852 lines
107 KiB
XML
852 lines
107 KiB
XML
<document id="6B65A1F411B3890AE0B1F51A0957F1BE" ID="10.11646/zootaxa.4250.4.2SLASH10731" ID-CLB-Dataset="33291" ID-DOI="10.11646/zootaxa.4250.4.2" ID-GBIF-Dataset="ce4e62c5-1d2a-478e-a0f3-1d2637e76f8e" ID-ISSN="1175-5326" ID-Zenodo-Dep="495277" ID-ZooBank="CE45B4DB-641B-4D37-AD35-93CCB384BE1D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1491462921247" checkinUser="plazi" docAuthor="Kompier, Tom" docDate="2017" docId="03FAA703ED3C463AFF5FF747A0BCFF2F" docLanguage="en" docName="zootaxa.4250.4.2SLASH10731.pdf" docOrigin="Zootaxa 4250 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lyriothemis bivittata Rambur 1842" docType="treatment" docVersion="9" lastPageNumber="325" masterDocId="FFC3DF7BED394631FFC8F307A445FFFA" masterDocTitle="The riddle of Lyriothemis bivittata (Rambur, 1842): Lyriothemis kameliyae spec. nov. (Odonata: Libellulidae)" masterLastPageNumber="326" masterPageNumber="315" pageNumber="320" updateTime="1698438726893" updateUser="ExternalLinkService">
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<mods:mods id="37A05CEEA71E7DD4A0CC929191958E1B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="97EC845647C004603CDFEEBFB4465687">
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<mods:title id="E89713526301E3B0C59F0DAD60EFDD16">The riddle of Lyriothemis bivittata (Rambur, 1842): Lyriothemis kameliyae spec. nov. (Odonata: Libellulidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="8F20A7309ABAC5D71A7CB1401A462F53" type="personal">
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<mods:roleTerm id="C1DC45D644649365060F3522DD3664C0">Author</mods:roleTerm>
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<mods:namePart id="776E1CC10572DA13425C29F6CE682F03">Kompier, Tom</mods:namePart>
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<mods:typeOfResource id="12BB16C335A4F78F663DF8E75601F3DD">text</mods:typeOfResource>
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<mods:title id="F0A3D95C0E535B42E619BDBB49F7EFBC">Zootaxa</mods:title>
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<mods:part id="04E303F29F21A5A6ECA75DE8033750E5">
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<mods:date id="6453FA2252AEFEC1764311F42480E80E">2017</mods:date>
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<mods:number id="EAA85332BAF188B2F1D92A0053E16DF2">4250</mods:number>
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<mods:detail id="FD0935E3DC3C58C0A0363FFCAFD624F7" type="issue">
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<mods:number id="83A1AA529B35B4EFBB756F364488C860">4</mods:number>
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<mods:end id="5114F067345DD988DC09182B892D2E1F">326</mods:end>
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<mods:identifier id="591429831A7B4088089273DD73C14EE4" type="DOI">10.11646/zootaxa.4250.4.2</mods:identifier>
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</mods:mods>
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<treatment id="03FAA703ED3C463AFF5FF747A0BCFF2F" ID-DOI="http://doi.org/10.5281/zenodo.5689900" ID-GBIF-Taxon="128343990" ID-Zenodo-Dep="5689900" LSID="urn:lsid:plazi:treatment:03FAA703ED3C463AFF5FF747A0BCFF2F" httpUri="http://treatment.plazi.org/id/03FAA703ED3C463AFF5FF747A0BCFF2F" lastPageId="11" lastPageNumber="325" pageId="5" pageNumber="320">
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<subSubSection id="C349459EED3C4634FF5FF747A632FBA0" box="[151,631,1087,1114]" pageId="5" pageNumber="320" type="nomenclature">
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<paragraph id="8BEC1615ED3C4634FF5FF747A632FBA0" blockId="5.[151,631,1087,1147]" box="[151,631,1087,1114]" pageId="5" pageNumber="320">
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<heading id="D0A4A179ED3C4634FF5FF747A632FBA0" bold="true" box="[151,631,1087,1114]" fontSize="11" level="1" pageId="5" pageNumber="320" reason="1">
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<emphasis id="B927CA07ED3C4634FF5FF747A632FBA0" bold="true" box="[151,631,1087,1114]" pageId="5" pageNumber="320">
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2.
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<taxonomicName id="4C536D96ED3C4634FF7CF738A632FBA0" ID-CoL="3WTKV" authority="Rambur, 1842" authorityName="Rambur" authorityYear="1842" box="[180,631,1087,1114]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="320" phylum="Arthropoda" rank="species" species="bivittata">
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<emphasis id="B927CA07ED3C4634FF7CF738A5E9FBA3" bold="true" box="[180,428,1087,1113]" italics="true" pageId="5" pageNumber="320">Lyriothemis bivittata</emphasis>
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(
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<bibRefCitation id="EFC26BE4ED3C4634FE73F747A62AFBA0" author="Rambur" box="[443,623,1088,1114]" pageId="5" pageNumber="320" refString="Rambur, M. P. (1842) Histoire naturelle des insectes. Nevropteres. Libraire Encycl. de Roret, Paris, xvii + 534 pp." type="book" year="1842">Rambur, 1842</bibRefCitation>
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)
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C349459EED3C4637FF5FF765A01FFB87" lastPageId="6" lastPageNumber="321" pageId="5" pageNumber="320" type="description">
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<paragraph id="8BEC1615ED3C4634FF5FF765A601FB81" blockId="5.[151,631,1087,1147]" box="[151,580,1122,1147]" pageId="5" pageNumber="320">
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(
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<figureCitation id="13680A90ED3C4634FF57F765A4ADFB81" box="[159,232,1122,1147]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="5" pageNumber="320">Figs. 1</figureCitation>
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A, 1D, 2A, 2D, 4C, 5B, 6A, 7)
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</paragraph>
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<paragraph id="8BEC1615ED3C4634FF5FF7ADA700FAF1" blockId="5.[151,1436,1194,2011]" pageId="5" pageNumber="320">
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<emphasis id="B927CA07ED3C4634FF5FF7ADA610FB39" bold="true" box="[151,597,1194,1219]" pageId="5" pageNumber="320">Specimens used for description. Male</emphasis>
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: Huu Lien Nature Reserve,
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<collectingRegion id="4997D8F7ED3C4634FC5CF7ACA02BFB39" box="[916,1134,1194,1219]" country="Vietnam" name="Lang Son" pageId="5" pageNumber="320">
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Lang
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<collectingRegion id="4997D8F7ED3C4634FC1DF7ACA02BFB39" box="[981,1134,1194,1219]" inferredBySuffix="true" pageId="5" pageNumber="320">Son Province</collectingRegion>
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</collectingRegion>
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, northern
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<collectingCountry id="F3445685ED3C4634FB17F7ADA17BFB39" box="[1247,1342,1194,1219]" name="Vietnam" pageId="5" pageNumber="320">Vietnam</collectingCountry>
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(
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<geoCoordinate id="EE6770D2ED3C4634FA85F7ADA4EBFB1C" direction="north" orientation="latitude" pageId="5" pageNumber="320" precision="55" value="21.662">21.662 N</geoCoordinate>
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,
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<geoCoordinate id="EE6770D2ED3C4634FF76F7C9A57CFB1D" box="[190,313,1230,1255]" direction="east" orientation="longitude" pageId="5" pageNumber="320" precision="55" value="106.373">106.373 E</geoCoordinate>
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),
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<date id="FFED30D5ED3C4634FE9BF7C9A59BFB1D" box="[339,478,1230,1255]" pageId="5" pageNumber="320" value="2016-07-23">23.VII.2016</date>
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, T. Kompier leg;
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<emphasis id="B927CA07ED3C4634FD7FF7C9A74BFB1D" bold="true" box="[695,782,1230,1255]" pageId="5" pageNumber="320">Female</emphasis>
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: Xuan Son National Park,
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<collectingRegion id="4997D8F7ED3C4634FB9EF7C9A168FB1D" box="[1110,1325,1230,1255]" country="Vietnam" name="Phu Tho" pageId="5" pageNumber="320">
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Phu
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<collectingRegion id="4997D8F7ED3C4634FB45F7C9A168FB1D" box="[1165,1325,1230,1255]" inferredBySuffix="true" pageId="5" pageNumber="320">Tho Province</collectingRegion>
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</collectingRegion>
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, northern
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<collectingCountry id="F3445685ED3C4634FF5FF7F5A4B3FAF1" box="[151,246,1266,1291]" name="Vietnam" pageId="5" pageNumber="320">Vietnam</collectingCountry>
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(
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<geoCoordinate id="EE6770D2ED3C4634FECCF7F4A535FAF0" box="[260,368,1267,1291]" direction="north" orientation="latitude" pageId="5" pageNumber="320" precision="55" value="21.11">21.110 N</geoCoordinate>
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,
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<geoCoordinate id="EE6770D2ED3C4634FEB3F7F4A5B4FAF1" box="[379,497,1267,1291]" direction="east" orientation="longitude" pageId="5" pageNumber="320" precision="55" value="104.994">104.994 E</geoCoordinate>
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),
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<date id="FFED30D5ED3C4634FDCEF7F5A6D5FAF1" box="[518,656,1266,1291]" pageId="5" pageNumber="320" value="2016-07-31">31.VII.2016</date>
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, same collector.
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</paragraph>
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<paragraph id="8BEC1615ED3C4634FF0FF611A5E9FA61" blockId="5.[151,1436,1194,2011]" pageId="5" pageNumber="320">
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<emphasis id="B927CA07ED3C4634FF0FF611A5FBFAD5" bold="true" box="[199,446,1302,1327]" pageId="5" pageNumber="320">Description of male</emphasis>
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(
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<figureCitation id="13680A90ED3C4634FE1CF611A661FAD5" box="[468,548,1302,1327]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="5" pageNumber="320">Figs. 1</figureCitation>
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A, 5B).
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<emphasis id="B927CA07ED3C4634FD59F611A68BFAD4" box="[657,718,1302,1326]" italics="true" pageId="5" pageNumber="320">Head</emphasis>
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. Labium yellowish white, with brown inner edge. Labrum, anteclypeus, postclypeus and antefrons white, postfrons metallic dark green, corners somewhat rounded. Vertex metallic green, with two small tubercles, antennae black, occiput orange-brown. Compound eyes blackish grey, chestnut-brown dorsally.
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</paragraph>
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<paragraph id="8BEC1615ED3C4634FF0FF6A1A009FA45" blockId="5.[151,1436,1194,2011]" box="[199,1100,1446,1471]" pageId="5" pageNumber="320">
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<emphasis id="B927CA07ED3C4634FF0FF6A1A57CFA44" box="[199,313,1446,1470]" italics="true" pageId="5" pageNumber="320">Prothorax</emphasis>
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. Black, but center of posterior lobe pale brown and anterior lobe pale.
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</paragraph>
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<paragraph id="8BEC1615ED3C4634FF0FF6CCA490F8B1" blockId="5.[151,1436,1194,2011]" pageId="5" pageNumber="320">
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<emphasis id="B927CA07ED3C4634FF0FF6CCA505FA19" box="[199,320,1483,1507]" italics="true" pageId="5" pageNumber="320">Synthorax.</emphasis>
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(
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<figureCitation id="13680A90ED3C4634FE87F6CDA5CBFA19" box="[335,398,1482,1507]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="5" pageNumber="320">Fig. 1</figureCitation>
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A) Blackish brown with markings as follows: dorsal side uniform warm brown, but close to antehumeral suture blackish brown like sides. Mesepimeron with an oblong yellow spot on ventral half, metepimeron almost completely yellow.
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<emphasis id="B927CA07ED3C4634FDB0F514A6E8F9D0" box="[632,685,1555,1578]" italics="true" pageId="5" pageNumber="320">Legs</emphasis>
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black. Femora with short spines growing progressively longer distally, tibiae with longer black spines.
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<emphasis id="B927CA07ED3C4634FD96F530A6E4F9B4" box="[606,673,1591,1614]" italics="true" pageId="5" pageNumber="320">Wings</emphasis>
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. (
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<figureCitation id="13680A90ED3C4634FD71F531A6BDF9B5" box="[697,760,1590,1615]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="5" pageNumber="320">Fig. 1</figureCitation>
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A) Slightly tinted amber, somewhat stronger at apices. Two brown streaks above and below median space in both fore and hindwing as follows: First two cells between subcosta and radius blackish brown, third and fourth cell tinted brown and cubito-anal space blackish brown until first crossvein and brownish to second. Veins black. Membranule brown. Anal loop 19–21 celled. 2–3 bridge crossveins in all wings, last antenodal crossvein complete. Forewing with 2 cubito-anal crossveins, triangle 2- celled, subtriangle 3-celled. Hingwing with 2 cubito-anal crossveins, triangle 2-celled. Nodal index in forewing 9– 17:15–9 and in hindwing 10–12:13–10. Pterostigma black, covering 2.5-3 cells, slightly over 5 times longer than wide.
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</paragraph>
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<paragraph id="8BEC1615ED3C4637FF0FF451A7F4FEBA" blockId="5.[151,1436,1194,2011]" lastBlockId="6.[151,1437,151,1149]" lastPageId="6" lastPageNumber="321" pageId="5" pageNumber="320">
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<emphasis id="B927CA07ED3C4634FF0FF451A57DF894" box="[199,312,1878,1902]" italics="true" pageId="5" pageNumber="320">Abdomen.</emphasis>
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(
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<figureCitation id="13680A90ED3C4634FE82F451A5CEF895" box="[330,395,1878,1903]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="5" pageNumber="320">Fig. 1</figureCitation>
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A) Slightly broader than thorax in dorsal view, S3 broadest. S1 ochre-yellow. S2 dorsally bright red to somewhat above lateral carina, lower part orange, lateral carina thinly black. Hamulus (
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<figureCitation id="13680A90ED3C4634FB3CF47DA173F869" box="[1268,1334,1914,1939]" captionStart="FIGURE 5" captionStartId="6.[151,250,1662,1684]" captionTargetBox="[159,1412,1204,1644]" captionTargetId="figure@6.[159,1412,1204,1645]" captionTargetPageId="6" captionText="FIGURE 5. Hamuli in anteroventral view: (A) L. tricolor, Taiwan, courtesy Wen-Chi Yeh; (B) L. bivittata, Huu Lien NR, 23. VII. 2016; (C) L. kameliyae, paratype, Huu Lien NR, 24. V. 2014." httpUri="https://zenodo.org/record/495283/files/figure.png" pageId="5" pageNumber="320">Fig. 5</figureCitation>
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B) black, large, shaped like a rectangular dish, clearly depressed in center, with a backward-directed spine somewhat lateral of its inner posterior corner and a large curved incision slightly anterior of middle of inner edge. Orange genital lobe very slender, pointed and nearly 4 times as long as wide. S3 dorsally bright red to thinly black lateral carina, ventrally orange. S4–8 bright red dorsally, with almost imperceptibly black lateral, dorsal and posterior carina, ventrally orange. S9 red dorsally and orange ventrally, but with some darker blotches and black carinae. S10 black but dorsally red.
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<emphasis id="B927CA07ED3F4637FE9CF203A650FEE6" box="[340,533,260,284]" italics="true" pageId="6" pageNumber="321">Anal appendages</emphasis>
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. Cerci dark brown, twice length of S10, extending just beyond epiproct, pointed, subapically with 4 small ventral teeth. Epiproct of normal
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<typeStatus id="54E8A8B7ED3F4637FCECF22FA712FEBA" box="[804,855,296,320]" pageId="6" pageNumber="321">type</typeStatus>
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, brown.
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</paragraph>
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<paragraph id="8BEC1615ED3F4637FF0FF24AA01FFE9F" blockId="6.[151,1437,151,1149]" box="[199,1114,332,357]" pageId="6" pageNumber="321">Measurements (mm): Abdomen incl. appendages 32, hindwing 36, pterostigma 5.</paragraph>
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<paragraph id="8BEC1615ED3F4637FF0FF268A7FEFE0E" blockId="6.[151,1437,151,1149]" pageId="6" pageNumber="321">
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<emphasis id="B927CA07ED3F4637FF0FF268A580FE72" bold="true" box="[199,453,367,392]" pageId="6" pageNumber="321">Description of female</emphasis>
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(
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<figureCitation id="13680A90ED3F4637FE1CF268A65BFE72" box="[468,542,367,392]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="6" pageNumber="321">Figs. 1</figureCitation>
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D, 2A, 4C, 6A).
|
||
<emphasis id="B927CA07ED3F4637FD13F277A752FE72" box="[731,791,368,392]" italics="true" pageId="6" pageNumber="321">Head</emphasis>
|
||
. Labium black, but anterolateral corners whitish. Labrum black. Anteclypeus brownish white, postclypeus blackish brown. Antefrons, postfrons and vertex metallic dark green, antefrons with whitish edges along compound eyes. Vertex with two small tubercles, antennae brown, occiput orange brown. Compound eyes grayish black with chestnut top.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED3F4637FF0FF107A57FFDC6" blockId="6.[151,1437,151,1149]" pageId="6" pageNumber="321">
|
||
<emphasis id="B927CA07ED3F4637FF0FF107A57CFDE2" box="[199,313,512,536]" italics="true" pageId="6" pageNumber="321">Prothorax</emphasis>
|
||
. Black, but anterior lobe with anterior edge pale yellow and middle and posterior lobe with brown spot on center.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED3F4637FF0FF14FA758FC32" blockId="6.[151,1437,151,1149]" pageId="6" pageNumber="321">
|
||
<emphasis id="B927CA07ED3F4637FF0FF14FA505FD9A" box="[199,320,584,608]" italics="true" pageId="6" pageNumber="321">Synthorax.</emphasis>
|
||
(
|
||
<figureCitation id="13680A90ED3F4637FE9AF140A5DAFD9A" box="[338,415,583,608]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="6" pageNumber="321">Figs. 1</figureCitation>
|
||
D, 4C) Blackish brown with markings as follows: dorsal side uniform warm brown, but close to antehumeral suture blackish brown like sides. Mesepimeron with an oblong yellow band, more brownyellow on its dorsal half, metepisternum more warm brown towards its dorsal edge, metepimeron almost completely yellow.
|
||
<emphasis id="B927CA07ED3F4637FEBCF1B2A5ECFD36" box="[372,425,693,716]" italics="true" pageId="6" pageNumber="321">Legs</emphasis>
|
||
black. Femora with short spines growing progressively longer distally, tibiae with longer black spines.
|
||
<emphasis id="B927CA07ED3F4637FEE6F1DFA534FD15" box="[302,369,728,751]" italics="true" pageId="6" pageNumber="321">Wings</emphasis>
|
||
. (
|
||
<figureCitation id="13680A90ED3F4637FE41F1D0A591FD0A" box="[393,468,727,752]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="6" pageNumber="321">Figs. 1</figureCitation>
|
||
D, 2A) Tinted amber, strongest at apices. Two brown streaks above and below median space in both fore- and hindwing as follows: first three cells between subcosta and radius blackish brown, fourth to sixth cell tinted brown, cubito-anal space blackish brown until first crossvein. Veins black. Membranule brown. Anal loop 33–34 celled. 5 bridge crossveins in forewings,
|
||
<quantity id="4CABBBF0ED3F4637FCFFF043A73AFCA7" box="[823,895,836,861]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="6" pageNumber="321" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">2–3 in</quantity>
|
||
hindwings. Last antenodal crossvein complete. Forewing with 1 cubito-anal crossvein, triangle 3-celled, subtriangle 5–6 celled. Hindwing with 2 cubito-anal crossveins, triangle 2-celled. Nodal index in forewing 11–17:19–12 and in hindwing 11–13:14–13. Pterostigma dark brown, covering 2–3 cells, 6 times longer than wide.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED3F4637FF0FF0D3A02AFBA2" blockId="6.[151,1437,151,1149]" pageId="6" pageNumber="321">
|
||
<emphasis id="B927CA07ED3F4637FF0FF0D3A57DFC16" box="[199,312,980,1004]" italics="true" pageId="6" pageNumber="321">Abdomen.</emphasis>
|
||
(
|
||
<figureCitation id="13680A90ED3F4637FE80F0D3A5D1FC16" box="[328,404,980,1005]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="6" pageNumber="321">Figs. 1</figureCitation>
|
||
D, 2A, 4C) S1 ochre yellow. S2–7 red dorsally and yellow-orange ventrally, carinae thinly blackish. S8 dorsally red, but carinae somewhat thicker black, and ventrally black. S9 dorsally with a red triangle on either side of a black line over dorsal carina, remainder black. S10 and cerci black, slightly longer than S10. S8 somewhat dilated.
|
||
<emphasis id="B927CA07ED3F4637FEA2F747A64CFBA2" box="[362,521,1088,1112]" italics="true" pageId="6" pageNumber="321">Vulvar lamina</emphasis>
|
||
(
|
||
<figureCitation id="13680A90ED3F4637FDD0F738A61FFBA2" box="[536,602,1087,1112]" captionStart="FIGURE 6" captionStartId="8.[151,250,737,759]" captionTargetBox="[165,1417,196,718]" captionTargetId="figure@8.[165,1417,196,718]" captionTargetPageId="8" captionText="FIGURE 6. Vulvar lamina: (A) L. bivittata, Xuan Son NP, 31. VII. 2016; (B) L. kameliyae allotype." httpUri="https://zenodo.org/record/495284/files/figure.png" pageId="6" pageNumber="321">Fig. 6</figureCitation>
|
||
A) short, slightly curled, only narrowly incised.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED3F4637FF0FF762A01FFB87" blockId="6.[151,1437,151,1149]" box="[199,1114,1124,1149]" pageId="6" pageNumber="321">Measurements (mm): Abdomen incl. appendages 34, hindwing 41, pterostigma 6.</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF2C469DED3F4637FF5FF579A77AF949" httpUri="https://zenodo.org/record/495283/files/figure.png" pageId="6" pageNumber="321" targetBox="[159,1412,1204,1644]" targetPageId="6">
|
||
<paragraph id="8BEC1615ED3F4637FF5FF579A77AF949" blockId="6.[151,1436,1662,1715]" pageId="6" pageNumber="321">
|
||
<emphasis id="B927CA07ED3F4637FF5FF579A550F96E" bold="true" box="[151,277,1662,1684]" pageId="6" pageNumber="321">FIGURE 5.</emphasis>
|
||
Hamuli in anteroventral view: (A)
|
||
<taxonomicName id="4C536D96ED3F4637FD44F587A6B2F96E" box="[652,759,1662,1685]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="tricolor">
|
||
<emphasis id="B927CA07ED3F4637FD44F587A6B2F96E" box="[652,759,1662,1685]" italics="true" pageId="6" pageNumber="321">L. tricolor</emphasis>
|
||
</taxonomicName>
|
||
, Taiwan, courtesy Wen-Chi Yeh; (B)
|
||
<taxonomicName id="4C536D96ED3F4637FB4DF587A12CF96E" authority="Huu Lien" authorityName="Huu Lien" box="[1157,1385,1662,1685]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED3F4637FB4DF587A0B2F96E" box="[1157,1271,1662,1685]" italics="true" pageId="6" pageNumber="321">L. bivittata</emphasis>
|
||
, Huu Lien
|
||
</taxonomicName>
|
||
NR, 23.VII.2016; (C)
|
||
<taxonomicName id="4C536D96ED3F4637FE80F599A58FF949" box="[328,458,1693,1715]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED3F4637FE80F599A58FF949" box="[328,458,1693,1715]" italics="true" pageId="6" pageNumber="321">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
, paratype, Huu Lien NR, 24.V.2014.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C349459EED3F4636FF0FF403A7B1FBA2" lastPageId="7" lastPageNumber="322" pageId="6" pageNumber="321" type="diagnosis">
|
||
<paragraph id="8BEC1615ED3F4636FF0FF403A77DFE72" blockId="6.[151,1437,1796,2037]" lastBlockId="7.[151,1437,151,1832]" lastPageId="7" lastPageNumber="322" pageId="6" pageNumber="321">
|
||
<emphasis id="B927CA07ED3F4637FF0FF403A504F8E7" bold="true" box="[199,321,1796,1821]" pageId="6" pageNumber="321">Diagnosis.</emphasis>
|
||
Van der Poorten (2008) provided an overview of all known
|
||
<taxonomicName id="4C536D96ED3F4637FBE9F403A0E3F8E6" box="[1057,1190,1796,1820]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="321" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B927CA07ED3F4637FBE9F403A0E3F8E6" box="[1057,1190,1796,1820]" italics="true" pageId="6" pageNumber="321">Lyriothemis</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
<taxonomicName id="4C536D96ED3F4637FADFF403A54CF8BA" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="kameliyae" status="sp. nov.">
|
||
<emphasis id="B927CA07ED3F4637FADFF403A54CF8BA" italics="true" pageId="6" pageNumber="321">Lyriothemis kameliyae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A214777CED3F4637FEC7F42EA53AF8BA" box="[271,383,1833,1856]" pageId="6" pageNumber="321" rank="species">spec. nov.</taxonomicNameLabel>
|
||
can be separated from almost all other
|
||
<taxonomicName id="4C536D96ED3F4637FCF2F42FA7FAF8BA" box="[826,959,1832,1856]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="321" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B927CA07ED3F4637FCF2F42FA7FAF8BA" box="[826,959,1832,1856]" italics="true" pageId="6" pageNumber="321">Lyriothemis</emphasis>
|
||
</taxonomicName>
|
||
species on the basis of the combination of red abdomen (in the male), dark streaks at the wing base, and thoracic pattern with two yellow lateral markings. These characters are shared with
|
||
<taxonomicName id="4C536D96ED3F4637FDD0F477A741F872" box="[536,772,1904,1928]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED3F4637FDD0F477A741F872" box="[536,772,1904,1928]" italics="true" pageId="6" pageNumber="321">Lyriothemis bivittata</emphasis>
|
||
</taxonomicName>
|
||
, but it can be separated from that species on the basis of many clear differences, as described below. There are two other similar species. The first occurs also in
|
||
<collectingCountry id="F3445685ED3F4637FAF5F493A1D9F857" box="[1341,1436,1940,1965]" name="Vietnam" pageId="6" pageNumber="321">Vietnam</collectingCountry>
|
||
and is most likely another undescribed species (
|
||
<figureCitation id="13680A90ED3F4637FD0CF4BFA750F82A" box="[708,789,1976,2001]" captionStart="FIGURE 2" captionStartId="3.[151,250,1818,1840]" captionTargetBox="[185,1417,665,1777]" captionTargetId="figure@3.[174,1417,665,1780]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal scans: (A) ♀ Lyriothemis bivittata, Xuan Son NP, 31. VII. 2016; (B) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (C) ♂ L. kameliyae, paratype, Huu Lien NR, 24. V. 2014. Photos in natural habitat: (D) ♀ L. bivittata (not collected), Tam Dao NP, 18. VIII. 2013; (E) ♀ L. sp., Quang Binh Province, 20. V. 2016; (F) ♂ (not collected) L. kameliyae, Huu Lien NR, 21. IV. 2014." httpUri="https://zenodo.org/record/495280/files/figure.png" pageId="6" pageNumber="321">Figs. 2</figureCitation>
|
||
E, 4D). It can readily be separated by its distinct dorsal thoracic stripes, which are absent in
|
||
<taxonomicName id="4C536D96ED3F4637FD81F4DAA68DF80E" box="[585,712,2012,2036]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED3F4637FD81F4DAA68DF80E" box="[585,712,2012,2036]" italics="true" pageId="6" pageNumber="321">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
or vague and more rectangular in
|
||
<taxonomicName id="4C536D96ED3F4637FBA0F4DAA0BEF80E" box="[1128,1275,2012,2036]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="321" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED3F4637FBA0F4DAA0BEF80E" box="[1128,1275,2012,2036]" italics="true" pageId="6" pageNumber="321">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
. See also the discussion section. The other is
|
||
<taxonomicName id="4C536D96ED3E4636FE37F39FA711FF4A" authority="Ris, 1919" authorityName="Ris" authorityYear="1919" box="[511,852,151,176]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="322" phylum="Arthropoda" rank="species" species="tricolor">
|
||
<emphasis id="B927CA07ED3E4636FE37F39FA6A5FF4A" box="[511,736,152,176]" italics="true" pageId="7" pageNumber="322">Lyriothemis tricolor</emphasis>
|
||
Ris, 1919
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13680A90ED3E4636FCABF390A7EBFF4A" box="[867,942,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="7" pageNumber="322">Figs. 1</figureCitation>
|
||
C, 1F, 5A). That species is thought to range from
|
||
<collectingCountry id="F3445685ED3E4636FF1CF3BBA54BFF2F" box="[212,270,188,213]" name="India" pageId="7" pageNumber="322">India</collectingCountry>
|
||
in the west to the Ryukyu Islands of
|
||
<collectingCountry id="F3445685ED3E4636FD67F3BAA6AAFF2E" box="[687,751,189,212]" name="Japan" pageId="7" pageNumber="322">Japan</collectingCountry>
|
||
in the east, although currently not known from
|
||
<collectingCountry id="F3445685ED3E4636FACFF3BBA12CFF2F" box="[1287,1385,188,213]" name="Vietnam" pageId="7" pageNumber="322">Vietnam</collectingCountry>
|
||
, and can be separated on the basis of, amongst others, distinct yellow dorsal stripes on the thorax, a black line over its abdomen, its more orange abdominal color and differently shaped hamulus, of which the inner edge is completely straight without incision (
|
||
<figureCitation id="13680A90ED3E4636FE75F220A646FEBA" box="[445,515,295,320]" captionStart="FIGURE 6" captionStartId="8.[151,250,737,759]" captionTargetBox="[165,1417,196,718]" captionTargetId="figure@8.[165,1417,196,718]" captionTargetPageId="8" captionText="FIGURE 6. Vulvar lamina: (A) L. bivittata, Xuan Son NP, 31. VII. 2016; (B) L. kameliyae allotype." httpUri="https://zenodo.org/record/495284/files/figure.png" pageId="7" pageNumber="322">Fig. 6</figureCitation>
|
||
A). Note that contrary to the key in
|
||
<bibRefCitation id="EFC26BE4ED3E4636FC6BF22FA0DDFEBA" author="Fraser" box="[931,1176,295,320]" pageId="7" pageNumber="322" refString="Fraser, F. C. (1936) The Fauna of British India, including Ceylon and Burma. Odonata. Uol. III. Taylor & Francis, London, 448 pp." type="book" year="1936" yearSuffix="p">Fraser (1936, p. 265)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C536D96ED3E4636FB6DF22EA15FFEBA" box="[1189,1306,296,320]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="322" phylum="Arthropoda" rank="species" species="tricolor">
|
||
<emphasis id="B927CA07ED3E4636FB6DF22EA15FFEBA" box="[1189,1306,296,320]" italics="true" pageId="7" pageNumber="322">L. tricolor</emphasis>
|
||
</taxonomicName>
|
||
sometimes displays dark streaks at the wing base and has 3 rows of cells at the beginning of the discoidal field, rendering these characters useless for identification purposes (
|
||
<figureCitation id="13680A90ED3E4636FD54F268A6A2FE72" box="[668,743,367,392]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="7" pageNumber="322">Figs. 1</figureCitation>
|
||
C, 1F).
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED3E4636FF0FF293A730FCEF" blockId="7.[151,1437,151,1832]" pageId="7" pageNumber="322">
|
||
Males of
|
||
<taxonomicName id="4C536D96ED3E4636FEFDF292A583FE56" box="[309,454,404,428]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="322" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED3E4636FEFDF292A583FE56" box="[309,454,404,428]" italics="true" pageId="7" pageNumber="322">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
differ from
|
||
<taxonomicName id="4C536D96ED3E4636FD9EF292A697FE56" box="[598,722,404,428]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="322" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED3E4636FD9EF292A697FE56" box="[598,722,404,428]" italics="true" pageId="7" pageNumber="322">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
by having black labium and occiput, and more square-shaped postfrons marking, black prothorax and blackish-brown dorsum of the thorax, sometimes with hard to discern indication of dorsal stripes. The abdomen is as wide as the thorax, S1–2 are yellow with black markings, S8 is distinctly marked with black, S9–10 are all black, and the ventral side of the abdomen is yellow. Structurally they are distinctly smaller, with proportionally shorter pterostigma and differently shaped hamulus (
|
||
<figureCitation id="13680A90ED3E4636FB16F123A166FDC7" box="[1246,1315,548,573]" captionStart="FIGURE 6" captionStartId="8.[151,250,737,759]" captionTargetBox="[165,1417,196,718]" captionTargetId="figure@8.[165,1417,196,718]" captionTargetPageId="8" captionText="FIGURE 6. Vulvar lamina: (A) L. bivittata, Xuan Son NP, 31. VII. 2016; (B) L. kameliyae allotype." httpUri="https://zenodo.org/record/495284/files/figure.png" pageId="7" pageNumber="322">Fig. 6</figureCitation>
|
||
C) with vshaped incision and posterior tooth at inner corner. The differences in wing venation seem too variable to be of much use. The male of
|
||
<taxonomicName id="4C536D96ED3E4636FE7EF16AA673FD7E" box="[438,566,620,644]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="322" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED3E4636FE7EF16AA673FD7E" box="[438,566,620,644]" italics="true" pageId="7" pageNumber="322">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
by contrast has white labium and brown occiput, rounded corners to the postfrons marking, pale markings on anterior and posterior lobes of prothorax, and warm brown thoracic dorsum. The abdomen is wider than the thorax, S1–2 are ochre yellow to orange, unmarked, and S3–10 are red dorsally and orange ventrally. Structurally
|
||
<taxonomicName id="4C536D96ED3E4636FE2BF1DEA618FD0A" box="[483,605,728,752]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="322" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED3E4636FE2BF1DEA618FD0A" box="[483,605,728,752]" italics="true" pageId="7" pageNumber="322">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
is larger, with proportionally longer pterostigma and hamulus with curved incision and a small posterior tooth in different position (fig. 6B).
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED3E4636FF0FF018A7B1FBA2" blockId="7.[151,1437,151,1832]" pageId="7" pageNumber="322">
|
||
Females of
|
||
<taxonomicName id="4C536D96ED3E4636FE9DF026A5AFFCC2" box="[341,490,800,824]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="322" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED3E4636FE9DF026A5AFFCC2" box="[341,490,800,824]" italics="true" pageId="7" pageNumber="322">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
have black labium and occiput. Anterior lobe of prothorax with pale marking, remainder black. Dorsum of prothorax blackish brown with vague somewhat paler dorsal stripes. Abdomen yellow and pale orange to brownish orange, with more or less distinct longitudinal stripes, S8–10 all black, ventral side of S1–7 yellow. In
|
||
<taxonomicName id="4C536D96ED3E4636FE9EF08AA591FC5E" box="[342,468,908,932]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="322" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED3E4636FE9EF08AA591FC5E" box="[342,468,908,932]" italics="true" pageId="7" pageNumber="322">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
the labium has pale edges, occiput is brown, and the middle and posterior lobes of prothorax have brown markings. Dorsum of thorax is warm brown without dorsal stripes. S1 is ochre yellow and unmarked, S2–7 are dorsally red, ventrally orange. S8–9 are dorsally red with black markings, but black ventrally. S10 is black. Structurally
|
||
<taxonomicName id="4C536D96ED3E4636FE00F0FEA603FBEA" box="[456,582,1016,1040]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="322" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED3E4636FE00F0FEA603FBEA" box="[456,582,1016,1040]" italics="true" pageId="7" pageNumber="322">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
is much larger bodied, with relatively longer pterostigma and the vulvar lamina only narrowly incised. Finally, in both males and females of
|
||
<taxonomicName id="4C536D96ED3E4636FC47F71AA04CFBCE" box="[911,1033,1052,1076]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="322" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED3E4636FC47F71AA04CFBCE" box="[911,1033,1052,1076]" italics="true" pageId="7" pageNumber="322">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
the dark streaking at the base of the wings tends to be more extensive. The characters are summarized in
|
||
<tableCitation id="C6D123AEED3E4636FC52F738A7ABFBA2" box="[922,1006,1087,1112]" captionStart="TABLE 1" captionStartId="9.[151,239,152,175]" captionTargetBox="[151,1436,233,1163]" captionTargetPageId="9" captionText="TABLE 1. Comparison of main differences between males (left two columns) and females (right two columns) of L." httpUri="http://table.plazi.org/id/DF2C469DED304638FF5FF39FA1D9FF55" pageId="7" pageNumber="322" tableUuid="DF2C469DED304638FF5FF39FA1D9FF55">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349459EED3E4636FF0FF763A1CEF8D2" pageId="7" pageNumber="322" type="discussion">
|
||
<paragraph id="8BEC1615ED3E4636FF0FF763A03AFA1E" blockId="7.[151,1437,151,1832]" pageId="7" pageNumber="322">
|
||
<emphasis id="B927CA07ED3E4636FF0FF763A509FB87" bold="true" box="[199,332,1124,1149]" pageId="7" pageNumber="322">Discussion.</emphasis>
|
||
Thanks to the help of Professor Jean Legrand I was able to study photos of Rambur’s
|
||
<typeStatus id="54E8A8B7ED3E4636FAC3F762A17EFB87" box="[1291,1339,1125,1149]" pageId="7" pageNumber="322">type</typeStatus>
|
||
(
|
||
<figureCitation id="13680A90ED3E4636FA82F763A1CBFB86" box="[1354,1422,1124,1149]" captionStart="FIGURE 7" captionStartId="8.[151,250,1801,1823]" captionTargetBox="[156,1436,815,1775]" captionTargetId="figure@8.[156,1436,815,1775]" captionTargetPageId="8" captionText="FIGURE 7. Rambur’s type of L. bivittata, courtesy of Jean Legrand." httpUri="https://zenodo.org/record/495285/files/figure.png" pageId="7" pageNumber="322">Fig. 7</figureCitation>
|
||
). It displays warm brown dorsum to the thorax, pterostigmata approximately 6 times longer than wide, and unmarked abdomen with orange visible on S8. These characters enabled me to identify my Vietnamese specimens. In his description of specimens in Selys’, Martin’s, and British Museum’s collections
|
||
<bibRefCitation id="EFC26BE4ED3E4636FBB1F7C8A0B6FB12" author="Ris" box="[1145,1267,1231,1256]" pageId="7" pageNumber="322" refString="Ris, F. (1909) Libellulines 1. Collections zoologiques du Baron Edm. de Selys Longchamps, Catalogue systematique et descriptif, 9, 1 - 120." type="journal article" year="1909">Ris (1909)</bibRefCitation>
|
||
described two males from
|
||
<collectingCountry id="F3445685ED3E4636FEF7F7F3A5CBFAF7" box="[319,398,1268,1293]" name="Vietnam" pageId="7" pageNumber="322">Tonkin</collectingCountry>
|
||
(“a.
|
||
<specimenCount id="9D55DD9CED3E4636FE2FF7F3A64EFAF6" box="[487,523,1268,1292]" pageId="7" pageNumber="322" type="male">2♂</specimenCount>
|
||
”) that are characterized, amongst others, as follows: “Thorax vorne kupferbraun…Abdomen anfangs sehr breit…leuchtend karminrot…unterseite 1–8 gelbrot, 9–10 dunkel….” (Front of thorax copper-brown…Abdomen first very broad…bright red…ventral side S1–8 yellow-red, 9–10 dark). He notes abdomen
|
||
<quantity id="4CABBBF0ED3E4636FE8DF667A5F2FA82" box="[325,439,1376,1400]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="3.7" metricValueMin="3.3" pageId="7" pageNumber="322" unit="mm" value="35.0" valueMax="37.0" valueMin="33.0">33–37mm</quantity>
|
||
and pterostigma
|
||
<quantity id="4CABBBF0ED3E4636FDBFF658A6A1FA82" box="[631,740,1375,1400]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="7" pageNumber="322" unit="mm" value="4.75" valueMax="5.0" valueMin="4.5">4.5–5mm</quantity>
|
||
, respectively. According to him a third male (from Darjeeling) also belongs to this group on the basis of coloration. It measures 30 resp.
|
||
<quantity id="4CABBBF0ED3E4636FC14F683A06AFA67" box="[988,1071,1412,1437]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="322" unit="mm" value="4.5">4.5mm</quantity>
|
||
. Although the description of the hamulus mentions a triangular incision, the accompanying drawing shows a hamulus identical to that of the male of
|
||
<taxonomicName id="4C536D96ED3E4636FF5FF6CAA554FA1E" box="[151,273,1484,1508]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="322" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED3E4636FF5FF6CAA554FA1E" box="[151,273,1484,1508]" italics="true" pageId="7" pageNumber="322">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
here described from
|
||
<collectingCountry id="F3445685ED3E4636FDC8F6CBA626FA1F" box="[512,611,1484,1509]" name="Vietnam" pageId="7" pageNumber="322">Vietnam</collectingCountry>
|
||
. It is likely these males were indeed
|
||
<taxonomicName id="4C536D96ED3E4636FC35F6CAA032FA1E" box="[1021,1143,1484,1508]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="322" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED3E4636FC35F6CAA032FA1E" box="[1021,1143,1484,1508]" italics="true" pageId="7" pageNumber="322">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED3E4636FF0FF6F6A1CEF8D2" blockId="7.[151,1437,151,1832]" pageId="7" pageNumber="322">
|
||
In his description of the second group (“b.
|
||
<specimenCount id="9D55DD9CED3E4636FD71F6F7A698F9F2" box="[697,733,1520,1544]" pageId="7" pageNumber="322" type="male">3♂</specimenCount>
|
||
”) Ris does not further discuss the hamulus. It is not clear he noticed differences, nor is it evident to which of the two groups the hamulus in his drawing belongs. The three males under “b.
|
||
<specimenCount id="9D55DD9CED3E4636FE85F53FA535F9AA" box="[333,368,1592,1616]" pageId="7" pageNumber="322" type="male">3♂</specimenCount>
|
||
” he describes as “Thorax vorne ganz schwarz, eine bronzebrauner Schulterstreif nur undeutlich erkennbar…Abdomen schlanker, Segment 9–10 oben, 8–10 unten schwarz…vordere Hälfte van Segment 2 schwärzlich.” (Dorsum of thorax completely black, a bronze-brown shoulderstripe only just noticeable…Abdomen more slender, S9–10 black above, S8–10 black below…First half of S2 blackish.) Measurements are 28, 30, 32 for abdomen and 3, 3.5 and 3.5 for pterostigmata, respectively. The illustration of the wings of the specimen from Than-Moi (p. 112) shows a pterostigma that is
|
||
<date id="FFED30D5ED3E4636FD03F5EBA6A2F8FE" box="[715,743,1772,1796]" pageId="7" pageNumber="322">4x</date>
|
||
longer than wide. It seems that although he described the underside of the abdomen as bright red, these smaller males therefore most likely refer to
|
||
<taxonomicName id="4C536D96ED3E4636FB4DF416A151F8D2" box="[1157,1300,1808,1832]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="322" phylum="Arthropoda" rank="species" species="kameliyae" status="sp. nov.">
|
||
<emphasis id="B927CA07ED3E4636FB4DF416A151F8D2" box="[1157,1300,1808,1832]" italics="true" pageId="7" pageNumber="322">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A214777CED3E4636FAD3F416A1CEF8D2" box="[1307,1419,1809,1832]" pageId="7" pageNumber="322" rank="species">spec. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF2C469DED314639FF5FF1E6A0C7FD0D" box="[151,1154,737,759]" httpUri="https://zenodo.org/record/495284/files/figure.png" pageId="8" pageNumber="323" targetBox="[165,1417,196,718]" targetPageId="8">
|
||
<paragraph id="8BEC1615ED314639FF5FF1E6A0C7FD0D" blockId="8.[151,1154,737,759]" box="[151,1154,737,759]" pageId="8" pageNumber="323">
|
||
<emphasis id="B927CA07ED314639FF5FF1E6A557FD0C" bold="true" box="[151,274,737,759]" pageId="8" pageNumber="323">FIGURE 6.</emphasis>
|
||
Vulvar lamina: (A)
|
||
<taxonomicName id="4C536D96ED314639FE28F1E5A685FD0D" authority="Xuan Son" authorityName="Xuan Son" box="[480,704,737,759]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="323" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED314639FE28F1E5A5B0FD0D" box="[480,501,738,759]" italics="true" pageId="8" pageNumber="323">L.</emphasis>
|
||
<emphasis id="B927CA07ED314639FE32F1E6A614FD0D" box="[506,593,737,759]" italics="true" pageId="8" pageNumber="323">bivittata</emphasis>
|
||
<emphasis id="B927CA07ED314639FD99F1E6A610FD0D" box="[593,597,737,759]" italics="true" pageId="8" pageNumber="323">,</emphasis>
|
||
Xuan Son
|
||
</taxonomicName>
|
||
NP, 31.VII.2016; (B)
|
||
<taxonomicName id="4C536D96ED314639FC6AF1E5A067FD0D" box="[930,1058,737,759]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="323" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED314639FC6AF1E5A067FD0D" box="[930,1058,737,759]" italics="true" pageId="8" pageNumber="323">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
allotype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2C469DED314639FF5FF40EA717F8E5" box="[151,850,1801,1823]" httpUri="https://zenodo.org/record/495285/files/figure.png" pageId="8" pageNumber="323" targetBox="[156,1436,815,1775]" targetPageId="8">
|
||
<paragraph id="8BEC1615ED314639FF5FF40EA717F8E5" blockId="8.[151,850,1801,1823]" box="[151,850,1801,1823]" pageId="8" pageNumber="323">
|
||
<emphasis id="B927CA07ED314639FF5FF40EA557F8E4" bold="true" box="[151,274,1801,1823]" pageId="8" pageNumber="323">FIGURE 7.</emphasis>
|
||
Rambur’s type of
|
||
<emphasis id="B927CA07ED314639FE18F40DA600F8E5" box="[464,581,1801,1823]" italics="true" pageId="8" pageNumber="323">
|
||
<taxonomicName id="4C536D96ED314639FE18F40DA604F8E5" box="[464,577,1801,1823]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="323" phylum="Arthropoda" rank="species" species="bivittata">L. bivittata</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
courtesy of Jean Legrand.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2C469DED304638FF5FF39FA1D9FF55" ID-Table-UUID="DF2C469DED304638FF5FF39FA1D9FF55" box="[151,1436,152,175]" httpUri="http://table.plazi.org/id/DF2C469DED304638FF5FF39FA1D9FF55" pageId="9" pageNumber="324" targetBox="[151,1436,233,1163]" targetIsTable="true" targetPageId="9">
|
||
<paragraph id="8BEC1615ED304638FF5FF39FA1D9FF55" blockId="9.[151,1436,152,207]" box="[151,1436,152,175]" pageId="9" pageNumber="324">
|
||
<emphasis id="B927CA07ED304638FF5FF39FA549FF55" bold="true" box="[151,268,152,175]" pageId="9" pageNumber="324">TABLE 1.</emphasis>
|
||
Comparison of main differences between males (left two columns) and females (right two columns) of L.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C349459EED304638FF5FF3B0A105F9AA" pageId="9" pageNumber="324" type="reference_group">
|
||
<paragraph id="8BEC1615ED304638FF5FF3B0A696FF35" blockId="9.[151,1436,152,207]" box="[151,723,183,207]" pageId="9" pageNumber="324">
|
||
<taxonomicName id="4C536D96ED304638FF5FF3B0A546FF35" ID-CoL="9X3FH" box="[151,259,183,207]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="kameliyae" status="sp. nov.">
|
||
<emphasis id="B927CA07ED304638FF5FF3B0A546FF35" box="[151,259,183,207]" italics="true" pageId="9" pageNumber="324">kameliyae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A214777CED304638FEC2F3BFA531FF35" box="[266,372,184,207]" pageId="9" pageNumber="324" rank="species">spec. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C536D96ED304638FE6FF3BFA68AFF35" ID-CoL="3WTKV" authority="Rambur, 1842" authorityName="Rambur" authorityYear="1842" box="[423,719,183,207]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED304638FE6FF3BFA659FF35" box="[423,540,183,207]" italics="true" pageId="9" pageNumber="324">L. bivittata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC26BE4ED304638FDE2F3BFA682FF35" author="Rambur" box="[554,711,184,207]" pageId="9" pageNumber="324" refString="Rambur, M. P. (1842) Histoire naturelle des insectes. Nevropteres. Libraire Encycl. de Roret, Paris, xvii + 534 pp." type="book" year="1842">Rambur, 1842</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED304638FE45F3EDA0FEFB71" pageId="9" pageNumber="324">
|
||
<table id="F953E4B5ED30B9CEFF5FF3EEA1D9FB71" box="[151,1436,233,1163]" gridcols="5" gridrows="14" pageId="9" pageNumber="324">
|
||
<tr id="35631457ED30B9CEFF5FF3EEA1D9FEFB" box="[151,1436,233,257]" gridrow="0" pageId="9" pageNumber="324" rowspan-0="1">
|
||
<th id="76B27D2BED30B9CEFE45F3EEA639FEFB" box="[397,636,233,257]" gridcol="1" gridrow="0" pageId="9" pageNumber="324">
|
||
<taxonomicName id="4C536D96ED304638FE45F3EDA653FEFB" ID-CoL="9X3FH" box="[397,534,233,257]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED304638FE45F3EDA653FEFB" box="[397,534,233,257]" italics="true" pageId="9" pageNumber="324">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
♂
|
||
</th>
|
||
<th id="76B27D2BED30B9CEFD5CF3EEA723FEFB" box="[660,870,233,257]" gridcol="2" gridrow="0" pageId="9" pageNumber="324">
|
||
<taxonomicName id="4C536D96ED304638FD5CF3EDA74FFEFB" ID-CoL="3WTKV" box="[660,778,233,257]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED304638FD5CF3EDA74FFEFB" box="[660,778,233,257]" italics="true" pageId="9" pageNumber="324">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
♂
|
||
</th>
|
||
<th id="76B27D2BED30B9CEFC53F3EEA0CEFEFB" box="[923,1163,233,257]" gridcol="3" gridrow="0" pageId="9" pageNumber="324">
|
||
<taxonomicName id="4C536D96ED304638FC53F3EDA066FEFB" ID-CoL="9X3FH" box="[923,1059,233,257]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED304638FC53F3EDA066FEFB" box="[923,1059,233,257]" italics="true" pageId="9" pageNumber="324">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
♀
|
||
</th>
|
||
<th id="76B27D2BED30B9CEFB6AF3EEA1D9FEFB" box="[1186,1436,233,257]" gridcol="4" gridrow="0" pageId="9" pageNumber="324">
|
||
<taxonomicName id="4C536D96ED304638FB6AF3EDA153FEFB" ID-CoL="3WTKV" box="[1186,1302,233,257]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED304638FB6AF3EDA153FEFB" box="[1186,1302,233,257]" italics="true" pageId="9" pageNumber="324">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
♀
|
||
</th>
|
||
</tr>
|
||
<tr id="35631457ED30B9CEFF5FF212A1D9FED1" box="[151,1436,277,299]" gridrow="1" pageId="9" pageNumber="324">
|
||
<th id="76B27D2BED30B9CEFF5FF212A531FED1" box="[151,372,277,299]" gridcol="0" gridrow="1" pageId="9" pageNumber="324">labium</th>
|
||
<td id="76B27D2BED30B9CEFE45F212A639FED1" box="[397,636,277,299]" gridcol="1" gridrow="1" pageId="9" pageNumber="324">black</td>
|
||
<td id="76B27D2BED30B9CEFD5CF212A723FED1" box="[660,870,277,299]" gridcol="2" gridrow="1" pageId="9" pageNumber="324">yellowish white</td>
|
||
<td id="76B27D2BED30B9CEFC53F212A0CEFED1" box="[923,1163,277,299]" gridcol="3" gridrow="1" pageId="9" pageNumber="324">black</td>
|
||
<td id="76B27D2BED30B9CEFB6AF212A1D9FED1" box="[1186,1436,277,299]" gridcol="4" gridrow="1" pageId="9" pageNumber="324">black edged pale</td>
|
||
</tr>
|
||
<tr id="35631457ED30B9CEFF5FF239A1D9FEAE" box="[151,1436,318,340]" gridrow="2" pageId="9" pageNumber="324">
|
||
<th id="76B27D2BED30B9CEFF5FF239A531FEAE" box="[151,372,318,340]" gridcol="0" gridrow="2" pageId="9" pageNumber="324">occiput</th>
|
||
<td id="76B27D2BED30B9CEFE45F239A639FEAE" box="[397,636,318,340]" gridcol="1" gridrow="2" pageId="9" pageNumber="324">black</td>
|
||
<td id="76B27D2BED30B9CEFD5CF239A723FEAE" box="[660,870,318,340]" gridcol="2" gridrow="2" pageId="9" pageNumber="324">brown</td>
|
||
<td id="76B27D2BED30B9CEFC53F239A0CEFEAE" box="[923,1163,318,340]" gridcol="3" gridrow="2" pageId="9" pageNumber="324">black</td>
|
||
<td id="76B27D2BED30B9CEFB6AF239A1D9FEAE" box="[1186,1436,318,340]" gridcol="4" gridrow="2" pageId="9" pageNumber="324">brown</td>
|
||
</tr>
|
||
<tr id="35631457ED30B9CEFF5FF26FA1D9FE84" box="[151,1436,360,382]" gridrow="3" pageId="9" pageNumber="324" rowspan-3="1" rowspan-4="1">
|
||
<th id="76B27D2BED30B9CEFF5FF26FA531FE84" box="[151,372,360,382]" gridcol="0" gridrow="3" pageId="9" pageNumber="324">postfrons mark</th>
|
||
<td id="76B27D2BED30B9CEFE45F26FA639FE84" box="[397,636,360,382]" gridcol="1" gridrow="3" pageId="9" pageNumber="324">more square</td>
|
||
<td id="76B27D2BED30B9CEFD5CF26FA723FE84" box="[660,870,360,382]" gridcol="2" gridrow="3" pageId="9" pageNumber="324">more rounded</td>
|
||
</tr>
|
||
<tr id="35631457ED30B9CEFF5FF296A1D9FE1E" box="[151,1436,401,484]" gridrow="4" pageId="9" pageNumber="324">
|
||
<th id="76B27D2BED30B9CEFF5FF296A531FE1E" box="[151,372,401,484]" gridcol="0" gridrow="4" pageId="9" pageNumber="324">prothorax</th>
|
||
<td id="76B27D2BED30B9CEFE45F296A639FE1E" box="[397,636,401,484]" gridcol="1" gridrow="4" pageId="9" pageNumber="324">black</td>
|
||
<td id="76B27D2BED30B9CEFD5CF296A723FE1E" box="[660,870,401,484]" gridcol="2" gridrow="4" pageId="9" pageNumber="324">pale markings</td>
|
||
<td id="76B27D2BED30B9CEFC53F296A0CEFE1E" box="[923,1163,401,484]" gridcol="3" gridrow="4" pageId="9" pageNumber="324">black with pale mark on anterior lobe</td>
|
||
<td id="76B27D2BED30B9CEFB6AF296A1D9FE1E" box="[1186,1436,401,484]" gridcol="4" gridrow="4" pageId="9" pageNumber="324">also brown marks on middle and posterior lobe</td>
|
||
</tr>
|
||
<tr id="35631457ED30B9CEFF5FF2FFA1D9FDD6" box="[151,1436,504,556]" gridrow="5" pageId="9" pageNumber="324">
|
||
<th id="76B27D2BED30B9CEFF5FF2FFA531FDD6" box="[151,372,504,556]" gridcol="0" gridrow="5" pageId="9" pageNumber="324">dorsum of thorax</th>
|
||
<td id="76B27D2BED30B9CEFE45F2FFA639FDD6" box="[397,636,504,556]" gridcol="1" gridrow="5" pageId="9" pageNumber="324">blackish brown</td>
|
||
<td id="76B27D2BED30B9CEFD5CF2FFA723FDD6" box="[660,870,504,556]" gridcol="2" gridrow="5" pageId="9" pageNumber="324">warm brown</td>
|
||
<td id="76B27D2BED30B9CEFC53F2FFA0CEFDD6" box="[923,1163,504,556]" gridcol="3" gridrow="5" pageId="9" pageNumber="324">blackish brown with indistinct dorsal stripes</td>
|
||
<td id="76B27D2BED30B9CEFB6AF2FFA1D9FDD6" box="[1186,1436,504,556]" gridcol="4" gridrow="5" pageId="9" pageNumber="324">warm brown</td>
|
||
</tr>
|
||
<tr id="35631457ED30B9CEFF5FF147A1D9FD8E" box="[151,1436,576,628]" gridrow="6" pageId="9" pageNumber="324">
|
||
<th id="76B27D2BED30B9CEFF5FF147A531FD8E" box="[151,372,576,628]" gridcol="0" gridrow="6" pageId="9" pageNumber="324">S1–2</th>
|
||
<td id="76B27D2BED30B9CEFE45F147A639FD8E" box="[397,636,576,628]" gridcol="1" gridrow="6" pageId="9" pageNumber="324">yellow, marked with black</td>
|
||
<td id="76B27D2BED30B9CEFD5CF147A723FD8E" box="[660,870,576,628]" gridcol="2" gridrow="6" pageId="9" pageNumber="324">ochre to orange, unmarked</td>
|
||
<td id="76B27D2BED30B9CEFC53F147A0CEFD8E" box="[923,1163,576,628]" gridcol="3" gridrow="6" pageId="9" pageNumber="324">yellow, marked with black</td>
|
||
<td id="76B27D2BED30B9CEFB6AF147A1D9FD8E" box="[1186,1436,576,628]" gridcol="4" gridrow="6" pageId="9" pageNumber="324">ochre yellow and red</td>
|
||
</tr>
|
||
<tr id="35631457ED30B9CEFF5FF18FA1D9FD21" box="[151,1436,648,731]" gridrow="7" pageId="9" pageNumber="324">
|
||
<th id="76B27D2BED30B9CEFF5FF18FA531FD21" box="[151,372,648,731]" gridcol="0" gridrow="7" pageId="9" pageNumber="324">abdomen</th>
|
||
<td id="76B27D2BED30B9CEFE45F18FA639FD21" box="[397,636,648,731]" gridcol="1" gridrow="7" pageId="9" pageNumber="324">red, same width thorax, venter yellow</td>
|
||
<td id="76B27D2BED30B9CEFD5CF18FA723FD21" box="[660,870,648,731]" gridcol="2" gridrow="7" pageId="9" pageNumber="324">red,>width thorax, venter orange</td>
|
||
<td id="76B27D2BED30B9CEFC53F18FA0CEFD21" box="[923,1163,648,731]" gridcol="3" gridrow="7" pageId="9" pageNumber="324">yellow, pale orange to brown, often striped, venter yellow</td>
|
||
<td id="76B27D2BED30B9CEFB6AF18FA1D9FD21" box="[1186,1436,648,731]" gridcol="4" gridrow="7" pageId="9" pageNumber="324">red, unmarked, venter orange</td>
|
||
</tr>
|
||
<tr id="35631457ED30B9CEFF5FF1E9A1D9FCD9" box="[151,1436,750,803]" gridrow="8" pageId="9" pageNumber="324">
|
||
<th id="76B27D2BED30B9CEFF5FF1E9A531FCD9" box="[151,372,750,803]" gridcol="0" gridrow="8" pageId="9" pageNumber="324">S8–10</th>
|
||
<td id="76B27D2BED30B9CEFE45F1E9A639FCD9" box="[397,636,750,803]" gridcol="1" gridrow="8" pageId="9" pageNumber="324">S8 with black, S9–10 black</td>
|
||
<td id="76B27D2BED30B9CEFD5CF1E9A723FCD9" box="[660,870,750,803]" gridcol="2" gridrow="8" pageId="9" pageNumber="324">red</td>
|
||
<td id="76B27D2BED30B9CEFC53F1E9A0CEFCD9" box="[923,1163,750,803]" gridcol="3" gridrow="8" pageId="9" pageNumber="324">black</td>
|
||
<td id="76B27D2BED30B9CEFB6AF1E9A1D9FCD9" box="[1186,1436,750,803]" gridcol="4" gridrow="8" pageId="9" pageNumber="324">S8–9 with red, remainder black</td>
|
||
</tr>
|
||
<tr id="35631457ED30B9CEFF5FF031A1D9FC70" box="[151,1436,822,906]" gridrow="9" pageId="9" pageNumber="324" rowspan-3="1" rowspan-4="1">
|
||
<th id="76B27D2BED30B9CEFF5FF031A531FC70" box="[151,372,822,906]" gridcol="0" gridrow="9" pageId="9" pageNumber="324">hamulus</th>
|
||
<td id="76B27D2BED30B9CEFE45F031A639FC70" box="[397,636,822,906]" gridcol="1" gridrow="9" pageId="9" pageNumber="324">v-shaped incision and tooth at corner</td>
|
||
<td id="76B27D2BED30B9CEFD5CF031A723FC70" box="[660,870,822,906]" gridcol="2" gridrow="9" pageId="9" pageNumber="324">curved incision and tooth removed from corner</td>
|
||
</tr>
|
||
<tr id="35631457ED30B9CEFF5FF09AA1D9FC49" box="[151,1436,925,947]" gridrow="10" pageId="9" pageNumber="324" rowspan-1="1" rowspan-2="1">
|
||
<th id="76B27D2BED30B9CEFF5FF09AA531FC49" box="[151,372,925,947]" gridcol="0" gridrow="10" pageId="9" pageNumber="324">vulvar lamina</th>
|
||
<td id="76B27D2BED30B9CEFC53F09AA0CEFC49" box="[923,1163,925,947]" gridcol="3" gridrow="10" pageId="9" pageNumber="324">widely incised</td>
|
||
<td id="76B27D2BED30B9CEFB6AF09AA1D9FC49" box="[1186,1436,925,947]" gridcol="4" gridrow="10" pageId="9" pageNumber="324">narrowly separated</td>
|
||
</tr>
|
||
<tr id="35631457ED30B9CEFF5FF0C1A1D9FBE0" box="[151,1436,966,1050]" gridrow="11" pageId="9" pageNumber="324">
|
||
<th id="76B27D2BED30B9CEFF5FF0C1A531FBE0" box="[151,372,966,1050]" gridcol="0" gridrow="11" pageId="9" pageNumber="324">pterostigma mm</th>
|
||
<td id="76B27D2BED30B9CEFE45F0C1A639FBE0" box="[397,636,966,1050]" gridcol="1" gridrow="11" pageId="9" pageNumber="324">3–3.5 about 4x longer than wide</td>
|
||
<td id="76B27D2BED30B9CEFD5CF0C1A723FBE0" box="[660,870,966,1050]" gridcol="2" gridrow="11" pageId="9" pageNumber="324">5 about 5x longer than wide</td>
|
||
<td id="76B27D2BED30B9CEFC53F0C1A0CEFBE0" box="[923,1163,966,1050]" gridcol="3" gridrow="11" pageId="9" pageNumber="324">4 about 4x longer than wide</td>
|
||
<td id="76B27D2BED30B9CEFB6AF0C1A1D9FBE0" box="[1186,1436,966,1050]" gridcol="4" gridrow="11" pageId="9" pageNumber="324">6 about 6x longer than wide</td>
|
||
</tr>
|
||
<tr id="35631457ED30B9CEFF5FF72AA1D9FB98" box="[151,1436,1069,1122]" gridrow="12" pageId="9" pageNumber="324">
|
||
<th id="76B27D2BED30B9CEFF5FF72AA531FB98" box="[151,372,1069,1122]" gridcol="0" gridrow="12" pageId="9" pageNumber="324">extent of streaking at wing base</th>
|
||
<td id="76B27D2BED30B9CEFE45F72AA639FB98" box="[397,636,1069,1122]" gridcol="1" gridrow="12" pageId="9" pageNumber="324">3 cells</td>
|
||
<td id="76B27D2BED30B9CEFD5CF72AA723FB98" box="[660,870,1069,1122]" gridcol="2" gridrow="12" pageId="9" pageNumber="324">4–5 cells</td>
|
||
<td id="76B27D2BED30B9CEFC53F72AA0CEFB98" box="[923,1163,1069,1122]" gridcol="3" gridrow="12" pageId="9" pageNumber="324">3–4 cells</td>
|
||
<td id="76B27D2BED30B9CEFB6AF72AA1D9FB98" box="[1186,1436,1069,1122]" gridcol="4" gridrow="12" pageId="9" pageNumber="324">6 cells</td>
|
||
</tr>
|
||
<tr id="35631457ED30B9CEFF5FF772A1D9FB71" box="[151,1436,1141,1163]" gridrow="13" pageId="9" pageNumber="324">
|
||
<th id="76B27D2BED30B9CEFF5FF772A531FB71" box="[151,372,1141,1163]" gridcol="0" gridrow="13" pageId="9" pageNumber="324">abdomen mm</th>
|
||
<td id="76B27D2BED30B9CEFE45F772A639FB71" box="[397,636,1141,1163]" gridcol="1" gridrow="13" pageId="9" pageNumber="324">26.5–29</td>
|
||
<td id="76B27D2BED30B9CEFD5CF772A723FB71" box="[660,870,1141,1163]" gridcol="2" gridrow="13" pageId="9" pageNumber="324">32</td>
|
||
<td id="76B27D2BED30B9CEFC53F772A0CEFB71" box="[923,1163,1141,1163]" gridcol="3" gridrow="13" pageId="9" pageNumber="324">27–27.5</td>
|
||
<td id="76B27D2BED30B9CEFB6AF772A1D9FB71" box="[1186,1436,1141,1163]" gridcol="4" gridrow="13" pageId="9" pageNumber="324">34</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED304638FF3DF7C8A105F9AA" blockId="9.[151,1437,1231,2013]" pageId="9" pageNumber="324">
|
||
The five female specimens described by Ris are all somewhat different. Female “a.” (from Sibsagar,
|
||
<collectingRegion id="4997D8F7ED304638FF5FF7F3A4A0FAF6" box="[151,229,1268,1292]" country="India" name="Assam" pageId="9" pageNumber="324">Assam</collectingRegion>
|
||
) has yellowish brown dorsum to the thorax and bright red abdomen (although S7–10 are missing). The pterostigma is with>
|
||
<quantity id="4CABBBF0ED304638FE4FF61FA584FACA" box="[391,449,1304,1328]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="324" unit="mm" value="4.0">4mm</quantity>
|
||
possibly short for
|
||
<taxonomicName id="4C536D96ED304638FD5FF61EA756FACA" ID-CoL="3WTKV" box="[663,787,1304,1328]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED304638FD5FF61EA756FACA" box="[663,787,1304,1328]" italics="true" pageId="9" pageNumber="324">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
, but the coloration seems to suggest this species. Female “b.” (from Than-Moi,
|
||
<collectingCountry id="F3445685ED304638FE59F63BA5A7FAAF" box="[401,482,1340,1365]" name="Vietnam" pageId="9" pageNumber="324">Tonkin</collectingCountry>
|
||
) has similar dorsum, but with vague shoulder stripes (sic.) and S1 black dorsally, S2– 7 brownish and S8–10 black. Its abdomen is
|
||
<quantity id="4CABBBF0ED304638FD6AF667A6A9FA82" box="[674,748,1376,1400]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="9" pageNumber="324" unit="mm" value="29.0">29mm</quantity>
|
||
, pterostigma
|
||
<quantity id="4CABBBF0ED304638FC4FF667A781FA82" box="[903,964,1376,1400]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="324" unit="mm" value="4.0">4mm</quantity>
|
||
. This likely is
|
||
<taxonomicName id="4C536D96ED304638FBBAF666A146FA82" ID-CoL="9X3FH" box="[1138,1283,1376,1400]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED304638FBBAF666A146FA82" box="[1138,1283,1376,1400]" italics="true" pageId="9" pageNumber="324">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
. Female “c.” (from
|
||
<collectingCountry id="F3445685ED304638FF15F683A56BFA67" box="[221,302,1412,1437]" name="Vietnam" pageId="9" pageNumber="324">Tonkin</collectingCountry>
|
||
) has a darker thorax, but with a vague copper-brown stripe. S1 is black dorsally, S2 black with yellow spots, S3–7 are yellow with black dorsal and posterior carinae and black longitudinal stripe, S8–10 black. Below S1 is black, S2–7 are yellow and 8–10 are black. Its abdomen is 30, pterostigma is
|
||
<quantity id="4CABBBF0ED304638FB81F6CBA0C2FA1E" box="[1097,1159,1484,1508]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="324" unit="mm" value="4.0">4mm</quantity>
|
||
. This is unmistakenly
|
||
<taxonomicName id="4C536D96ED304638FA4DF6CAA54CF9F2" ID-CoL="9X3FH" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED304638FA4DF6CAA54CF9F2" italics="true" pageId="9" pageNumber="324">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
. Female “d.” (from
|
||
<collectingCountry id="F3445685ED304638FE21F6E8A67FF9F2" box="[489,570,1519,1544]" name="Vietnam" pageId="9" pageNumber="324">Tonkin</collectingCountry>
|
||
) has S1–8 bright red, S8 with black sides, S9–10 black. Its abdomen is
|
||
<quantity id="4CABBBF0ED304638FA9CF6F7A1D9F9F2" box="[1364,1436,1520,1544]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="9" pageNumber="324" unit="mm" value="33.0">33mm</quantity>
|
||
and pterostigma <
|
||
<quantity id="4CABBBF0ED304638FEAAF513A5DBF9D7" box="[354,414,1556,1581]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="324" unit="mm" value="5.0">5mm</quantity>
|
||
. This could be
|
||
<taxonomicName id="4C536D96ED304638FD81F512A686F9D6" ID-CoL="3WTKV" box="[585,707,1556,1580]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED304638FD81F512A686F9D6" box="[585,707,1556,1580]" italics="true" pageId="9" pageNumber="324">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
. Female “e.” (from
|
||
<collectingCountry id="F3445685ED304638FC6EF513A7B2F9D7" box="[934,1015,1556,1581]" name="Vietnam" pageId="9" pageNumber="324">Tonkin</collectingCountry>
|
||
) has S2–7 brown, S8–10 black and is again smaller, abdomen 29 and pterostigmata
|
||
<quantity id="4CABBBF0ED304638FD51F530A6AEF9AA" box="[665,747,1591,1616]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="324" unit="mm" value="4.5">4.5mm</quantity>
|
||
, respectively. This is another
|
||
<taxonomicName id="4C536D96ED304638FBFEF53EA083F9AA" ID-CoL="9X3FH" box="[1078,1222,1592,1616]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED304638FBFEF53EA083F9AA" box="[1078,1222,1592,1616]" italics="true" pageId="9" pageNumber="324">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
candidate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349459EED30463AFF0FF55AA0BCFF2F" lastPageId="11" lastPageNumber="326" pageId="9" pageNumber="324" type="description">
|
||
<paragraph id="8BEC1615ED304638FF0FF55AA591F88A" blockId="9.[151,1437,1231,2013]" pageId="9" pageNumber="324">
|
||
<bibRefCitation id="EFC26BE4ED304638FF0FF55AA524F98F" author="Fraser" box="[199,353,1628,1653]" pageId="9" pageNumber="324" refString="Fraser, F. C. (1936) The Fauna of British India, including Ceylon and Burma. Odonata. Uol. III. Taylor & Francis, London, 448 pp." type="book" year="1936">Fraser (1936)</bibRefCitation>
|
||
provided a more extensive description of
|
||
<taxonomicName id="4C536D96ED304638FC82F55AA782F98E" box="[842,967,1628,1652]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED304638FC82F55AA782F98E" box="[842,967,1628,1652]" italics="true" pageId="9" pageNumber="324">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
. His description of the male is based on large (
|
||
<quantity id="4CABBBF0ED304638FF2AF587A512F962" box="[226,343,1664,1688]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="3.7" metricValueMin="3.3" pageId="9" pageNumber="324" unit="mm" value="35.0" valueMax="37.0" valueMin="33.0">33–37mm</quantity>
|
||
) specimens and mentions bright yellow labium, prothorax with posterior lobe ferruginous at middle, bright red abdomen with black borders on S9–10, all suggesting
|
||
<taxonomicName id="4C536D96ED304638FC18F5A2A009F946" box="[976,1100,1700,1724]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED304638FC18F5A2A009F946" box="[976,1100,1700,1724]" italics="true" pageId="9" pageNumber="324">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
. The illustration of the wings on p. 264 shows a pterostigma
|
||
<date id="FFED30D5ED304638FE3DF5C0A654F91A" box="[501,529,1735,1760]" pageId="9" pageNumber="324">5x</date>
|
||
longer than wide and extensive dark streaks at the wing base, also supporting
|
||
<taxonomicName id="4C536D96ED304638FA4DF5CEA4B1F8FE" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED304638FA4DF5CEA4B1F8FE" italics="true" pageId="9" pageNumber="324">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
. His description of the female contains characters of at least two specimens from widely different origin (
|
||
<collectingRegion id="4997D8F7ED304638FF56F417A4A9F8D2" box="[158,236,1808,1832]" country="India" name="Assam" pageId="9" pageNumber="324">Assam</collectingRegion>
|
||
and
|
||
<collectingCountry id="F3445685ED304638FEECF417A51BF8D2" box="[292,350,1808,1832]" name="Laos" pageId="9" pageNumber="324">Laos</collectingCountry>
|
||
), but does not contain definite characteristics of
|
||
<taxonomicName id="4C536D96ED304638FC4BF416A057F8D2" box="[899,1042,1808,1832]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED304638FC4BF416A057F8D2" box="[899,1042,1808,1832]" italics="true" pageId="9" pageNumber="324">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
, or it must be the statement that in some cases S8–10 are black. It is possible that some of the characteristics he mentions do not belong to
|
||
<taxonomicName id="4C536D96ED304638FAE9F432A1DEF8B6" box="[1313,1435,1844,1868]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="324" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED304638FAE9F432A1DEF8B6" box="[1313,1435,1844,1868]" italics="true" pageId="9" pageNumber="324">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
but concern another species.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED30463BFF0FF47BA508FCC2" blockId="9.[151,1437,1231,2013]" lastBlockId="10.[151,1437,151,2013]" lastPageId="10" lastPageNumber="325" pageId="9" pageNumber="324">
|
||
<bibRefCitation id="EFC26BE4ED304638FF0FF47BA52BF86F" author="Asahina" box="[199,366,1916,1941]" pageId="9" pageNumber="324" refString="Asahina, S. (1969) South Vietnam Odonata taken by Mr. Y. Inoue. Japanese Journal of Zoology, 16 (1), 1 - 19." type="journal article" year="1969">Asahina (1969</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC26BE4ED304638FEB0F47BA5F7F86F" author="Asahina" box="[376,434,1916,1941]" pageId="9" pageNumber="324" refString="Asahina, S. (1996) Records of the northern Vietnamese Odonata taken by the expedition members from National Science Museum, Tokyo 3. Aeschnidae, Corduliidae and Libellulidae. Bulletin National Science Museum Tokyo, Series A, 22 (2), 69 - 80." type="journal article" year="1996">1996</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC26BE4ED304638FE76F47BA5B2F86E" author="Asahina" box="[446,503,1916,1940]" pageId="9" pageNumber="324" refString="Asahina, S. (1988) A list of the Odonata from Thailand Part XI, Libellulidae- 1. Tombo, 31, 9 - 26." type="journal article" year="1988">1988</bibRefCitation>
|
||
) provides us with something of a puzzle.
|
||
<bibRefCitation id="EFC26BE4ED304638FC00F47BA030F86E" author="Asahina" box="[968,1141,1916,1941]" pageId="9" pageNumber="324" refString="Asahina, S. (1988) A list of the Odonata from Thailand Part XI, Libellulidae- 1. Tombo, 31, 9 - 26." type="journal article" year="1988">Asahina (1988)</bibRefCitation>
|
||
features a Thai male from
|
||
<collectingRegion id="4997D8F7ED304638FF5FF498A567F842" box="[151,290,1951,1976]" country="Thailand" name="Chanthaburi" pageId="9" pageNumber="324">Chanthaburi</collectingRegion>
|
||
that is relatively large (abdomen
|
||
<quantity id="4CABBBF0ED304638FD68F4A7A6ACF842" box="[672,745,1952,1976]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="9" pageNumber="324" unit="mm" value="31.0">31mm</quantity>
|
||
) with entirely red abdomen. However, important parts of the description are not provided, and the illustration of the hamulus is very sketchy (and does not show the posterior tooth). It is possible this specimen is true
|
||
<taxonomicName id="4C536D96ED33463BFDACF39EA69BFF4A" box="[612,734,152,176]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED33463BFDACF39EA69BFF4A" box="[612,734,152,176]" italics="true" pageId="10" pageNumber="325">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
. It illustrates the hamuli of a male from
|
||
<collectingCountry id="F3445685ED33463BFB56F390A0A4FF4A" box="[1182,1249,151,176]" name="Nepal" pageId="10" pageNumber="325">Nepal</collectingCountry>
|
||
and a male from Thu Duc in southern
|
||
<collectingCountry id="F3445685ED33463BFE4AF3BBA5A4FF2F" box="[386,481,188,213]" name="Vietnam" pageId="10" pageNumber="325">Vietnam</collectingCountry>
|
||
but does not further describe these. The male from southern
|
||
<collectingCountry id="F3445685ED33463BFB4DF3BBA0A1FF2F" box="[1157,1252,188,213]" name="Vietnam" pageId="10" pageNumber="325">Vietnam</collectingCountry>
|
||
had earlier been described in his 1969 paper. The illustrations of the hamuli in his 1988 paper for both specimens suggest
|
||
<taxonomicName id="4C536D96ED33463BFA4DF3E6A54CFEE6" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED33463BFA4DF3E6A54CFEE6" italics="true" pageId="10" pageNumber="325">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
. Likewise, the description of the male specimens from southern
|
||
<collectingCountry id="F3445685ED33463BFC27F203A00BFEE7" box="[1007,1102,260,285]" name="Vietnam" pageId="10" pageNumber="325">Vietnam</collectingCountry>
|
||
in his 1969 paper (p. 13–14) refers to “front side of pterothorax uniformly dark brownish” and abdomen length
|
||
<quantity id="4CABBBF0ED33463BFB4EF220A15AFEBA" box="[1158,1311,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="2.775" metricValueMax="2.95" metricValueMin="2.6" pageId="10" pageNumber="325" unit="mm" value="27.75" valueMax="29.5" valueMin="26.0">26—29.5mm</quantity>
|
||
. This also indicates
|
||
<taxonomicName id="4C536D96ED33463BFECCF24AA5D0FE9E" box="[260,405,332,356]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED33463BFECCF24AA5D0FE9E" box="[260,405,332,356]" italics="true" pageId="10" pageNumber="325">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
. The depiction of the hamuli in this (1969) paper shows the posterior tooth in a different position from that in
|
||
<bibRefCitation id="EFC26BE4ED33463BFE58F268A601FE72" author="Asahina" box="[400,580,367,392]" pageId="10" pageNumber="325" refString="Asahina, S. (1988) A list of the Odonata from Thailand Part XI, Libellulidae- 1. Tombo, 31, 9 - 26." type="journal article" year="1988">Asahina (1988)</bibRefCitation>
|
||
. Either one of the illustrations must be incorrect, or a different individual (from
|
||
<collectingRegion id="4997D8F7ED33463BFF16F292A565FE57" box="[222,288,405,429]" country="Thailand" name="Trang" pageId="10" pageNumber="325">Trang</collectingRegion>
|
||
Bom in southern
|
||
<collectingCountry id="F3445685ED33463BFE21F293A609FE57" box="[489,588,404,429]" name="Vietnam" pageId="10" pageNumber="325">Vietnam</collectingCountry>
|
||
?) is illustrated in his 1969 paper. If the illustration is correct, the identity of the concerned specimen is best left undecided. It is also not likely that it is
|
||
<emphasis id="B927CA07ED33463BFC2CF2BEA067FE2A" box="[996,1058,441,464]" italics="true" pageId="10" pageNumber="325">L. sp.</emphasis>
|
||
(see below), of which at least the female has bright yellow markings on the dorsum of the thorax. The 4 Thai females (also from
|
||
<collectingRegion id="4997D8F7ED33463BFAC0F2DBA1D1FE0F" box="[1288,1428,476,501]" country="Thailand" name="Chanthaburi" pageId="10" pageNumber="325">Chanthaburi</collectingRegion>
|
||
)
|
||
<bibRefCitation id="EFC26BE4ED33463BFF5FF2F8A50DFDE2" author="Asahina" box="[151,328,511,536]" pageId="10" pageNumber="325" refString="Asahina, S. (1988) A list of the Odonata from Thailand Part XI, Libellulidae- 1. Tombo, 31, 9 - 26." type="journal article" year="1988">Asahina (1988)</bibRefCitation>
|
||
describes are relatively small,
|
||
<quantity id="4CABBBF0ED33463BFD7FF2F8A76EFDE2" box="[695,811,511,536]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="10" pageNumber="325" unit="mm" value="27.0" valueMax="29.0" valueMin="25.0">25–29mm</quantity>
|
||
, and have shining black labium and chocolate brown thorax. Their abdomen is described as orange yellow when young, reddish brown when aged, with black edges on S8–9 and black S10. Although somewhat of a mix, it is well possible these females belong to
|
||
<taxonomicName id="4C536D96ED33463BFB18F14EA124FD9A" box="[1232,1377,584,608]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED33463BFB18F14EA124FD9A" box="[1232,1377,584,608]" italics="true" pageId="10" pageNumber="325">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
. The female depicted in
|
||
<bibRefCitation id="EFC26BE4ED33463BFEBDF16BA6DFFD7F" author="Asahina" box="[373,666,620,645]" pageId="10" pageNumber="325" refString="Asahina, S. (1996) Records of the northern Vietnamese Odonata taken by the expedition members from National Science Museum, Tokyo 3. Aeschnidae, Corduliidae and Libellulidae. Bulletin National Science Museum Tokyo, Series A, 22 (2), 69 - 80." type="journal article" year="1996" yearSuffix="p">Asahina (1996, p. 76–77)</bibRefCitation>
|
||
from Cuc Phuong in
|
||
<collectingRegion id="4997D8F7ED33463BFC51F16BA03AFD7F" box="[921,1151,620,645]" country="Vietnam" name="Ninh Binh" pageId="10" pageNumber="325">Ninh Binh Province</collectingRegion>
|
||
looks very much like
|
||
<taxonomicName id="4C536D96ED33463BFA4DF16AA54CFD52" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED33463BFA4DF16AA54CFD52" italics="true" pageId="10" pageNumber="325">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
. It has yellowish abdomen with black stripes and very dark dorsum to the thorax. The streaks at the wing bases seem short for
|
||
<taxonomicName id="4C536D96ED33463BFE48F1B2A5BFFD36" box="[384,506,692,716]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED33463BFE48F1B2A5BFFD36" box="[384,506,692,716]" italics="true" pageId="10" pageNumber="325">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
too and more in line with
|
||
<taxonomicName id="4C536D96ED33463BFCE8F1B2A7EAFD36" box="[800,943,692,716]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED33463BFCE8F1B2A7EAFD36" box="[800,943,692,716]" italics="true" pageId="10" pageNumber="325">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
. But the large size (abd.
|
||
<quantity id="4CABBBF0ED33463BFB0CF1B3A148FD36" box="[1220,1293,692,716]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="10" pageNumber="325" unit="mm" value="39.0">39mm</quantity>
|
||
) is puzzling. It seems plausible this is a mistake. Lastly, the immature female mentioned in
|
||
<bibRefCitation id="EFC26BE4ED33463BFBD4F1D0A08EFD0A" author="Asahina" box="[1052,1227,727,752]" pageId="10" pageNumber="325" refString="Asahina, S. (1969) South Vietnam Odonata taken by Mr. Y. Inoue. Japanese Journal of Zoology, 16 (1), 1 - 19." type="journal article" year="1969">Asahina (1969)</bibRefCitation>
|
||
is, based on wing pattern, size, coloration of thorax and labium a good candidate for
|
||
<taxonomicName id="4C536D96ED33463BFC6AF1FAA071FCEE" box="[930,1076,764,788]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED33463BFC6AF1FAA071FCEE" box="[930,1076,764,788]" italics="true" pageId="10" pageNumber="325">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
, although he does not mention distinct striping.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED33463BFF0FF043A54AFC5E" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="325">
|
||
<bibRefCitation id="EFC26BE4ED33463BFF0FF043A519FCA6" author="Lahiri" box="[199,348,836,861]" pageId="10" pageNumber="325" refString="Lahiri, A. R. (1987) Studies on the odonate fauna of Meghalaya. Records of the Zoological Survey of India, Occasional Paper 99, 1 - 402." type="journal article" year="1987">Lahiri (1987)</bibRefCitation>
|
||
reported a possible
|
||
<taxonomicName id="4C536D96ED33463BFDF4F042A6F2FCA6" box="[572,695,836,860]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED33463BFDF4F042A6F2FCA6" box="[572,695,836,860]" italics="true" pageId="10" pageNumber="325">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
female from Meghalaya in India. Judging from the size (abdomen
|
||
<quantity id="4CABBBF0ED33463BFF5FF06FA49BFC7A" box="[151,222,872,896]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="10" pageNumber="325" unit="mm" value="33.0">33mm</quantity>
|
||
and hindwing
|
||
<quantity id="4CABBBF0ED33463BFE4CF06FA588FC7A" box="[388,461,872,896]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="10" pageNumber="325" unit="mm" value="42.0">42mm</quantity>
|
||
) in combination with the extensive dark streaking at the wing base, it is certainly not
|
||
<taxonomicName id="4C536D96ED33463BFA4DF06EA54CFC5E" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED33463BFA4DF06EA54CFC5E" italics="true" pageId="10" pageNumber="325">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED33463BFF0FF0B7A6E3FC16" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="325">
|
||
<bibRefCitation id="EFC26BE4ED33463BFF0FF0B7A53DFC32" author="Do" box="[199,376,944,968]" pageId="10" pageNumber="325" refString="Do, M. C., Bui, M. H., Nguyen, T. H. & Phan, Q. T. (2011) Anisoptera of Cuc Phuong National Park, North Vietnam. International Dragonfly Fund, Report 33, 1 - 18." type="book chapter" year="2011">
|
||
Do
|
||
<emphasis id="B927CA07ED33463BFF3AF0B6A569FC32" box="[242,300,944,968]" italics="true" pageId="10" pageNumber="325">et al.</emphasis>
|
||
(2011
|
||
</bibRefCitation>
|
||
, p. 15) also recorded
|
||
<taxonomicName id="4C536D96ED33463BFDBAF0B6A6ABFC32" box="[626,750,944,968]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED33463BFDBAF0B6A6ABFC32" box="[626,750,944,968]" italics="true" pageId="10" pageNumber="325">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
from Cuc Phuong National Park, but it is obvious from the photo provided that this is in fact
|
||
<taxonomicName id="4C536D96ED33463BFDD8F0D2A6DAFC16" box="[528,671,980,1004]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED33463BFDD8F0D2A6DAFC16" box="[528,671,980,1004]" italics="true" pageId="10" pageNumber="325">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED33463BFF0FF0FFA0C0FB3E" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="325">
|
||
Many of these old descriptions cannot be conclusively attributed to either species, although some can. It is evident that both species had been lumped into one for over a century. It is thus necessary to check specimens in collections to ascertain their true identity. The male
|
||
<taxonomicName id="4C536D96ED33463BFD38F746A728FBA2" box="[752,877,1088,1112]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED33463BFD38F746A728FBA2" box="[752,877,1088,1112]" italics="true" pageId="10" pageNumber="325">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
specimens made available on the website of the British Museum of Natural History (www.data.nhm.ac.uk; BMNH(E)1201870 & 1201871) are clearly true
|
||
<taxonomicName id="4C536D96ED33463BFA4DF762A4B1FB5A" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED33463BFA4DF762A4B1FB5A" italics="true" pageId="10" pageNumber="325">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
. Other females from
|
||
<collectingRegion id="4997D8F7ED33463BFE20F780A637FB5A" box="[488,626,1159,1184]" country="Thailand" name="Chanthaburi" pageId="10" pageNumber="325">Chanthaburi</collectingRegion>
|
||
and one female from
|
||
<collectingRegion id="4997D8F7ED33463BFCA5F780A04FFB5A" box="[877,1034,1159,1184]" country="Thailand" name="Kanchanaburi" pageId="10" pageNumber="325">Kanchanaburi</collectingRegion>
|
||
not discussed here can for instance be found in the collection of Bro. Amnuay Pinratana in
|
||
<collectingRegion id="4997D8F7ED33463BFCCFF7ABA729FB3F" box="[775,876,1196,1221]" country="Thailand" name="Krung Thep Mahanakhon" pageId="10" pageNumber="325">Bangkok</collectingRegion>
|
||
(M. Hämäläinen
|
||
<emphasis id="B927CA07ED33463BFBFAF7AAA030FB3E" box="[1074,1141,1196,1220]" italics="true" pageId="10" pageNumber="325">in litt.</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED33463BFF0FF7C8A69CFA3A" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="325">
|
||
Within
|
||
<collectingCountry id="F3445685ED33463BFED3F7C8A53FFB12" box="[283,378,1231,1256]" name="Vietnam" pageId="10" pageNumber="325">Vietnam</collectingCountry>
|
||
there is one more
|
||
<taxonomicName id="4C536D96ED33463BFD83F7D7A695FB12" box="[587,720,1232,1256]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B927CA07ED33463BFD83F7D7A695FB12" box="[587,720,1232,1256]" italics="true" pageId="10" pageNumber="325">Lyriothemis</emphasis>
|
||
</taxonomicName>
|
||
species that looks like both
|
||
<taxonomicName id="4C536D96ED33463BFBD8F7D6A0E5FB12" box="[1040,1184,1232,1256]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED33463BFBD8F7D6A0E5FB12" box="[1040,1184,1232,1256]" italics="true" pageId="10" pageNumber="325">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C536D96ED33463BFB1FF7D6A114FB12" box="[1239,1361,1232,1256]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED33463BFB1FF7D6A114FB12" box="[1239,1361,1232,1256]" italics="true" pageId="10" pageNumber="325">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
and is somewhat intermediate in size. The male still eludes capture. The female (
|
||
<figureCitation id="13680A90ED33463BFC21F7F3A07CFAF6" box="[1001,1081,1268,1293]" captionStart="FIGURE 2" captionStartId="3.[151,250,1818,1840]" captionTargetBox="[185,1417,665,1777]" captionTargetId="figure@3.[174,1417,665,1780]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal scans: (A) ♀ Lyriothemis bivittata, Xuan Son NP, 31. VII. 2016; (B) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (C) ♂ L. kameliyae, paratype, Huu Lien NR, 24. V. 2014. Photos in natural habitat: (D) ♀ L. bivittata (not collected), Tam Dao NP, 18. VIII. 2013; (E) ♀ L. sp., Quang Binh Province, 20. V. 2016; (F) ♂ (not collected) L. kameliyae, Huu Lien NR, 21. IV. 2014." httpUri="https://zenodo.org/record/495280/files/figure.png" pageId="10" pageNumber="325">Figs. 2</figureCitation>
|
||
E, 4D) shares the lateral thorax pattern and dark streaks on the wing bases with both species, and a bright red unmarked abdomen (S1–7) with
|
||
<taxonomicName id="4C536D96ED33463BFA4DF61EA4B1FAAE" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED33463BFA4DF61EA4B1FAAE" italics="true" pageId="10" pageNumber="325">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
. But it has all black S8–10 and two bright yellow dorsal stripes on an otherwise blackish brown dorsum of its thorax. Although these dorsal stripes are similar to those of
|
||
<taxonomicName id="4C536D96ED33463BFC9FF666A78FFA82" box="[855,970,1376,1400]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="tricolor">
|
||
<emphasis id="B927CA07ED33463BFC9FF666A78FFA82" box="[855,970,1376,1400]" italics="true" pageId="10" pageNumber="325">L. tricolor</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13680A90ED33463BFC11F658A061FA82" box="[985,1060,1375,1400]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[216,1358,229,1915]" captionTargetId="figure@2.[212,1376,181,1915]" captionTargetPageId="2" captionText="FIGURE 1. Lateral scans: (A) ♂ Lyriothemis bivittata, Huu Lien NR, 23. VII. 2016; (B) ♂ L. kameliyae, holotype; (C) ♂ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh); (D) ♀ L. bivittata, Xuan Son NP, 31. VII. 2016; (E) ♀ L. kameliyae, paratype, Xuan Son NP, 31. VII. 2016; (F) ♀ L. tricolor, Taiwan (courtesy of Wen-Chi Yeh)." httpUri="https://zenodo.org/record/495278/files/figure.png" pageId="10" pageNumber="325">Figs. 1</figureCitation>
|
||
C, 1F), the abdomen pattern does not fit that species. Until its male is found, this species remains an enigma. In addition to the specimen illustrated here I have recorded it also from
|
||
<collectingRegion id="4997D8F7ED33463BFDC3F6AEA690FA3A" box="[523,725,1447,1472]" country="Vietnam" name="Ha Tinh" pageId="10" pageNumber="325">Ha Tinh Province</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEC1615ED33463AFF0FF6CBA0BCFF2F" blockId="10.[151,1437,151,2013]" lastBlockId="11.[151,1436,151,213]" lastPageId="11" lastPageNumber="326" pageId="10" pageNumber="325">
|
||
<materialsCitation id="3B3B1C48ED33463BFF0FF6CBA156F86F" ID-GBIF-Occurrence="1455704849" collectorName="Huu Lien Nature Reserve & At & In & Cuc Phuong National Park & Tam Dao National Park & Sebastien Delonglee & Both & April. & September & August. The & Van der Poorten & Asia & Lao" country="Nepal" elevation="1000" location="northern Vietnam" pageId="10" pageNumber="325" specimenCount="1" stateProvince="Yunnan">
|
||
<emphasis id="B927CA07ED33463BFF0FF6CBA624FA1F" bold="true" box="[199,609,1484,1509]" pageId="10" pageNumber="325">Note on behavior and occurrence.</emphasis>
|
||
Males and females of both species inhabit tropical rainforest habitats in mountainous areas, where they occur in very low densities. They often perch on branches in areas of tree fall, or otherwise sunny open spots inside the forest. Some individuals were observed striking pools of water on hot afternoons, likely to cool off.
|
||
<taxonomicName id="4C536D96ED33463BFE27F53EA6C4F9AA" box="[495,641,1592,1616]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED33463BFE27F53EA6C4F9AA" box="[495,641,1592,1616]" italics="true" pageId="10" pageNumber="325">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
has been verified from reserves in
|
||
<location id="8E8C40CEED33463BFBD2F530A0A1F9AA" LSID="urn:lsid:plazi:treatment:03FAA703ED3C463AFF5FF747A0BCFF2F:8E8C40CEED33463BFBD2F530A0A1F9AA" box="[1050,1252,1591,1616]" country="Nepal" name="northern Vietnam" pageId="10" pageNumber="325" stateProvince="Yunnan">northern Vietnam</location>
|
||
, especially with karst mountains. In Xuan Son National Park it was found at approximately
|
||
<quantity id="4CABBBF0ED33463BFBCCF55BA016F98E" box="[1028,1107,1628,1652]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="325" unit="m" value="1000.0">
|
||
<elevation id="007EF126ED33463BFBCCF55BA016F98E" box="[1028,1107,1628,1652]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="325" unit="m" value="1000.0">1000m</elevation>
|
||
</quantity>
|
||
asl, but in
|
||
<collectorName id="26A673C3ED33463BFB11F55AA4B4F962" pageId="10" pageNumber="325">Huu Lien Nature Reserve</collectorName>
|
||
the elevation is only 200–300 meters.
|
||
<collectorName id="26A673C3ED33463BFD0BF587A69AF962" box="[707,735,1664,1688]" pageId="10" pageNumber="325">At</collectorName>
|
||
both these locations it occurs together with
|
||
<emphasis id="B927CA07ED33463BFB3AF586A13CF962" box="[1266,1401,1664,1688]" italics="true" pageId="10" pageNumber="325">
|
||
<taxonomicName id="4C536D96ED33463BFB3AF586A130F962" box="[1266,1397,1664,1688]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="bivittata">L. bivittata</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<collectorName id="26A673C3ED33463BFA4CF586A1DEF962" box="[1412,1435,1665,1688]" pageId="10" pageNumber="325">In</collectorName>
|
||
addition to the other records mentioned previously,
|
||
<taxonomicName id="4C536D96ED33463BFD34F5A2A7D5F946" box="[764,912,1700,1724]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED33463BFD34F5A2A7D5F946" box="[764,912,1700,1724]" italics="true" pageId="10" pageNumber="325">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
has also been photographed at
|
||
<collectorName id="26A673C3ED33463BFAC5F5A3A571F91A" pageId="10" pageNumber="325">Cuc Phuong National Park</collectorName>
|
||
and
|
||
<collectorName id="26A673C3ED33463BFEB9F5CEA6C7F91A" box="[369,642,1735,1760]" pageId="10" pageNumber="325">Tam Dao National Park</collectorName>
|
||
by
|
||
<collectorName id="26A673C3ED33463BFD7AF5C0A7DEF91A" box="[690,923,1735,1760]" pageId="10" pageNumber="325">Sébastien Delonglée</collectorName>
|
||
(pers. comm.).
|
||
<collectorName id="26A673C3ED33463BFB9DF5C0A0CEF91A" box="[1109,1163,1735,1760]" pageId="10" pageNumber="325">Both</collectorName>
|
||
species appear in early
|
||
<collectorName id="26A673C3ED33463BFF5FF5EBA49FF8FF" box="[151,218,1772,1797]" pageId="10" pageNumber="325">April.</collectorName>
|
||
<taxonomicName id="4C536D96ED33463BFF29F5EAA518F8FE" box="[225,349,1772,1796]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED33463BFF29F5EAA518F8FE" box="[225,349,1772,1796]" italics="true" pageId="10" pageNumber="325">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
has been recorded at least into
|
||
<collectorName id="26A673C3ED33463BFD01F5EBA704F8FF" box="[713,833,1772,1797]" pageId="10" pageNumber="325">September</collectorName>
|
||
and L.
|
||
<taxonomicName id="4C536D96ED33463BFC53F5EBA048F8FE" box="[923,1037,1772,1796]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED33463BFC53F5EBA048F8FE" box="[923,1037,1772,1796]" italics="true" pageId="10" pageNumber="325">kameliyae</emphasis>
|
||
</taxonomicName>
|
||
into early
|
||
<collectorName id="26A673C3ED33463BFB42F5EBA152F8FF" box="[1162,1303,1772,1797]" pageId="10" pageNumber="325">August. The</collectorName>
|
||
range of
|
||
<taxonomicName id="4C536D96ED33463BFA4DF5EAA54CF8D2" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED33463BFA4DF5EAA54CF8D2" italics="true" pageId="10" pageNumber="325">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
possibly extends westward to
|
||
<collectingCountry id="F3445685ED33463BFDA8F408A6E6F8D2" box="[608,675,1807,1832]" name="Nepal" pageId="10" pageNumber="325">Nepal</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFC26BE4ED33463BFD7AF408A71CF8D2" author="Asahina" box="[690,857,1807,1832]" pageId="10" pageNumber="325" refString="Asahina, S. (1988) A list of the Odonata from Thailand Part XI, Libellulidae- 1. Tombo, 31, 9 - 26." type="journal article" year="1988">Asahina, 1988</bibRefCitation>
|
||
), but it certainly occurs in the south-east of
|
||
<collectingRegion id="4997D8F7ED33463BFA8FF417A1D9F8DD" box="[1351,1436,1808,1831]" country="China" name="Yunnan" pageId="10" pageNumber="325">Yunnan</collectingRegion>
|
||
in
|
||
<collectingCountry id="F3445685ED33463BFF0DF433A54EF8B7" box="[197,267,1844,1869]" name="China" pageId="10" pageNumber="325">China</collectingCountry>
|
||
, judging from photos of both a male and female published on http://nc.kl.edu.tw/bbs/ showthread.php?t=45178.
|
||
<collectorName id="26A673C3ED33463BFE09F45FA630F895" box="[449,629,1879,1904]" pageId="10" pageNumber="325">Van der Poorten</collectorName>
|
||
(2008) called
|
||
<taxonomicName id="4C536D96ED33463BFCD0F45EA7D7F88A" box="[792,914,1880,1904]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED33463BFCD0F45EA7D7F88A" box="[792,914,1880,1904]" italics="true" pageId="10" pageNumber="325">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
widespread in the southern part of continental
|
||
<collectorName id="26A673C3ED33463BFF5FF47BA488F86F" box="[151,205,1916,1941]" pageId="10" pageNumber="325">Asia</collectorName>
|
||
, known from
|
||
<collectingCountry id="F3445685ED33463BFEB9F47BA5B2F86F" box="[369,503,1916,1941]" name="Bangladesh" pageId="10" pageNumber="325">Bangladesh</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3445685ED33463BFDCCF47BA60FF86F" box="[516,586,1916,1941]" name="Nepal" pageId="10" pageNumber="325">Nepal</collectingCountry>
|
||
, NE
|
||
<collectingCountry id="F3445685ED33463BFD4FF47BA686F86F" box="[647,707,1916,1941]" name="India" pageId="10" pageNumber="325">India</collectingCountry>
|
||
,
|
||
<collectorName id="26A673C3ED33463BFD19F47AA6B8F86E" box="[721,765,1917,1940]" pageId="10" pageNumber="325">Lao</collectorName>
|
||
PDR,
|
||
<collectingCountry id="F3445685ED33463BFC85F47AA031F86E" box="[845,1140,1916,1941]" isEnumeration="true" pageId="10" pageNumber="325">Myanmar, Thailand, Laos</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3445685ED33463BFB78F47BA156F86F" box="[1200,1299,1916,1941]" name="Vietnam" pageId="10" pageNumber="325">Vietnam</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B3B1C48ED33463BFAD7F47AA047F843" ID-GBIF-Occurrence="1455704851" country="China" location="Guizhou" pageId="10" pageNumber="325" specimenCount="1" stateProvince="Guizhou">
|
||
In addition Haomiao
|
||
<bibRefCitation id="EFC26BE4ED33463BFECFF4A7A5E7F842" author="Zhang" box="[263,418,1952,1977]" pageId="10" pageNumber="325" refString="Zhang, H. (2011) Karst Forest Odonata from Southern Guizhou, China. International Dragonfly Fund, Report 37, 1 - 35." type="book chapter" year="2011">Zhang (2011)</bibRefCitation>
|
||
reported
|
||
<taxonomicName id="4C536D96ED33463BFDDBF4A6A6D4F842" box="[531,657,1952,1976]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="325" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED33463BFDDBF4A6A6D4F842" box="[531,657,1952,1976]" italics="true" pageId="10" pageNumber="325">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
from Southern
|
||
<collectingRegion id="4997D8F7ED33463BFC82F4A7A7E8F843" box="[842,941,1952,1977]" country="China" name="Guizhou" pageId="10" pageNumber="325">Guizhou</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3445685ED33463BFC73F4A7A047F843" box="[955,1026,1952,1977]" name="China" pageId="10" pageNumber="325">China</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B3B1C48ED33463AFBC6F4A7A0BCFF2F" ID-GBIF-Occurrence="1455704850" country="India" lastPageId="11" lastPageNumber="326" location="Vietnam" pageId="10" pageNumber="325" specimenCount="1">
|
||
Although it likely occurs over this complete range, given the specimens from
|
||
<collectingCountry id="F3445685ED33463BFDB0F4C3A6F7F827" box="[632,690,1988,2013]" name="India" pageId="10" pageNumber="325">India</collectingCountry>
|
||
on the one hand, and its occurrence in the north of
|
||
<location id="8E8C40CEED33463BFB39F4C3A115F827" LSID="urn:lsid:plazi:treatment:03FAA703ED3C463AFF5FF747A0BCFF2F:8E8C40CEED33463BFB39F4C3A115F827" box="[1265,1360,1988,2013]" country="India" name="Vietnam" pageId="10" pageNumber="325">Vietnam</location>
|
||
on the other, making it plausible it also occurs in
|
||
<collectingCountry id="F3445685ED32463AFDA5F390A6F6FF4A" box="[621,691,151,176]" name="China" pageId="11" pageNumber="326">China</collectingCountry>
|
||
, it is advisable to re-examine all specimens in collections. It can be concluded that
|
||
<taxonomicName id="4C536D96ED32463AFE8AF3BAA5F9FF2E" box="[322,444,188,212]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="326" phylum="Arthropoda" rank="species" species="bivittata">
|
||
<emphasis id="B927CA07ED32463AFE8AF3BAA5F9FF2E" box="[322,444,188,212]" italics="true" pageId="11" pageNumber="326">L. bivittata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C536D96ED32463AFE3BF3BAA6C6FF2E" box="[499,643,188,212]" class="Insecta" family="Libellulidae" genus="Lyriothemis" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="326" phylum="Arthropoda" rank="species" species="kameliyae">
|
||
<emphasis id="B927CA07ED32463AFE3BF3BAA6C6FF2E" box="[499,643,188,212]" italics="true" pageId="11" pageNumber="326">L. kameliyae</emphasis>
|
||
</taxonomicName>
|
||
at least co-occur over large parts of their known ranges.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |