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<document id="69A9F79157A61B564758D4F3422DC57F" ID-DOI="10.5252/z2016n4a1" ID-ISSN="1638-9387" ID-Zenodo-Dep="4578278" ID-ZooBank="urn:lsid:zoobank.org:pub:9E796669-C345-42D6-B0F9-95288DB701EE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614790673595" checkinUser="felipe" docAuthor="Desutter-Grandcolas, Laure, Anso, Jérémy &amp; Jourdan, Hervé" docDate="2016" docId="039187CEC606712AFED5F94D17889D3D" docLanguage="en" docName="Zoosystema.38.4.405-452.pdf" docOrigin="Zoosystema 38 (4)" docSource="http://dx.doi.org/10.5252/z2016n4a1" docStyle="DocumentStyle:0AF8C315773078909029C6FC3CC05C6C.3:Zoosystema.2015-2017.journal_article" docStyleId="0AF8C315773078909029C6FC3CC05C6C" docStyleName="Zoosystema.2015-2017.journal_article" docStyleVersion="3" docTitle="Agnotecous Saussure 1878" docType="treatment" docVersion="3" lastPageNumber="444" masterDocId="FFA8FFB6C621710DFFC6FFBF14609B36" masterDocTitle="Crickets of New Caledonia (Insecta, Orthoptera, Grylloidea): a key to genera, with diagnoses of extant genera and descriptions of new taxa" masterLastPageNumber="452" masterPageNumber="405" pageNumber="444" updateTime="1698879331528" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="573E91654ECBC1ABCB3991FAD157F68B">Crickets of New Caledonia (Insecta, Orthoptera, Grylloidea): a key to genera, with diagnoses of extant genera and descriptions of new taxa</mods:title>
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<mods:namePart id="A39F726DFE7C969690E99338653A5E9E">Desutter-Grandcolas, Laure</mods:namePart>
<mods:affiliation id="F2A15D59414BE9EA615948A60C654875">Institut de Systématique, Évolution et Biodiversité, ISYEB - UMR 7205 CNRS, MNHN, UPMC, EPHE, Muséum national dHistoire naturelle, Sorbonne Universités, case postale 50, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) desutter @ mnhn. fr</mods:affiliation>
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<mods:namePart id="78CF58A10452323C3926C9819B44F55E">Anso, Jérémy</mods:namePart>
<mods:affiliation id="C9D17FF557F6284CB91ECAADC1E4654D">Institut de Systématique, Évolution et Biodiversité, ISYEB - UMR 7205 CNRS, MNHN, UPMC, EPHE, Muséum national dHistoire naturelle, Sorbonne Universités, case postale 50, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) and Institut méditerranéen de Biodiversité et dÉcologie marine et continentale (IMBE), Aix-Marseille Université, UMR CNRS, IRD, UAPV, Centre IRD Nouméa, BP A 5, 98848 Nouméa Cedex, (Nouvelle-Calédonie)</mods:affiliation>
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<mods:namePart id="6F484DF730E1ACB036456E4606D0FEB9">Jourdan, Hervé</mods:namePart>
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<paragraph id="8B8736D8C606712AFED5F94D16189C3A" blockId="39.[275,632,1778,1804]" box="[275,632,1778,1804]" pageId="39" pageNumber="444">
<heading id="D0CF81B4C606712AFED5F94D16189C3A" box="[275,632,1778,1804]" centered="true" fontSize="9" level="2" pageId="39" pageNumber="444" reason="2">
Genus
<taxonomicName id="4C384D5BC606712AFEA6F94D16189C3A" authority="Saussure, 1878" authorityName="Saussure" authorityYear="1878" box="[352,632,1778,1804]" class="Insecta" family="Haglotettigoniidae" genus="Agnotecous" kingdom="Animalia" order="Orthoptera" pageId="39" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B94CEACAC606712AFEA6F94D15B09C3A" box="[352,464,1778,1804]" italics="true" pageId="39" pageNumber="444">Agnotecous</emphasis>
<bibRefCitation id="EFA94B29C606712AFE11F94D16189C3A" author="SAUSSURE H. &amp; DE" box="[471,632,1778,1804]" pageId="39" pageNumber="444" pagination="369 - 702" refId="ref36461" refString="SAUSSURE H. DE 1878. - Melanges orthopterologiques. VIe fascicule. Gryllides (2 e partie). Memoires de la Societe de Physique et d'Histoire naturelle de Geneve 25: 369 - 702." type="journal article" year="1878">Saussure, 1878</bibRefCitation>
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</paragraph>
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<paragraph id="8B8736D8C606712AFF42F89615DC9C76" blockId="39.[132,775,1832,1925]" box="[132,444,1832,1857]" pageId="39" pageNumber="444">
<treatmentCitationGroup id="AB2811F6C606712AFF42F89615DC9C76" box="[132,444,1832,1857]" pageId="39" pageNumber="444">
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<taxonomicName id="4C384D5BC606712AFF42F89615D89C76" authority="Saussure, 1878: 546" authorityName="Saussure" authorityPageNumber="546" authorityYear="1878" box="[132,440,1832,1857]" class="Insecta" family="Haglotettigoniidae" genus="Agnotecous" kingdom="Animalia" order="Orthoptera" pageId="39" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B94CEACAC606712AFF42F89614899C76" box="[132,233,1833,1856]" italics="true" pageId="39" pageNumber="444">Agnotecous</emphasis>
<bibRefCitation id="EFA94B29C606712AFF36F89615D89C76" author="SAUSSURE H. &amp; DE" box="[240,440,1832,1857]" pageId="39" pageNumber="444" pagination="369 - 702" refId="ref36461" refString="SAUSSURE H. DE 1878. - Melanges orthopterologiques. VIe fascicule. Gryllides (2 e partie). Memoires de la Societe de Physique et d'Histoire naturelle de Geneve 25: 369 - 702." type="journal article" year="1878">Saussure, 1878: 546</bibRefCitation>
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.
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</paragraph>
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<typeStatus id="5483887AC606712AFF42F8EC14D59C5E" box="[132,181,1875,1898]" pageId="39" pageNumber="444">TYPE</typeStatus>
SPECIES. —
<taxonomicName id="4C384D5BC606712AFEF4F8ED16F59C5C" authority="Saussure, 1878" authorityName="Saussure" authorityYear="1878" box="[306,661,1874,1898]" class="Insecta" family="Haglotettigoniidae" genus="Agnotecous" kingdom="Animalia" order="Orthoptera" pageId="39" pageNumber="444" phylum="Arthropoda" rank="species" species="tapinopus">
<emphasis id="B94CEACAC606712AFEF4F8ED159A9C5C" box="[306,506,1874,1898]" italics="true" pageId="39" pageNumber="444">Agnotecous tapinopus</emphasis>
<bibRefCitation id="EFA94B29C606712AFDC4F8ED16F59C5C" author="SAUSSURE H. &amp; DE" box="[514,661,1874,1898]" pageId="39" pageNumber="444" pagination="369 - 702" refId="ref36461" refString="SAUSSURE H. DE 1878. - Melanges orthopterologiques. VIe fascicule. Gryllides (2 e partie). Memoires de la Societe de Physique et d'Histoire naturelle de Geneve 25: 369 - 702." type="journal article" year="1878">Saussure, 1878</bibRefCitation>
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by original monotypy.
</paragraph>
<paragraph id="8B8736D8C606712AFF42F822152C9CDF" blockId="39.[132,775,1948,2025]" pageId="39" pageNumber="444">
ILLUSTRATIONS. — Habitus:
<figureCitation id="13032A5DC606712AFE63F82316639C82" box="[421,515,1948,1972]" captionStart="FIG" captionStartId="38.[133,144,1939,1957]" captionTargetBox="[132,1446,215,1906]" captionTargetId="figure-26@38.[694,1259,776,1342]" captionTargetPageId="38" captionText="FIG. 15. — New Caledonian Euscyrtinae Gorochov,1985 (A), Oecanthinae Blanchard,1845 (B) and Eneopterinae Saussure,1874 (C-F) (Gryllidae):A, Proturana sp., from Sarramea, by day (Photo PG); B, Oecanthus rufescens Serville, 1839, from Col des Roussettes, by night (Photo JA); C, Cardiodactylus novaeguineae (de Haan, 1842), from Lifou (Loyalty Islands), by night (Photo PG); D, Agnotecous sp., from Col de Mouirange, by day (Photo LDG); E, Lebinthus lifouensis Desutter- Grandcolas, 1997, from Lifou, by day (Photo PG); F, Pixibinthus sonicus Anso &amp; Robillard, 2016, from Col de la Madeleine, by day (Photo JA). Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4578317" httpUri="https://zenodo.org/record/4578317/files/figure.png" pageId="39" pageNumber="444">Fig. 15D</figureCitation>
. Male genitalia: Desutter-
<bibRefCitation id="EFA94B29C606712AFF42F80815CF9CF9" author="GRANDCOLAS L. &amp; ROBILLARD T." box="[132,431,1975,1999]" pageId="39" pageNumber="444" pagination="65 - 92" refId="ref34790" refString="DESUTTER- GRANDCOLAS L. &amp; ROBILLARD T. 2006. - Phylogenetic systematics and evolution of Agnotecous in New Caledonia (Orthoptera: Grylloidea, Eneopteridae). Systematic Entomology 31: 65 - 92. https: // doi. org / 10.1111 / j. 1365 - 3113.2005.00299. x" type="journal article" year="2006">Grandcolas &amp; Robillard 2006</bibRefCitation>
: figs 4-6. Calling songs:
<bibRefCitation id="EFA94B29C606712AFD6DF808148C9CDF" author="ROBILLARD T. &amp; NATTIER R. &amp; GRANDCOLAS L." pageId="39" pageNumber="444" pagination="17 - 35" refId="ref36362" refString="ROBILLARD T., NATTIER R. &amp; DESUTTER- GRANDCOLAS L. 2010. - New species of the New Caledonian endemic genus Agnotecous (Orthoptera, Grylloidea, Eneopterinae, Lebinthini). Zootaxa 2559: 17 - 35." type="journal article" year="2010">
Robillard
<emphasis id="B94CEACAC606712AFF42F86D14D39CDF" box="[132,179,2001,2025]" italics="true" pageId="39" pageNumber="444">et al.</emphasis>
2010
</bibRefCitation>
: figs 3, 7.
</paragraph>
</subSubSection>
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<paragraph id="8B8736D8C606712AFCEBFF67109599A0" blockId="39.[811,1457,215,1244]" pageId="39" pageNumber="444">
DIAGNOSIS. — After
<bibRefCitation id="EFA94B29C606712AFC36FF6810F29BD9" author="OTTE D. &amp; ALEXANDER R. D. &amp; CADE W." box="[1008,1170,215,239]" pageId="39" pageNumber="444" pagination="375 - 457" refId="ref35864" refString="OTTE D., ALEXANDER R. D. &amp; CADE W. 1987. - The Crickets of New Caledonia (Gryllidae). Proceedings of the Academy of Natural Sciences of Philadelphia 139: 375 - 457." type="journal article" year="1987">
Otte
<emphasis id="B94CEACAC606712AFBE7FF67102D9BD9" box="[1057,1101,215,239]" italics="true" pageId="39" pageNumber="444">et al.</emphasis>
(1987)
</bibRefCitation>
, Desutter-Grandcolas (1997c) and Desutter-Grandcolas &amp; Robillard (2006). Medium to large species with brown coloration, without HWs and with short HWs in both males and females. Head flat dorsally. Fastigium very wide, 1.5 to 2 × as large as scape. Ocelli set as a wide and flat triangle; median ocellus vertical and subapical on fastigium. Eyes not prominent. Scapes small, slightly longer than wide. Joint 5 of maxillary palpi short, regularly widened distad, truncate obliquely at apex. Pronotum distinctively long and wide; DD somewhat wider than long, narrowed anteriorly. TI with an oval outer tympanum and a slit-like inner tympanum; with three apical spurs.TII with four apical spurs. TIII with four subapical spurs on each side; inner subapical spurs longer and strongly curved; with three inner and three outer apical spurs, median spur the longest on each side; median outer spur twice as long as other outer spurs.TIII serrulated over its whole length. Basitarsomeres III with two rows of dorsal spines; inner row with one or few spines. Cerci moderately long.
</paragraph>
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<emphasis id="B94CEACAC606712AFCEBFD2617079987" bold="true" box="[813,871,665,689]" pageId="39" pageNumber="444">Male.</emphasis>
Metanotum without glandular structures. FWs short, not reaching abdomen mid length, truncate dorsally; lateral field very high, longer than dorsal field, and partly covering body dorsum. Venation: dorsal field with complete stridulum, with one or no harp vein, and a reduced mirror, lost in apical reticulation; lateral field with numerous parallel veins. Subgenital plate long and rounded; apex narrow and acute; lateral margin inflated, probably glandular. Male genitalia: Pseudepiphallic sclerite very elongate, tubular; pseudepiphallic apex most often elongate as a prominent median process, and with two distal lobes; rami short, extending pseudepiphallic sclerite; pseudepiphallic parameres small and little differentiate; ectophallic fold simple and very short; ectophallic apodemes long, thin and almost parallel; dorsal cavity wide, low and distinctly plicated transversally; endophallic sclerite located at level of ectophallic arc; endophallic apodeme crest-like on dorsal cavity.
</paragraph>
<paragraph id="8B8736D8C606712AFCEBFBFF107F9FEA" blockId="39.[811,1457,215,1244]" pageId="39" pageNumber="444">
<emphasis id="B94CEACAC606712AFCEBFBFF17199F6E" bold="true" box="[813,889,1088,1112]" pageId="39" pageNumber="444">Female.</emphasis>
FWs short, not reaching tergite II distal margin; most often not overlapping; dorsal and lateral fields with strong, longitudinal parallel veins, and variable transverse veinlets. Subgenital plate small, transverse. Ovipositor straight. Female genitalia with a very small copulatory papilla; apex sclerotized, distinctly separate from half-ringed basal sclerite.
</paragraph>
<paragraph id="8B8736D8C606712AFCEBFB4117CE9E7C" blockId="39.[813,1456,1277,1354]" pageId="39" pageNumber="444">
CALLING SONGS. — Long echemes emitted with a dominant frequencies of 15 to 18 kHz, carried by the second harmonic (
<bibRefCitation id="EFA94B29C606712AFABBFAA717C09E7C" author="ANSO J. &amp; BARRABE L. &amp; GRANDCOLAS L. &amp; JOURDAN H. &amp; GRANDCOLAS P. &amp; DONG J. &amp; ROBILLARD T." pageId="39" pageNumber="444" pagination="0150920" refId="ref33517" refString="ANSO J., BARRABE L., DESUTTER- GRANDCOLAS L., JOURDAN H., GRANDCOLAS P., DONG J. &amp; ROBILLARD T. 2016 b. - Old lineage on an old island: Pixibinthus, a new cricket genus from New Caledonia shed light on Gryllid diversification in a hotspot of biodiversity. Plos One 11 (3): e 0150920. https: // doi. org / 10.1371 / journal. pone. 0150920" type="journal article" year="2016">
Anso
<emphasis id="B94CEACAC606712AFCEBFA8C173C9E7C" box="[813,860,1330,1354]" italics="true" pageId="39" pageNumber="444">et al.</emphasis>
2016b
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).
</paragraph>
</subSubSection>
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<paragraph id="8B8736D8C606712AFCEBFAD411419EAB" blockId="39.[811,1456,1387,1437]" pageId="39" pageNumber="444">
DISTRIBUTION. — Endemic to
<collectingCountry id="F32F7648C606712AFB8EFAD410BB9EB5" box="[1096,1243,1387,1411]" name="New Caledonia" pageId="39" pageNumber="444">New Caledonia</collectingCountry>
. Mostly forest-dwelling species known throughout the whole Grande Terre.
</paragraph>
<paragraph id="8B8736D8C606712AFCEBFA0117889D3D" blockId="39.[813,1457,1470,1547]" pageId="39" pageNumber="444">HABITAT. — Forest-dwelling species which usually hide in leaf litter, and are active at night either in the leaf litter or perched on understorey plants.</paragraph>
</subSubSection>
</treatment>
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