523 lines
94 KiB
XML
523 lines
94 KiB
XML
<document id="0E06A9BF29AEBC640BEB4EDAD4414B98" ID-DOI="10.5852/ejt.2014.76" ID-GBIF-Dataset="6f3572d0-0a33-42fa-94d1-88028a541cdb" ID-ISSN="2118-9773" ID-Zenodo-Dep="3835020" ID-ZooBank="01BEFFB2-2E53-4B1E-A7BC-36702313BBF8" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1589939423581" checkinUser="tatiana" docAuthor="Okanishi, Masanori & Fujita, Toshihiko" docDate="2014" docId="03917D3BD907F77AB76B7834FD470935" docLanguage="en" docName="ejt-2014-76.pdf.imf" docOrigin="European Journal of Taxonomy 76 (76)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Asterostegus sabineae Okanishi & Fujita 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="9" masterDocId="FFA80543D904F772B50F7C50FFB80E14" masterDocTitle="A taxonomic review of the genus Asterostegus (Echinodermata: Ophiuroidea), with the description of a new species" masterLastPageNumber="18" masterPageNumber="1" pageNumber="4" updateTime="1737120617184" updateUser="carolina" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FE238E4715737579EE26F4FD47CE83DD">A taxonomic review of the genus Asterostegus (Echinodermata: Ophiuroidea), with the description of a new species</mods:title>
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<mods:namePart id="0A66331CCCFC3804A9D6C128B1250061">Okanishi, Masanori</mods:namePart>
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<mods:namePart id="F652529529708394692A790282EEBDA8">Fujita, Toshihiko</mods:namePart>
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<mods:affiliation id="4CDECF760945A3F0DD0F11F8F28FE118">National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki, 305 - 0005 Japan Corresponding author.</mods:affiliation>
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<treatment id="03917D3BD907F77AB76B7834FD470935" ID-DOI="http://doi.org/10.5281/zenodo.3851753" ID-GBIF-Taxon="164241191" ID-Zenodo-Dep="3851753" LSID="urn:lsid:plazi:treatment:03917D3BD907F77AB76B7834FD470935" httpUri="http://treatment.plazi.org/id/03917D3BD907F77AB76B7834FD470935" lastPageId="8" lastPageNumber="9" pageId="3" pageNumber="4" scope_class="Ophiuroidea" scope_family="Euryalidae" scope_phylum="Echinodermata" scope_superFamily="Euryalidea">
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<taxonomicName id="4C38B7AED907F771B76B7834FCD60A6B" ID-CoL="5W7VR" authority="Okanishi & Fujita, 2014" authorityName="Okanishi & Fujita" authorityYear="2014" box="[612,878,1124,1151]" class="Ophiuroidea" family="Euryalidae" genus="Asterostegus" kingdom="Animalia" order="Euryalida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="species" species="sabineae" status="sp. nov.">
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<emphasis id="B94C103FD907F771B76B7834FCD60A6B" bold="true" box="[612,878,1124,1151]" italics="true" pageId="3" pageNumber="4">Asterostegus sabineae</emphasis>
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<taxonomicNameLabel id="A27FAD44D907F771B6797835FC680A6B" box="[886,976,1125,1151]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="1303D0A8D907F771B7E678FBFCF20AD2" box="[745,842,1195,1222]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="4.[189,232,1787,1813]" captionStartId-1="5.[189,232,1892,1918]" captionStartId-2="6.[189,232,1858,1884]" captionStartId-3="7.[189,232,1858,1884]" captionTargetBox-0="[278,1305,265,1748]" captionTargetBox-1="[188,1397,406,1852]" captionTargetBox-2="[188,1398,509,1814]" captionTargetBox-3="[192,1396,755,1814]" captionTargetId-0="figure@4.[277,1305,265,1748]" captionTargetId-1="figure@5.[188,1399,404,1852]" captionTargetId-2="figure@6.[188,1398,507,1814]" captionTargetId-3="figure@7.[189,1398,752,1814]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="Fig. 2. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H). A. Aboral disc and proximal portion of arms. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc and proximal portion of arms. E. Jaws. F. Oral periphery of disc and proximal portion of arm. G. Lateral disc. H. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle." captionText-1="Fig. 3. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) (A–F). A. Aboral basal portion of arm. B. Aboral middle portion of arm. C. Aboral and lateral distal portion of arm. D. Oral proximal portion of arm. E. Oral middle portion of arm. F. Lateral distal portion of arm. Abbreviations: AS = arm spine, LAP = lateral arm plate, Ten = tentacles, Tu = tubercle." captionText-2="Fig. 4. Asterostegus sabineae sp. nov., paratype (SMNH-Type-8536), SEM photographs. A-D. Vertebrae from middle portion of arm: distal view (A), aboral view (B), oral view (C), lateral view (D). E-H. Vertebrae from distal portion of arm: distal view (E), aboral view (F), oral view (G), lateral view (H). I. Lateral arm plate from middle portion of arm. Abbreviations: K = knobs, MO = muscle opening, OB = oral bridge." captionText-3="Fig. 5. Asterostegus sabineae sp. nov., paratype (SMNH-Type-8536), SEM photographs. A. Lateral arm plate from middle portion of arm, abradial view. B. Lateral arm plate from distal portion of arm, adradial view. C. Lateral arm plate from distal portion of arm, oral view. D-F. Arm spines: from proximal portion of arm (D), middle portion of arm (E), distal portion of arm, an arc indicates lamina (F). Abbreviations: MO = muscle opening, NO = nerve opening, ST = secondary tooth." figureDoi-0="http://doi.org/10.5281/zenodo.3835024" figureDoi-1="http://doi.org/10.5281/zenodo.3835026" figureDoi-2="http://doi.org/10.5281/zenodo.3835028" figureDoi-3="http://doi.org/10.5281/zenodo.3835030" httpUri-0="https://zenodo.org/record/3835024/files/figure.png" httpUri-1="https://zenodo.org/record/3835026/files/figure.png" httpUri-2="https://zenodo.org/record/3835028/files/figure.png" httpUri-3="https://zenodo.org/record/3835030/files/figure.png" pageId="3" pageNumber="4">Figs 2-5</figureCitation>
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<taxonomicName id="4C38B7AED907F771B5B278BDFEEA0B13" ID-CoL="7NRJH" authority="Mortensen, 1933" box="[189,338,1261,1287]" class="Ophiuroidea" family="Euryalidae" genus="Asterostegus" kingdom="Animalia" order="Euryalida" pageId="3" pageNumber="4" phylum="Echinodermata" rank="genus">
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<emphasis id="B94C103FD907F771B5B278BDFEEA0B13" box="[189,338,1261,1287]" italics="true" pageId="3" pageNumber="4">Asterostegus</emphasis>
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sp. –
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<treatmentCitation id="0A99EA3CD907F771B49778BCFC9F0B13" author="Okanishi M. & Fujita T." box="[408,807,1260,1287]" page="568" pageId="3" pageNumber="4" year="2013">
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<bibRefCitation id="EFA9B1DCD907F771B49778BCFD540B13" author="Okanishi M. & Fujita T." box="[408,748,1260,1287]" pageId="3" pageNumber="4" pagination="566 - 580" refId="ref6762" refString="Okanishi M. & Fujita T. 2013. Molecular phylogeny based on increased number of species and genes revealed more robust family-level systematics of the order Euryalida (Echinodermata: Ophiuroidea). Molecular Phylogenetics and Evolution 69: 566 - 580. http: // dx. doi. org / 10.1016 / j. ympev. 2013.07.021" type="journal article" year="2013">Okanishi & Fujita 2013: 568</bibRefCitation>
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, 575
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, fig. 1, table 1.
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<subSubSection id="C3229FA6D907F771B5B2797DFBC00B8D" pageId="3" pageNumber="4" type="diagnosis">
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<emphasis id="B94C103FD907F771B5B2797DFE8E0B53" bold="true" box="[189,310,1325,1351]" pageId="3" pageNumber="4">Diagnosis</emphasis>
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</heading>
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<paragraph id="8B87CC2DD907F771B5B2790CFBC00B8D" blockId="3.[189,1398,1372,1434]" pageId="3" pageNumber="4">
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Tubercles
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<emphasis id="B94C103FD907F771B437790DFEE30B62" box="[312,347,1373,1398]" italics="true" pageId="3" pageNumber="4">ca.</emphasis>
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long cover entire radial shields. On proximal portion of arm, one (rarely two) tubercle on each lateral bar. One oral interradial plate present on each interradius.
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<subSubSection id="C3229FA6D907F771B5B27991FD9A0839" pageId="3" pageNumber="4" type="etymology">
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The specific name is for Dr. Sabine Stöhr of the Swedish Museum of Natural History, eminent Swedish researcher of the
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.
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material
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:
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(
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),
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<specimenCount id="9D3E07A4D907F771B65E7AD3FC340889" box="[849,908,1667,1693]" pageId="3" pageNumber="4" type="generic">three</specimenCount>
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(
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<specimenCode id="DB9E6456D907F771B1117AD3FAB80889" box="[1054,1280,1667,1693]" pageId="3" pageNumber="4">SMNH-Type-8333</specimenCode>
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) and
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(
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<specimenCode id="DB9E6456D907F771B43C7AF6FD9E08D5" box="[307,550,1702,1729]" pageId="3" pageNumber="4">SMNH-Type-85436</specimenCode>
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), all ethanol preserved specimens, off
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|
||
,
|
||
<collectingMunicipality id="6BE35657D907F771B19D7AF6FAA708D4" box="[1170,1311,1702,1728]" pageId="3" pageNumber="4">South Cape</collectingMunicipality>
|
||
,
|
||
<location id="8EE79AF6D907F771B0227AF6FEBE08F7" LSID="urn:lsid:plazi:treatment:03917D3BD907F77AB76B7834FD470935:8EE79AF6D907F771B0227AF6FEBE08F7" country="Madagascar" municipality="South Cape" name="Sainte Marie" pageId="3" pageNumber="4">Sainte Marie</location>
|
||
, 26˚16.35’S, 45˚10.45’E,
|
||
<quantity id="4CC061C8D907F771B73C7A99FD0C08F0" box="[563,692,1737,1764]" metricMagnitude="2" metricUnit="m" metricValue="4.41" metricValueMax="4.7299999999999995" metricValueMin="4.09" pageId="3" pageNumber="4" unit="m" value="441.0" valueMax="473.0" valueMin="409.0">
|
||
<elevation id="00152B1ED907F771B73C7A99FD0C08F0" box="[563,692,1737,1764]" metricMagnitude="2" metricUnit="m" metricValue="4.41" metricValueMax="4.7299999999999995" metricValueMin="4.09" pageId="3" pageNumber="4" unit="m" value="441.0" valueMax="473.0" valueMin="409.0">409-473 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FF86EAEDD907F771B7B07A99FCE408F7" box="[703,860,1737,1764]" pageId="3" pageNumber="4" value="2010-05-14">
|
||
<collectingDate id="EFC21305D907F771B7B07A99FCE408F7" box="[703,860,1737,1764]" pageId="3" pageNumber="4" value="2010-05-14">14 May 2010</collectingDate>
|
||
</date>
|
||
, collected during the cruise Atimo Vatae 2010, St. CP3616.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3229FA6D907F77AB5B27B7DFF550D85" lastPageId="8" lastPageNumber="9" pageId="3" pageNumber="4" type="description">
|
||
<paragraph id="8B87CC2DD907F771B5B27B7DFEF60953" blockId="3.[189,334,1837,1863]" box="[189,334,1837,1863]" pageId="3" pageNumber="4">
|
||
<heading id="D0CF7B41D907F771B5B27B7DFEF60953" bold="true" box="[189,334,1837,1863]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
|
||
<emphasis id="B94C103FD907F771B5B27B7DFEF60953" bold="true" box="[189,334,1837,1863]" pageId="3" pageNumber="4">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B87CC2DD907F774B5B27B0DFCFD0FC1" blockId="3.[189,1399,1884,1946]" lastBlockId="6.[189,1399,267,470]" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4">
|
||
MEASUREMENTS.
|
||
<typeStatus id="5483728FD907F771B4717B0CFE530962" box="[382,491,1884,1910]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
|
||
(MNHN IE-2013-8012): disc diameter
|
||
<quantity id="4CC061C8D907F771B6C37B0CFB810963" box="[972,1081,1884,1911]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="3" pageNumber="4" unit="mm" value="11.7">11.7 mm</quantity>
|
||
, arm length
|
||
<quantity id="4CC061C8D907F771B1DD7B0CFA800963" box="[1234,1336,1884,1911]" metricMagnitude="-1" metricUnit="m" metricValue="1.18" pageId="3" pageNumber="4" unit="mm" value="118.0">118 mm</quantity>
|
||
. One
|
||
<typeStatus id="5483728FD907F771B5B27BD0FE9A098E" box="[189,290,1920,1946]" pageId="3" pageNumber="4" type="paratype">paratype</typeStatus>
|
||
(SMNH-Type-8536): disc diameter
|
||
<quantity id="4CC061C8D907F771B7C17B2FFC82098E" box="[718,826,1919,1946]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="3" pageNumber="4" unit="mm" value="13.3">13.3 mm</quantity>
|
||
, arm length at least
|
||
<quantity id="4CC061C8D907F771B12B7B2FFB35098E" box="[1060,1165,1919,1946]" metricMagnitude="-2" metricUnit="m" metricValue="5.54" pageId="3" pageNumber="4" unit="mm" value="55.4">55.4 mm</quantity>
|
||
(arms convoluted). DISC. Disc circular in shape. Radial shields and their surrounds tumid (
|
||
<figureCitation id="1303D0A8D901F777B6F17D5BFBE50F32" box="[1022,1117,267,294]" captionStart="Fig" captionStartId="4.[189,232,1787,1813]" captionTargetBox="[278,1305,265,1748]" captionTargetId="figure@4.[277,1305,265,1748]" captionTargetPageId="4" captionText="Fig. 2. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H). A. Aboral disc and proximal portion of arms. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc and proximal portion of arms. E. Jaws. F. Oral periphery of disc and proximal portion of arm. G. Lateral disc. H. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835024" httpUri="https://zenodo.org/record/3835024/files/figure.png" pageId="5" pageNumber="6">Fig. 2A</figureCitation>
|
||
). Aboral surface of disc covered by skin and two to four domed tubercles in a line on each radial shield (
|
||
<figureCitation id="1303D0A8D901F777B15E7D7FFB730F5D" box="[1105,1227,303,329]" captionStart="Fig" captionStartId="4.[189,232,1787,1813]" captionTargetBox="[278,1305,265,1748]" captionTargetId="figure@4.[277,1305,265,1748]" captionTargetPageId="4" captionText="Fig. 2. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H). A. Aboral disc and proximal portion of arms. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc and proximal portion of arms. E. Jaws. F. Oral periphery of disc and proximal portion of arm. G. Lateral disc. H. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835024" httpUri="https://zenodo.org/record/3835024/files/figure.png" pageId="5" pageNumber="6">Fig. 2A-C</figureCitation>
|
||
). Tubercles
|
||
<emphasis id="B94C103FD901F777B05B7D60FACF0F5D" box="[1364,1399,304,329]" italics="true" pageId="5" pageNumber="6">ca.</emphasis>
|
||
1.0-
|
||
<quantity id="4CC061C8D901F777B5E77D02FEFA0F78" box="[232,322,338,364]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="6" unit="mm" value="1.2">1.2 mm</quantity>
|
||
long on disc periphery,
|
||
<emphasis id="B94C103FD901F777B7527D03FDC70F78" box="[605,639,339,364]" italics="true" pageId="5" pageNumber="6">ca.</emphasis>
|
||
<quantity id="4CC061C8D901F777B7887D02FD590F78" box="[647,737,338,364]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="6" unit="mm" value="0.8">0.8 mm</quantity>
|
||
long at disc center (
|
||
<figureCitation id="1303D0A8D901F777B6C07D02FB930F78" box="[975,1067,338,364]" captionStart="Fig" captionStartId="4.[189,232,1787,1813]" captionTargetBox="[278,1305,265,1748]" captionTargetId="figure@4.[277,1305,265,1748]" captionTargetPageId="4" captionText="Fig. 2. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H). A. Aboral disc and proximal portion of arms. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc and proximal portion of arms. E. Jaws. F. Oral periphery of disc and proximal portion of arm. G. Lateral disc. H. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835024" httpUri="https://zenodo.org/record/3835024/files/figure.png" pageId="5" pageNumber="6">Fig. 2B</figureCitation>
|
||
). Radial shields,
|
||
<emphasis id="B94C103FD901F777B1FD7D03FAAC0F78" box="[1266,1300,339,364]" italics="true" pageId="5" pageNumber="6">ca.</emphasis>
|
||
<quantity id="4CC061C8D901F777B0127D02FACE0F78" box="[1309,1398,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="6" unit="mm" value="7.5">7.5 mm</quantity>
|
||
long,
|
||
<quantity id="4CC061C8D902F774B40C7D5BFE370F32" box="[259,399,267,294]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="2.7" metricValueMin="0.7" pageId="6" pageNumber="7" unit="mm" value="1.7" valueMax="2.7" valueMin="0.7">0.7-2.7 mm</quantity>
|
||
wide, completely covered by skin and tubercles (
|
||
<figureCitation id="1303D0A8D902F774B6FF7D5BFBD60F31" box="[1008,1134,267,293]" captionStart="Fig" captionStartId="4.[189,232,1787,1813]" captionTargetBox="[278,1305,265,1748]" captionTargetId="figure@4.[277,1305,265,1748]" captionTargetPageId="4" captionText="Fig. 2. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H). A. Aboral disc and proximal portion of arms. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc and proximal portion of arms. E. Jaws. F. Oral periphery of disc and proximal portion of arm. G. Lateral disc. H. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835024" httpUri="https://zenodo.org/record/3835024/files/figure.png" pageId="6" pageNumber="7">Fig. 2B-C</figureCitation>
|
||
). Oral surface of disc covered by skin (
|
||
<figureCitation id="1303D0A8D902F774B48C7D7EFE450F5D" box="[387,509,302,329]" captionStart="Fig" captionStartId="4.[189,232,1787,1813]" captionTargetBox="[278,1305,265,1748]" captionTargetId="figure@4.[277,1305,265,1748]" captionTargetPageId="4" captionText="Fig. 2. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H). A. Aboral disc and proximal portion of arms. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc and proximal portion of arms. E. Jaws. F. Oral periphery of disc and proximal portion of arm. G. Lateral disc. H. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835024" httpUri="https://zenodo.org/record/3835024/files/figure.png" pageId="6" pageNumber="7">Fig. 2D-E</figureCitation>
|
||
). More than three triangular teeth form vertical row on dental plate (
|
||
<figureCitation id="1303D0A8D902F774B0007D7EFAD20F5D" box="[1295,1386,302,329]" captionStart="Fig" captionStartId="4.[189,232,1787,1813]" captionTargetBox="[278,1305,265,1748]" captionTargetId="figure@4.[277,1305,265,1748]" captionTargetPageId="4" captionText="Fig. 2. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H). A. Aboral disc and proximal portion of arms. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc and proximal portion of arms. E. Jaws. F. Oral periphery of disc and proximal portion of arm. G. Lateral disc. H. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835024" httpUri="https://zenodo.org/record/3835024/files/figure.png" pageId="6" pageNumber="7">Fig. 2E</figureCitation>
|
||
). One oral interradial plate present on distal side of adoral shield (
|
||
<figureCitation id="1303D0A8D902F774B6BC7D02FBAA0F78" box="[947,1042,338,364]" captionStart="Fig" captionStartId="4.[189,232,1787,1813]" captionTargetBox="[278,1305,265,1748]" captionTargetId="figure@4.[277,1305,265,1748]" captionTargetPageId="4" captionText="Fig. 2. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H). A. Aboral disc and proximal portion of arms. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc and proximal portion of arms. E. Jaws. F. Oral periphery of disc and proximal portion of arm. G. Lateral disc. H. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835024" httpUri="https://zenodo.org/record/3835024/files/figure.png" pageId="6" pageNumber="7">Fig. 2H</figureCitation>
|
||
). Lateral interradial surface of disc nearly vertical, covered by skin and granule-shaped external ossicles,
|
||
<emphasis id="B94C103FD902F774B1367D26FBE40F9B" box="[1081,1116,374,399]" italics="true" pageId="6" pageNumber="7">ca.</emphasis>
|
||
<quantity id="4CC061C8D902F774B16A7D25FB400F9B" box="[1125,1272,373,400]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="4.0" metricValueMin="1.5" pageId="6" pageNumber="7" unit="mm" value="275.0" valueMax="400.0" valueMin="150.0">150-400 µm</quantity>
|
||
long (
|
||
<figureCitation id="1303D0A8D902F774B0487D25FF5D0FA6" captionStart="Fig" captionStartId="4.[189,232,1787,1813]" captionTargetBox="[278,1305,265,1748]" captionTargetId="figure@4.[277,1305,265,1748]" captionTargetPageId="4" captionText="Fig. 2. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H). A. Aboral disc and proximal portion of arms. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc and proximal portion of arms. E. Jaws. F. Oral periphery of disc and proximal portion of arm. G. Lateral disc. H. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835024" httpUri="https://zenodo.org/record/3835024/files/figure.png" pageId="6" pageNumber="7">Fig. 2H</figureCitation>
|
||
). Two genital slits in each interradius, 3.0 mm long and
|
||
<quantity id="4CC061C8D902F774B6867DC8FC5D0FA7" box="[905,997,408,435]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="7" unit="mm" value="0.4">0.4 mm</quantity>
|
||
wide (
|
||
<figureCitation id="1303D0A8D902F774B1377DC8FB210FA6" box="[1080,1177,408,434]" captionStart="Fig" captionStartId="4.[189,232,1787,1813]" captionTargetBox="[278,1305,265,1748]" captionTargetId="figure@4.[277,1305,265,1748]" captionTargetPageId="4" captionText="Fig. 2. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H). A. Aboral disc and proximal portion of arms. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc and proximal portion of arms. E. Jaws. F. Oral periphery of disc and proximal portion of arm. G. Lateral disc. H. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835024" httpUri="https://zenodo.org/record/3835024/files/figure.png" pageId="6" pageNumber="7">Fig. 2H</figureCitation>
|
||
). One madreporite present on distal side of oral interradial plate (
|
||
<figureCitation id="1303D0A8D902F774B7D67DEBFC8F0FC1" box="[729,823,443,469]" captionStart="Fig" captionStartId="4.[189,232,1787,1813]" captionTargetBox="[278,1305,265,1748]" captionTargetId="figure@4.[277,1305,265,1748]" captionTargetPageId="4" captionText="Fig. 2. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H). A. Aboral disc and proximal portion of arms. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc and proximal portion of arms. E. Jaws. F. Oral periphery of disc and proximal portion of arm. G. Lateral disc. H. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835024" httpUri="https://zenodo.org/record/3835024/files/figure.png" pageId="6" pageNumber="7">Fig. 2H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF479CA5D900F776B5B27AABFC9209FC" ID-DOI="http://doi.org/10.5281/zenodo.3835024" ID-Zenodo-Dep="3835024" httpUri="https://zenodo.org/record/3835024/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1787,1813]" targetBox="[278,1305,265,1748]" targetPageId="4">
|
||
<paragraph id="8B87CC2DD900F776B5B27AABFC9209FC" blockId="4.[189,1399,1787,2024]" pageId="4" pageNumber="5">
|
||
<emphasis id="B94C103FD900F776B5B27AABFEB60901" bold="true" box="[189,270,1787,1813]" pageId="4" pageNumber="5">Fig. 2.</emphasis>
|
||
<taxonomicName id="4C38B7AED900F776B4167AABFD980901" authority="Okanishi & Fujita, 2014" authorityName="Okanishi & Fujita" authorityYear="2014" box="[281,544,1787,1813]" class="Ophiuroidea" family="Euryalidae" genus="Asterostegus" kingdom="Animalia" order="Euryalida" pageId="4" pageNumber="5" phylum="Echinodermata" rank="species" species="sabineae" status="sp. nov.">
|
||
<emphasis id="B94C103FD900F776B4167AABFD980901" box="[281,544,1787,1813]" italics="true" pageId="4" pageNumber="5">Asterostegus sabineae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27FAD44D900F776B7247AABFD340901" box="[555,652,1787,1813]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H).
|
||
<emphasis id="B94C103FD900F776B4887B4EFE24092C" bold="true" box="[391,412,1822,1848]" pageId="4" pageNumber="5">A</emphasis>
|
||
. Aboral disc and proximal portion of arms.
|
||
<emphasis id="B94C103FD900F776B6A47B4EFC07092C" bold="true" box="[939,959,1822,1848]" pageId="4" pageNumber="5">B</emphasis>
|
||
. Aboral periphery of disc.
|
||
<emphasis id="B94C103FD900F776B00E7B4EFAAE092C" bold="true" box="[1281,1302,1822,1848]" pageId="4" pageNumber="5">C</emphasis>
|
||
. Aboral central disc.
|
||
<emphasis id="B94C103FD900F776B45E7B11FEDE094F" bold="true" box="[337,358,1857,1883]" pageId="4" pageNumber="5">D</emphasis>
|
||
. Oral disc and proximal portion of arms.
|
||
<emphasis id="B94C103FD900F776B6457B11FCE6094F" bold="true" box="[842,862,1857,1883]" pageId="4" pageNumber="5">E</emphasis>
|
||
. Jaws.
|
||
<emphasis id="B94C103FD900F776B6BC7B11FC7D094F" bold="true" box="[947,965,1857,1883]" pageId="4" pageNumber="5">F</emphasis>
|
||
. Oral periphery of disc and proximal portion of arm.
|
||
<emphasis id="B94C103FD900F776B4727B34FE2C096A" bold="true" box="[381,404,1892,1918]" pageId="4" pageNumber="5">G</emphasis>
|
||
. Lateral disc.
|
||
<emphasis id="B94C103FD900F776B74C7B35FDE2096B" bold="true" box="[579,602,1893,1919]" pageId="4" pageNumber="5">H</emphasis>
|
||
. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF479CA5D901F777B5B27B34FC2809FC" ID-DOI="http://doi.org/10.5281/zenodo.3835026" ID-Zenodo-Dep="3835026" httpUri="https://zenodo.org/record/3835026/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1892,1918]" targetBox="[188,1397,406,1852]" targetPageId="5">
|
||
<paragraph id="8B87CC2DD901F777B5B27B34FC2809FC" blockId="5.[189,1399,1892,2025]" pageId="5" pageNumber="6">
|
||
<emphasis id="B94C103FD901F777B5B27B34FEB2096A" bold="true" box="[189,266,1892,1918]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
|
||
<taxonomicName id="4C38B7AED901F777B41F7B35FDAC096B" authority="Okanishi & Fujita, 2014" authorityName="Okanishi & Fujita" authorityYear="2014" box="[272,532,1893,1919]" class="Ophiuroidea" family="Euryalidae" genus="Asterostegus" kingdom="Animalia" order="Euryalida" pageId="5" pageNumber="6" phylum="Echinodermata" rank="species" species="sabineae" status="sp. nov.">
|
||
<emphasis id="B94C103FD901F777B41F7B35FDAC096B" box="[272,532,1893,1919]" italics="true" pageId="5" pageNumber="6">Asterostegus sabineae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27FAD44D901F777B7147B35FDCF096B" box="[539,631,1893,1919]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (MNHN IE-2013-8012) (A–F).
|
||
<emphasis id="B94C103FD901F777B16A7B34FBC2096A" bold="true" box="[1125,1146,1892,1918]" pageId="5" pageNumber="6">A</emphasis>
|
||
. Aboral basal portion of arm.
|
||
<emphasis id="B94C103FD901F777B4147BD8FE9709B6" bold="true" box="[283,303,1928,1954]" pageId="5" pageNumber="6">B</emphasis>
|
||
. Aboral middle portion of arm.
|
||
<emphasis id="B94C103FD901F777B7A67BD8FD0609B6" bold="true" box="[681,702,1928,1954]" pageId="5" pageNumber="6">C</emphasis>
|
||
. Aboral and lateral distal portion of arm.
|
||
<emphasis id="B94C103FD901F777B1A37BD8FB7909B6" bold="true" box="[1196,1217,1928,1954]" pageId="5" pageNumber="6">D</emphasis>
|
||
. Oral proximal portion of arm.
|
||
<emphasis id="B94C103FD901F777B47C7BFBFE3F09D1" bold="true" box="[371,391,1963,1989]" pageId="5" pageNumber="6">E</emphasis>
|
||
. Oral middle portion of arm.
|
||
<emphasis id="B94C103FD901F777B7D17BFBFD4809D1" bold="true" box="[734,752,1963,1989]" pageId="5" pageNumber="6">F</emphasis>
|
||
. Lateral distal portion of arm. Abbreviations: AS = arm spine, LAP = lateral arm plate, Ten = tentacles, Tu = tubercle.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF479CA5D902F774B5B27B12FEFD09FD" ID-DOI="http://doi.org/10.5281/zenodo.3835028" ID-Zenodo-Dep="3835028" httpUri="https://zenodo.org/record/3835028/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1858,1884]" targetBox="[188,1398,509,1814]" targetPageId="6">
|
||
<paragraph id="8B87CC2DD902F774B5B27B12FEFD09FD" blockId="6.[189,1399,1858,2025]" pageId="6" pageNumber="7">
|
||
<emphasis id="B94C103FD902F774B5B27B12FEB10948" bold="true" box="[189,265,1858,1884]" pageId="6" pageNumber="7">Fig. 4.</emphasis>
|
||
<taxonomicName id="4C38B7AED902F774B4007B13FDA90949" authority="Okanishi & Fujita, 2014" authorityName="Okanishi & Fujita" authorityYear="2014" box="[271,529,1859,1885]" class="Ophiuroidea" family="Euryalidae" genus="Asterostegus" kingdom="Animalia" order="Euryalida" pageId="6" pageNumber="7" phylum="Echinodermata" rank="species" species="sabineae" status="sp. nov.">
|
||
<emphasis id="B94C103FD902F774B4007B13FDA90949" box="[271,529,1859,1885]" italics="true" pageId="6" pageNumber="7">Asterostegus sabineae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27FAD44D902F774B7187B13FDCB0949" box="[535,627,1859,1885]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype (SMNH-Type-8536), SEM photographs.
|
||
<emphasis id="B94C103FD902F774B1CA7B12FB410948" bold="true" box="[1221,1273,1858,1884]" pageId="6" pageNumber="7">A-D</emphasis>
|
||
. Vertebrae from middle portion of arm: distal view (A), aboral view (B), oral view (C), lateral view (D).
|
||
<emphasis id="B94C103FD902F774B0347B36FAD70994" bold="true" box="[1339,1391,1894,1920]" pageId="6" pageNumber="7">E-H</emphasis>
|
||
. Vertebrae from distal portion of arm: distal view (E), aboral view (F), oral view (G), lateral view (H).
|
||
<emphasis id="B94C103FD902F774B06B7BD9FAD709B7" bold="true" box="[1380,1391,1929,1955]" pageId="6" pageNumber="7">I</emphasis>
|
||
. Lateral arm plate from middle portion of arm. Abbreviations: K = knobs, MO = muscle opening, OB = oral bridge.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B87CC2DD903F77AB5B27D5CFD5C0F5D" blockId="7.[189,1399,267,681]" lastBlockId="8.[189,1399,267,329]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
|
||
ARMS. Arms simple, five in number, with no abrupt gap in width due to presence of gonads in proximal portion of arms (
|
||
<figureCitation id="1303D0A8D903F775B48F7D7EFE5A0F5D" box="[384,482,302,329]" captionStart="Fig" captionStartId="4.[189,232,1787,1813]" captionTargetBox="[278,1305,265,1748]" captionTargetId="figure@4.[277,1305,265,1748]" captionTargetPageId="4" captionText="Fig. 2. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H). A. Aboral disc and proximal portion of arms. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc and proximal portion of arms. E. Jaws. F. Oral periphery of disc and proximal portion of arm. G. Lateral disc. H. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835024" httpUri="https://zenodo.org/record/3835024/files/figure.png" pageId="7" pageNumber="8">Figs 2A</figureCitation>
|
||
;
|
||
<figureCitation id="1303D0A8D903F775B4E17D7FFDAD0F5D" box="[494,533,303,329]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[188,1397,406,1852]" captionTargetId="figure@5.[188,1399,404,1852]" captionTargetPageId="5" captionText="Fig. 3. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) (A–F). A. Aboral basal portion of arm. B. Aboral middle portion of arm. C. Aboral and lateral distal portion of arm. D. Oral proximal portion of arm. E. Oral middle portion of arm. F. Lateral distal portion of arm. Abbreviations: AS = arm spine, LAP = lateral arm plate, Ten = tentacles, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835026" httpUri="https://zenodo.org/record/3835026/files/figure.png" pageId="7" pageNumber="8">3A</figureCitation>
|
||
). Arms tapering gradually distally. Proximal portion of arms 3.0 mm wide and
|
||
<quantity id="4CC061C8D903F775B5E17D02FEF00F78" box="[238,328,338,364]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="8" unit="mm" value="3.7">3.7 mm</quantity>
|
||
high, oblong in cross-section. Aboral surface arched and oral surface flattened from middle to distal portion of arms. Proximal portion of the arms covered by skin and one (rarely two) domed tubercles,
|
||
<emphasis id="B94C103FD903F775B4377DC9FEE30FA6" box="[312,347,409,434]" italics="true" pageId="7" pageNumber="8">ca.</emphasis>
|
||
1.0-
|
||
<quantity id="4CC061C8D903F775B49B7DC8FE490FA7" box="[404,497,408,435]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="8" unit="mm" value="1.3">1.3 mm</quantity>
|
||
long (
|
||
<figureCitation id="1303D0A8D903F775B74E7DC8FD1A0FA6" box="[577,674,408,434]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[188,1397,406,1852]" captionTargetId="figure@5.[188,1399,404,1852]" captionTargetPageId="5" captionText="Fig. 3. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) (A–F). A. Aboral basal portion of arm. B. Aboral middle portion of arm. C. Aboral and lateral distal portion of arm. D. Oral proximal portion of arm. E. Oral middle portion of arm. F. Lateral distal portion of arm. Abbreviations: AS = arm spine, LAP = lateral arm plate, Ten = tentacles, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835026" httpUri="https://zenodo.org/record/3835026/files/figure.png" pageId="7" pageNumber="8">Fig. 3A</figureCitation>
|
||
), on each lateral bar. If two tubercles present on lateral bar, both are smaller,
|
||
<emphasis id="B94C103FD903F775B4877DECFE130FC1" box="[392,427,444,469]" italics="true" pageId="7" pageNumber="8">ca.</emphasis>
|
||
<quantity id="4CC061C8D903F775B4BD7DEBFDB60FC2" box="[434,526,443,470]" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="8" unit="mm" value="800.0">800 µm</quantity>
|
||
long (
|
||
<figureCitation id="1303D0A8D903F775B7567DEBFD000FC1" box="[601,696,443,469]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[188,1397,406,1852]" captionTargetId="figure@5.[188,1399,404,1852]" captionTargetPageId="5" captionText="Fig. 3. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) (A–F). A. Aboral basal portion of arm. B. Aboral middle portion of arm. C. Aboral and lateral distal portion of arm. D. Oral proximal portion of arm. E. Oral middle portion of arm. F. Lateral distal portion of arm. Abbreviations: AS = arm spine, LAP = lateral arm plate, Ten = tentacles, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835026" httpUri="https://zenodo.org/record/3835026/files/figure.png" pageId="7" pageNumber="8">Fig. 3A</figureCitation>
|
||
). From middle to distal portion of arms, tubercles gradually decrease in size distally, becoming absent near arm tips (
|
||
<figureCitation id="1303D0A8D903F775B66B7D8EFC580FED" box="[868,992,478,505]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[188,1397,406,1852]" captionTargetId="figure@5.[188,1399,404,1852]" captionTargetPageId="5" captionText="Fig. 3. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) (A–F). A. Aboral basal portion of arm. B. Aboral middle portion of arm. C. Aboral and lateral distal portion of arm. D. Oral proximal portion of arm. E. Oral middle portion of arm. F. Lateral distal portion of arm. Abbreviations: AS = arm spine, LAP = lateral arm plate, Ten = tentacles, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835026" httpUri="https://zenodo.org/record/3835026/files/figure.png" pageId="7" pageNumber="8">Fig. 3B-C</figureCitation>
|
||
). Oral surface of arms covered by skin (
|
||
<figureCitation id="1303D0A8D903F775B5F17E52FEC10C08" box="[254,377,514,540]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[188,1397,406,1852]" captionTargetId="figure@5.[188,1399,404,1852]" captionTargetPageId="5" captionText="Fig. 3. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) (A–F). A. Aboral basal portion of arm. B. Aboral middle portion of arm. C. Aboral and lateral distal portion of arm. D. Oral proximal portion of arm. E. Oral middle portion of arm. F. Lateral distal portion of arm. Abbreviations: AS = arm spine, LAP = lateral arm plate, Ten = tentacles, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835026" httpUri="https://zenodo.org/record/3835026/files/figure.png" pageId="7" pageNumber="8">Fig. 3D-E</figureCitation>
|
||
). First to third tentacle pores lacking arm spines; two arm spines from fourth pore (
|
||
<figureCitation id="1303D0A8D903F775B0487E52FF580C2B" captionStart="Fig" captionStartId="4.[189,232,1787,1813]" captionTargetBox="[278,1305,265,1748]" captionTargetId="figure@4.[277,1305,265,1748]" captionTargetPageId="4" captionText="Fig. 2. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H). A. Aboral disc and proximal portion of arms. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc and proximal portion of arms. E. Jaws. F. Oral periphery of disc and proximal portion of arm. G. Lateral disc. H. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835024" httpUri="https://zenodo.org/record/3835024/files/figure.png" pageId="7" pageNumber="8">Fig. 2F</figureCitation>
|
||
). In proximal third of arms, outer arm spines
|
||
<emphasis id="B94C103FD903F775B60A7E76FC900C2B" box="[773,808,550,575]" italics="true" pageId="7" pageNumber="8">ca.</emphasis>
|
||
one quarter to half as long as corresponding arm segment, and inner arm spines slightly shorter than outer spines (
|
||
<figureCitation id="1303D0A8D903F775B6B67E18FBA00C76" box="[953,1048,584,610]" captionStart="Fig" captionStartId="4.[189,232,1787,1813]" captionTargetBox="[278,1305,265,1748]" captionTargetId="figure@4.[277,1305,265,1748]" captionTargetPageId="4" captionText="Fig. 2. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H). A. Aboral disc and proximal portion of arms. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc and proximal portion of arms. E. Jaws. F. Oral periphery of disc and proximal portion of arm. G. Lateral disc. H. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835024" httpUri="https://zenodo.org/record/3835024/files/figure.png" pageId="7" pageNumber="8">Figs 2F</figureCitation>
|
||
;
|
||
<figureCitation id="1303D0A8D903F775B12B7E18FBF30C76" box="[1060,1099,584,610]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[188,1397,406,1852]" captionTargetId="figure@5.[188,1399,404,1852]" captionTargetPageId="5" captionText="Fig. 3. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) (A–F). A. Aboral basal portion of arm. B. Aboral middle portion of arm. C. Aboral and lateral distal portion of arm. D. Oral proximal portion of arm. E. Oral middle portion of arm. F. Lateral distal portion of arm. Abbreviations: AS = arm spine, LAP = lateral arm plate, Ten = tentacles, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835026" httpUri="https://zenodo.org/record/3835026/files/figure.png" pageId="7" pageNumber="8">3D</figureCitation>
|
||
). In middle of arms inner and outer arm spines subequal,
|
||
<emphasis id="B94C103FD903F775B7227E3CFDE80C91" box="[557,592,620,645]" italics="true" pageId="7" pageNumber="8">ca.</emphasis>
|
||
two-thirds as long as corresponding arm segment (
|
||
<figureCitation id="1303D0A8D903F775B1AA7E3BFABB0C91" box="[1189,1283,619,645]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[188,1397,406,1852]" captionTargetId="figure@5.[188,1399,404,1852]" captionTargetPageId="5" captionText="Fig. 3. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) (A–F). A. Aboral basal portion of arm. B. Aboral middle portion of arm. C. Aboral and lateral distal portion of arm. D. Oral proximal portion of arm. E. Oral middle portion of arm. F. Lateral distal portion of arm. Abbreviations: AS = arm spine, LAP = lateral arm plate, Ten = tentacles, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835026" httpUri="https://zenodo.org/record/3835026/files/figure.png" pageId="7" pageNumber="8">Fig. 3D</figureCitation>
|
||
). In distal third of arms, arm spines hook-shaped (
|
||
<figureCitation id="1303D0A8D903F775B7837EDEFD5D0CBD" box="[652,741,654,681]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[188,1397,406,1852]" captionTargetId="figure@5.[188,1399,404,1852]" captionTargetPageId="5" captionText="Fig. 3. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) (A–F). A. Aboral basal portion of arm. B. Aboral middle portion of arm. C. Aboral and lateral distal portion of arm. D. Oral proximal portion of arm. E. Oral middle portion of arm. F. Lateral distal portion of arm. Abbreviations: AS = arm spine, LAP = lateral arm plate, Ten = tentacles, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835026" httpUri="https://zenodo.org/record/3835026/files/figure.png" pageId="7" pageNumber="8">Fig. 3F</figureCitation>
|
||
). Inner and outer spines subequal,
|
||
<emphasis id="B94C103FD903F775B1767EDFFB240CBC" box="[1145,1180,655,680]" italics="true" pageId="7" pageNumber="8">ca.</emphasis>
|
||
two-thirds as long as corresponding arm segment (
|
||
<figureCitation id="1303D0A8D90CF77AB73C7D5BFD730F32" box="[563,715,267,294]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[188,1397,406,1852]" captionTargetId="figure@5.[188,1399,404,1852]" captionTargetPageId="5" captionText="Fig. 3. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) (A–F). A. Aboral basal portion of arm. B. Aboral middle portion of arm. C. Aboral and lateral distal portion of arm. D. Oral proximal portion of arm. E. Oral middle portion of arm. F. Lateral distal portion of arm. Abbreviations: AS = arm spine, LAP = lateral arm plate, Ten = tentacles, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835026" httpUri="https://zenodo.org/record/3835026/files/figure.png" pageId="8" pageNumber="9">Fig. 3C, E-F</figureCitation>
|
||
). Lateral arm plates concealed by skin and visible at distal arm tips where skin becomes thinner (
|
||
<figureCitation id="1303D0A8D90CF77AB7737D7EFD6F0F5D" box="[636,727,302,329]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[188,1397,406,1852]" captionTargetId="figure@5.[188,1399,404,1852]" captionTargetPageId="5" captionText="Fig. 3. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) (A–F). A. Aboral basal portion of arm. B. Aboral middle portion of arm. C. Aboral and lateral distal portion of arm. D. Oral proximal portion of arm. E. Oral middle portion of arm. F. Lateral distal portion of arm. Abbreviations: AS = arm spine, LAP = lateral arm plate, Ten = tentacles, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835026" httpUri="https://zenodo.org/record/3835026/files/figure.png" pageId="8" pageNumber="9">Fig. 3E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF479CA5D903F775B5B27B12FC6E09FD" ID-DOI="http://doi.org/10.5281/zenodo.3835030" ID-Zenodo-Dep="3835030" httpUri="https://zenodo.org/record/3835030/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1858,1884]" targetBox="[192,1396,755,1814]" targetPageId="7">
|
||
<paragraph id="8B87CC2DD903F775B5B27B12FC6E09FD" blockId="7.[189,1399,1858,2026]" pageId="7" pageNumber="8">
|
||
<emphasis id="B94C103FD903F775B5B27B12FEB10949" bold="true" box="[189,265,1858,1885]" pageId="7" pageNumber="8">Fig. 5.</emphasis>
|
||
<taxonomicName id="4C38B7AED903F775B41F7B13FDAB0949" authority="Okanishi & Fujita, 2014" authorityName="Okanishi & Fujita" authorityYear="2014" box="[272,531,1859,1885]" class="Ophiuroidea" family="Euryalidae" genus="Asterostegus" kingdom="Animalia" order="Euryalida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="species" species="sabineae" status="sp. nov.">
|
||
<emphasis id="B94C103FD903F775B41F7B13FDAB0949" box="[272,531,1859,1885]" italics="true" pageId="7" pageNumber="8">Asterostegus sabineae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27FAD44D903F775B7157B13FDCE0949" box="[538,630,1859,1885]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype (SMNH-Type-8536), SEM photographs.
|
||
<emphasis id="B94C103FD903F775B1C37B12FB590948" bold="true" box="[1228,1249,1858,1884]" pageId="7" pageNumber="8">A</emphasis>
|
||
. Lateral arm plate from middle portion of arm, abradial view.
|
||
<emphasis id="B94C103FD903F775B7FE7B36FCBD0994" bold="true" box="[753,773,1894,1920]" pageId="7" pageNumber="8">B</emphasis>
|
||
. Lateral arm plate from distal portion of arm, adradial view.
|
||
<emphasis id="B94C103FD903F775B40D7BD9FEAF09B7" bold="true" box="[258,279,1929,1955]" pageId="7" pageNumber="8">C</emphasis>
|
||
. Lateral arm plate from distal portion of arm, oral view.
|
||
<emphasis id="B94C103FD903F775B6AD7BD9FC6B09B7" bold="true" box="[930,979,1929,1955]" pageId="7" pageNumber="8">D-F</emphasis>
|
||
. Arm spines: from proximal portion of arm (D), middle portion of arm (E), distal portion of arm, an arc indicates lamina (F). Abbreviations: MO = muscle opening, NO = nerve opening, ST = secondary tooth.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B87CC2DD90CF77AB5B27D20FAFD0F9E" blockId="8.[189,1349,368,394]" box="[189,1349,368,394]" pageId="8" pageNumber="9">
|
||
COLOUR. Body creamy white, arm spines brown (
|
||
<figureCitation id="1303D0A8D90CF77AB7F57D20FC1C0F9E" box="[762,932,368,394]" captionStart="Fig" captionStartId="4.[189,232,1787,1813]" captionTargetBox="[278,1305,265,1748]" captionTargetId="figure@4.[277,1305,265,1748]" captionTargetPageId="4" captionText="Fig. 2. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H). A. Aboral disc and proximal portion of arms. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc and proximal portion of arms. E. Jaws. F. Oral periphery of disc and proximal portion of arm. G. Lateral disc. H. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835024" httpUri="https://zenodo.org/record/3835024/files/figure.png" pageId="8" pageNumber="9">Figs 2A, G, H</figureCitation>
|
||
;
|
||
<figureCitation id="1303D0A8D90CF77AB6BE7D20FC430F9E" box="[945,1019,368,394]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[188,1397,406,1852]" captionTargetId="figure@5.[188,1399,404,1852]" captionTargetPageId="5" captionText="Fig. 3. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) (A–F). A. Aboral basal portion of arm. B. Aboral middle portion of arm. C. Aboral and lateral distal portion of arm. D. Oral proximal portion of arm. E. Oral middle portion of arm. F. Lateral distal portion of arm. Abbreviations: AS = arm spine, LAP = lateral arm plate, Ten = tentacles, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835026" httpUri="https://zenodo.org/record/3835026/files/figure.png" pageId="8" pageNumber="9">3B, D</figureCitation>
|
||
). Colour in life is unknown.
|
||
</paragraph>
|
||
<paragraph id="8B87CC2DD90CF77AB5B27DE2FE2E0C68" blockId="8.[189,1399,433,636]" pageId="8" pageNumber="9">
|
||
OSSICLE MORPHOLOGY. Ossicles separated from one
|
||
<typeStatus id="5483728FD90CF77AB62F7DE2FC300FD8" box="[800,904,434,460]" pageId="8" pageNumber="9" type="paratype">paratype</typeStatus>
|
||
, SMNH-Type-8536. Vertebrae. Vertebrae with streptospondylous articulations (
|
||
<figureCitation id="1303D0A8D90CF77AB7747D85FD440FFB" box="[635,764,469,495]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[188,1398,509,1814]" captionTargetId="figure@6.[188,1398,507,1814]" captionTargetPageId="6" captionText="Fig. 4. Asterostegus sabineae sp. nov., paratype (SMNH-Type-8536), SEM photographs. A-D. Vertebrae from middle portion of arm: distal view (A), aboral view (B), oral view (C), lateral view (D). E-H. Vertebrae from distal portion of arm: distal view (E), aboral view (F), oral view (G), lateral view (H). I. Lateral arm plate from middle portion of arm. Abbreviations: K = knobs, MO = muscle opening, OB = oral bridge." figureDoi="http://doi.org/10.5281/zenodo.3835028" httpUri="https://zenodo.org/record/3835028/files/figure.png" pageId="8" pageNumber="9">Fig. 4A, E</figureCitation>
|
||
), oral bridges on oral side (
|
||
<figureCitation id="1303D0A8D90CF77AB14A7D85FB7E0FFB" box="[1093,1222,469,495]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[188,1398,509,1814]" captionTargetId="figure@6.[188,1398,507,1814]" captionTargetPageId="6" captionText="Fig. 4. Asterostegus sabineae sp. nov., paratype (SMNH-Type-8536), SEM photographs. A-D. Vertebrae from middle portion of arm: distal view (A), aboral view (B), oral view (C), lateral view (D). E-H. Vertebrae from distal portion of arm: distal view (E), aboral view (F), oral view (G), lateral view (H). I. Lateral arm plate from middle portion of arm. Abbreviations: K = knobs, MO = muscle opening, OB = oral bridge." figureDoi="http://doi.org/10.5281/zenodo.3835028" httpUri="https://zenodo.org/record/3835028/files/figure.png" pageId="8" pageNumber="9">Fig. 4C, G</figureCitation>
|
||
) and knobs on lateral side (
|
||
<figureCitation id="1303D0A8D90CF77AB45E7DA8FE6E0C06" box="[337,470,504,530]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[188,1398,509,1814]" captionTargetId="figure@6.[188,1398,507,1814]" captionTargetPageId="6" captionText="Fig. 4. Asterostegus sabineae sp. nov., paratype (SMNH-Type-8536), SEM photographs. A-D. Vertebrae from middle portion of arm: distal view (A), aboral view (B), oral view (C), lateral view (D). E-H. Vertebrae from distal portion of arm: distal view (E), aboral view (F), oral view (G), lateral view (H). I. Lateral arm plate from middle portion of arm. Abbreviations: K = knobs, MO = muscle opening, OB = oral bridge." figureDoi="http://doi.org/10.5281/zenodo.3835028" httpUri="https://zenodo.org/record/3835028/files/figure.png" pageId="8" pageNumber="9">Fig. 4D, H</figureCitation>
|
||
), and not inclining from proximal-aboral side to distal-oral side of arms (
|
||
<figureCitation id="1303D0A8D90CF77AB0487DA8FEE90C22" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[188,1398,509,1814]" captionTargetId="figure@6.[188,1398,507,1814]" captionTargetPageId="6" captionText="Fig. 4. Asterostegus sabineae sp. nov., paratype (SMNH-Type-8536), SEM photographs. A-D. Vertebrae from middle portion of arm: distal view (A), aboral view (B), oral view (C), lateral view (D). E-H. Vertebrae from distal portion of arm: distal view (E), aboral view (F), oral view (G), lateral view (H). I. Lateral arm plate from middle portion of arm. Abbreviations: K = knobs, MO = muscle opening, OB = oral bridge." figureDoi="http://doi.org/10.5281/zenodo.3835028" httpUri="https://zenodo.org/record/3835028/files/figure.png" pageId="8" pageNumber="9">Fig. 4B, D, F, H</figureCitation>
|
||
) throughout the arms. These features indicate the new species’ affiliation to the family
|
||
<taxonomicName id="4C38B7AED90CF77AB5B27E6EFEF80C4C" authorityName="Gray" authorityYear="1840" box="[189,320,574,600]" class="Ophiuroidea" family="Euryalidae" kingdom="Animalia" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="family">Euryalidae</taxonomicName>
|
||
. Knobs on proximal portion of arms elongated, becoming rounded toward distal portion of arms (
|
||
<figureCitation id="1303D0A8D90CF77AB4097E31FE300C68" box="[262,392,609,636]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[188,1398,509,1814]" captionTargetId="figure@6.[188,1398,507,1814]" captionTargetPageId="6" captionText="Fig. 4. Asterostegus sabineae sp. nov., paratype (SMNH-Type-8536), SEM photographs. A-D. Vertebrae from middle portion of arm: distal view (A), aboral view (B), oral view (C), lateral view (D). E-H. Vertebrae from distal portion of arm: distal view (E), aboral view (F), oral view (G), lateral view (H). I. Lateral arm plate from middle portion of arm. Abbreviations: K = knobs, MO = muscle opening, OB = oral bridge." figureDoi="http://doi.org/10.5281/zenodo.3835028" httpUri="https://zenodo.org/record/3835028/files/figure.png" pageId="8" pageNumber="9">Fig. 4D, H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B87CC2DD90CF77AB5B27EF3FF550D85" blockId="8.[189,1399,675,913]" pageId="8" pageNumber="9">
|
||
LATERAL ARM PLATES. Lateral arm plates in middle portion of arms with two pairs of muscle and nerve openings, each of them associated with arm spine articulation (
|
||
<figureCitation id="1303D0A8D90CF77AB6927E96FC4C0CF4" box="[925,1012,710,736]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[188,1398,509,1814]" captionTargetId="figure@6.[188,1398,507,1814]" captionTargetPageId="6" captionText="Fig. 4. Asterostegus sabineae sp. nov., paratype (SMNH-Type-8536), SEM photographs. A-D. Vertebrae from middle portion of arm: distal view (A), aboral view (B), oral view (C), lateral view (D). E-H. Vertebrae from distal portion of arm: distal view (E), aboral view (F), oral view (G), lateral view (H). I. Lateral arm plate from middle portion of arm. Abbreviations: K = knobs, MO = muscle opening, OB = oral bridge." figureDoi="http://doi.org/10.5281/zenodo.3835028" httpUri="https://zenodo.org/record/3835028/files/figure.png" pageId="8" pageNumber="9">Figs 4I</figureCitation>
|
||
,
|
||
<figureCitation id="1303D0A8D90CF77AB10F7E96FBFD0CF5" box="[1024,1093,710,737]" captionStart="Fig" captionStartId="7.[189,232,1858,1884]" captionTargetBox="[192,1396,755,1814]" captionTargetId="figure@7.[189,1398,752,1814]" captionTargetPageId="7" captionText="Fig. 5. Asterostegus sabineae sp. nov., paratype (SMNH-Type-8536), SEM photographs. A. Lateral arm plate from middle portion of arm, abradial view. B. Lateral arm plate from distal portion of arm, adradial view. C. Lateral arm plate from distal portion of arm, oral view. D-F. Arm spines: from proximal portion of arm (D), middle portion of arm (E), distal portion of arm, an arc indicates lamina (F). Abbreviations: MO = muscle opening, NO = nerve opening, ST = secondary tooth." figureDoi="http://doi.org/10.5281/zenodo.3835030" httpUri="https://zenodo.org/record/3835030/files/figure.png" pageId="8" pageNumber="9">5A-C</figureCitation>
|
||
). Dorsal and ventral lobes beside muscle openings contact each other, a tubercle structure present between each muscle and nerve opening (
|
||
<figureCitation id="1303D0A8D90CF77AB4257F5CFE380D33" box="[298,384,780,807]" captionStart="Fig" captionStartId="6.[189,232,1858,1884]" captionTargetBox="[188,1398,509,1814]" captionTargetId="figure@6.[188,1398,507,1814]" captionTargetPageId="6" captionText="Fig. 4. Asterostegus sabineae sp. nov., paratype (SMNH-Type-8536), SEM photographs. A-D. Vertebrae from middle portion of arm: distal view (A), aboral view (B), oral view (C), lateral view (D). E-H. Vertebrae from distal portion of arm: distal view (E), aboral view (F), oral view (G), lateral view (H). I. Lateral arm plate from middle portion of arm. Abbreviations: K = knobs, MO = muscle opening, OB = oral bridge." figureDoi="http://doi.org/10.5281/zenodo.3835028" httpUri="https://zenodo.org/record/3835028/files/figure.png" pageId="8" pageNumber="9">Figs 4I</figureCitation>
|
||
,
|
||
<figureCitation id="1303D0A8D90CF77AB4857F5CFE0A0D33" box="[394,434,780,807]" captionStart="Fig" captionStartId="7.[189,232,1858,1884]" captionTargetBox="[192,1396,755,1814]" captionTargetId="figure@7.[189,1398,752,1814]" captionTargetPageId="7" captionText="Fig. 5. Asterostegus sabineae sp. nov., paratype (SMNH-Type-8536), SEM photographs. A. Lateral arm plate from middle portion of arm, abradial view. B. Lateral arm plate from distal portion of arm, adradial view. C. Lateral arm plate from distal portion of arm, oral view. D-F. Arm spines: from proximal portion of arm (D), middle portion of arm (E), distal portion of arm, an arc indicates lamina (F). Abbreviations: MO = muscle opening, NO = nerve opening, ST = secondary tooth." figureDoi="http://doi.org/10.5281/zenodo.3835030" httpUri="https://zenodo.org/record/3835030/files/figure.png" pageId="8" pageNumber="9">5A</figureCitation>
|
||
). Arm spines club-shaped, both
|
||
<quantity id="4CC061C8D90CF77AB62A7F5CFC380D33" box="[805,896,780,807]" metricMagnitude="-1" metricUnit="m" metricValue="6.5" pageId="8" pageNumber="9" unit="mm" value="650.0">650 µm</quantity>
|
||
long and wide on proximal portion of arms (
|
||
<figureCitation id="1303D0A8D90CF77AB5C97F60FE9A0D5E" box="[198,290,815,842]" captionStart="Fig" captionStartId="7.[189,232,1858,1884]" captionTargetBox="[192,1396,755,1814]" captionTargetId="figure@7.[189,1398,752,1814]" captionTargetPageId="7" captionText="Fig. 5. Asterostegus sabineae sp. nov., paratype (SMNH-Type-8536), SEM photographs. A. Lateral arm plate from middle portion of arm, abradial view. B. Lateral arm plate from distal portion of arm, adradial view. C. Lateral arm plate from distal portion of arm, oral view. D-F. Arm spines: from proximal portion of arm (D), middle portion of arm (E), distal portion of arm, an arc indicates lamina (F). Abbreviations: MO = muscle opening, NO = nerve opening, ST = secondary tooth." figureDoi="http://doi.org/10.5281/zenodo.3835030" httpUri="https://zenodo.org/record/3835030/files/figure.png" pageId="8" pageNumber="9">Fig. 5D</figureCitation>
|
||
),
|
||
<quantity id="4CC061C8D90CF77AB4387F7FFE2A0D5E" box="[311,402,815,842]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="9" unit="mm" value="450.0">450 µm</quantity>
|
||
long and
|
||
<quantity id="4CC061C8D90CF77AB70C7F7FFDE60D5E" box="[515,606,815,842]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="mm" value="500.0">500 µm</quantity>
|
||
wide on middle portion of arms (
|
||
<figureCitation id="1303D0A8D90CF77AB6EE7F60FB830D5E" box="[993,1083,815,842]" captionStart="Fig" captionStartId="7.[189,232,1858,1884]" captionTargetBox="[192,1396,755,1814]" captionTargetId="figure@7.[189,1398,752,1814]" captionTargetPageId="7" captionText="Fig. 5. Asterostegus sabineae sp. nov., paratype (SMNH-Type-8536), SEM photographs. A. Lateral arm plate from middle portion of arm, abradial view. B. Lateral arm plate from distal portion of arm, adradial view. C. Lateral arm plate from distal portion of arm, oral view. D-F. Arm spines: from proximal portion of arm (D), middle portion of arm (E), distal portion of arm, an arc indicates lamina (F). Abbreviations: MO = muscle opening, NO = nerve opening, ST = secondary tooth." figureDoi="http://doi.org/10.5281/zenodo.3835030" httpUri="https://zenodo.org/record/3835030/files/figure.png" pageId="8" pageNumber="9">Fig. 5E</figureCitation>
|
||
), hook-shaped,
|
||
<emphasis id="B94C103FD90CF77AB1FC7F60FAB70D5D" box="[1267,1295,816,841]" italics="true" pageId="8" pageNumber="9">ca</emphasis>
|
||
.
|
||
<quantity id="4CC061C8D90CF77AB0137F60FACF0D5E" box="[1308,1399,816,842]" metricMagnitude="-1" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="9" unit="mm" value="230.0">230 µm</quantity>
|
||
long on distal portion of arms, with one secondary tooth and smooth lamina on proximal portion (
|
||
<figureCitation id="1303D0A8D90CF77AB0487F03FF580D85" captionStart="Fig" captionStartId="7.[189,232,1858,1884]" captionTargetBox="[192,1396,755,1814]" captionTargetId="figure@7.[189,1398,752,1814]" captionTargetPageId="7" captionText="Fig. 5. Asterostegus sabineae sp. nov., paratype (SMNH-Type-8536), SEM photographs. A. Lateral arm plate from middle portion of arm, abradial view. B. Lateral arm plate from distal portion of arm, adradial view. C. Lateral arm plate from distal portion of arm, oral view. D-F. Arm spines: from proximal portion of arm (D), middle portion of arm (E), distal portion of arm, an arc indicates lamina (F). Abbreviations: MO = muscle opening, NO = nerve opening, ST = secondary tooth." figureDoi="http://doi.org/10.5281/zenodo.3835030" httpUri="https://zenodo.org/record/3835030/files/figure.png" pageId="8" pageNumber="9">Fig. 5F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3229FA6D90CF77AB5B27FE7FB380A15" box="[189,1152,951,1025]" pageId="8" pageNumber="9" type="distribution">
|
||
<paragraph id="8B87CC2DD90CF77AB5B27FE7FEEE0DC5" blockId="8.[189,342,951,977]" box="[189,342,951,977]" pageId="8" pageNumber="9">
|
||
<heading id="D0CF7B41D90CF77AB5B27FE7FEEE0DC5" bold="true" box="[189,342,951,977]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
|
||
<emphasis id="B94C103FD90CF77AB5B27FE7FEEE0DC5" bold="true" box="[189,342,951,977]" pageId="8" pageNumber="9">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B87CC2DD90CF77AB5B27FB6FB380A15" blockId="8.[189,1152,998,1025]" box="[189,1152,998,1025]" pageId="8" pageNumber="9">
|
||
Only known from off
|
||
<collectingCountry id="F32F8CBDD90CF77AB4CF7FB6FDE80A14" box="[448,592,998,1024]" name="Madagascar" pageId="8" pageNumber="9">Madagascar</collectingCountry>
|
||
,
|
||
<quantity id="4CC061C8D90CF77AB7547FB6FD630A15" box="[603,731,998,1025]" metricMagnitude="2" metricUnit="m" metricValue="4.41" metricValueMax="4.7299999999999995" metricValueMin="4.09" pageId="8" pageNumber="9" unit="m" value="441.0" valueMax="473.0" valueMin="409.0">409-473 m</quantity>
|
||
depth (
|
||
<typeStatus id="5483728FD90CF77AB63C7FB6FCDF0A14" box="[819,871,998,1024]" pageId="8" pageNumber="9">type</typeStatus>
|
||
locality, present study).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3229FA6D90CF77AB5B27878FD470935" pageId="8" pageNumber="9" type="discussion">
|
||
<paragraph id="8B87CC2DD90CF77AB5B27878FE970A56" blockId="8.[189,303,1064,1090]" box="[189,303,1064,1090]" pageId="8" pageNumber="9">
|
||
<heading id="D0CF7B41D90CF77AB5B27878FE970A56" bold="true" box="[189,303,1064,1090]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
|
||
<emphasis id="B94C103FD90CF77AB5B27878FE970A56" bold="true" box="[189,303,1064,1090]" pageId="8" pageNumber="9">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B87CC2DD90CF77AB5B27806FAA80AA3" blockId="8.[189,1398,1110,1207]" pageId="8" pageNumber="9">
|
||
The new species falls within
|
||
<taxonomicName id="4C38B7AED90CF77AB7177806FD150A64" authorityName="Mortensen" authorityYear="1933" box="[536,685,1110,1136]" class="Ophiuroidea" family="Euryalidae" genus="Asterostegus" kingdom="Animalia" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B94C103FD90CF77AB7177806FD150A64" box="[536,685,1110,1136]" italics="true" pageId="8" pageNumber="9">Asterostegus</emphasis>
|
||
</taxonomicName>
|
||
by virtue of having simple arms, tubercles on radial shields and aboral arms, oral interradial plates, triangular teeth and granule-shaped oral papillae, an oral bridge on oral side of vertebrae, and smooth lamina of distal arm spines (
|
||
<bibRefCitation id="EFA9B1DCD90CF77AB6CB78CCFB800AA2" author="Fell H. B." box="[964,1080,1180,1206]" pageId="8" pageNumber="9" pagination="1 - 44" refId="ref6365" refString="Fell H. B. 1960. Synoptic keys to the genera of Ophiuroidea. Zoology Publications from Victoria University of Wellington 26: 1 - 44." type="journal article" year="1960">Fell 1960</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA9B1DCD90CF77AB14B78CCFABC0AA3" author="McKnight D. G." box="[1092,1284,1180,1207]" pageId="8" pageNumber="9" pagination="385 - 389" refId="ref6488" refString="McKnight D. G. 2003. A new species of Asterostegus (Echinodermata: Ophiuroidea) from the Cook Islands, South Pacific Ocean. Species Diversity 8: 385 - 389." type="journal article" year="2003">McKnight 2003</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B87CC2DD90CF77AB5B2788EFE750BFA" blockId="8.[189,1399,1246,1518]" pageId="8" pageNumber="9">
|
||
<taxonomicName id="4C38B7AED90CF77AB5B2788EFE7A0AEC" authority="Okanishi & Fujita, 2014" authorityName="Okanishi & Fujita" authorityYear="2014" box="[189,450,1246,1272]" class="Ophiuroidea" family="Euryalidae" genus="Asterostegus" kingdom="Animalia" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="sabineae" status="sp. nov.">
|
||
<emphasis id="B94C103FD90CF77AB5B2788EFE7A0AEC" box="[189,450,1246,1272]" italics="true" pageId="8" pageNumber="9">Asterostegus sabineae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27FAD44D90CF77AB4C5788EFD9E0AEC" box="[458,550,1246,1272]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be distinguished from the other two species as follows: two to four tubercles on the radial shields of the present new species form a line on each radial shield (
|
||
<figureCitation id="1303D0A8D90CF77AB0067951FAD10B0F" box="[1289,1385,1281,1307]" captionStart="Fig" captionStartId="4.[189,232,1787,1813]" captionTargetBox="[278,1305,265,1748]" captionTargetId="figure@4.[277,1305,265,1748]" captionTargetPageId="4" captionText="Fig. 2. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H). A. Aboral disc and proximal portion of arms. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc and proximal portion of arms. E. Jaws. F. Oral periphery of disc and proximal portion of arm. G. Lateral disc. H. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835024" httpUri="https://zenodo.org/record/3835024/files/figure.png" pageId="8" pageNumber="9">Fig. 2A</figureCitation>
|
||
), whereas the tubercles of
|
||
<taxonomicName id="4C38B7AED90CF77AB4EE7975FDFC0B2A" authorityName="McKnight" authorityYear="2003" box="[481,580,1317,1343]" class="Ophiuroidea" family="Euryalidae" genus="Asterostegus" kingdom="Animalia" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="maini">
|
||
<emphasis id="B94C103FD90CF77AB4EE7975FDFC0B2A" box="[481,580,1317,1343]" italics="true" pageId="8" pageNumber="9">A. maini</emphasis>
|
||
</taxonomicName>
|
||
number
|
||
<emphasis id="B94C103FD90CF77AB7A37975FD700B2A" box="[684,712,1317,1342]" italics="true" pageId="8" pageNumber="9">ca</emphasis>
|
||
. 10 and are scattered on the periphery of the radial shields only (
|
||
<figureCitation id="1303D0A8D90CF77AB40E7917FEE60B76" box="[257,350,1351,1378]" captionStart="Fig" captionStartId="10.[187,230,1824,1850]" captionTargetBox="[238,1340,266,1779]" captionTargetId="figure@10.[238,1341,263,1783]" captionTargetPageId="10" captionText="Fig. 6. Asterostegus maini McKnight, 2003 (NIWA 7564, H-733). A. Aboral view. B. Oral view. C. Aboral periphery of disc. D. Aboral central disc. E. Jaws. F. Oral periphery of disc. G. Lateral disc. H. Aboral proximal portion of arm. I. Aboral middle portion of arm. J. Oral proximal portion of arm. K. Lateral middle portion of arm. Arrows indicate positions where oral interradial plates should exist (F, G). Abbreviations: ADS = adoral shield, AS = arm spine, GS = genital slit, LAP = lateral arm plate, Te = teeth, Ten = tentacle, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835032" httpUri="https://zenodo.org/record/3835032/files/figure.png" pageId="8" pageNumber="9">Fig. 6C</figureCitation>
|
||
). Those of
|
||
<taxonomicName id="4C38B7AED90CF77AB4EC7918FD2D0B75" authorityName="Mortensen" authorityYear="1933" box="[483,661,1351,1378]" class="Ophiuroidea" family="Euryalidae" genus="Asterostegus" kingdom="Animalia" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="tuberculatus">
|
||
<emphasis id="B94C103FD90CF77AB4EC7918FD2D0B75" box="[483,661,1351,1378]" italics="true" pageId="8" pageNumber="9">A. tuberculatus</emphasis>
|
||
</taxonomicName>
|
||
number 5-15 and are scattered over the entire radial shields as well as on the interradial areas and periphery of the aboral disc (
|
||
<figureCitation id="1303D0A8D90CF77AB6B2793BFBA50B91" box="[957,1053,1387,1413]" captionStart="Fig" captionStartId="12.[189,232,1790,1816]" captionTargetBox="[254,1322,268,1748]" captionTargetId="figure@12.[249,1324,265,1751]" captionTargetPageId="12" captionText="Fig. 7. Asterostegus tuberculatus Mortensen, 1933 (SMNH 123461). A. Aboral disc and proximal portion of arm. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc. E. Jaws. F. Oral periphery of disc. G. Oral periphery of disc, skin removed to observe internal ossicles, oral interradial plates of the same row are connected by black bars. H. Lateral disc, an arrow indicates madreporite. I. Aboral proximal portion of arm. Abbreviations: ADS = adoral shield, GEO = granule-shaped external ossicle, GS = genital slit, LB = lateral bar, OIP = oral interradial plate, OP = oral papillae, Te = teeth, Ten = tentacle, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835034" httpUri="https://zenodo.org/record/3835034/files/figure.png" pageId="8" pageNumber="9">Fig. 7A</figureCitation>
|
||
). The size of tubercles of the new species on the periphery of the radial shield is
|
||
<emphasis id="B94C103FD90CF77AB61579DFFC8E0BBC" box="[794,822,1423,1448]" italics="true" pageId="8" pageNumber="9">ca</emphasis>
|
||
. 1.0-
|
||
<quantity id="4CC061C8D90CF77AB67C79DEFC760BBC" box="[883,974,1422,1448]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="9" unit="mm" value="1.2">1.2 mm</quantity>
|
||
long (
|
||
<figureCitation id="1303D0A8D90CF77AB11579DEFBCF0BBC" box="[1050,1143,1422,1448]" captionStart="Fig" captionStartId="4.[189,232,1787,1813]" captionTargetBox="[278,1305,265,1748]" captionTargetId="figure@4.[277,1305,265,1748]" captionTargetPageId="4" captionText="Fig. 2. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H). A. Aboral disc and proximal portion of arms. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc and proximal portion of arms. E. Jaws. F. Oral periphery of disc and proximal portion of arm. G. Lateral disc. H. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835024" httpUri="https://zenodo.org/record/3835024/files/figure.png" pageId="8" pageNumber="9">Fig. 2B</figureCitation>
|
||
). In contrast, those of
|
||
<taxonomicName id="4C38B7AED90CF77AB5B279E2FE9A0BDF" authorityName="McKnight" authorityYear="2003" box="[189,290,1458,1484]" class="Ophiuroidea" family="Euryalidae" genus="Asterostegus" kingdom="Animalia" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="maini">
|
||
<emphasis id="B94C103FD90CF77AB5B279E2FE9A0BDF" box="[189,290,1458,1484]" italics="true" pageId="8" pageNumber="9">A. maini</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C38B7AED90CF77AB45179E2FDAA0BDF" authorityName="Mortensen" authorityYear="1933" box="[350,530,1457,1484]" class="Ophiuroidea" family="Euryalidae" genus="Asterostegus" kingdom="Animalia" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="tuberculatus">
|
||
<emphasis id="B94C103FD90CF77AB45179E2FDAA0BDF" box="[350,530,1457,1484]" italics="true" pageId="8" pageNumber="9">A. tuberculatus</emphasis>
|
||
</taxonomicName>
|
||
in the same position are
|
||
<emphasis id="B94C103FD90CF77AB64F79E2FCE40BDF" box="[832,860,1458,1483]" italics="true" pageId="8" pageNumber="9">ca</emphasis>
|
||
.
|
||
<quantity id="4CC061C8D90CF77AB66379E1FC470BD8" box="[876,1023,1457,1484]" metricMagnitude="-1" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="8" pageNumber="9" unit="mm" value="275.0" valueMax="300.0" valueMin="250.0">250-300 µm</quantity>
|
||
(
|
||
<figureCitation id="1303D0A8D90CF77AB11F79E1FBD40BD8" box="[1040,1132,1457,1484]" captionStart="Fig" captionStartId="10.[187,230,1824,1850]" captionTargetBox="[238,1340,266,1779]" captionTargetId="figure@10.[238,1341,263,1783]" captionTargetPageId="10" captionText="Fig. 6. Asterostegus maini McKnight, 2003 (NIWA 7564, H-733). A. Aboral view. B. Oral view. C. Aboral periphery of disc. D. Aboral central disc. E. Jaws. F. Oral periphery of disc. G. Lateral disc. H. Aboral proximal portion of arm. I. Aboral middle portion of arm. J. Oral proximal portion of arm. K. Lateral middle portion of arm. Arrows indicate positions where oral interradial plates should exist (F, G). Abbreviations: ADS = adoral shield, AS = arm spine, GS = genital slit, LAP = lateral arm plate, Te = teeth, Ten = tentacle, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835032" httpUri="https://zenodo.org/record/3835032/files/figure.png" pageId="8" pageNumber="9">Fig. 6C</figureCitation>
|
||
) and
|
||
<emphasis id="B94C103FD90CF77AB1BE79E2FB750BDF" box="[1201,1229,1458,1483]" italics="true" pageId="8" pageNumber="9">ca</emphasis>
|
||
.
|
||
<quantity id="4CC061C8D90CF77AB1D279E1FA820BD8" box="[1245,1338,1457,1484]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="9" unit="mm" value="350.0">350 µm</quantity>
|
||
long (
|
||
<figureCitation id="1303D0A8D90CF77AB5C97984FE9C0BFA" box="[198,292,1492,1518]" captionStart="Fig" captionStartId="12.[189,232,1790,1816]" captionTargetBox="[254,1322,268,1748]" captionTargetId="figure@12.[249,1324,265,1751]" captionTargetPageId="12" captionText="Fig. 7. Asterostegus tuberculatus Mortensen, 1933 (SMNH 123461). A. Aboral disc and proximal portion of arm. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc. E. Jaws. F. Oral periphery of disc. G. Oral periphery of disc, skin removed to observe internal ossicles, oral interradial plates of the same row are connected by black bars. H. Lateral disc, an arrow indicates madreporite. I. Aboral proximal portion of arm. Abbreviations: ADS = adoral shield, GEO = granule-shaped external ossicle, GS = genital slit, LB = lateral bar, OIP = oral interradial plate, OP = oral papillae, Te = teeth, Ten = tentacle, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835034" httpUri="https://zenodo.org/record/3835034/files/figure.png" pageId="8" pageNumber="9">Fig. 7A</figureCitation>
|
||
), respectively.
|
||
</paragraph>
|
||
<paragraph id="8B87CC2DD90CF77AB5B27A46FEE90862" blockId="8.[189,1399,1558,1654]" pageId="8" pageNumber="9">
|
||
The new species has one or two tubercles on the top of each lateral bar in the arms proximally (
|
||
<figureCitation id="1303D0A8D90CF77AB0487A46FF5C0847" captionStart="Fig" captionStartId="4.[189,232,1787,1813]" captionTargetBox="[278,1305,265,1748]" captionTargetId="figure@4.[277,1305,265,1748]" captionTargetPageId="4" captionText="Fig. 2. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) and one paratype (SMNH- Type-8536) (H). A. Aboral disc and proximal portion of arms. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc and proximal portion of arms. E. Jaws. F. Oral periphery of disc and proximal portion of arm. G. Lateral disc. H. Lateral disc, skin removed to observe internal ossicles, an arrow indicates the madreporite; aboral proximal portion of arm. Abbreviations: ADS = adoral shield, AS = Arm spine, GEO = granule-shaped external ossicles, Go = gonads, GS= genital slit, OIP= oral interradial plate, Te = teeth, TS = tentacle sheath, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835024" httpUri="https://zenodo.org/record/3835024/files/figure.png" pageId="8" pageNumber="9">Fig. 2H</figureCitation>
|
||
), while
|
||
<taxonomicName id="4C38B7AED90CF77AB44F7A69FE1B0847" authorityName="McKnight" authorityYear="2003" box="[320,419,1593,1619]" class="Ophiuroidea" family="Euryalidae" genus="Asterostegus" kingdom="Animalia" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="maini">
|
||
<emphasis id="B94C103FD90CF77AB44F7A69FE1B0847" box="[320,419,1593,1619]" italics="true" pageId="8" pageNumber="9">A. maini</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C38B7AED90CF77AB4D57A69FD340847" authorityName="Mortensen" authorityYear="1933" box="[474,652,1593,1619]" class="Ophiuroidea" family="Euryalidae" genus="Asterostegus" kingdom="Animalia" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="tuberculatus">
|
||
<emphasis id="B94C103FD90CF77AB4D57A69FD340847" box="[474,652,1593,1619]" italics="true" pageId="8" pageNumber="9">A. tuberculatus</emphasis>
|
||
</taxonomicName>
|
||
have three to five (
|
||
<figureCitation id="1303D0A8D90CF77AB6657A69FC7F0840" box="[874,967,1593,1620]" captionStart="Fig" captionStartId="10.[187,230,1824,1850]" captionTargetBox="[238,1340,266,1779]" captionTargetId="figure@10.[238,1341,263,1783]" captionTargetPageId="10" captionText="Fig. 6. Asterostegus maini McKnight, 2003 (NIWA 7564, H-733). A. Aboral view. B. Oral view. C. Aboral periphery of disc. D. Aboral central disc. E. Jaws. F. Oral periphery of disc. G. Lateral disc. H. Aboral proximal portion of arm. I. Aboral middle portion of arm. J. Oral proximal portion of arm. K. Lateral middle portion of arm. Arrows indicate positions where oral interradial plates should exist (F, G). Abbreviations: ADS = adoral shield, AS = arm spine, GS = genital slit, LAP = lateral arm plate, Te = teeth, Ten = tentacle, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835032" httpUri="https://zenodo.org/record/3835032/files/figure.png" pageId="8" pageNumber="9">Fig. 6H</figureCitation>
|
||
), and two to three tubercles (
|
||
<figureCitation id="1303D0A8D90CF77AB0187A69FAD00847" box="[1303,1384,1593,1619]" captionStart="Fig" captionStartId="12.[189,232,1790,1816]" captionTargetBox="[254,1322,268,1748]" captionTargetId="figure@12.[249,1324,265,1751]" captionTargetPageId="12" captionText="Fig. 7. Asterostegus tuberculatus Mortensen, 1933 (SMNH 123461). A. Aboral disc and proximal portion of arm. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc. E. Jaws. F. Oral periphery of disc. G. Oral periphery of disc, skin removed to observe internal ossicles, oral interradial plates of the same row are connected by black bars. H. Lateral disc, an arrow indicates madreporite. I. Aboral proximal portion of arm. Abbreviations: ADS = adoral shield, GEO = granule-shaped external ossicle, GS = genital slit, LB = lateral bar, OIP = oral interradial plate, OP = oral papillae, Te = teeth, Ten = tentacle, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835034" httpUri="https://zenodo.org/record/3835034/files/figure.png" pageId="8" pageNumber="9">Fig. 7I</figureCitation>
|
||
), respectively.
|
||
</paragraph>
|
||
<paragraph id="8B87CC2DD90CF77AB5B27ACDFD470935" blockId="8.[189,1399,1693,1826]" pageId="8" pageNumber="9">
|
||
The number of oral interradial plates of the new species is one on each interradius (
|
||
<figureCitation id="1303D0A8D90CF77AB1C87ACDFA9908AC" box="[1223,1313,1693,1720]" captionStart="Fig" captionStartId="5.[189,232,1892,1918]" captionTargetBox="[188,1397,406,1852]" captionTargetId="figure@5.[188,1399,404,1852]" captionTargetPageId="5" captionText="Fig. 3. Asterostegus sabineae sp. nov., holotype (MNHN IE-2013-8012) (A–F). A. Aboral basal portion of arm. B. Aboral middle portion of arm. C. Aboral and lateral distal portion of arm. D. Oral proximal portion of arm. E. Oral middle portion of arm. F. Lateral distal portion of arm. Abbreviations: AS = arm spine, LAP = lateral arm plate, Ten = tentacles, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835026" httpUri="https://zenodo.org/record/3835026/files/figure.png" pageId="8" pageNumber="9">Fig. 3F</figureCitation>
|
||
) while there are three to five plates in
|
||
<taxonomicName id="4C38B7AED90CF77AB7317A91FC3808CF" authority="(McKnight 2003)" baseAuthorityName="McKnight" baseAuthorityYear="2003" box="[574,896,1729,1755]" class="Ophiuroidea" family="Euryalidae" genus="Asterostegus" kingdom="Animalia" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="maini">
|
||
<emphasis id="B94C103FD90CF77AB7317A91FD1D08CF" box="[574,677,1729,1755]" italics="true" pageId="8" pageNumber="9">A. maini</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA9B1DCD90CF77AB7B87A91FCC008CF" author="McKnight D. G." box="[695,888,1729,1755]" pageId="8" pageNumber="9" pagination="385 - 389" refId="ref6488" refString="McKnight D. G. 2003. A new species of Asterostegus (Echinodermata: Ophiuroidea) from the Cook Islands, South Pacific Ocean. Species Diversity 8: 385 - 389." type="journal article" year="2003">McKnight 2003</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and five to eight in
|
||
<taxonomicName id="4C38B7AED90CF77AB1717A91FA8C08CF" authorityName="Mortensen" authorityYear="1933" box="[1150,1332,1729,1755]" class="Ophiuroidea" family="Euryalidae" genus="Asterostegus" kingdom="Animalia" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="tuberculatus">
|
||
<emphasis id="B94C103FD90CF77AB1717A91FA8C08CF" box="[1150,1332,1729,1755]" italics="true" pageId="8" pageNumber="9">A. tuberculatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1303D0A8D90CF77AB0487A91FF5A08EA" captionStart="Fig" captionStartId="12.[189,232,1790,1816]" captionTargetBox="[254,1322,268,1748]" captionTargetId="figure@12.[249,1324,265,1751]" captionTargetPageId="12" captionText="Fig. 7. Asterostegus tuberculatus Mortensen, 1933 (SMNH 123461). A. Aboral disc and proximal portion of arm. B. Aboral periphery of disc. C. Aboral central disc. D. Oral disc. E. Jaws. F. Oral periphery of disc. G. Oral periphery of disc, skin removed to observe internal ossicles, oral interradial plates of the same row are connected by black bars. H. Lateral disc, an arrow indicates madreporite. I. Aboral proximal portion of arm. Abbreviations: ADS = adoral shield, GEO = granule-shaped external ossicle, GS = genital slit, LB = lateral bar, OIP = oral interradial plate, OP = oral papillae, Te = teeth, Ten = tentacle, Tu = tubercle." figureDoi="http://doi.org/10.5281/zenodo.3835034" httpUri="https://zenodo.org/record/3835034/files/figure.png" pageId="8" pageNumber="9">Fig. 7G</figureCitation>
|
||
) (
|
||
<bibRefCitation id="EFA9B1DCD90CF77AB5F27AB4FE6D08EB" author="Mortensen T." box="[253,469,1764,1791]" pageId="8" pageNumber="9" pagination="215 - 400" refId="ref6544" refString="Mortensen T. 1933 b. Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916. LXV. Echinoderms of South Africa (Asteroidea and Ophiuroidea). Meddelelser fra Dansk naturhistorisk Forening 93: 215 - 400." type="journal article" year="1933">Mortensen 1933b</bibRefCitation>
|
||
). The oral interradial plates of the latter two species form rows (see also the description of
|
||
<taxonomicName id="4C38B7AED90CF77AB4697B58FDA10935" authorityName="Mortensen" authorityYear="1933" box="[358,537,1799,1826]" class="Ophiuroidea" family="Euryalidae" genus="Asterostegus" kingdom="Animalia" order="Euryalida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="tuberculatus">
|
||
<emphasis id="B94C103FD90CF77AB4697B58FDA10935" box="[358,537,1799,1826]" italics="true" pageId="8" pageNumber="9">A. tuberculatus</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="EFA9B1DCD90CF77AB73E7B57FD4B0935" author="McKnight D. G." box="[561,755,1799,1825]" pageId="8" pageNumber="9" pagination="385 - 389" refId="ref6488" refString="McKnight D. G. 2003. A new species of Asterostegus (Echinodermata: Ophiuroidea) from the Cook Islands, South Pacific Ocean. Species Diversity 8: 385 - 389." type="journal article" year="2003">McKnight 2003</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |