826,1849]" pageId="3" pageNumber="2" type="female">♀</specimenCount>
|
||
<collectorName id="26CD5330FFE60576FDE5F88DFD68043E" box="[544,645,1826,1849]" pageId="3" pageNumber="2">Bao Loc</collectorName>
|
||
,
|
||
<collectingRegion id="49FCF804FFE60576FD54F88DFCB7043E" box="[657,858,1826,1850]" country="Vietnam" name="Lam Dong" pageId="3" pageNumber="2">Lam Dong Prov.</collectingRegion>
|
||
S.
|
||
<collectingCountry id="F32F7676FFE60576FC44F88DFC02043D" box="[897,1007,1826,1850]" name="Vietnam" pageId="3" pageNumber="2">Vietnam</collectingCountry>
|
||
.
|
||
<date id="FF861026FFE60576FC3FF88DFBBE043D" box="[1018,1107,1826,1850]" pageId="3" pageNumber="2" value="2014-10">
|
||
<collectingDate id="EFC2E9CEFFE60576FC3FF88DFBBE043D" box="[1018,1107,1826,1850]" pageId="3" pageNumber="2" value="2014-10">X.2014</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8736E6FFE10571FF05FF6DFCE2021D" blockId="4.[192,1440,193,282]" pageId="4" pageNumber="3">
|
||
<emphasis id="B94CEAF4FFE10571FF05FF6DFE5003DE" bold="true" box="[192,445,193,218]" pageId="4" pageNumber="3">Type depositories.</emphasis>
|
||
The
|
||
<typeStatus id="54838844FFE10571FE3CFF6DFD8803DD" box="[505,613,194,218]" pageId="4" pageNumber="3" type="holotype">holotype</typeStatus>
|
||
,
|
||
<typeStatus id="54838844FFE10571FDB5FF6DFD3F03DD" box="[624,722,194,218]" pageId="4" pageNumber="3" type="allotype">allotype</typeStatus>
|
||
and two additional
|
||
<typeStatus id="54838844FFE10571FC01FF6DFBD303DD" box="[964,1086,194,218]" pageId="4" pageNumber="3" type="paratype">paratypes</typeStatus>
|
||
will be entrusted to
|
||
<emphasis id="B94CEAF4FFE10571FAF2FF6DFA7403DD" bold="true" box="[1335,1433,194,218]" pageId="4" pageNumber="3">
|
||
<collectionCode id="ED29AE23FFE10571FAF2FF6DFA7403DD" box="[1335,1433,194,218]" country="USA" httpUri="http://grbio.org/cool/q3kg-qwj4" name="John May Museum of Natural History" pageId="4" pageNumber="3">MNHM</collectionCode>
|
||
</emphasis>
|
||
. Some
|
||
<typeStatus id="54838844FFE10571FECEFF4DFE6803FD" box="[267,389,226,250]" pageId="4" pageNumber="3" type="paratype">paratypes</typeStatus>
|
||
will be deposited in the author’s collection and also in collections of Messrs. Massimo Prandi, Kazuho Kobayashi and Miss Kate Ford.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C322656DFFE10571FF05FEEEFC07025D" box="[192,1002,321,346]" pageId="4" pageNumber="3" type="distribution">
|
||
<paragraph id="8B8736E6FFE10571FF05FEEEFC07025D" blockId="4.[192,1002,321,346]" box="[192,1002,321,346]" pageId="4" pageNumber="3">
|
||
<emphasis id="B94CEAF4FFE10571FF05FEEEFE99025E" bold="true" box="[192,372,321,345]" pageId="4" pageNumber="3">Distribution.</emphasis>
|
||
<collectingRegion id="49FCF804FFE10571FEBEFEEDFE16025D" box="[379,507,322,346]" country="Vietnam" name="Lam Dong" pageId="4" pageNumber="3">Lam Dong</collectingRegion>
|
||
and
|
||
<collectingRegion id="49FCF804FFE10571FDFDFEEDFD20025D" box="[568,717,322,346]" country="Vietnam" name="Binh Thuan" pageId="4" pageNumber="3">Binh Thuan</collectingRegion>
|
||
provinces, S.
|
||
<collectingCountry id="F32F7676FFE10571FCBDFEEDFC0B025D" box="[888,998,322,346]" name="Vietnam" pageId="4" pageNumber="3">Vietnam</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C322656DFFE10571FF05FE2DFDAD02BD" pageId="4" pageNumber="3" type="etymology">
|
||
<paragraph id="8B8736E6FFE10571FF05FE2DFDAD02BD" blockId="4.[192,1440,386,442]" pageId="4" pageNumber="3">
|
||
<emphasis id="B94CEAF4FFE10571FF05FE2DFEB1029D" bold="true" box="[192,348,386,410]" pageId="4" pageNumber="3">Etymology.</emphasis>
|
||
This new subspecies is named after Mr. Massimo Prandi from
|
||
<collectingCountry id="F32F7676FFE10571FB92FE2DFB79029D" box="[1111,1172,386,410]" name="Italy" pageId="4" pageNumber="3">Italy</collectingCountry>
|
||
, who provided invaluable assistance in my research.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |