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<document id="6D51DA868F15E99F9EAD7E351D7502CB" ID-DOI="10.5281/zenodo.178859" ID-GBIF-Dataset="82336eb7-1f40-4679-ba3b-e2143e2c18b1" ID-ISSN="1175-5326" ID-Zenodo-Dep="178859" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459926551010" checkinUser="plazi" docAuthor="Liao, Yulin, Pawson, David L. &amp; Liu, Wei" docDate="2007" docId="0391879FFFF99A3D21A4FB0A0342FD5A" docLanguage="en" docName="zt01608p034.pdf" docOrigin="Zootaxa 1608" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Phyllophorus (Phyllophorus) maculatus Liao, Pawson &amp; Liu, 2007, new species" docType="treatment" docVersion="8" lastPageNumber="34" masterDocId="FFA8FFE7FFF89A3E2133FFD80733FFC8" masterDocTitle="Phyllophorus (Phyllophorus) maculatus, a new species of sea cucumber from the Yellow Sea (Echinodermata: Holothuroidea: Dendrochirotida)" masterLastPageNumber="34" masterPageNumber="31" pageNumber="32" updateTime="1698229592983" updateUser="plazi">
<mods:mods id="EB7F149C3D16A1C39048FEA56E572829" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F63488B1587AB58E6A108EF135EC6995">
<mods:title id="7EBAABFBC357CA530D649351637617B5">Phyllophorus (Phyllophorus) maculatus, a new species of sea cucumber from the Yellow Sea (Echinodermata: Holothuroidea: Dendrochirotida)</mods:title>
</mods:titleInfo>
<mods:name id="4AD52BE2D62B80ADF7B8CECC123EF840" type="personal">
<mods:role id="1CD754ABD9CEC198AF7A344FC9268926">
<mods:roleTerm id="9FA04093614FFFD53ED8B4B9DB20CAE6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="43AD5F80076938F061154917BCBB7197">Liao, Yulin</mods:namePart>
</mods:name>
<mods:name id="039317841E78CB9C24666C7EA2630A63" type="personal">
<mods:role id="B77436D415158B34479C7533E21BD97D">
<mods:roleTerm id="3858B004ACC7D28E880BC11962B1F2CA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="156FA18EA376BA82696C758BD9C155CB">Pawson, David L.</mods:namePart>
</mods:name>
<mods:name id="06B2FA041B7C02C3BEB2A728B90CA37E" type="personal">
<mods:role id="99612275E195ADE960111A2A72EC5B40">
<mods:roleTerm id="CC46F8E3AAFFCBDD07A970532CF669F1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="72DD0CC689846FF34393858C0F053CD6">Liu, Wei</mods:namePart>
</mods:name>
<mods:typeOfResource id="3FAA800B1E595A69C952F0BD1C517806">text</mods:typeOfResource>
<mods:relatedItem id="4766270887EE6F1C3BCE1A75EB3BE657" type="host">
<mods:titleInfo id="A7D58247A227ADA4A26C1F769FF83717">
<mods:title id="BE63F52EF3A4012964E8C13C454FA413">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="03DC2CE48DFAE8A49CF3FAF6FE01A8A0">
<mods:date id="33E602F43A19AA60F3D56DB1909B88BF">2007</mods:date>
<mods:detail id="E004479385317FE50F86BA947EC08589" type="volume">
<mods:number id="EF4EC786A5D5E9ACC7F20E7030414273">1608</mods:number>
</mods:detail>
<mods:extent id="D52C920FBC40243CFA759608B9720D1F" unit="page">
<mods:start id="7AB4FB64A3D88A9BD6B14EC96B9E6FCF">31</mods:start>
<mods:end id="EF84188867758C28ED52ABEDADA1B4A4">34</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F83079966CF5BE491E5ED42491E1C642">journal article</mods:classification>
<mods:identifier id="799EB7D298277C8FC222D04D3E43288B" type="DOI">10.5281/zenodo.178859</mods:identifier>
<mods:identifier id="91F20515F81D7C290CEFDFE881E09A78" type="GBIF-Dataset">82336eb7-1f40-4679-ba3b-e2143e2c18b1</mods:identifier>
<mods:identifier id="E8D14A570D60B1D8BB184C27E37E5058" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="62308B845AA5EBB387F67EEB0B209BE9" type="Zenodo-Dep">178859</mods:identifier>
</mods:mods>
<treatment id="0391879FFFF99A3D21A4FB0A0342FD5A" ID-DOI="http://doi.org/10.5281/zenodo.6244158" ID-GBIF-Taxon="119374357" ID-Zenodo-Dep="6244158" LSID="urn:lsid:plazi:treatment:0391879FFFF99A3D21A4FB0A0342FD5A" httpUri="http://treatment.plazi.org/id/0391879FFFF99A3D21A4FB0A0342FD5A" lastPageId="3" lastPageNumber="34" pageId="1" pageNumber="32">
<subSubSection id="C3226502FFF99A3F21A4FB0A0612FAC7" pageId="1" pageNumber="32" type="nomenclature">
<paragraph id="8B873689FFF99A3F21A4FB0A0428FB24" blockId="1.[151,795,1234,1295]" box="[151,795,1234,1260]" pageId="1" pageNumber="32">
<heading id="D0CF81E5FFF99A3F21A4FB0A0428FB24" bold="true" box="[151,795,1234,1260]" fontSize="11" level="1" pageId="1" pageNumber="32" reason="1">
<emphasis id="B94CEA9BFFF99A3F21A4FB0A0428FB24" bold="true" box="[151,795,1234,1260]" pageId="1" pageNumber="32">
<taxonomicName id="4C384D0AFFF99A3F21A4FB0A054FFB24" ID-CoL="6VG75" box="[151,636,1234,1260]" class="Holothuroidea" family="Phyllophoridae" genus="Phyllophorus" kingdom="Animalia" order="Dendrochirotida" pageId="1" pageNumber="32" phylum="Echinodermata" rank="species" species="maculatus" status="sp. nov." subGenus="Phyllophorus">
<emphasis id="B94CEA9BFFF99A3F21A4FB0A054FFB24" bold="true" box="[151,636,1234,1260]" italics="true" pageId="1" pageNumber="32">Phyllophorus (Phyllophorus) maculatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27F57E0FFF99A3F23B8FB0A0428FB24" box="[651,795,1234,1260]" pageId="1" pageNumber="32" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B873689FFF99A3F21A4FB2D0612FAC7" blockId="1.[151,795,1234,1295]" box="[151,289,1269,1295]" pageId="1" pageNumber="32">
<figureCitation id="13032A0CFFF99A3F21A4FB2D0612FAC7" box="[151,289,1269,1295]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[151,255,1977,2001]" captionStartId-1="2.[151,255,1702,1726]" captionTargetBox-0="[314,1274,1559,1958]" captionTargetBox-1="[313,1271,659,1702]" captionTargetId-0="figure@1.[314,1274,1559,1958]" captionTargetId-1="figure@2.[313,1273,657,1746]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Phyllophorus (Phyllophorus) maculatus new species. Left, lateral view of Holotype, total length 80 mm. Right, calcareous ring of a Paratype, length of ring 20 mm." captionText-1="FIGURE 2. Phyllophorus (Phyllophorus) maculatus new species. A, tables from posterior end of body; B, rods from tentacles; C, endplate from tube foot; D, rosettes from introvert." httpUri-0="https://zenodo.org/record/178860/files/figure.png" httpUri-1="https://zenodo.org/record/178861/files/figure.png" pageId="1" pageNumber="32">Figures 1–2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3226502FFF99A3F21A4FAEA06E3FA24" pageId="1" pageNumber="32" type="diagnosis">
<paragraph id="8B873689FFF99A3F21A4FAEA06E3FA24" blockId="1.[151,1436,1330,1516]" pageId="1" pageNumber="32">
<emphasis id="B94CEA9BFFF99A3F21A4FAEA062AFA84" bold="true" box="[151,281,1330,1356]" pageId="1" pageNumber="32">Diagnosis:</emphasis>
Medium-sized, up to
<quantity id="4CC09B6CFFF99A3F2314FAEA0549FA84" box="[551,634,1330,1356]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="32" unit="mm" value="80.0">80 mm</quantity>
long and
<quantity id="4CC09B6CFFF99A3F23C2FAEA0476FA84" box="[753,837,1330,1356]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="32" unit="mm" value="20.0">20 mm</quantity>
in diameter. Body cylindrical, posterior end more or less tapering. Tube feet small, scattered on body wall, more numerous ventrally. Tentacles
<quantity id="4CC09B6CFFF99A3F25DDFA820219FABC" box="[1262,1322,1370,1396]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="32" unit="in" value="20.0">20 in</quantity>
three circles, 10+5+5. Calcareous ring with medium-length posterior projections on radials, each composed of about 10 small pieces. Ossicles in form of four-pillared tables with low spires, scarce in body wall, present only in posterior region near anus.
</paragraph>
</subSubSection>
<caption id="DF476601FFF99A3F21A4F861043FF839" httpUri="https://zenodo.org/record/178860/files/figure.png" pageId="1" pageNumber="32" targetBox="[314,1274,1559,1958]" targetPageId="1">
<paragraph id="8B873689FFF99A3F21A4F861043FF839" blockId="1.[151,1436,1977,2033]" pageId="1" pageNumber="32">
<emphasis id="B94CEA9BFFF99A3F21A4F8610628F819" bold="true" box="[151,283,1977,2001]" pageId="1" pageNumber="32">FIGURE 1.</emphasis>
<taxonomicName id="4C384D0AFFF99A3F2018F86205E8F819" box="[299,731,1978,2001]" class="Holothuroidea" family="Phyllophoridae" genus="Phyllophorus" kingdom="Animalia" order="Dendrochirotida" pageId="1" pageNumber="32" phylum="Echinodermata" rank="species" species="maculatus" status="sp. nov." subGenus="Phyllophorus">
<emphasis id="B94CEA9BFFF99A3F2018F86205E8F819" box="[299,731,1978,2001]" italics="true" pageId="1" pageNumber="32">Phyllophorus (Phyllophorus) maculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57E0FFF99A3F23D1F8620454F819" box="[738,871,1978,2001]" pageId="1" pageNumber="32" rank="species">new species</taxonomicNameLabel>
. Left, lateral view of Holotype, total length 80mm. Right, calcareous ring of a Paratype, length of ring 20 mm.
</paragraph>
</caption>
<subSubSection id="C3226502FFFA9A3C21F2FF4F038BFE02" pageId="2" pageNumber="33" type="materials_examined">
<paragraph id="8B873689FFFA9A3C21F2FF4F038BFE02" blockId="2.[151,1437,151,658]" pageId="2" pageNumber="33">
<emphasis id="B94CEA9BFFFA9A3C21F2FF4F068EFF79" bold="true" box="[193,445,151,177]" pageId="2" pageNumber="33">Material Examined:</emphasis>
<collectionCode id="ED29AE4CFFFA9A3C20FFFF400559FF7A" box="[460,618,152,178]" pageId="2" pageNumber="33">
<typeStatus id="5483882BFFFA9A3C20FFFF400559FF7A" box="[460,618,152,178]" pageId="2" pageNumber="33" type="holotype">HOLOTYPE</typeStatus>
</collectionCode>
,
<collectionCode id="ED29AE4CFFFA9A3C2346FF4005FEFF7A" box="[629,717,152,178]" httpUri="http://grbio.org/cool/dg3j-1xzp" name="Institute of Oceanology, Chinese Academy of Scineces" pageId="2" pageNumber="33">IOCAS</collectionCode>
<collectionCode id="ED29AE4CFFFA9A3C23E7FF4005C3FF7A" box="[724,752,152,178]" pageId="2" pageNumber="33">E-</collectionCode>
1110, Yellow Sea, 36ºN, 120ºE,
<date id="FF861049FFFA9A3C255BFF40027CFF7A" box="[1128,1359,152,178]" pageId="2" pageNumber="33" value="2002-09-16">
16 September
<quantity id="4CC09B6CFFFA9A3C2422FF40027CFF7A" box="[1297,1359,152,178]" metricMagnitude="3" metricUnit="m" metricValue="2.00232" pageId="2" pageNumber="33" unit="m" value="2002.32">2002</quantity>
</date>
, 32 m, muddy sand bottom.
<collectionCode id="ED29AE4CFFFA9A3C20A4FF18050AFF12" box="[407,569,192,218]" pageId="2" pageNumber="33">
<typeStatus id="5483882BFFFA9A3C20A4FF18050AFF12" box="[407,569,192,218]" pageId="2" pageNumber="33" type="paratype">PARATYPES</typeStatus>
</collectionCode>
<collectionCode id="ED29AE4CFFFA9A3C237BFF180593FF12" box="[584,672,192,218]" httpUri="http://grbio.org/cool/dg3j-1xzp" name="Institute of Oceanology, Chinese Academy of Scineces" pageId="2" pageNumber="33">IOCAS</collectionCode>
<collectionCode id="ED29AE4CFFFA9A3C239BFF1805F7FF12" box="[680,708,192,218]" pageId="2" pageNumber="33">E-</collectionCode>
1111, Yellow Sea, 36ºN,
<geoCoordinate id="EE0C504EFFFA9A3C22C2FF180352FF12" box="[1009,1121,192,218]" direction="east" orientation="longitude" pageId="2" pageNumber="33" precision="925" value="120.5">120º30’E</geoCoordinate>
,
<date id="FF861049FFFA9A3C255DFF180205FF12" box="[1134,1334,192,218]" pageId="2" pageNumber="33" value="2000-10-19">
19 October
<quantity id="4CC09B6CFFFA9A3C25CBFF180205FF12" box="[1272,1334,192,218]" metricMagnitude="3" metricUnit="m" metricValue="2.0003" pageId="2" pageNumber="33" unit="m" value="2000.3">2000</quantity>
</date>
, 30 m,
<specimenCount id="9D3EFD00FFFA9A3C24BEFF180626FECA" pageId="2" pageNumber="33" type="generic">2 specimens</specimenCount>
;
<collectionCode id="ED29AE4CFFFA9A3C2010FF300648FECA" box="[291,379,232,258]" httpUri="http://grbio.org/cool/dg3j-1xzp" name="Institute of Oceanology, Chinese Academy of Scineces" pageId="2" pageNumber="33">IOCAS</collectionCode>
<collectionCode id="ED29AE4CFFFA9A3C20B6FF300692FECA" box="[389,417,232,258]" pageId="2" pageNumber="33">E-</collectionCode>
1112, Yellow Sea,
<geoCoordinate id="EE0C504EFFFA9A3C23B7FF3005C0FECA" box="[644,755,232,258]" direction="north" orientation="latitude" pageId="2" pageNumber="33" precision="925" value="35.5">35º 30’N</geoCoordinate>
,
<geoCoordinate id="EE0C504EFFFA9A3C2231FF300441FECA" box="[770,882,232,258]" direction="east" orientation="longitude" pageId="2" pageNumber="33" precision="925" value="121.5">121º30’E</geoCoordinate>
,
<date id="FF861049FFFA9A3C22B1FF300315FECA" box="[898,1062,232,258]" pageId="2" pageNumber="33" value="2001-05-19">
19 May
<quantity id="4CC09B6CFFFA9A3C22DAFF300315FECA" box="[1001,1062,232,258]" metricMagnitude="3" metricUnit="m" metricValue="2.00132" pageId="2" pageNumber="33" unit="m" value="2001.32">2001</quantity>
</date>
, 32 m,
<specimenCount id="9D3EFD00FFFA9A3C25B6FF300223FECA" box="[1157,1296,232,258]" pageId="2" pageNumber="33" type="generic">1 specimen</specimenCount>
;
<collectionCode id="ED29AE4CFFFA9A3C242DFF300245FECA" box="[1310,1398,232,258]" httpUri="http://grbio.org/cool/dg3j-1xzp" name="Institute of Oceanology, Chinese Academy of Scineces" pageId="2" pageNumber="33">IOCAS</collectionCode>
<collectionCode id="ED29AE4CFFFA9A3C24B3FF3002AFFECA" box="[1408,1436,232,258]" pageId="2" pageNumber="33">E-</collectionCode>
1113, Yellow Sea,
<geoCoordinate id="EE0C504EFFFA9A3C2046FEC806E9FEE2" box="[373,474,272,298]" direction="north" orientation="latitude" pageId="2" pageNumber="33" precision="925" value="35.5">35º30’N</geoCoordinate>
,
<geoCoordinate id="EE0C504EFFFA9A3C20D4FEC8056BFEE2" box="[487,600,272,298]" direction="east" orientation="longitude" pageId="2" pageNumber="33" precision="925" value="121.5">121º30’E</geoCoordinate>
,
<date id="FF861049FFFA9A3C2356FEC80435FEE2" box="[613,774,272,298]" pageId="2" pageNumber="33" value="2003-06-15">
15 June
<quantity id="4CC09B6CFFFA9A3C23FAFEC80435FEE2" box="[713,774,272,298]" metricMagnitude="3" metricUnit="m" metricValue="2.00333" pageId="2" pageNumber="33" unit="m" value="2003.33">2003</quantity>
</date>
, 33 m,
<specimenCount id="9D3EFD00FFFA9A3C2253FEC804DAFEE2" box="[864,1001,272,298]" pageId="2" pageNumber="33" type="generic">1 specimen</specimenCount>
;
<collectionCode id="ED29AE4CFFFA9A3C22CCFEC80364FEE2" box="[1023,1111,272,298]" httpUri="http://grbio.org/cool/dg3j-1xzp" name="Institute of Oceanology, Chinese Academy of Scineces" pageId="2" pageNumber="33">IOCAS</collectionCode>
<collectionCode id="ED29AE4CFFFA9A3C256CFEC80348FEE2" box="[1119,1147,272,298]" pageId="2" pageNumber="33">E-</collectionCode>
1114, Yellow Sea, 36ºN, 121ºE,
<date id="FF861049FFFA9A3C21DEFEE006BEFE9A" box="[237,397,312,338]" pageId="2" pageNumber="33" value="2004-06-15">
15 June
<quantity id="4CC09B6CFFFA9A3C207CFEE006BEFE9A" box="[335,397,312,338]" metricMagnitude="3" metricUnit="m" metricValue="2.00432" pageId="2" pageNumber="33" unit="m" value="2004.32">2004</quantity>
</date>
, 32 m,
<specimenCount id="9D3EFD00FFFA9A3C20D0FEE0055FFE9A" box="[483,620,312,338]" pageId="2" pageNumber="33" type="generic">1 specimen</specimenCount>
;
<collectionCode id="ED29AE4CFFFA9A3C23B3FEE005EBFE9A" box="[640,728,312,338]" httpUri="http://grbio.org/cool/dg3j-1xzp" name="Institute of Oceanology, Chinese Academy of Scineces" pageId="2" pageNumber="33">IOCAS</collectionCode>
<collectionCode id="ED29AE4CFFFA9A3C23ECFEE005C8FE9A" box="[735,763,312,338]" pageId="2" pageNumber="33">E-</collectionCode>
1115, Yellow Sea,
<geoCoordinate id="EE0C504EFFFA9A3C22E5FEE00370FE9A" box="[982,1091,312,338]" direction="north" orientation="latitude" pageId="2" pageNumber="33" precision="925" value="35.5">35º 30’N</geoCoordinate>
, 121º E,
<date id="FF861049FFFA9A3C259EFEE0027FFE9A" box="[1197,1356,312,338]" pageId="2" pageNumber="33" value="2003-06-15">
15 June
<quantity id="4CC09B6CFFFA9A3C243CFEE0027FFE9A" box="[1295,1356,312,338]" metricMagnitude="3" metricUnit="m" metricValue="2.0033499999999997" pageId="2" pageNumber="33" unit="m" value="2003.35">2003</quantity>
</date>
, 35 m,
<specimenCount id="9D3EFD00FFFA9A3C21A4FEB8062CFEB2" box="[151,287,352,378]" pageId="2" pageNumber="33" type="generic">1 specimen</specimenCount>
;
<collectionCode id="ED29AE4CFFFA9A3C2019FEB806B1FEB2" box="[298,386,352,378]" httpUri="http://grbio.org/cool/dg3j-1xzp" name="Institute of Oceanology, Chinese Academy of Scineces" pageId="2" pageNumber="33">IOCAS</collectionCode>
<collectionCode id="ED29AE4CFFFA9A3C20B9FEB80695FEB2" box="[394,422,352,378]" pageId="2" pageNumber="33">E-</collectionCode>
1116, Yellow Sea,
<geoCoordinate id="EE0C504EFFFA9A3C234CFEB805DFFEB2" box="[639,748,352,378]" direction="north" orientation="latitude" pageId="2" pageNumber="33" precision="925" value="35.5">35º 30’N</geoCoordinate>
,
<geoCoordinate id="EE0C504EFFFA9A3C23CBFEB8045CFEB2" box="[760,879,352,378]" direction="east" orientation="longitude" pageId="2" pageNumber="33" precision="925" value="120.5">120º 30’E</geoCoordinate>
,
<date id="FF861049FFFA9A3C224FFEB8030BFEB2" box="[892,1080,352,378]" pageId="2" pageNumber="33" value="2001-08-11">
11 August
<quantity id="4CC09B6CFFFA9A3C22C8FEB8030BFEB2" box="[1019,1080,352,378]" metricMagnitude="3" metricUnit="m" metricValue="2.0012999999999996" pageId="2" pageNumber="33" unit="m" value="2001.3">2001</quantity>
</date>
, 30 m,
<specimenCount id="9D3EFD00FFFA9A3C25BDFEB80226FEB2" box="[1166,1301,352,378]" pageId="2" pageNumber="33" type="generic">1 specimen</specimenCount>
;
<collectionCode id="ED29AE4CFFFA9A3C2412FEB8024AFEB2" box="[1313,1401,352,378]" httpUri="http://grbio.org/cool/dg3j-1xzp" name="Institute of Oceanology, Chinese Academy of Scineces" pageId="2" pageNumber="33">IOCAS</collectionCode>
<collectionCode id="ED29AE4CFFFA9A3C24B2FEB802AEFEB2" box="[1409,1437,352,378]" pageId="2" pageNumber="33">E-</collectionCode>
1117, Yellow Sea, 36ºN, 121ºE,
<date id="FF861049FFFA9A3C2315FE5005FEFE6A" box="[550,717,392,418]" pageId="2" pageNumber="33" value="2003-06-15">
15 June
<quantity id="4CC09B6CFFFA9A3C23A3FE5005FEFE6A" box="[656,717,392,418]" metricMagnitude="3" metricUnit="m" metricValue="2.00322" pageId="2" pageNumber="33" unit="m" value="2003.22">2003</quantity>
</date>
, 22 m,
<specimenCount id="9D3EFD00FFFA9A3C221CFE50048FFE6A" box="[815,956,392,418]" pageId="2" pageNumber="33" type="generic">1 specimen</specimenCount>
;
<collectionCode id="ED29AE4CFFFA9A3C22FFFE500317FE6A" box="[972,1060,392,418]" httpUri="http://grbio.org/cool/dg3j-1xzp" name="Institute of Oceanology, Chinese Academy of Scineces" pageId="2" pageNumber="33">IOCAS</collectionCode>
<collectionCode id="ED29AE4CFFFA9A3C251CFE500378FE6A" box="[1071,1099,392,418]" pageId="2" pageNumber="33">E-</collectionCode>
1118, Yellow Sea,
<geoCoordinate id="EE0C504EFFFA9A3C2401FE5002A4FE6A" box="[1330,1431,392,418]" direction="north" orientation="latitude" pageId="2" pageNumber="33" precision="925" value="35.5">35º30’N</geoCoordinate>
,
<geoCoordinate id="EE0C504EFFFA9A3C21A4FE680634FE02" box="[151,263,432,458]" direction="east" orientation="longitude" pageId="2" pageNumber="33" precision="925" value="121.5">121º30’E</geoCoordinate>
,
<date id="FF861049FFFA9A3C2020FE680681FE02" box="[275,434,432,458]" pageId="2" pageNumber="33" value="2001-05-19">
19 May
<quantity id="4CC09B6CFFFA9A3C2046FE680681FE02" box="[373,434,432,458]" metricMagnitude="3" metricUnit="m" metricValue="2.00145" pageId="2" pageNumber="33" unit="m" value="2001.45">2001</quantity>
</date>
, 45 m,
<specimenCount id="9D3EFD00FFFA9A3C233BFE6805A1FE02" box="[520,658,432,458]" pageId="2" pageNumber="33" type="generic">1 specimen</specimenCount>
. All specimens collected with a Petersen grab.
</paragraph>
</subSubSection>
<subSubSection id="C3226502FFFA9A3D21F5FE0F034AFE02" lastPageId="3" lastPageNumber="34" pageId="2" pageNumber="33" type="description">
<paragraph id="8B873689FFFA9A3C21F5FE0F0401FD5A" blockId="2.[151,1437,151,658]" pageId="2" pageNumber="33">
<emphasis id="B94CEA9BFFFA9A3C21F5FE0F0653FE39" bold="true" box="[198,352,471,497]" pageId="2" pageNumber="33">Description:</emphasis>
<typeStatus id="5483882BFFFA9A3C2043FE0006EEFE3A" box="[368,477,472,498]" pageId="2" pageNumber="33" type="holotype">Holotype</typeStatus>
(
<figureCitation id="13032A0CFFFA9A3C20DEFE000563FE3A" box="[493,592,472,498]" captionStart="FIGURE 1" captionStartId="1.[151,255,1977,2001]" captionTargetBox="[314,1274,1559,1958]" captionTargetId="figure@1.[314,1274,1559,1958]" captionTargetPageId="1" captionText="FIGURE 1. Phyllophorus (Phyllophorus) maculatus new species. Left, lateral view of Holotype, total length 80 mm. Right, calcareous ring of a Paratype, length of ring 20 mm." httpUri="https://zenodo.org/record/178860/files/figure.png" pageId="2" pageNumber="33">Figure 1</figureCitation>
)
<quantity id="4CC09B6CFFFA9A3C2350FE000585FE3A" box="[611,694,472,498]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="33" unit="mm" value="80.0">80 mm</quantity>
long, diameter at mid-body
<quantity id="4CC09B6CFFFA9A3C2535FE00036EFE3A" box="[1030,1117,472,498]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="33" unit="mm" value="20.0">20 mm</quantity>
. Body cylindrical, slightly curved, with anterior end rounded, posterior end more or less tapering. Tentacles retracted; anus terminal, surrounded by five minute papillae. Tentacles 20, apparently in three circles of 10+5+5. Tube feet small, numerous, scattered all over body, more numerous ventrally than dorsally. Body wall thick, soft. Color in alcohol whitish or cream, with numerous black bands or patches.
</paragraph>
<caption id="DF476601FFFA9A3C21A4F97E0472F915" httpUri="https://zenodo.org/record/178861/files/figure.png" pageId="2" pageNumber="33" targetBox="[313,1271,659,1702]" targetPageId="2">
<paragraph id="8B873689FFFA9A3C21A4F97E0472F915" blockId="2.[151,1436,1702,1757]" pageId="2" pageNumber="33">
<emphasis id="B94CEA9BFFFA9A3C21A4F97E0629F976" bold="true" box="[151,282,1702,1726]" pageId="2" pageNumber="33">FIGURE 2.</emphasis>
<taxonomicName id="4C384D0AFFFA9A3C2019F97F05EAF976" box="[298,729,1703,1726]" class="Holothuroidea" family="Phyllophoridae" genus="Phyllophorus" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="33" phylum="Echinodermata" rank="species" species="maculatus" status="sp. nov." subGenus="Phyllophorus">
<emphasis id="B94CEA9BFFFA9A3C2019F97F05EAF976" box="[298,729,1703,1726]" italics="true" pageId="2" pageNumber="33">Phyllophorus (Phyllophorus) maculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27F57E0FFFA9A3C23D3F97E0457F975" box="[736,868,1702,1725]" pageId="2" pageNumber="33" rank="species">new species</taxonomicNameLabel>
. A, tables from posterior end of body; B, rods from tentacles; C, endplate from tube foot; D, rosettes from introvert.
</paragraph>
</caption>
<paragraph id="8B873689FFFA9A3C21FBF8D70273F839" blockId="2.[151,1436,1807,2033]" pageId="2" pageNumber="33">
Ossicles in body wall scarce or absent. No ossicles in anterior and middle part of body, except for endplates in tube feet (
<figureCitation id="13032A0CFFFA9A3C2045F8EF06EBF899" box="[374,472,1847,1873]" captionStart="FIGURE 1" captionStartId="1.[151,255,1977,2001]" captionTargetBox="[314,1274,1559,1958]" captionTargetId="figure@1.[314,1274,1559,1958]" captionTargetPageId="1" captionText="FIGURE 1. Phyllophorus (Phyllophorus) maculatus new species. Left, lateral view of Holotype, total length 80 mm. Right, calcareous ring of a Paratype, length of ring 20 mm." httpUri="https://zenodo.org/record/178860/files/figure.png" pageId="2" pageNumber="33">Figure 1</figureCitation>
C). Ossicles present only in vicinity of anus, as tables (
<figureCitation id="13032A0CFFFA9A3C2554F8EF03FFF899" box="[1127,1228,1847,1873]" captionStart="FIGURE 2" captionStartId="2.[151,255,1702,1726]" captionTargetBox="[313,1271,659,1702]" captionTargetId="figure@2.[313,1273,657,1746]" captionTargetPageId="2" captionText="FIGURE 2. Phyllophorus (Phyllophorus) maculatus new species. A, tables from posterior end of body; B, rods from tentacles; C, endplate from tube foot; D, rosettes from introvert." httpUri="https://zenodo.org/record/178861/files/figure.png" pageId="2" pageNumber="33">Figure 2</figureCitation>
A) with low fourpillared spires. Disk perforated by four large central holes and 8–12 small peripheral holes; edge of disc sinuous, with small peripheral knobs. Spire end in crown of eight spines. Disk diameter 74–179μm (average 120μm), spire height 51–63 μm. In tentacles, delicate supporting rods (
<figureCitation id="13032A0CFFFA9A3C22EAF877030EF801" box="[985,1085,1967,1993]" captionStart="FIGURE 2" captionStartId="2.[151,255,1702,1726]" captionTargetBox="[313,1271,659,1702]" captionTargetId="figure@2.[313,1273,657,1746]" captionTargetPageId="2" captionText="FIGURE 2. Phyllophorus (Phyllophorus) maculatus new species. A, tables from posterior end of body; B, rods from tentacles; C, endplate from tube foot; D, rosettes from introvert." httpUri="https://zenodo.org/record/178861/files/figure.png" pageId="2" pageNumber="33">Figure 2</figureCitation>
B) of length 68–310μm (average 180μm), often with one or two perforations at extremities. In introvert, rosettes only (
<figureCitation id="13032A0CFFFA9A3C2585F80F0228F839" box="[1206,1307,2007,2033]" captionStart="FIGURE 2" captionStartId="2.[151,255,1702,1726]" captionTargetBox="[313,1271,659,1702]" captionTargetId="figure@2.[313,1273,657,1746]" captionTargetPageId="2" captionText="FIGURE 2. Phyllophorus (Phyllophorus) maculatus new species. A, tables from posterior end of body; B, rods from tentacles; C, endplate from tube foot; D, rosettes from introvert." httpUri="https://zenodo.org/record/178861/files/figure.png" pageId="2" pageNumber="33">Figure 2</figureCitation>
D).
</paragraph>
<paragraph id="8B873689FFFB9A3D21FDFF4F034AFE02" blockId="3.[151,1437,151,658]" pageId="3" pageNumber="34">
<emphasis id="B94CEA9BFFFB9A3D21FDFF4F0666FF79" bold="true" box="[206,341,151,177]" pageId="3" pageNumber="34">
<typeStatus id="5483882BFFFB9A3D21FDFF4F0663FF79" box="[206,336,151,177]" pageId="3" pageNumber="34" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Nine
<typeStatus id="5483882BFFFB9A3D2099FF400529FF7A" box="[426,538,152,178]" pageId="3" pageNumber="34" type="paratype">paratypes</typeStatus>
collected near or at same station as
<typeStatus id="5483882BFFFB9A3D22E3FF400308FF7A" box="[976,1083,152,178]" pageId="3" pageNumber="34" type="holotype">holotype</typeStatus>
. Two
<typeStatus id="5483882BFFFB9A3D25BEFF4003CEFF7A" box="[1165,1277,152,178]" pageId="3" pageNumber="34" type="paratype">paratypes</typeStatus>
lack ossicles altogether; collectors noted that these two specimens had been placed in formalin for several days; presumably ossicles dissolved due to acidic conditions. In one
<typeStatus id="5483882BFFFB9A3D2273FF300496FECA" box="[832,933,232,258]" pageId="3" pageNumber="34" type="paratype">paratype</typeStatus>
tube feet arranged in five double rows in radii, and apparently absent from interradii; other features identical with those of the
<typeStatus id="5483882BFFFB9A3D25AAFEC80230FEE2" box="[1177,1283,272,298]" pageId="3" pageNumber="34" type="holotype">holotype</typeStatus>
. Calcareous ring (
<figureCitation id="13032A0CFFFB9A3D21E6FEE00604FE9A" box="[213,311,312,338]" captionStart="FIGURE 1" captionStartId="1.[151,255,1977,2001]" captionTargetBox="[314,1274,1559,1958]" captionTargetId="figure@1.[314,1274,1559,1958]" captionTargetPageId="1" captionText="FIGURE 1. Phyllophorus (Phyllophorus) maculatus new species. Left, lateral view of Holotype, total length 80 mm. Right, calcareous ring of a Paratype, length of ring 20 mm." httpUri="https://zenodo.org/record/178860/files/figure.png" pageId="3" pageNumber="34">Figure 1</figureCitation>
) large, about
<quantity id="4CC09B6CFFFB9A3D20EBFEE00519FE9A" box="[472,554,312,338]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="34" unit="mm" value="20.0">20 mm</quantity>
in length, including posterior projections, ring composed of five compound radial and five single interradial plates. Radials divided anteriorly into two unequal halves by a deep notch; posterior projections each composed of 8–10 small pieces. Interradials lower, more or less triangular, anterior point attenuate, posterior base broad, rounded. Polian vesicle and stone canal single.
</paragraph>
</subSubSection>
<subSubSection id="C3226502FFFB9A3D21F5FE0F06F3FDD2" pageId="3" pageNumber="34" type="etymology">
<paragraph id="8B873689FFFB9A3D21F5FE0F06F3FDD2" blockId="3.[151,1437,151,658]" pageId="3" pageNumber="34">
<emphasis id="B94CEA9BFFFB9A3D21F5FE0F0664FE39" bold="true" box="[198,343,471,497]" pageId="3" pageNumber="34">Etymology:</emphasis>
The species name is derived from Latin
<emphasis id="B94CEA9BFFFB9A3D2271FE0004ABFE39" box="[834,920,472,497]" italics="true" pageId="3" pageNumber="34">macula</emphasis>
, in reference to the blotches of color on the body wall of this species.
</paragraph>
</subSubSection>
<subSubSection id="C3226502FFFB9A3D21F6FDFF0342FD5A" pageId="3" pageNumber="34" type="discussion">
<paragraph id="8B873689FFFB9A3D21F6FDFF0342FD5A" blockId="3.[151,1437,151,658]" pageId="3" pageNumber="34">
<emphasis id="B94CEA9BFFFB9A3D21F6FDFF0672FD89" bold="true" box="[197,321,551,577]" pageId="3" pageNumber="34">Remarks:</emphasis>
It is assumed that this is a burrowing species, like most of its relatives. This new species differs from its consubgeners in possessing body wall tables with short spires, the tables present only in the vicinity of the anus; in the shape of the calcareous ring; and in the color of the body.
</paragraph>
</subSubSection>
</treatment>
</document>
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