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<document id="B49E56DEC6957B3E9ADA2142D6C3D844" ID-DOI="10.1080/00222930110088734" ID-ISSN="1464-5262" ID-Zenodo-Dep="4748152" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1620673573293" checkinUser="carolina" docAuthor="Shimomura, M &amp; Mawatari, SF" docDate="2002" docId="03FA87ADFF9FFFD3FE48FB166E26F97E" docLanguage="en" docName="JNATHIST.36.2103-2126.pdf.imf" docOrigin="Journal of Natural History 36 (17)" docStyle="DocumentStyle:B3570C21952C5CF44265488F0CE432D6.2:JNatHist.2003-2004.journal_article.2cover" docStyleId="B3570C21952C5CF44265488F0CE432D6" docStyleName="JNatHist.2003-2004.journal_article.2cover" docStyleVersion="2" docTitle="Heterosignum elegans Shimomura &amp; Mawatari 2002, sp. nov." docType="treatment" docVersion="4" lastPageNumber="2112" masterDocId="FFC3FFD5FF98FFD9FFF5FFCA6F4AFFCC" masterDocTitle="A revision of Heterosignum (Asellota, Isopoda, Paramunnidae) with descriptions of three new species" masterLastPageNumber="2126" masterPageNumber="2103" pageNumber="2109" updateTime="1698915214474" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="98C8E27F72AA4088723E616C23DF5E87" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E91FA35D3D046B756417F908A076AC8B">
<mods:title id="0FBB767F9E56EBD50AEDD74300DEE69F">A revision of Heterosignum (Asellota, Isopoda, Paramunnidae) with descriptions of three new species</mods:title>
</mods:titleInfo>
<mods:name id="FE32AB5B437982060BCFA7924AA96C99" type="personal">
<mods:role id="93157F5832076217BF84DD122FD5890D">
<mods:roleTerm id="3398D3797B9AD8F46A969AFC44F96788">Author</mods:roleTerm>
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<mods:namePart id="3B7D168D59B92C997CD39685342CC9FD">Shimomura, M</mods:namePart>
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<mods:namePart id="F83EC1E1F5D980A956C9CEE1E13A7BF4">Mawatari, SF</mods:namePart>
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<mods:typeOfResource id="16E4EBC1D984D9C6770D833BF0D23251">text</mods:typeOfResource>
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<mods:titleInfo id="47E5D77D8CEDBF209D7097C27AA2C1E8">
<mods:title id="BBD37110414DDC56D800D3B5E5D632BF">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="4FC39FFF66FF5B50270D8D2535031C1A">
<mods:date id="09397270782C3F2D363A18CA3230133F">2002</mods:date>
<mods:detail id="109EF133CD3EFDD6C0C71B1D11EDE115" type="pubDate">
<mods:number id="02406AAFE43EAB9BD5C81C6F84A7203D">2002-12-31</mods:number>
</mods:detail>
<mods:detail id="93DA005F6DF22E89043DAC20A0E147A9" type="volume">
<mods:number id="6E25A0AB34632E0F7D2ED6F119964D66">36</mods:number>
</mods:detail>
<mods:detail id="C1414D0B079EB3C615468E78A13F1187" type="issue">
<mods:number id="6A04CAD1DEC57A6B128282312F2A0D61">17</mods:number>
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<mods:start id="6EB6A414F9B2CB34047D0A2DA0F9E970">2103</mods:start>
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<mods:identifier id="B4C1B393CD8CAB0241D50369D814A92A" type="DOI">10.1080/00222930110088734</mods:identifier>
<mods:identifier id="479AF7C6C3F21650917DCD6F64B9F19A" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="A32D3968654A2B0679F2B6387BB40D0B" type="Zenodo-Dep">4748152</mods:identifier>
</mods:mods>
<treatment id="03FA87ADFF9FFFD3FE48FB166E26F97E" ID-DOI="http://doi.org/10.5281/zenodo.4748055" ID-Zenodo-Dep="4748055" LSID="urn:lsid:plazi:treatment:03FA87ADFF9FFFD3FE48FB166E26F97E" httpUri="http://treatment.plazi.org/id/03FA87ADFF9FFFD3FE48FB166E26F97E" lastPageId="10" lastPageNumber="2112" pageId="7" pageNumber="2109">
<subSubSection id="C3496530FF9FFFDEFE48FB166DFBFAD8" pageId="7" pageNumber="2109" type="nomenclature">
<paragraph id="8BEC36BBFF9FFFDEFE48FB166C5FFB38" blockId="7.[445,789,1244,1301]" box="[445,789,1244,1268]" pageId="7" pageNumber="2109">
<heading id="D0A481D7FF9FFFDEFE48FB166C5FFB38" box="[445,789,1244,1268]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="2109" reason="3">
<taxonomicName id="4C534D38FF9FFFDEFE48FB166DFBFB38" ID-CoL="6LRYS" authority="Shimomura &amp; Mawatari, 2002" authorityName="Shimomura &amp; Mawatari" authorityYear="2002" box="[445,689,1244,1268]" class="Malacostraca" family="Paramunnidae" genus="Heterosignum" kingdom="Animalia" order="Isopoda" pageId="7" pageNumber="2109" phylum="Arthropoda" rank="species" species="elegans" status="sp. nov.">
<emphasis id="B927EAA9FF9FFFDEFE48FB166DFBFB38" bold="true" box="[445,689,1244,1268]" italics="true" pageId="7" pageNumber="2109">Heterosignum elegans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21457D2FF9FFFDEFD4FFB176C5FFB38" box="[698,789,1245,1268]" pageId="7" pageNumber="2109" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BEC36BBFF9FFFDEFDD7FB366DFBFAD8" blockId="7.[445,789,1244,1301]" box="[546,689,1276,1301]" pageId="7" pageNumber="2109">(gures 46)</paragraph>
</subSubSection>
<subSubSection id="C3496530FF9FFFD3FF79FAF66E26F97E" lastPageId="10" lastPageNumber="2112" pageId="7" pageNumber="2109" type="materials_examined">
<paragraph id="8BEC36BBFF9FFFDEFF79FAF66E15FA98" blockId="7.[140,1094,1340,1555]" box="[140,351,1340,1364]" pageId="7" pageNumber="2109">
<emphasis id="B927EAA9FF9FFFDEFF79FAF66E15FA98" box="[140,351,1340,1364]" italics="true" pageId="7" pageNumber="2109">Material examined</emphasis>
</paragraph>
<paragraph id="8BEC36BBFF9FFFDEFF46FA966E76F9DF" blockId="7.[140,1094,1340,1555]" pageId="7" pageNumber="2109">
<materialsCitation id="3B3B3CE6FF9FFFDEFF46FA966E72F9DF" collectingDate="1999-11-18" collectionCode="V" country="Japan" elevation="115" latitude="45.7" location="Toyoshio-maru" longLatPrecision="677" longitude="45.13" pageId="7" pageNumber="2109" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yamaguchi" typeStatus="holotype">
TR/
<collectionCode id="ED42AE7EFF9FFFDEFF11FA966FBDFABF" box="[228,247,1372,1395]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="2109" type="Museum">V</collectionCode>
<location id="8E8C6060FF9FFFDEFEFDFA966EFEFAB8" LSID="urn:lsid:plazi:treatment:03FA87ADFF9FFFD3FE48FB166E26F97E:8E8C6060FF9FFFDEFEFDFA966EFEFAB8" box="[264,436,1372,1396]" country="Japan" latitude="45.7" longLatPrecision="677" longitude="45.13" name="Toyoshio-maru" pageId="7" pageNumber="2109" stateProvince="Yamaguchi">Toyoshio-maru</location>
, St. 10, Umoti-no-Se, oV
<collectingRegion id="4997F859FF9FFFDEFD1BFA966CA0FAB8" box="[750,1002,1372,1396]" country="Japan" name="Yamaguchi" pageId="7" pageNumber="2109">Yamaguchi Prefecture</collectingRegion>
,
<collectingCountry id="F344762BFF9FFFDEFC09FA966B74FABF" box="[1020,1086,1372,1395]" name="Japan" pageId="7" pageNumber="2109">Japan</collectingCountry>
, 34°
<geoCoordinate id="EE67507CFF9FFFDEFF44FAB66E41FA5F" box="[177,267,1404,1428]" degrees="45.70" direction="north" orientation="latitude" pageId="7" pageNumber="2109" precision="555" value="45.7">45.70°N</geoCoordinate>
, 130°
<geoCoordinate id="EE67507CFF9FFFDEFEBAFAB66EEFFA5F" box="[335,421,1404,1428]" degrees="45.13" direction="east" orientation="longitude" pageId="7" pageNumber="2109" precision="555" value="45.13">45.13°E</geoCoordinate>
,
<quantity id="4CAB9B5EFF9FFFDEFE42FAB66EB1FA5F" box="[439,507,1404,1428]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="7" pageNumber="2109" unit="m" value="115.0">
<elevation id="007ED188FF9FFFDEFE42FAB66EB1FA5F" box="[439,507,1404,1428]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="7" pageNumber="2109" unit="m" value="115.0">115 m</elevation>
</quantity>
, sand and mud,
<collectingMethod id="52124EACFF9FFFDEFD3BFAB66C09FA5F" box="[718,835,1404,1428]" pageId="7" pageNumber="2109">sledge net</collectingMethod>
(mesh size
<quantity id="4CAB9B5EFF9FFFDEFC23FAB66B7DFA58" box="[982,1079,1404,1428]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="2109" unit="mm" value="0.33">0.33 mm</quantity>
),
<date id="FFED107BFF9FFFDEFF79FA566E33FA78" box="[140,377,1436,1460]" pageId="7" pageNumber="2109" value="1999-11-18">
<collectingDate id="EFA9E993FF9FFFDEFF79FA566E33FA78" box="[140,377,1436,1460]" pageId="7" pageNumber="2109" value="1999-11-18">18 November 1999</collectingDate>
</date>
:
<typeStatus id="54E88819FF9FFFDEFE61FA566EBCFA78" box="[404,502,1436,1460]" pageId="7" pageNumber="2109" type="holotype">holotype</typeStatus>
, male,
<quantity id="4CAB9B5EFF9FFFDEFD96FA566DFFFA7F" box="[611,693,1436,1460]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="2109" unit="mm" value="1.1">1.1 mm</quantity>
(ZIHU-01907);
<typeStatus id="54E88819FF9FFFDEFC78FA566CB3FA78" box="[909,1017,1436,1460]" pageId="7" pageNumber="2109" type="paratype">paratypes</typeStatus>
,
<specimenCount id="9D55FD32FF9FFFDEFBE3FA566F81FA18" pageId="7" pageNumber="2109" type="male">four males</specimenCount>
,
<quantity id="4CAB9B5EFF9FFFDEFF14FA766E78FA1F" box="[225,306,1468,1492]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="2109" unit="mm" value="1.1">1.1 mm</quantity>
(ZIHU-01908-10),
<quantity id="4CAB9B5EFF9FFFDEFDD1FA766D3FFA1F" box="[548,629,1468,1492]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="2109" unit="mm" value="1.2">1.2 mm</quantity>
(ZIHU-01911),
<specimenCount id="9D55FD32FF9FFFDEFCB1FA766C95FA18" box="[836,991,1468,1492]" pageId="7" pageNumber="2109" type="female">three females</specimenCount>
,
<quantity id="4CAB9B5EFF9FFFDEFC01FA766B0CFA1F" box="[1012,1094,1468,1492]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="2109" unit="mm" value="1.4">1.4 mm</quantity>
(ZIHU-01912),
<quantity id="4CAB9B5EFF9FFFDEFE94FA116EFBFA3F" box="[353,433,1499,1523]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="2109" unit="mm" value="1.2">1.2 mm</quantity>
(ZIHU-01913),
<quantity id="4CAB9B5EFF9FFFDEFD5FFA116DB6FA3F" box="[682,764,1499,1523]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="2109" unit="mm" value="1.1">1.1 mm</quantity>
(ZIHU-01914), 1.0 mm (ZIHU-01915)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEC36BBFF9FFFDEFF79F9F66E40F99F" blockId="7.[140,1095,1596,1714]" box="[140,266,1596,1619]" pageId="7" pageNumber="2109">
<emphasis id="B927EAA9FF9FFFDEFF79F9F66E40F99F" box="[140,266,1596,1619]" italics="true" pageId="7" pageNumber="2109">Description</emphasis>
</paragraph>
<paragraph id="8BEC36BBFF9FFFD1FF46F9916CF8FD1A" blockId="7.[140,1095,1596,1714]" lastBlockId="8.[140,1095,160,1714]" lastPageId="8" lastPageNumber="2110" pageId="7" pageNumber="2109">
<emphasis id="B927EAA9FF9FFFDEFF46F9916FBFF9BF" box="[179,245,1627,1651]" italics="true" pageId="7" pageNumber="2109">Male.</emphasis>
Body (gure 4A) about 2.6 times as long as maximum width (except spine-like processes), widest at pereonite 3. Head broad, with six dorsal setae, with a pair of long preorbital lobes; frontal margin of head convex, with one or two pairs of setae; labrum surpassing head anteriorly; posterior margin of head convex. Eyestalks slender, very long, arising from posterior half of head, reaching proximal two-thirds of peduncular article 2 of antenna 1. Pereonites 17 with scattered small dorsal and lateral setae. Pereonite 1 laterally rounded, without lateral spine-like processes; pereonites 2 and 3 with a pair of long single lateral spine-like processes; pereonites 4 and 6 with a pair of slightly backwardly curved single lateral spine-like processes; pereonites 5 and 7 with a pair of short single lateral spine-like processes. Pereonite 1 shortest; pereonite 2 twice as long as pereonite 1; pereonite 3 longest; pereonite 4 slightly shorter than pereonite 3; pereonite 5 half as long as pereonite 4; pereonite 6 longer than pereonite 5; pereonite 7 as long as pereonite 6. Pereonites 13 increasing in width (except spine-like processes); pereonites 47 decreasing in width (except spine-like processes). Coxal plates dorsally invisible on all pereonites. Pleonite 1 small, with a pair of dorsal setae. Pleotelson 1.7 times as long as broad, with scattered small dorsal setae, consisting of two parts; anterior part of pleotelson long, cylindrical, about two- fths as long as posterior; posterior part of pleotelson bulbous, with six pairs of lateral teeth. Uropod (gure 4C) biramous, cylindrical; ventral ramus, 2.6 times as long as broad, bearing six apical and two lateral setae; dorsal ramus one-third length of ventral ramus, with two apical setae.
</paragraph>
<paragraph id="8BEC36BBFF90FFD1FF46FD146D94FC59" blockId="8.[140,1095,160,1714]" pageId="8" pageNumber="2110">Antenna 1 (gure 4D) with three-articulate peduncle and three-articulate agellum. Peduncular article 1 shorter than article 2, with two lateral and two mesial setae; article 2 longest, narrower than article 1, with nine distal setae; article 3 about one-third length of article 2. Flagellum article 1 shortest, with one distolateral seta; article 2 longer than article 1, without setae; article 3 as long as article 2, distally with one long, three short setae and one aesthetasc.</paragraph>
<paragraph id="8BEC36BBFF90FFD1FF46FC546D62FB58" blockId="8.[140,1095,160,1714]" pageId="8" pageNumber="2110">Antenna 2 (gure 4E) with peduncle composed of six articles: article 1 not visible dorsally; article 2 widest, longer than broad, mesially curved, without setae; article 3 longer than article 1, with convex proximal protrusion bearing single seta, and with three lateral, one mesial and one ventrodistal setae; article 4 shortest, with two long mesial and one ventrodistal setae; article 5 twice as long as article 4, with two lateral, one mesial and one ventrodistal setae; article 6 longest, with eight distal and one proximomesial setae. Flagellum longer than peduncular article 6, composed of eight articles, each with a few setae.</paragraph>
<paragraph id="8BEC36BBFF90FFD1FF46FB566B56FB38" blockId="8.[140,1095,160,1714]" pageId="8" pageNumber="2110">Left mandible (gure 4F) incisor with four cusps; lacinia mobilis with four teeth; molar process with two setae. Right mandible (gure 4G) lacking lacinia mobilis; incisor with ve cusps; spine row with one spine; molar process with two setae.</paragraph>
<paragraph id="8BEC36BBFF90FFD1FF46FB366E5CFABF" blockId="8.[140,1095,160,1714]" pageId="8" pageNumber="2110">Maxilla 1 (gure 5A) with inner lobe bearing four apical and four lateral setae; outer lobe with nine apical and two mesial setae. Maxilla 2 (gure 5B) with inner lobe bearing six apical and three mesial setae; outer two lobes each with two apical setae.</paragraph>
<paragraph id="8BEC36BBFF90FFD1FF46FAB66CEAF99F" blockId="8.[140,1095,160,1714]" pageId="8" pageNumber="2110">Maxilliped (gure 5C) with narrow palp composed of ve articles; article 1 broader than long, without setae; article two trapezoidal, slightly wider than article 1, with two mesial setae; article 3 longest, with three mesial and one distolateral setae; article 4 slightly shorter than article 3, with two distomesial setae; article 5 narrowest, with three apical and one mesial setae; endite rectangular, bearing three pectinate and two simple setae distally, and two mesial coupling hooks; epipod broad, subtriangular, reaching third palp article, with rounded apex.</paragraph>
<paragraph id="8BEC36BBFF90FFD3FF46F9916F8CFE9B" blockId="8.[140,1095,160,1714]" lastBlockId="10.[140,1095,160,1300]" lastPageId="10" lastPageNumber="2112" pageId="8" pageNumber="2110">Pereopod 1 (gure 5D) shorter than pereopods 27: basis the longest article, 1.6 times as long as ischium, with two ventral and one dorsal setae; ischium twice as long as merus, with one blunt robust projection, and with one ventral seta; merus trapezoidal, dorsodistally with one simple and one robust setae, and with three ventrodistal setae; carpus ovate, ventrally with two simple, two sensory robust setae and seven acute teeth; propodus ovate, ventrally with one simple, two robust sensory setae and two teeth, and dorsally with one simple seta; dactylus narrowest, shorter than propodus, with four distal setae, one slender unguis and one acute accessory claw.</paragraph>
<caption id="DF2C6633FF91FFD0FF79F9C96B02F9A4" ID-DOI="http://doi.org/10.5281/zenodo.4748166" ID-Zenodo-Dep="4748166" httpUri="https://zenodo.org/record/4748166/files/figure.png" pageId="9" pageNumber="2111" startId="9.[140,156,1539,1560]" targetBox="[183,1049,155,1508]" targetPageId="9">
<paragraph id="8BEC36BBFF91FFD0FF79F9C96B02F9A4" blockId="9.[140,1096,1536,1641]" pageId="9" pageNumber="2111">
FIG. 4.
<taxonomicName id="4C534D38FF91FFD0FF12F9C96E9AF9D4" authority="Shimomura &amp; Mawatari, 2002" authorityName="Shimomura &amp; Mawatari" authorityYear="2002" box="[231,464,1538,1560]" class="Malacostraca" family="Paramunnidae" genus="Heterosignum" kingdom="Animalia" order="Isopoda" pageId="9" pageNumber="2111" phylum="Arthropoda" rank="species" species="elegans" status="sp. nov.">
<emphasis id="B927EAA9FF91FFD0FF12F9C96E9AF9D4" box="[231,464,1538,1560]" italics="true" pageId="9" pageNumber="2111">Heterosignum elegans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21457D2FF91FFD0FE13F9C96D0FF9D4" box="[486,581,1539,1560]" pageId="9" pageNumber="2111" rank="species">sp. nov.</taxonomicNameLabel>
: (A) holotype male; (B) paratype female (ZIHU-01912); (C, F, G) paratype male (ZIHU-01908); (D, E) paratype female (ZIHU-01914). (A) Habitus, dorsal; (B) habitus, dorsal; (C) left uropod; (D) right antenna 1; (E) right antenna 2; (F) left mandible; (G) right mandible. Scales=0.1 mm.
</paragraph>
</caption>
<paragraph id="8BEC36BBFF92FFD3FF46FE956CA0FD9B" blockId="10.[140,1095,160,1300]" pageId="10" pageNumber="2112">Pereopods 27 (gure 5E, F) subequal in shape and length; bases as long as ischia, with one or two dorsal and one or two ventral setae; ischia twice as long as meri, with two dorsal and one or two ventral setae; meri trapezoidal, with one dorsodistal and two or three ventral setae; carpi as long as bases, with two or three robust sensory and one or two simple setae ventrally, two or three simple setae dorsally; propodi narrower than carpi, with two or three robust sensory and one simple setae ventrally, three simple setae dorsally; dactyli shortest, narrowest, distally with three simple setae, one slender unguis and one minute accessory claw.</paragraph>
<paragraph id="8BEC36BBFF92FFD3FF46FD946EDEFC59" blockId="10.[140,1095,160,1300]" pageId="10" pageNumber="2112">Pleopod 1 (gure 5G) twice as long as broad, with two pairs of ventrolateral setae and two pairs of apical setae, with a pair of short lateral projections, each projection bearing ve setae. Pleopod 2 (gure 6A) with narrow protopod and convex lateral margin, with 15 lateral and three ventral setae; endopod surpassing apex of protopod, with slender, long, curved second article; exopod stout. Pleopod 3 (gure 6B) with endopod bearing three stout plumose setae apically; exopod composed of two articles, as long as endopod, narrower than endopod; article 2 with one terminal seta. Pleopod 4 (gure 6C) with exopod, slender, tapering to pointed apex, without setae; endopod broad. Pleopod 5 (gure 6D) ovate, uniramous, twice as long as broad.</paragraph>
<paragraph id="8BEC36BBFF92FFD3FF46FC546E5CFAD8" blockId="10.[140,1095,160,1300]" pageId="10" pageNumber="2112">
<emphasis id="B927EAA9FF92FFD3FF46FC546E40FC7A" box="[179,266,926,950]" italics="true" pageId="10" pageNumber="2112">Female.</emphasis>
Similar to male in morphology of all pereonal appendages. Body (gure 4B) broader than in male, about 2.2 times as long as maximum width (except lateral spine-like processes), widest at pereonite 3. Pereonite 1 without single lateral spine-like processes; pereonites 24 with a pair of long, single lateral spine-like processes; pereonites 5 and 7 with a pair of short, single lateral spine-like processes; pereonite 6 with a pair of slightly backwardly curved long single lateral spine-like processes. Pereonite 1 shortest; pereonite 2 twice as long as pereonite 1; pereonite 3 as long as pereonite 2; pereonite 4 longest; pereonite 5 about one-third as long as pereonite 4; pereonite 6 length of pereonite 5; pereonite 7 as long as pereonite 6. Pereonites 13 increasing in width; pereonites 47 decreasing in width. Operculum (gure 6E) ovate, 1.4 times as long as broad, with a fringe of lateral setae, without apical setae.
</paragraph>
<paragraph id="8BEC36BBFF92FFD3FF79FAF66FA4FA98" blockId="10.[140,1095,1340,1714]" box="[140,238,1340,1364]" pageId="10" pageNumber="2112">
<emphasis id="B927EAA9FF92FFD3FF79FAF66FA4FA98" box="[140,238,1340,1364]" italics="true" pageId="10" pageNumber="2112">Remarks</emphasis>
</paragraph>
<paragraph id="8BEC36BBFF92FFD3FF46FA966E26F97E" blockId="10.[140,1095,1340,1714]" pageId="10" pageNumber="2112">
The absence of dorsal spines on pereonites links
<taxonomicName id="4C534D38FF92FFD3FD19FA976CA8FAB8" authority="Shimomura &amp; Mawatari, 2002" authorityName="Shimomura &amp; Mawatari" authorityYear="2002" box="[748,994,1372,1396]" class="Malacostraca" family="Paramunnidae" genus="Heterosignum" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="2112" phylum="Arthropoda" rank="species" species="elegans" status="sp. nov.">
<emphasis id="B927EAA9FF92FFD3FD19FA976CA8FAB8" box="[748,994,1372,1396]" italics="true" pageId="10" pageNumber="2112">Heterosignum elegans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21457D2FF92FFD3FC19FA966B0CFABF" box="[1004,1094,1372,1395]" pageId="10" pageNumber="2112" rank="species">sp. nov.</taxonomicNameLabel>
to
<taxonomicName id="4C534D38FF92FFD3FF59FAB76EBCFA58" authority="(Gamo, 1976)" baseAuthorityName="Gamo" baseAuthorityYear="1976" box="[172,502,1404,1428]" class="Malacostraca" family="Paramunnidae" genus="Heterosignum" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="2112" phylum="Arthropoda" rank="species" species="mutsuensis">
<emphasis id="B927EAA9FF92FFD3FF59FAB76E03FA58" box="[172,329,1405,1428]" italics="true" pageId="10" pageNumber="2112">H. mutsuensis</emphasis>
(
<bibRefCitation id="EFC24B4AFF92FFD3FEAFFAB66EA2FA58" author="GAMO, S." box="[346,488,1404,1428]" pageId="10" pageNumber="2112" pagination="39 - 45" refId="ref10218" refString="GAMO, S., 1976, Heterosignum mutsuensis gen. nov., sp. nov., a new Isopod Crustacea from Mutsu Bay, Northern Japan (Paraselloidea, Munnidae), Proceedings of the Japanese Society of Systematic Zoology, 12, 39 - 45." type="journal article" year="1976">Gamô, 1976</bibRefCitation>
)
</taxonomicName>
. The present new species is distinguished, however, from
<taxonomicName id="4C534D38FF92FFD3FF3FFA576E2EFA78" baseAuthorityName="Gamo" baseAuthorityYear="1976" box="[202,356,1437,1460]" class="Malacostraca" family="Paramunnidae" genus="Heterosignum" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="2112" phylum="Arthropoda" rank="species" species="mutsuensis">
<emphasis id="B927EAA9FF92FFD3FF3FFA576E2EFA78" box="[202,356,1437,1460]" italics="true" pageId="10" pageNumber="2112">H. mutsuensis</emphasis>
</taxonomicName>
by the following features (those of
<taxonomicName id="4C534D38FF92FFD3FD01FA576CC4FA78" baseAuthorityName="Gamo" baseAuthorityYear="1976" box="[756,910,1437,1460]" class="Malacostraca" family="Paramunnidae" genus="Heterosignum" kingdom="Animalia" order="Isopoda" pageId="10" pageNumber="2112" phylum="Arthropoda" rank="species" species="mutsuensis">
<emphasis id="B927EAA9FF92FFD3FD01FA576CC4FA78" box="[756,910,1437,1460]" italics="true" pageId="10" pageNumber="2112">H. mutsuensis</emphasis>
</taxonomicName>
in parentheses): a pair of preorbital lobes are present on head (absent); eyestalks are long and reaching to the proximal two-thirds of peduncle article 2 of antenna 2 (relatively short and reaching to the proximal half); cylindrical anterior part of pleotelson is very long (short); pleopod 1 is armed with two pairs of apical setae (three pairs of apical setae), two pairs of ventrolateral setae (without setae) and a pair of lateral projections, each projection bearing ve apical setae (nine pairs of apical setae); second article of pleopod 2 is very long (short); and no apical setae on the operculum (some apical setae).
</paragraph>
</subSubSection>
</treatment>
</document>