286 lines
65 KiB
XML
286 lines
65 KiB
XML
<document id="43734B1CB57841F6603C47E0771B86B1" ID-DOI="10.35463/j.apr.2021.02.01" ID-ISSN="1842-371x" ID-Zenodo-Dep="12583322" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719585668405" checkinUser="felipe" docAuthor="SCHLAGINTWEIT, FELIX & RASHIDI, KOOROSH" docDate="2021" docId="03B887C22B484E6FFCACD6B8B71ADA5C" docLanguage="en" docName="ActaPalaeontolRom.17.2.3-13.pdf" docOrigin="Acta Palaeontologica Romaniae 17 (2)" docSource="http://dx.doi.org/10.35463/j.apr.2021.02.01" docStyle="DocumentStyle:CA474AAD92E97C533D63E0C5C0F159F1.1:ActaPalaeontolRom.2019-.journal_article" docStyleId="CA474AAD92E97C533D63E0C5C0F159F1" docStyleName="ActaPalaeontolRom.2019-.journal_article" docStyleVersion="1" docTitle="Gusicella minima SCHLAGINTWEIT & RASHIDI 2021, comb. nov." docType="treatment" docVersion="2" lastPageNumber="12" masterDocId="FF81FFBA2B4F4E66FF9CD04CB245D838" masterDocTitle="Dictyoconella Henson, 1948, Upper Cretaceous Larger Benthic Foraminifera: A Taxonomic Revision With The Establishment Of Gusicella Gen. Nov. (Type-Species Dictyoconella Minima Henson)" masterLastPageNumber="13" masterPageNumber="3" pageNumber="10" updateTime="1719587089633" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9B821ECFF5485AF233611288074029EE">Dictyoconella Henson, 1948, Upper Cretaceous Larger Benthic Foraminifera: A Taxonomic Revision With The Establishment Of Gusicella Gen. Nov. (Type-Species Dictyoconella Minima Henson)</mods:title>
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<mods:namePart id="6DA73197EE8B7BF87A4CC1032D868440">SCHLAGINTWEIT, FELIX</mods:namePart>
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<mods:namePart id="7A653C9582125B186D7B1E915921444D">RASHIDI, KOOROSH</mods:namePart>
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<mods:title id="75C0D5A33969530AC3A8D56E8B905DC7">Acta Palaeontologica Romaniae</mods:title>
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<mods:date id="209D17278E8E11BC52DE2F8AFBB539C0">2021</mods:date>
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<treatment id="03B887C22B484E6FFCACD6B8B71ADA5C" LSID="urn:lsid:plazi:treatment:03B887C22B484E6FFCACD6B8B71ADA5C" httpUri="http://treatment.plazi.org/id/03B887C22B484E6FFCACD6B8B71ADA5C" lastPageId="9" lastPageNumber="12" pageId="7" pageNumber="10">
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<subSubSection id="C30B655F2B484E61FCACD6B8B75CDF33" box="[816,1305,1780,1804]" pageId="7" pageNumber="10" type="nomenclature">
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<paragraph id="8BAE36D42B484E61FCACD6B8B75CDF33" blockId="7.[816,1305,1780,1804]" box="[816,1305,1780,1804]" pageId="7" pageNumber="10">
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<heading id="D0E681B82B484E61FCACD6B8B75CDF33" box="[816,1305,1780,1804]" fontSize="10" level="2" pageId="7" pageNumber="10" reason="5">
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<taxonomicName id="4C114D572B484E61FCACD6B8B6DFDF34" ID-CoL="82PFG" authority="(Henson, 1948)" authorityName="SCHLAGINTWEIT & RASHIDI" authorityYear="2021" baseAuthorityName="SCHLAGINTWEIT & RASHIDI" baseAuthorityYear="1948" box="[816,1178,1780,1804]" class="Globothalamea" family="Orbitolinidae" genus="Gusicella" kingdom="Chromista" order="Loftusiida" pageId="7" pageNumber="10" phylum="Foraminifera" rank="species" species="minima" status="comb. nov.">
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<emphasis id="B965EAC62B484E61FCACD6B8B1A8DF34" box="[816,1005,1780,1804]" italics="true" pageId="7" pageNumber="10">Gusicella minima</emphasis>
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(
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<bibRefCitation id="EF804B252B484E61FC67D6B8B6D7DF34" author="Henson, F. R. S." box="[1019,1170,1780,1804]" pageId="7" pageNumber="10" pagination="1 - 127" refId="ref5977" refString="Henson, F. R. S., 1948. Larger imperforate Foraminifera of south-western Asia. Families Lituolidae, Orbitolinidae and Meandropsinidae. London, Monograph British Museum (Natural History): 1 - 127." type="book chapter" year="1948">Henson, 1948</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A25657BD2B484E61FB3DD6B8B75CDF33" box="[1185,1305,1780,1804]" pageId="7" pageNumber="10" rank="species">comb. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30B655F2B484E6FFCACD759B3C1DF57" lastPageId="9" lastPageNumber="12" pageId="7" pageNumber="10" type="description">
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<paragraph id="8BAE36D42B484E61FCACD759B189DF15" blockId="7.[816,972,1813,1837]" box="[816,972,1813,1837]" pageId="7" pageNumber="10">
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<figureCitation id="132A2A512B484E61FCACD759B13CDF15" box="[816,889,1813,1837]" captionStart="Fig" captionStartId="1.[240,275,1053,1074]" captionTargetBox="[251,1385,154,1015]" captionTargetId="figure-864@1.[251,1385,154,1015]" captionTargetPageId="1" captionText="Fig. 2 Microfacies with Dictyoconella complanata Henson (a-b) and Gusicella minima (Henson) gen. et comb. nov. (c- d) from the Maastrichtian Tarbur Formation of the Naghan (a-b) and Mandegan sections (c-d), SW Iran (Zagros zone). Abbreviations: D = Dictyoconella, Di = Dicyclina, G = Gyroconulina, Gu = Gusicella, L = Loftusia, O = Omphalocyclus, P = Pseudonummoloculina, Ps = Pseudocymopolia, T = Tarburina. Thin sections: 2Ng 81 (a), 2Ng 81-4 (b), Rt 109 (c), Rt 105 (d)." figureDoi="http://doi.org/10.5281/zenodo.12583326" httpUri="https://zenodo.org/record/12583326/files/figure.png" pageId="7" pageNumber="10">Figs. 2</figureCitation>
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c-d, 7-9
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</paragraph>
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<paragraph id="8BAE36D42B484E61FCACD71BB193DFA8" blockId="7.[816,1437,1879,2068]" pageId="7" pageNumber="10">
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*1948
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<taxonomicName id="4C114D572B484E61FCE2D71BB628DF57" authorityName="SCHLAGINTWEIT & RASHIDI" authorityYear="2021" box="[894,1133,1879,1903]" class="Globothalamea" family="Orbitolinidae" genus="Dictyoconella" kingdom="Chromista" order="Loftusiida" pageId="7" pageNumber="10" phylum="Foraminifera" rank="species" species="minima" status="sp. nov.">
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<emphasis id="B965EAC62B484E61FCE2D71BB628DF57" box="[894,1133,1879,1903]" italics="true" pageId="7" pageNumber="10">Dictyoconella minima</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25657BD2B484E61FBEBD714B6F1DF57" box="[1143,1204,1880,1903]" pageId="7" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
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– Henson, p. 25, pl. 11, fig. 3, 8-10.
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</paragraph>
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<paragraph id="8BAE36D42B484E61FCACD7D5B69BDFEA" blockId="7.[816,1437,1879,2068]" pageId="7" pageNumber="10">
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1998
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<taxonomicName id="4C114D572B484E61FCF1D7D5B6F7DF89" authority="Henson" authorityName="Henson" authorityYear="1948" box="[877,1202,1945,1969]" class="Globothalamea" family="Orbitolinidae" genus="Dictyoconella" kingdom="Chromista" order="Loftusiida" pageId="7" pageNumber="10" phylum="Foraminifera" rank="species" species="minima">
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<emphasis id="B965EAC62B484E61FCF1D7D5B61FDF89" box="[877,1114,1945,1969]" italics="true" pageId="7" pageNumber="10">Dictyoconella minima</emphasis>
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Henson
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</taxonomicName>
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– Whittaker et al., pl. 4, fig. 3, pl. 5, figs. 1-2, pl. 48, figs. 1-4.
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</paragraph>
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<paragraph id="8BAE36D42B484E61FCACD797B61CD02C" blockId="7.[816,1437,1879,2068]" pageId="7" pageNumber="10">
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2008
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<taxonomicName id="4C114D572B484E61FCEFD797B686DFCB" authority="Henson" authorityName="Henson" authorityYear="1948" box="[883,1219,2011,2035]" class="Globothalamea" family="Orbitolinidae" genus="Dictyoconella" kingdom="Chromista" order="Loftusiida" pageId="7" pageNumber="10" phylum="Foraminifera" rank="species" species="minima">
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<emphasis id="B965EAC62B484E61FCEFD797B620DFCB" box="[883,1125,2011,2035]" italics="true" pageId="7" pageNumber="10">Dictyoconella minima</emphasis>
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Henson
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</taxonomicName>
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– Schlagintweit et al., p.34, fig. 3a-b pars, 5-6.
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</paragraph>
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<caption id="DF6E665C2B474E6EFF7BD6FDB638DFEB" ID-DOI="http://doi.org/10.5281/zenodo.12583342" ID-Zenodo-Dep="12583342" httpUri="https://zenodo.org/record/12583342/files/figure.png" pageId="8" pageNumber="11" startId="8.[231,266,1713,1734]" targetBox="[231,1408,152,1671]" targetPageId="8" targetType="figure">
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<paragraph id="8BAE36D42B474E6EFF7BD6FDB638DFEB" blockId="8.[231,1409,1713,2003]" pageId="8" pageNumber="11">
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<emphasis id="B965EAC62B474E6EFF7BD6FDB364DEFE" bold="true" box="[231,289,1713,1734]" pageId="8" pageNumber="11">Fig. 9</emphasis>
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<taxonomicName id="4C114D572B474E6EFEB4D6FDB076DEFF" authority="(Henson)" authorityName="SCHLAGINTWEIT & RASHIDI" authorityYear="2021" baseAuthorityName="SCHLAGINTWEIT & RASHIDI" baseAuthorityYear="1948" box="[296,563,1713,1735]" class="Globothalamea" family="Orbitolinidae" genus="Gusicella" kingdom="Chromista" order="Loftusiida" pageId="8" pageNumber="11" phylum="Foraminifera" rank="species" species="minima" status="gen. et comb. nov.">
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<emphasis id="B965EAC62B474E6EFEB4D6FDB396DEFF" box="[296,467,1713,1735]" italics="true" pageId="8" pageNumber="11">Gusicella minima</emphasis>
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(Henson)
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</taxonomicName>
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<taxonomicNameLabel id="A25657BD2B474E6EFDA6D6FEB0AADEFF" box="[570,751,1713,1735]" pageId="8" pageNumber="11" rank="species">gen. et comb. nov.</taxonomicNameLabel>
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from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section:
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<emphasis id="B965EAC62B474E6EFEF1D683B382DEDC" bold="true" box="[365,455,1743,1764]" pageId="8" pageNumber="11">c–f, h–k)</emphasis>
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; Mandegan section:
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<emphasis id="B965EAC62B474E6EFD0DD683B0A6DEDC" bold="true" box="[657,739,1743,1764]" pageId="8" pageNumber="11">a–b, g, l</emphasis>
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).
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<emphasis id="B965EAC62B474E6EFD6BD683B146DEDC" bold="true" box="[759,771,1743,1764]" pageId="8" pageNumber="11">a</emphasis>
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subaxial section of an assumed microspheric specimen.
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<emphasis id="B965EAC62B474E6EFABBD683B70EDEDC" bold="true" box="[1319,1355,1743,1764]" pageId="8" pageNumber="11">b–c</emphasis>
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axial sections showing initial spire with acute margin beneath the apex.
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<emphasis id="B965EAC62B474E6EFCF4D6A1B130DF3A" bold="true" box="[872,885,1773,1794]" pageId="8" pageNumber="11">d</emphasis>
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oblique section showing the marginal trough between the marginal and central zones as well as a circular row of marginal apertures between.
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<emphasis id="B965EAC62B474E6EFBA1D747B60DDF18" bold="true" box="[1085,1096,1803,1824]" pageId="8" pageNumber="11">e</emphasis>
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tangential section showing subepidermal network; septa in the lower part marked by white dash lines.
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<emphasis id="B965EAC62B474E6EFC06D765B1E7DF06" bold="true" box="[922,930,1833,1854]" pageId="8" pageNumber="11">f</emphasis>
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subaxial section of a possibly microspheric specimen.
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<emphasis id="B965EAC62B474E6EFEB9D70AB374DF63" bold="true" box="[293,305,1862,1883]" pageId="8" pageNumber="11">g</emphasis>
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transverse section.
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<emphasis id="B965EAC62B474E6EFE6AD70AB046DF63" bold="true" box="[502,515,1862,1883]" pageId="8" pageNumber="11">h</emphasis>
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subaxial section.
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<emphasis id="B965EAC62B474E6EFD2AD70AB0F8DF63" bold="true" box="[694,701,1862,1883]" pageId="8" pageNumber="11">i</emphasis>
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tangential section showing aligned main beams.
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<emphasis id="B965EAC62B474E6EFB38D70AB6E9DF63" bold="true" box="[1188,1196,1862,1883]" pageId="8" pageNumber="11">j</emphasis>
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subaxial section of a possibly microspheric specimen.
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<emphasis id="B965EAC62B474E6EFDB4D728B070DF41" bold="true" box="[552,565,1892,1913]" pageId="8" pageNumber="11">k</emphasis>
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tangential section.
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<emphasis id="B965EAC62B474E6EFD6ED728B0BCDF41" bold="true" box="[754,761,1892,1913]" pageId="8" pageNumber="11">l</emphasis>
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slightly oblique transverse section showing several orders of vertical partitions (beams, intercalary beams), marginal trough, and pillared central zone. Abbreviations: m.f. = marginal foramen, m.t. = marginal through, pi = pillar. Thin sections: Rt 105 (
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<emphasis id="B965EAC62B474E6EFCE5D7ECB1C0DF8D" bold="true" box="[889,901,1952,1973]" pageId="8" pageNumber="11">a</emphasis>
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), 2Ng 17 (
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<emphasis id="B965EAC62B474E6EFC6FD7ECB1BBDF8D" bold="true" box="[1011,1022,1952,1973]" pageId="8" pageNumber="11">c</emphasis>
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), Rt 96 (
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<emphasis id="B965EAC62B474E6EFBC5D7ECB623DF8D" bold="true" box="[1113,1126,1952,1973]" pageId="8" pageNumber="11">b</emphasis>
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), 2Ng 167 (
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<emphasis id="B965EAC62B474E6EFB7DD7ECB6ABDF8D" bold="true" box="[1249,1262,1952,1973]" pageId="8" pageNumber="11">d</emphasis>
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), 2Ng 174 (
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<emphasis id="B965EAC62B474E6EFAF4D7ECB736DF8D" bold="true" box="[1384,1395,1952,1973]" pageId="8" pageNumber="11">e</emphasis>
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), 2Ng 118-1 (
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<emphasis id="B965EAC62B474E6EFEC1D7F1B320DFEA" bold="true" box="[349,357,1981,2002]" pageId="8" pageNumber="11">f</emphasis>
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), Rt 108-2b (
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<emphasis id="B965EAC62B474E6EFE7BD7F1B3B6DFEA" bold="true" box="[487,499,1981,2002]" pageId="8" pageNumber="11">g</emphasis>
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), 2Ng 169 (
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<emphasis id="B965EAC62B474E6EFDF4D7F1B030DFEA" bold="true" box="[616,629,1981,2002]" pageId="8" pageNumber="11">h</emphasis>
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), 2Ng 191 (
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<emphasis id="B965EAC62B474E6EFD77D7F1B0B7DFEA" bold="true" box="[747,754,1981,2002]" pageId="8" pageNumber="11">i</emphasis>
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), 2Ng 176 (
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<emphasis id="B965EAC62B474E6EFCFBD7F1B12ADFEA" bold="true" box="[871,879,1981,2002]" pageId="8" pageNumber="11">j</emphasis>
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), 2Ng 146 (
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<emphasis id="B965EAC62B474E6EFC78D7F1B1B4DFEA" bold="true" box="[996,1009,1981,2002]" pageId="8" pageNumber="11">k</emphasis>
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), Rt 111-1 (
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<emphasis id="B965EAC62B474E6EFBF5D7F1B635DFEA" bold="true" box="[1129,1136,1981,2002]" pageId="8" pageNumber="11">l</emphasis>
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).
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</paragraph>
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</caption>
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<paragraph id="8BAE36D42B464E6FFF0BD0D2B3F9DE7D" blockId="9.[151,772,158,2068]" pageId="9" pageNumber="12">
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<emphasis id="B965EAC62B464E6FFF0BD0D2B361D88E" bold="true" box="[151,292,158,182]" pageId="9" pageNumber="12">Description:</emphasis>
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Test medium conical (apical angle from 50 to 90 degrees); both sides and cone base slightly convex. Assumed microspheric specimens broader and larger (
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<figureCitation id="132A2A512B464E6FFF03D14DB2A8D921" box="[159,237,257,281]" captionStart="Fig" captionStartId="6.[206,241,1674,1695]" captionTargetBox="[219,1374,159,1632]" captionTargetId="figure-36@6.[213,1374,159,1632]" captionTargetPageId="6" captionText="Fig. 8 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section: a–b, j, l; Mandegan section: c–i, k, m–p). a Subaxial section. b, i, l Oblique sections. c–e, m Deep tangential sections displaying primary septules (beams) continuous (aligned) from one chamber to the next. f Shallow tangential section displaying subepidermal network of beams and rafters. g, j, l, o–p Different transverse sections, some slightly oblique. Note five marginal foramina arranged in a radial circle (inside yellow marked rectangle in o). Note oval test outline in j. Subaxial section. Note alternation of pillars by stairway-like arrangement of foramina (yellow line) and opaque fused pillars/secondary deposits in the central zone. k Detail from h showing marginal foramina arranged in vertical lines between successive chambers. n Tangential-oblique section. Abbreviations: b = beam, ib = intercalary beam, f = foramen, m.f. = marginal foramen, m.t. = marginal through, pi = pillar, s = septum. Thin sections: Ng 196 (a), Ng 192 (b), Rt 108-3 (c–e, g, n), Rt 104 (f), Rt 96 (h, k), Rt 108-6 (i), Ng 187 (j), Ng 186 (l), Rt 107 (m), Rt 102-2 (o), Rt 104 (p)." figureDoi="http://doi.org/10.5281/zenodo.12583338" httpUri="https://zenodo.org/record/12583338/files/figure.png" pageId="9" pageNumber="12">Fig. 8a</figureCitation>
|
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,
|
||
<figureCitation id="132A2A512B464E6FFF65D14DB318D921" box="[249,349,257,281]" captionStart="Fig" captionStartId="8.[231,266,1713,1734]" captionTargetBox="[231,1408,152,1671]" captionTargetId="figure-34@8.[231,1408,152,1671]" captionTargetPageId="8" captionText="Fig. 9 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section: c–f, h–k); Mandegan section: a–b, g, l). a subaxial section of an assumed microspheric specimen. b–c axial sections showing initial spire with acute margin beneath the apex. d oblique section showing the marginal trough between the marginal and central zones as well as a circular row of marginal apertures between. e tangential section showing subepidermal network; septa in the lower part marked by white dash lines. f subaxial section of a possibly microspheric specimen. g transverse section. h subaxial section. i tangential section showing aligned main beams. j subaxial section of a possibly microspheric specimen. k tangential section. l slightly oblique transverse section showing several orders of vertical partitions (beams, intercalary beams), marginal trough, and pillared central zone. Abbreviations: m.f. = marginal foramen, m.t. = marginal through, pi = pillar. Thin sections: Rt 105 (a), 2Ng 17 (c), Rt 96 (b), 2Ng 167 (d), 2Ng 174 (e), 2Ng 118-1 (f), Rt 108-2b (g), 2Ng 169 (h), 2Ng 191 (i), 2Ng 176 (j), 2Ng 146 (k), Rt 111-1 (l)." figureDoi="http://doi.org/10.5281/zenodo.12583342" httpUri="https://zenodo.org/record/12583342/files/figure.png" pageId="9" pageNumber="12">9a, 9f, 9j</figureCitation>
|
||
). Transverse sections circular (
|
||
<figureCitation id="132A2A512B464E6FFD31D14DB0BED921" box="[685,763,257,281]" captionStart="Fig" captionStartId="6.[206,241,1674,1695]" captionTargetBox="[219,1374,159,1632]" captionTargetId="figure-36@6.[213,1374,159,1632]" captionTargetPageId="6" captionText="Fig. 8 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section: a–b, j, l; Mandegan section: c–i, k, m–p). a Subaxial section. b, i, l Oblique sections. c–e, m Deep tangential sections displaying primary septules (beams) continuous (aligned) from one chamber to the next. f Shallow tangential section displaying subepidermal network of beams and rafters. g, j, l, o–p Different transverse sections, some slightly oblique. Note five marginal foramina arranged in a radial circle (inside yellow marked rectangle in o). Note oval test outline in j. Subaxial section. Note alternation of pillars by stairway-like arrangement of foramina (yellow line) and opaque fused pillars/secondary deposits in the central zone. k Detail from h showing marginal foramina arranged in vertical lines between successive chambers. n Tangential-oblique section. Abbreviations: b = beam, ib = intercalary beam, f = foramen, m.f. = marginal foramen, m.t. = marginal through, pi = pillar, s = septum. Thin sections: Ng 196 (a), Ng 192 (b), Rt 108-3 (c–e, g, n), Rt 104 (f), Rt 96 (h, k), Rt 108-6 (i), Ng 187 (j), Ng 186 (l), Rt 107 (m), Rt 102-2 (o), Rt 104 (p)." figureDoi="http://doi.org/10.5281/zenodo.12583338" httpUri="https://zenodo.org/record/12583338/files/figure.png" pageId="9" pageNumber="12">Fig. 8p</figureCitation>
|
||
) or ovoidal-compressed (
|
||
<figureCitation id="132A2A512B464E6FFE3AD16EB3B0D902" box="[422,501,290,314]" captionStart="Fig" captionStartId="5.[149,184,1697,1718]" captionTargetBox="[149,1438,157,1655]" captionTargetId="figure-41@5.[149,1438,157,1655]" captionTargetPageId="5" captionText="Fig. 7 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Mandegan section: b, l; Naghan section: a, c–k). a Oblique transverse section. b Oblique section cutting the marginal (upper part) and central zones (lower part) displaying irregularly distributed fused pillars/secondary deposits. c, k Subaxial sections. d–g, Axial sections showing initial spire. h–i Details (from e and g) of the initial spire. j Tangential section. Slightly oblique transverse sections showing subdivision of the marginal zone and circle of marginal apertures (left and above). Abbreviations: m.f. = marginal foramen, m.t. = marginal trough, pr = protoconch, s.d. = secondary deposits/fused pillars. Thin sections: 2Ng 17 (a, k), Rt 100 (b), Ng 180-1 (c), 2Ng 179-1 (d), 2Ng 168 (e, h), 2Ng 175 (f), 2Ng 174 (g, i), 2Ng 177 (j), Rt 113 (l)." figureDoi="http://doi.org/10.5281/zenodo.12583336" httpUri="https://zenodo.org/record/12583336/files/figure.png" pageId="9" pageNumber="12">Fig. 7l</figureCitation>
|
||
,
|
||
<figureCitation id="132A2A512B464E6FFD9BD16EB05AD902" box="[519,543,290,314]" captionStart="Fig" captionStartId="6.[206,241,1674,1695]" captionTargetBox="[219,1374,159,1632]" captionTargetId="figure-36@6.[213,1374,159,1632]" captionTargetPageId="6" captionText="Fig. 8 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section: a–b, j, l; Mandegan section: c–i, k, m–p). a Subaxial section. b, i, l Oblique sections. c–e, m Deep tangential sections displaying primary septules (beams) continuous (aligned) from one chamber to the next. f Shallow tangential section displaying subepidermal network of beams and rafters. g, j, l, o–p Different transverse sections, some slightly oblique. Note five marginal foramina arranged in a radial circle (inside yellow marked rectangle in o). Note oval test outline in j. Subaxial section. Note alternation of pillars by stairway-like arrangement of foramina (yellow line) and opaque fused pillars/secondary deposits in the central zone. k Detail from h showing marginal foramina arranged in vertical lines between successive chambers. n Tangential-oblique section. Abbreviations: b = beam, ib = intercalary beam, f = foramen, m.f. = marginal foramen, m.t. = marginal through, pi = pillar, s = septum. Thin sections: Ng 196 (a), Ng 192 (b), Rt 108-3 (c–e, g, n), Rt 104 (f), Rt 96 (h, k), Rt 108-6 (i), Ng 187 (j), Ng 186 (l), Rt 107 (m), Rt 102-2 (o), Rt 104 (p)." figureDoi="http://doi.org/10.5281/zenodo.12583338" httpUri="https://zenodo.org/record/12583338/files/figure.png" pageId="9" pageNumber="12">8j</figureCitation>
|
||
). The initial part is characterized by a small spire of few chambers arranged in half a whorl that is closely attaching (almost parallel) to the side test wall (
|
||
<figureCitation id="132A2A512B464E6FFEE2D1C9B047D9A5" box="[382,514,389,414]" captionStart="Fig" captionStartId="5.[149,184,1697,1718]" captionTargetBox="[149,1438,157,1655]" captionTargetId="figure-41@5.[149,1438,157,1655]" captionTargetPageId="5" captionText="Fig. 7 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Mandegan section: b, l; Naghan section: a, c–k). a Oblique transverse section. b Oblique section cutting the marginal (upper part) and central zones (lower part) displaying irregularly distributed fused pillars/secondary deposits. c, k Subaxial sections. d–g, Axial sections showing initial spire. h–i Details (from e and g) of the initial spire. j Tangential section. Slightly oblique transverse sections showing subdivision of the marginal zone and circle of marginal apertures (left and above). Abbreviations: m.f. = marginal foramen, m.t. = marginal trough, pr = protoconch, s.d. = secondary deposits/fused pillars. Thin sections: 2Ng 17 (a, k), Rt 100 (b), Ng 180-1 (c), 2Ng 179-1 (d), 2Ng 168 (e, h), 2Ng 175 (f), 2Ng 174 (g, i), 2Ng 177 (j), Rt 113 (l)." figureDoi="http://doi.org/10.5281/zenodo.12583336" httpUri="https://zenodo.org/record/12583336/files/figure.png" pageId="9" pageNumber="12">Fig. 7d, g, i</figureCitation>
|
||
,
|
||
<figureCitation id="132A2A512B464E6FFD8CD1C9B06ED9A5" box="[528,555,389,413]" captionStart="Fig" captionStartId="8.[231,266,1713,1734]" captionTargetBox="[231,1408,152,1671]" captionTargetId="figure-34@8.[231,1408,152,1671]" captionTargetPageId="8" captionText="Fig. 9 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section: c–f, h–k); Mandegan section: a–b, g, l). a subaxial section of an assumed microspheric specimen. b–c axial sections showing initial spire with acute margin beneath the apex. d oblique section showing the marginal trough between the marginal and central zones as well as a circular row of marginal apertures between. e tangential section showing subepidermal network; septa in the lower part marked by white dash lines. f subaxial section of a possibly microspheric specimen. g transverse section. h subaxial section. i tangential section showing aligned main beams. j subaxial section of a possibly microspheric specimen. k tangential section. l slightly oblique transverse section showing several orders of vertical partitions (beams, intercalary beams), marginal trough, and pillared central zone. Abbreviations: m.f. = marginal foramen, m.t. = marginal through, pi = pillar. Thin sections: Rt 105 (a), 2Ng 17 (c), Rt 96 (b), 2Ng 167 (d), 2Ng 174 (e), 2Ng 118-1 (f), Rt 108-2b (g), 2Ng 169 (h), 2Ng 191 (i), 2Ng 176 (j), 2Ng 146 (k), Rt 111-1 (l)." figureDoi="http://doi.org/10.5281/zenodo.12583342" httpUri="https://zenodo.org/record/12583342/files/figure.png" pageId="9" pageNumber="12">9b</figureCitation>
|
||
). The acute apex is tilted, marking the early development in a downwards turning to the cone base (e.g.
|
||
<figureCitation id="132A2A512B464E6FFE49D184B065D9D8" box="[469,544,456,480]" captionStart="Fig" captionStartId="5.[149,184,1697,1718]" captionTargetBox="[149,1438,157,1655]" captionTargetId="figure-41@5.[149,1438,157,1655]" captionTargetPageId="5" captionText="Fig. 7 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Mandegan section: b, l; Naghan section: a, c–k). a Oblique transverse section. b Oblique section cutting the marginal (upper part) and central zones (lower part) displaying irregularly distributed fused pillars/secondary deposits. c, k Subaxial sections. d–g, Axial sections showing initial spire. h–i Details (from e and g) of the initial spire. j Tangential section. Slightly oblique transverse sections showing subdivision of the marginal zone and circle of marginal apertures (left and above). Abbreviations: m.f. = marginal foramen, m.t. = marginal trough, pr = protoconch, s.d. = secondary deposits/fused pillars. Thin sections: 2Ng 17 (a, k), Rt 100 (b), Ng 180-1 (c), 2Ng 179-1 (d), 2Ng 168 (e, h), 2Ng 175 (f), 2Ng 174 (g, i), 2Ng 177 (j), Rt 113 (l)." figureDoi="http://doi.org/10.5281/zenodo.12583336" httpUri="https://zenodo.org/record/12583336/files/figure.png" pageId="9" pageNumber="12">
|
||
Fig.
|
||
<quantity id="4CE99B312B464E6FFD9AD184B065D9D8" box="[518,544,456,480]" metricMagnitude="-3" metricUnit="kg" metricValue="7.0" pageId="9" pageNumber="12" unit="g" value="7.0">7g</quantity>
|
||
</figureCitation>
|
||
). The megalospheric embryo appears as a single subspherical chamber (protoconch) showing short septules at its upper part facing the cone base (
|
||
<figureCitation id="132A2A512B464E6FFE88D267B313DA7B" box="[276,342,555,579]" captionStart="Fig" captionStartId="5.[149,184,1697,1718]" captionTargetBox="[149,1438,157,1655]" captionTargetId="figure-41@5.[149,1438,157,1655]" captionTargetPageId="5" captionText="Fig. 7 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Mandegan section: b, l; Naghan section: a, c–k). a Oblique transverse section. b Oblique section cutting the marginal (upper part) and central zones (lower part) displaying irregularly distributed fused pillars/secondary deposits. c, k Subaxial sections. d–g, Axial sections showing initial spire. h–i Details (from e and g) of the initial spire. j Tangential section. Slightly oblique transverse sections showing subdivision of the marginal zone and circle of marginal apertures (left and above). Abbreviations: m.f. = marginal foramen, m.t. = marginal trough, pr = protoconch, s.d. = secondary deposits/fused pillars. Thin sections: 2Ng 17 (a, k), Rt 100 (b), Ng 180-1 (c), 2Ng 179-1 (d), 2Ng 168 (e, h), 2Ng 175 (f), 2Ng 174 (g, i), 2Ng 177 (j), Rt 113 (l)." figureDoi="http://doi.org/10.5281/zenodo.12583336" httpUri="https://zenodo.org/record/12583336/files/figure.png" pageId="9" pageNumber="12">Fig. 7</figureCitation>
|
||
g-i). The chambers are subdivided into marginal and central zones. The exoskeleton consists of horizontal (rafters) and radial vertical partitions (beams) subdividing the chamber margins. There are one to three rafters in the marginal zone (e.g.
|
||
<figureCitation id="132A2A512B464E6FFD98D2E3B017DAFF" box="[516,594,687,711]" captionStart="Fig" captionStartId="5.[149,184,1697,1718]" captionTargetBox="[149,1438,157,1655]" captionTargetId="figure-41@5.[149,1438,157,1655]" captionTargetPageId="5" captionText="Fig. 7 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Mandegan section: b, l; Naghan section: a, c–k). a Oblique transverse section. b Oblique section cutting the marginal (upper part) and central zones (lower part) displaying irregularly distributed fused pillars/secondary deposits. c, k Subaxial sections. d–g, Axial sections showing initial spire. h–i Details (from e and g) of the initial spire. j Tangential section. Slightly oblique transverse sections showing subdivision of the marginal zone and circle of marginal apertures (left and above). Abbreviations: m.f. = marginal foramen, m.t. = marginal trough, pr = protoconch, s.d. = secondary deposits/fused pillars. Thin sections: 2Ng 17 (a, k), Rt 100 (b), Ng 180-1 (c), 2Ng 179-1 (d), 2Ng 168 (e, h), 2Ng 175 (f), 2Ng 174 (g, i), 2Ng 177 (j), Rt 113 (l)." figureDoi="http://doi.org/10.5281/zenodo.12583336" httpUri="https://zenodo.org/record/12583336/files/figure.png" pageId="9" pageNumber="12">Fig. 7e</figureCitation>
|
||
). In case of two rafters, the upper one (in direction to the apex) is twice as long as the lower one. In case of three rafters, the middle one is roughly twice as long as the others. There are two to
|
||
<specimenCount id="9D17FD5D2B464E6FFF28D378B326DB74" box="[180,355,820,844]" count="3" pageId="9" pageNumber="12" type="intercalary">three intercalary</specimenCount>
|
||
beams between the longer main partitions (beams) (
|
||
<figureCitation id="132A2A512B464E6FFED8D319B3DFDB55" box="[324,410,853,877]" captionStart="Fig" captionStartId="6.[206,241,1674,1695]" captionTargetBox="[219,1374,159,1632]" captionTargetId="figure-36@6.[213,1374,159,1632]" captionTargetPageId="6" captionText="Fig. 8 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section: a–b, j, l; Mandegan section: c–i, k, m–p). a Subaxial section. b, i, l Oblique sections. c–e, m Deep tangential sections displaying primary septules (beams) continuous (aligned) from one chamber to the next. f Shallow tangential section displaying subepidermal network of beams and rafters. g, j, l, o–p Different transverse sections, some slightly oblique. Note five marginal foramina arranged in a radial circle (inside yellow marked rectangle in o). Note oval test outline in j. Subaxial section. Note alternation of pillars by stairway-like arrangement of foramina (yellow line) and opaque fused pillars/secondary deposits in the central zone. k Detail from h showing marginal foramina arranged in vertical lines between successive chambers. n Tangential-oblique section. Abbreviations: b = beam, ib = intercalary beam, f = foramen, m.f. = marginal foramen, m.t. = marginal through, pi = pillar, s = septum. Thin sections: Ng 196 (a), Ng 192 (b), Rt 108-3 (c–e, g, n), Rt 104 (f), Rt 96 (h, k), Rt 108-6 (i), Ng 187 (j), Ng 186 (l), Rt 107 (m), Rt 102-2 (o), Rt 104 (p)." figureDoi="http://doi.org/10.5281/zenodo.12583338" httpUri="https://zenodo.org/record/12583338/files/figure.png" pageId="9" pageNumber="12">Fig. 8p</figureCitation>
|
||
,
|
||
<figureCitation id="132A2A512B464E6FFE30D319B38FDB55" box="[428,458,853,877]" captionStart="Fig" captionStartId="8.[231,266,1713,1734]" captionTargetBox="[231,1408,152,1671]" captionTargetId="figure-34@8.[231,1408,152,1671]" captionTargetPageId="8" captionText="Fig. 9 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section: c–f, h–k); Mandegan section: a–b, g, l). a subaxial section of an assumed microspheric specimen. b–c axial sections showing initial spire with acute margin beneath the apex. d oblique section showing the marginal trough between the marginal and central zones as well as a circular row of marginal apertures between. e tangential section showing subepidermal network; septa in the lower part marked by white dash lines. f subaxial section of a possibly microspheric specimen. g transverse section. h subaxial section. i tangential section showing aligned main beams. j subaxial section of a possibly microspheric specimen. k tangential section. l slightly oblique transverse section showing several orders of vertical partitions (beams, intercalary beams), marginal trough, and pillared central zone. Abbreviations: m.f. = marginal foramen, m.t. = marginal through, pi = pillar. Thin sections: Rt 105 (a), 2Ng 17 (c), Rt 96 (b), 2Ng 167 (d), 2Ng 174 (e), 2Ng 118-1 (f), Rt 108-2b (g), 2Ng 169 (h), 2Ng 191 (i), 2Ng 176 (j), 2Ng 146 (k), Rt 111-1 (l)." figureDoi="http://doi.org/10.5281/zenodo.12583342" httpUri="https://zenodo.org/record/12583342/files/figure.png" pageId="9" pageNumber="12">9d</figureCitation>
|
||
). The latter are continuous from one chamber to the next (
|
||
<figureCitation id="132A2A512B464E6FFE75D33AB02CDBB6" box="[489,617,886,910]" captionStart="Fig" captionStartId="6.[206,241,1674,1695]" captionTargetBox="[219,1374,159,1632]" captionTargetId="figure-36@6.[213,1374,159,1632]" captionTargetPageId="6" captionText="Fig. 8 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section: a–b, j, l; Mandegan section: c–i, k, m–p). a Subaxial section. b, i, l Oblique sections. c–e, m Deep tangential sections displaying primary septules (beams) continuous (aligned) from one chamber to the next. f Shallow tangential section displaying subepidermal network of beams and rafters. g, j, l, o–p Different transverse sections, some slightly oblique. Note five marginal foramina arranged in a radial circle (inside yellow marked rectangle in o). Note oval test outline in j. Subaxial section. Note alternation of pillars by stairway-like arrangement of foramina (yellow line) and opaque fused pillars/secondary deposits in the central zone. k Detail from h showing marginal foramina arranged in vertical lines between successive chambers. n Tangential-oblique section. Abbreviations: b = beam, ib = intercalary beam, f = foramen, m.f. = marginal foramen, m.t. = marginal through, pi = pillar, s = septum. Thin sections: Ng 196 (a), Ng 192 (b), Rt 108-3 (c–e, g, n), Rt 104 (f), Rt 96 (h, k), Rt 108-6 (i), Ng 187 (j), Ng 186 (l), Rt 107 (m), Rt 102-2 (o), Rt 104 (p)." figureDoi="http://doi.org/10.5281/zenodo.12583338" httpUri="https://zenodo.org/record/12583338/files/figure.png" pageId="9" pageNumber="12">
|
||
Fig. 8c,
|
||
<quantity id="4CE99B312B464E6FFDD8D33AB02CDBB6" box="[580,617,886,910]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="12" unit="m" value="8.0">8m</quantity>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="132A2A512B464E6FFDE9D33AB0CFDBB6" box="[629,650,886,910]" captionStart="Fig" captionStartId="8.[231,266,1713,1734]" captionTargetBox="[231,1408,152,1671]" captionTargetId="figure-34@8.[231,1408,152,1671]" captionTargetPageId="8" captionText="Fig. 9 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section: c–f, h–k); Mandegan section: a–b, g, l). a subaxial section of an assumed microspheric specimen. b–c axial sections showing initial spire with acute margin beneath the apex. d oblique section showing the marginal trough between the marginal and central zones as well as a circular row of marginal apertures between. e tangential section showing subepidermal network; septa in the lower part marked by white dash lines. f subaxial section of a possibly microspheric specimen. g transverse section. h subaxial section. i tangential section showing aligned main beams. j subaxial section of a possibly microspheric specimen. k tangential section. l slightly oblique transverse section showing several orders of vertical partitions (beams, intercalary beams), marginal trough, and pillared central zone. Abbreviations: m.f. = marginal foramen, m.t. = marginal through, pi = pillar. Thin sections: Rt 105 (a), 2Ng 17 (c), Rt 96 (b), 2Ng 167 (d), 2Ng 174 (e), 2Ng 118-1 (f), Rt 108-2b (g), 2Ng 169 (h), 2Ng 191 (i), 2Ng 176 (j), 2Ng 146 (k), Rt 111-1 (l)." figureDoi="http://doi.org/10.5281/zenodo.12583342" httpUri="https://zenodo.org/record/12583342/files/figure.png" pageId="9" pageNumber="12">9i</figureCitation>
|
||
). They extend to the undivided marginal through marking the boundary to the central zone (
|
||
<figureCitation id="132A2A512B464E6FFE44D3F4B060DBE8" box="[472,549,952,976]" captionStart="Fig" captionStartId="5.[149,184,1697,1718]" captionTargetBox="[149,1438,157,1655]" captionTargetId="figure-41@5.[149,1438,157,1655]" captionTargetPageId="5" captionText="Fig. 7 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Mandegan section: b, l; Naghan section: a, c–k). a Oblique transverse section. b Oblique section cutting the marginal (upper part) and central zones (lower part) displaying irregularly distributed fused pillars/secondary deposits. c, k Subaxial sections. d–g, Axial sections showing initial spire. h–i Details (from e and g) of the initial spire. j Tangential section. Slightly oblique transverse sections showing subdivision of the marginal zone and circle of marginal apertures (left and above). Abbreviations: m.f. = marginal foramen, m.t. = marginal trough, pr = protoconch, s.d. = secondary deposits/fused pillars. Thin sections: 2Ng 17 (a, k), Rt 100 (b), Ng 180-1 (c), 2Ng 179-1 (d), 2Ng 168 (e, h), 2Ng 175 (f), 2Ng 174 (g, i), 2Ng 177 (j), Rt 113 (l)." figureDoi="http://doi.org/10.5281/zenodo.12583336" httpUri="https://zenodo.org/record/12583336/files/figure.png" pageId="9" pageNumber="12">Fig. 7a</figureCitation>
|
||
,
|
||
<figureCitation id="132A2A512B464E6FFDAED3F4B00ADBE8" box="[562,591,952,976]" captionStart="Fig" captionStartId="8.[231,266,1713,1734]" captionTargetBox="[231,1408,152,1671]" captionTargetId="figure-34@8.[231,1408,152,1671]" captionTargetPageId="8" captionText="Fig. 9 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section: c–f, h–k); Mandegan section: a–b, g, l). a subaxial section of an assumed microspheric specimen. b–c axial sections showing initial spire with acute margin beneath the apex. d oblique section showing the marginal trough between the marginal and central zones as well as a circular row of marginal apertures between. e tangential section showing subepidermal network; septa in the lower part marked by white dash lines. f subaxial section of a possibly microspheric specimen. g transverse section. h subaxial section. i tangential section showing aligned main beams. j subaxial section of a possibly microspheric specimen. k tangential section. l slightly oblique transverse section showing several orders of vertical partitions (beams, intercalary beams), marginal trough, and pillared central zone. Abbreviations: m.f. = marginal foramen, m.t. = marginal through, pi = pillar. Thin sections: Rt 105 (a), 2Ng 17 (c), Rt 96 (b), 2Ng 167 (d), 2Ng 174 (e), 2Ng 118-1 (f), Rt 108-2b (g), 2Ng 169 (h), 2Ng 191 (i), 2Ng 176 (j), 2Ng 146 (k), Rt 111-1 (l)." figureDoi="http://doi.org/10.5281/zenodo.12583342" httpUri="https://zenodo.org/record/12583342/files/figure.png" pageId="9" pageNumber="12">9d</figureCitation>
|
||
). Here, the septa slightly bend upwards in direction to the apex forming a “buttress on marginal ridge”
|
||
<emphasis id="B965EAC62B464E6FFE4ED3B7B048DC2A" box="[466,525,1019,1042]" italics="true" pageId="9" pageNumber="12">sensu</emphasis>
|
||
<bibRefCitation id="EF804B252B464E6FFD89D3B6B0EBDC2A" author="Davies, L. M." box="[533,686,1018,1042]" pageId="9" pageNumber="12" pagination="485 - 505" refId="ref5785" refString="Davies, L. M., 1930. The genus Dictyoconus and its allies: A review of the group, together with a description of three new species from the Lower Eocene beds of northern Baluchistan. Transactions of the Royal Society of Edinburgh 56 part II (no. 20): 485 - 505." type="journal article" year="1930">Davies (1930)</bibRefCitation>
|
||
as seen in axial sections (
|
||
<figureCitation id="132A2A512B464E6FFEFAD457B3F8DC0B" box="[358,445,1051,1075]" captionStart="Fig" captionStartId="6.[206,241,1674,1695]" captionTargetBox="[219,1374,159,1632]" captionTargetId="figure-36@6.[213,1374,159,1632]" captionTargetPageId="6" captionText="Fig. 8 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section: a–b, j, l; Mandegan section: c–i, k, m–p). a Subaxial section. b, i, l Oblique sections. c–e, m Deep tangential sections displaying primary septules (beams) continuous (aligned) from one chamber to the next. f Shallow tangential section displaying subepidermal network of beams and rafters. g, j, l, o–p Different transverse sections, some slightly oblique. Note five marginal foramina arranged in a radial circle (inside yellow marked rectangle in o). Note oval test outline in j. Subaxial section. Note alternation of pillars by stairway-like arrangement of foramina (yellow line) and opaque fused pillars/secondary deposits in the central zone. k Detail from h showing marginal foramina arranged in vertical lines between successive chambers. n Tangential-oblique section. Abbreviations: b = beam, ib = intercalary beam, f = foramen, m.f. = marginal foramen, m.t. = marginal through, pi = pillar, s = septum. Thin sections: Ng 196 (a), Ng 192 (b), Rt 108-3 (c–e, g, n), Rt 104 (f), Rt 96 (h, k), Rt 108-6 (i), Ng 187 (j), Ng 186 (l), Rt 107 (m), Rt 102-2 (o), Rt 104 (p)." figureDoi="http://doi.org/10.5281/zenodo.12583338" httpUri="https://zenodo.org/record/12583338/files/figure.png" pageId="9" pageNumber="12">Fig. 8k</figureCitation>
|
||
). Shallow-tangential sections display a pattern of subrounded alveolar compartments (subepidermal network) (
|
||
<figureCitation id="132A2A512B464E6FFE32D411B3BEDC4D" box="[430,507,1117,1141]" captionStart="Fig" captionStartId="5.[149,184,1697,1718]" captionTargetBox="[149,1438,157,1655]" captionTargetId="figure-41@5.[149,1438,157,1655]" captionTargetPageId="5" captionText="Fig. 7 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Mandegan section: b, l; Naghan section: a, c–k). a Oblique transverse section. b Oblique section cutting the marginal (upper part) and central zones (lower part) displaying irregularly distributed fused pillars/secondary deposits. c, k Subaxial sections. d–g, Axial sections showing initial spire. h–i Details (from e and g) of the initial spire. j Tangential section. Slightly oblique transverse sections showing subdivision of the marginal zone and circle of marginal apertures (left and above). Abbreviations: m.f. = marginal foramen, m.t. = marginal trough, pr = protoconch, s.d. = secondary deposits/fused pillars. Thin sections: 2Ng 17 (a, k), Rt 100 (b), Ng 180-1 (c), 2Ng 179-1 (d), 2Ng 168 (e, h), 2Ng 175 (f), 2Ng 174 (g, i), 2Ng 177 (j), Rt 113 (l)." figureDoi="http://doi.org/10.5281/zenodo.12583336" httpUri="https://zenodo.org/record/12583336/files/figure.png" pageId="9" pageNumber="12">Fig. 7j</figureCitation>
|
||
,
|
||
<figureCitation id="132A2A512B464E6FFD97D411B061DC4D" box="[523,548,1117,1141]" captionStart="Fig" captionStartId="6.[206,241,1674,1695]" captionTargetBox="[219,1374,159,1632]" captionTargetId="figure-36@6.[213,1374,159,1632]" captionTargetPageId="6" captionText="Fig. 8 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section: a–b, j, l; Mandegan section: c–i, k, m–p). a Subaxial section. b, i, l Oblique sections. c–e, m Deep tangential sections displaying primary septules (beams) continuous (aligned) from one chamber to the next. f Shallow tangential section displaying subepidermal network of beams and rafters. g, j, l, o–p Different transverse sections, some slightly oblique. Note five marginal foramina arranged in a radial circle (inside yellow marked rectangle in o). Note oval test outline in j. Subaxial section. Note alternation of pillars by stairway-like arrangement of foramina (yellow line) and opaque fused pillars/secondary deposits in the central zone. k Detail from h showing marginal foramina arranged in vertical lines between successive chambers. n Tangential-oblique section. Abbreviations: b = beam, ib = intercalary beam, f = foramen, m.f. = marginal foramen, m.t. = marginal through, pi = pillar, s = septum. Thin sections: Ng 196 (a), Ng 192 (b), Rt 108-3 (c–e, g, n), Rt 104 (f), Rt 96 (h, k), Rt 108-6 (i), Ng 187 (j), Ng 186 (l), Rt 107 (m), Rt 102-2 (o), Rt 104 (p)." figureDoi="http://doi.org/10.5281/zenodo.12583338" httpUri="https://zenodo.org/record/12583338/files/figure.png" pageId="9" pageNumber="12">8f</figureCitation>
|
||
,
|
||
<figureCitation id="132A2A512B464E6FFDA8D411B008DC4D" box="[564,589,1117,1141]" captionStart="Fig" captionStartId="8.[231,266,1713,1734]" captionTargetBox="[231,1408,152,1671]" captionTargetId="figure-34@8.[231,1408,152,1671]" captionTargetPageId="8" captionText="Fig. 9 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section: c–f, h–k); Mandegan section: a–b, g, l). a subaxial section of an assumed microspheric specimen. b–c axial sections showing initial spire with acute margin beneath the apex. d oblique section showing the marginal trough between the marginal and central zones as well as a circular row of marginal apertures between. e tangential section showing subepidermal network; septa in the lower part marked by white dash lines. f subaxial section of a possibly microspheric specimen. g transverse section. h subaxial section. i tangential section showing aligned main beams. j subaxial section of a possibly microspheric specimen. k tangential section. l slightly oblique transverse section showing several orders of vertical partitions (beams, intercalary beams), marginal trough, and pillared central zone. Abbreviations: m.f. = marginal foramen, m.t. = marginal through, pi = pillar. Thin sections: Rt 105 (a), 2Ng 17 (c), Rt 96 (b), 2Ng 167 (d), 2Ng 174 (e), 2Ng 118-1 (f), Rt 108-2b (g), 2Ng 169 (h), 2Ng 191 (i), 2Ng 176 (j), 2Ng 146 (k), Rt 111-1 (l)." figureDoi="http://doi.org/10.5281/zenodo.12583342" httpUri="https://zenodo.org/record/12583342/files/figure.png" pageId="9" pageNumber="12">9e</figureCitation>
|
||
). At their distal ends the primary beams are moderately thickening. The marginal chamberlets are tapering distally terminating inwards with a marginal foramen, oriented at about 45 degrees to the main axis. In transverse sections they form a circular ring (
|
||
<figureCitation id="132A2A512B464E6FFEA1D54FB3C3DD23" box="[317,390,1283,1307]" captionStart="Fig" captionStartId="5.[149,184,1697,1718]" captionTargetBox="[149,1438,157,1655]" captionTargetId="figure-41@5.[149,1438,157,1655]" captionTargetPageId="5" captionText="Fig. 7 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Mandegan section: b, l; Naghan section: a, c–k). a Oblique transverse section. b Oblique section cutting the marginal (upper part) and central zones (lower part) displaying irregularly distributed fused pillars/secondary deposits. c, k Subaxial sections. d–g, Axial sections showing initial spire. h–i Details (from e and g) of the initial spire. j Tangential section. Slightly oblique transverse sections showing subdivision of the marginal zone and circle of marginal apertures (left and above). Abbreviations: m.f. = marginal foramen, m.t. = marginal trough, pr = protoconch, s.d. = secondary deposits/fused pillars. Thin sections: 2Ng 17 (a, k), Rt 100 (b), Ng 180-1 (c), 2Ng 179-1 (d), 2Ng 168 (e, h), 2Ng 175 (f), 2Ng 174 (g, i), 2Ng 177 (j), Rt 113 (l)." figureDoi="http://doi.org/10.5281/zenodo.12583336" httpUri="https://zenodo.org/record/12583336/files/figure.png" pageId="9" pageNumber="12">Fig. 7l</figureCitation>
|
||
,
|
||
<figureCitation id="132A2A512B464E6FFE0ED54FB39DDD23" box="[402,472,1283,1307]" captionStart="Fig" captionStartId="6.[206,241,1674,1695]" captionTargetBox="[219,1374,159,1632]" captionTargetId="figure-36@6.[213,1374,159,1632]" captionTargetPageId="6" captionText="Fig. 8 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section: a–b, j, l; Mandegan section: c–i, k, m–p). a Subaxial section. b, i, l Oblique sections. c–e, m Deep tangential sections displaying primary septules (beams) continuous (aligned) from one chamber to the next. f Shallow tangential section displaying subepidermal network of beams and rafters. g, j, l, o–p Different transverse sections, some slightly oblique. Note five marginal foramina arranged in a radial circle (inside yellow marked rectangle in o). Note oval test outline in j. Subaxial section. Note alternation of pillars by stairway-like arrangement of foramina (yellow line) and opaque fused pillars/secondary deposits in the central zone. k Detail from h showing marginal foramina arranged in vertical lines between successive chambers. n Tangential-oblique section. Abbreviations: b = beam, ib = intercalary beam, f = foramen, m.f. = marginal foramen, m.t. = marginal through, pi = pillar, s = septum. Thin sections: Ng 196 (a), Ng 192 (b), Rt 108-3 (c–e, g, n), Rt 104 (f), Rt 96 (h, k), Rt 108-6 (i), Ng 187 (j), Ng 186 (l), Rt 107 (m), Rt 102-2 (o), Rt 104 (p)." figureDoi="http://doi.org/10.5281/zenodo.12583338" httpUri="https://zenodo.org/record/12583338/files/figure.png" pageId="9" pageNumber="12">8k, 8o</figureCitation>
|
||
). Foramina can form a continuous line running parallel to the outer test surface as seen in axial sections (
|
||
<figureCitation id="132A2A512B464E6FFE15D509B393DD65" box="[393,470,1349,1373]" captionStart="Fig" captionStartId="5.[149,184,1697,1718]" captionTargetBox="[149,1438,157,1655]" captionTargetId="figure-41@5.[149,1438,157,1655]" captionTargetPageId="5" captionText="Fig. 7 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Mandegan section: b, l; Naghan section: a, c–k). a Oblique transverse section. b Oblique section cutting the marginal (upper part) and central zones (lower part) displaying irregularly distributed fused pillars/secondary deposits. c, k Subaxial sections. d–g, Axial sections showing initial spire. h–i Details (from e and g) of the initial spire. j Tangential section. Slightly oblique transverse sections showing subdivision of the marginal zone and circle of marginal apertures (left and above). Abbreviations: m.f. = marginal foramen, m.t. = marginal trough, pr = protoconch, s.d. = secondary deposits/fused pillars. Thin sections: 2Ng 17 (a, k), Rt 100 (b), Ng 180-1 (c), 2Ng 179-1 (d), 2Ng 168 (e, h), 2Ng 175 (f), 2Ng 174 (g, i), 2Ng 177 (j), Rt 113 (l)." figureDoi="http://doi.org/10.5281/zenodo.12583336" httpUri="https://zenodo.org/record/12583336/files/figure.png" pageId="9" pageNumber="12">Fig. 7e</figureCitation>
|
||
,
|
||
<figureCitation id="132A2A512B464E6FFE7ED509B3B8DD65" box="[482,509,1349,1373]" captionStart="Fig" captionStartId="6.[206,241,1674,1695]" captionTargetBox="[219,1374,159,1632]" captionTargetId="figure-36@6.[213,1374,159,1632]" captionTargetPageId="6" captionText="Fig. 8 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Naghan section: a–b, j, l; Mandegan section: c–i, k, m–p). a Subaxial section. b, i, l Oblique sections. c–e, m Deep tangential sections displaying primary septules (beams) continuous (aligned) from one chamber to the next. f Shallow tangential section displaying subepidermal network of beams and rafters. g, j, l, o–p Different transverse sections, some slightly oblique. Note five marginal foramina arranged in a radial circle (inside yellow marked rectangle in o). Note oval test outline in j. Subaxial section. Note alternation of pillars by stairway-like arrangement of foramina (yellow line) and opaque fused pillars/secondary deposits in the central zone. k Detail from h showing marginal foramina arranged in vertical lines between successive chambers. n Tangential-oblique section. Abbreviations: b = beam, ib = intercalary beam, f = foramen, m.f. = marginal foramen, m.t. = marginal through, pi = pillar, s = septum. Thin sections: Ng 196 (a), Ng 192 (b), Rt 108-3 (c–e, g, n), Rt 104 (f), Rt 96 (h, k), Rt 108-6 (i), Ng 187 (j), Ng 186 (l), Rt 107 (m), Rt 102-2 (o), Rt 104 (p)." figureDoi="http://doi.org/10.5281/zenodo.12583338" httpUri="https://zenodo.org/record/12583338/files/figure.png" pageId="9" pageNumber="12">8a</figureCitation>
|
||
). Certain portions of the central zone are filled with opaque micritic masses (fused pillars and/or secondary deposits) that obliterate the original structure (
|
||
<figureCitation id="132A2A512B464E6FFEA5D5E5B339DDF9" box="[313,380,1449,1473]" captionStart="Fig" captionStartId="5.[149,184,1697,1718]" captionTargetBox="[149,1438,157,1655]" captionTargetId="figure-41@5.[149,1438,157,1655]" captionTargetPageId="5" captionText="Fig. 7 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Mandegan section: b, l; Naghan section: a, c–k). a Oblique transverse section. b Oblique section cutting the marginal (upper part) and central zones (lower part) displaying irregularly distributed fused pillars/secondary deposits. c, k Subaxial sections. d–g, Axial sections showing initial spire. h–i Details (from e and g) of the initial spire. j Tangential section. Slightly oblique transverse sections showing subdivision of the marginal zone and circle of marginal apertures (left and above). Abbreviations: m.f. = marginal foramen, m.t. = marginal trough, pr = protoconch, s.d. = secondary deposits/fused pillars. Thin sections: 2Ng 17 (a, k), Rt 100 (b), Ng 180-1 (c), 2Ng 179-1 (d), 2Ng 168 (e, h), 2Ng 175 (f), 2Ng 174 (g, i), 2Ng 177 (j), Rt 113 (l)." figureDoi="http://doi.org/10.5281/zenodo.12583336" httpUri="https://zenodo.org/record/12583336/files/figure.png" pageId="9" pageNumber="12">Fig. 7</figureCitation>
|
||
b-c, 8h). The pillars of the central zone are comparably narrowly spaced and alternate between consecutive chambers (
|
||
<figureCitation id="132A2A512B464E6FFE41D5A7B058DE3B" box="[477,541,1515,1539]" captionStart="Fig" captionStartId="5.[149,184,1697,1718]" captionTargetBox="[149,1438,157,1655]" captionTargetId="figure-41@5.[149,1438,157,1655]" captionTargetPageId="5" captionText="Fig. 7 Gusicella minima (Henson) gen. et comb. nov. from the late Maastrichtian of the Tarbur Formation, SW Iran (Mandegan section: b, l; Naghan section: a, c–k). a Oblique transverse section. b Oblique section cutting the marginal (upper part) and central zones (lower part) displaying irregularly distributed fused pillars/secondary deposits. c, k Subaxial sections. d–g, Axial sections showing initial spire. h–i Details (from e and g) of the initial spire. j Tangential section. Slightly oblique transverse sections showing subdivision of the marginal zone and circle of marginal apertures (left and above). Abbreviations: m.f. = marginal foramen, m.t. = marginal trough, pr = protoconch, s.d. = secondary deposits/fused pillars. Thin sections: 2Ng 17 (a, k), Rt 100 (b), Ng 180-1 (c), 2Ng 179-1 (d), 2Ng 168 (e, h), 2Ng 175 (f), 2Ng 174 (g, i), 2Ng 177 (j), Rt 113 (l)." figureDoi="http://doi.org/10.5281/zenodo.12583336" httpUri="https://zenodo.org/record/12583336/files/figure.png" pageId="9" pageNumber="12">Fig. 7</figureCitation>
|
||
c-d, 8a, h, 9f). In the central part, the foramina present a cribrate distribution and are vertically arranged.
|
||
</paragraph>
|
||
<paragraph id="8BAE36D42B464E6FFF0BD603B347DEBF" blockId="9.[151,772,158,2068]" pageId="9" pageNumber="12">
|
||
<emphasis id="B965EAC62B464E6FFF0BD603B35EDE5F" bold="true" box="[151,283,1615,1639]" pageId="9" pageNumber="12">Dimensions</emphasis>
|
||
(in mm; data from
|
||
<bibRefCitation id="EF804B252B464E6FFD9CD603B0DEDE5E" author="Henson, F. R. S." box="[512,667,1614,1638]" pageId="9" pageNumber="12" pagination="1 - 127" refId="ref5977" refString="Henson, F. R. S., 1948. Larger imperforate Foraminifera of south-western Asia. Families Lituolidae, Orbitolinidae and Meandropsinidae. London, Monograph British Museum (Natural History): 1 - 127." type="book chapter" year="1948">Henson, 1948</bibRefCitation>
|
||
, between brackets):
|
||
</paragraph>
|
||
<paragraph id="8BAE36D42B464E6FFF0BD6DDB324DED2" blockId="9.[151,772,158,2068]" pageId="9" pageNumber="12">
|
||
Test diameter (D):
|
||
<quantity id="4CE99B312B464E6FFEFFD6DCB3A6DE90" box="[355,483,1680,1704]" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.9" metricValueMin="0.7" pageId="9" pageNumber="12" unit="mm" value="1.2999999999999998" valueMax="1.9" valueMin="0.7">0.7–1.9 mm</quantity>
|
||
(
|
||
<quantity id="4CE99B312B464E6FFE6DD6DCB031DE90" box="[497,628,1680,1704]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.8" metricValueMin="0.7" pageId="9" pageNumber="12" unit="mm" value="1.25" valueMax="1.8" valueMin="0.7">0.7–1.8 mm</quantity>
|
||
). Note: specimens with a test diameter>
|
||
<quantity id="4CE99B312B464E6FFE51D6FDB065DEF1" box="[461,544,1713,1737]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="12" unit="mm" value="1.5">1.5 mm</quantity>
|
||
appear to be miscrospheric specimens.
|
||
</paragraph>
|
||
<paragraph id="8BAE36D42B464E6FFF0BD6B8B074DF34" blockId="9.[151,772,158,2068]" box="[151,561,1780,1804]" pageId="9" pageNumber="12">
|
||
Test height (H): 0.8–1.4 (1.0–
|
||
<quantity id="4CE99B312B464E6FFE4AD6B8B06DDF33" box="[470,552,1780,1804]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="12" unit="mm" value="1.5">1.5 mm</quantity>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BAE36D42B464E6FFF0BD759B374DF15" blockId="9.[151,772,158,2068]" box="[151,305,1813,1837]" pageId="9" pageNumber="12">D/H: 0.8–1.35</paragraph>
|
||
<paragraph id="8BAE36D42B464E6FFF0BD77AB3C1DF57" blockId="9.[151,772,158,2068]" pageId="9" pageNumber="12">
|
||
Numbers of chambers per
|
||
<quantity id="4CE99B312B464E6FFE26D77AB3BFDF76" box="[442,506,1846,1870]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="12" unit="mm" value="1.0">1 mm</quantity>
|
||
of the axial length: 12– 16, mostly 15–16 (15)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30B655F2B464E6FFF0BD734B71ADA5C" pageId="9" pageNumber="12" type="discussion">
|
||
<paragraph id="8BAE36D42B464E6FFF0BD734B71ADA5C" blockId="9.[151,772,158,2068]" lastBlockId="9.[816,1437,158,612]" pageId="9" pageNumber="12">
|
||
<emphasis id="B965EAC62B464E6FFF0BD734B38FDFA8" bold="true" box="[151,458,1912,1936]" pageId="9" pageNumber="12">Remarks on stratigraphy:</emphasis>
|
||
<bibRefCitation id="EF804B252B464E6FFE49D735B03CDFA8" author="Henson, F. R. S." box="[469,633,1912,1936]" pageId="9" pageNumber="12" pagination="1 - 127" refId="ref5977" refString="Henson, F. R. S., 1948. Larger imperforate Foraminifera of south-western Asia. Families Lituolidae, Orbitolinidae and Meandropsinidae. London, Monograph British Museum (Natural History): 1 - 127." type="book chapter" year="1948">Henson (1948)</bibRefCitation>
|
||
reported
|
||
<taxonomicName id="4C114D572B464E6FFD75D735B2A2DF89" authorityName="SCHLAGINTWEIT & RASHIDI" authorityYear="2021" baseAuthorityName="SCHLAGINTWEIT & RASHIDI" baseAuthorityYear="1948" class="Globothalamea" family="Orbitolinidae" genus="Gusicella" kingdom="Chromista" order="Loftusiida" pageId="9" pageNumber="12" phylum="Foraminifera" rank="species" species="minima">
|
||
<emphasis id="B965EAC62B464E6FFD75D735B0B9DFA8" box="[745,764,1913,1936]" italics="true" pageId="9" pageNumber="12">D</emphasis>
|
||
.
|
||
<emphasis id="B965EAC62B464E6FFF0BD7D6B2A2DF89" box="[151,231,1946,1969]" italics="true" pageId="9" pageNumber="12">minima</emphasis>
|
||
</taxonomicName>
|
||
“most probably” from the late Cenomanian or Turonian of
|
||
<collectingCountry id="F30676442B464E6FFEB5D7F6B321DFEA" box="[297,356,1978,2002]" name="Qatar" pageId="9" pageNumber="12">Qatar</collectingCountry>
|
||
(Dukhan no. 2 and 3 wells; Wasia Group, see figs.
|
||
<quantity id="4CE99B312B464E6FFECCD797B3D7DFCB" box="[336,402,2011,2035]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="5.08" metricValueMin="2.54" pageId="9" pageNumber="12" unit="in" value="1.5" valueMax="2.0" valueMin="1.0">1-2 in</quantity>
|
||
Sugden and Standring, 1975). In addition, Henson (op. cit, p. 26) noted “a single specimen, possibly of this species” from the Maastrichtian of
|
||
<collectingCountry id="F30676442B464E6FFCACD08CB11AD8EF" box="[816,863,192,215]" name="Iraq" pageId="9" pageNumber="12">Iraq</collectingCountry>
|
||
. For the Dukhan oil field of
|
||
<collectingCountry id="F30676442B464E6FFB0BD0F3B690D8EF" box="[1175,1237,191,215]" name="Qatar" pageId="9" pageNumber="12">Qatar</collectingCountry>
|
||
,
|
||
<bibRefCitation id="EF804B252B464E6FFB7DD0F3B183D8C0" author="Hewaidy, A. G. & Al-Hitmi, H. A." pageId="9" pageNumber="12" pagination="1 - 24" refId="ref5922" refString="Hewaidy, A. G. A, Al-Hitmi, H. A., 1994. Cretaceous- Early Eocene biostratigraphy of the Dukhan oil field, west Qatar, Arabian Gulf. M. E. R. C. Ain Shams University, Earth Science Series, 8: 1 - 24." type="journal article" year="1994">Hewaidy and Al-Hitmi (1994)</bibRefCitation>
|
||
established a Late Cenomanian “
|
||
<taxonomicName id="4C114D572B464E6FFAC6D0ADB1ACD921" authorityName="Henson" authorityYear="1948" class="Globothalamea" family="Orbitolinidae" genus="Dictyoconella" kingdom="Chromista" order="Loftusiida" pageId="9" pageNumber="12" phylum="Foraminifera" rank="species" species="minima">
|
||
<emphasis id="B965EAC62B464E6FFAC6D0ADB1ACD921" italics="true" pageId="9" pageNumber="12">Dictyoconella minima</emphasis>
|
||
</taxonomicName>
|
||
total range Zone”. The “zonal taxon” however, has not been illustrated and therefore, cannot be commented. Within the framework of our observations in the Cenomanian Sarvak Formation of
|
||
<collectingCountry id="F30676442B464E6FFB7ED129B748D944" box="[1250,1293,357,380]" name="Iraq" pageId="9" pageNumber="12">Iraq</collectingCountry>
|
||
(e.g.
|
||
<bibRefCitation id="EF804B252B464E6FFAC8D128B6F0D9A5" author="Yazdi-Moghadam, M. & Schlagintweit, F." pageId="9" pageNumber="12" pagination="104380" refId="ref6842" refString="Yazdi-Moghadam, M., Schlagintweit, F., 2020. Persiconus sarvaki gen. et sp. nov., a new complex orbitolinid (Foraminifera) from the Cenomanian of the Sarvak Formation (SW Iran, Zagros Zone). Cretaceous Research, 109: article 104380." type="journal article" year="2020">Yazdi-Moghadam and Schlagintweit, 2020</bibRefCitation>
|
||
), no specimens of
|
||
<taxonomicName id="4C114D572B464E6FFA1FD1CAB1C5D987" authorityName="Henson" authorityYear="1948" class="Globothalamea" family="Orbitolinidae" genus="Dictyoconella" kingdom="Chromista" order="Loftusiida" pageId="9" pageNumber="12" phylum="Foraminifera" rank="species" species="minima">
|
||
<emphasis id="B965EAC62B464E6FFA1FD1CAB1C5D987" italics="true" pageId="9" pageNumber="12">G. minima</emphasis>
|
||
</taxonomicName>
|
||
have been observed. There are specimens of an orbitolinid displaying a pillared central zone and a complex exoskeleton that needs further taxonomic study and evaluation, and it is different from
|
||
<taxonomicName id="4C114D572B464E6FFB5BD246B778DA1A" authorityName="Henson" authorityYear="1948" box="[1223,1341,522,546]" class="Globothalamea" family="Orbitolinidae" genus="Dictyoconella" kingdom="Chromista" order="Loftusiida" pageId="9" pageNumber="12" phylum="Foraminifera" rank="species" species="minima">
|
||
<emphasis id="B965EAC62B464E6FFB5BD246B778DA1A" box="[1223,1341,522,546]" italics="true" pageId="9" pageNumber="12">G. minima</emphasis>
|
||
</taxonomicName>
|
||
. So far, only the occurrence in the Maastrichtian is well constrained and older occurrences require further study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |