390 lines
52 KiB
XML
390 lines
52 KiB
XML
<document id="12D40A9BBC2D9D109273291C44E7F243" ID-CLB-Dataset="20988" ID-DOI="10.5852/ejt.2020.713" ID-GBIF-Dataset="1d5fdead-d612-4c66-b482-4f051f990351" ID-Zenodo-Dep="4020752" ID-ZooBank="A6CB58F5-1ECC-47F0-AA07-798844AF80A7" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1599662866733" checkinUser="valdenar" docAuthor="Antić, Dragan Ž. & Reip, Hans S." docDate="2020" docId="03A963622846FFB22E461DA90C44FE2A" docLanguage="en" docName="ejt-713_antic_reip.pdf" docOrigin="European Journal of Taxonomy 713" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Leucogeorgia borealis Antić & Reip 2020, sp. nov." docType="treatment" docUuid="D8CFFCD0-8D33-42A0-B7FD-E8FDE92E99E2" docUuidSource="ZooBank" docVersion="11" lastPageNumber="32" masterDocId="FF901B1A285DFFAD2C24183E0F08FFEE" masterDocTitle="The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)" masterLastPageNumber="106" masterPageNumber="1" pageNumber="28" updateTime="1698845858502" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4515E6C2BAA2BC2A0895866AB305A6FB">The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)</mods:title>
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<mods:roleTerm id="E8ED2AC5D09469DC881C98E9AE644F2A">Author</mods:roleTerm>
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<mods:namePart id="313AF865F01864A877EFEA6B28AAD750">Antić, Dragan Ž.</mods:namePart>
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<mods:affiliation id="F63899666394DC38544DCA4C01EA72BD">University of Belgrade - Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia. Department of Soil Zoology, Senckenberg Museum of Natural History Görlitz, Am Museum 1, 02826 Görlitz, Germany. Thüringer Höhlenverein e. V., Johann-Sebastian-Bach-Str. 16 a, 98529 Suhl, Germany.</mods:affiliation>
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<mods:nameIdentifier id="0AD1EF56740A6A10DA9D080C3AC31470" type="email">dragan.antic@bio.bg.ac.rs</mods:nameIdentifier>
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<mods:namePart id="BE3F77F66CBDFE9812FD9C123433A1E5">Reip, Hans S.</mods:namePart>
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<mods:nameIdentifier id="698EA2D1396D3FA2586B53067D544B9E" type="email">reip@myriapoden-info.de</mods:nameIdentifier>
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<mods:typeOfResource id="AD0440794608468A14332C99E564C56F">text</mods:typeOfResource>
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<mods:title id="2C12AA33B2545BFD5AAFF7BDA88BF575">European Journal of Taxonomy</mods:title>
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<mods:part id="14B341BF323A63BE69E599BB1ADE38E1">
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<mods:date id="9B289E4E7E04209305F5ADA54515896F">2020</mods:date>
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<mods:number id="5A62044A1E9A40698A6466DE2BB94F7F">2020-09-07</mods:number>
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<mods:detail id="E2FD6E77A0E35D34E13726F439C34DE6" type="volume">
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<mods:number id="F86A8E7F1C3CE5999950CF0534D06774">713</mods:number>
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<mods:identifier id="69F1844182DFA46328280A4877404D29" type="DOI">10.5852/ejt.2020.713</mods:identifier>
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<treatment id="03A963622846FFB22E461DA90C44FE2A" ID-DOI="http://doi.org/10.5281/zenodo.4335830" ID-GBIF-Taxon="167523049" ID-Zenodo-Dep="4335830" LSID="urn:lsid:zoobank.org:act:D8CFFCD0-8D33-42A0-B7FD-E8FDE92E99E2" httpUri="http://treatment.plazi.org/id/03A963622846FFB22E461DA90C44FE2A" lastPageId="31" lastPageNumber="32" pageId="27" pageNumber="28">
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<subSubSection id="C31A81FF2846FFB62E461DA90CDAFA5C" box="[610,978,1431,1458]" pageId="27" pageNumber="28" type="nomenclature">
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<paragraph id="8BBFD2742846FFB62E461DA90CDAFA5C" blockId="27.[356,1231,1431,1528]" box="[610,978,1431,1458]" pageId="27" pageNumber="28">
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<heading id="D0F765182846FFB62E461DA90CDAFA5C" box="[610,978,1431,1458]" centered="true" fontSize="11" level="2" pageId="27" pageNumber="28" reason="2">
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<taxonomicName id="4C00A9F72846FFB62E461DA90C78FA5C" ID-CoL="849PM" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[610,880,1431,1458]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
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<emphasis id="B9740E662846FFB62E461DA90C78FA5C" bold="true" box="[610,880,1431,1458]" italics="true" pageId="27" pageNumber="28">Leucogeorgia borealis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2846FFB62F531DA60CDAFA5C" box="[887,978,1432,1458]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A81FF2846FFB62D401D840C80FA16" pageId="27" pageNumber="28" type="description">
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<paragraph id="8BBFD2742846FFB62D401D840BC7FA3A" blockId="27.[356,1231,1431,1528]" box="[356,1231,1466,1492]" pageId="27" pageNumber="28">
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<uri id="FF91DE762846FFB62D401D840BC7FA3A" box="[356,1231,1466,1492]" pageId="27" pageNumber="28">
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urn:lsid:zoobank.org:act:
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<uuid id="FFA6E8A12846FFB62EA71D840BC7FA3A" box="[643,1231,1466,1492]" pageId="27" pageNumber="28">D8CFFCD0-8D33-42A0-B7FD-E8FDE92E99E2</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BBFD2742846FFB62E881DE30C80FA16" blockId="27.[356,1231,1431,1528]" box="[684,904,1501,1528]" pageId="27" pageNumber="28">
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<figureCitation id="133BCEF12846FFB62E881DE30C3AFA19" box="[684,818,1501,1527]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="28.[189,232,1894,1920]" captionStartId-1="30.[189,232,1832,1858]" captionStartId-2="31.[189,232,1929,1955]" captionTargetBox-0="[281,1306,265,1866]" captionTargetBox-1="[189,1399,370,1804]" captionTargetBox-2="[202,1379,487,1902]" captionTargetId-0="figure@28.[281,1306,265,1866]" captionTargetId-1="figure@30.[189,1399,370,1804]" captionTargetId-2="figure@31.[189,1399,471,1904]" captionTargetPageId-0="28" captionTargetPageId-1="30" captionTargetPageId-2="31" captionText-0="Fig. 16. Leucogeorgia borealis sp. nov., types from Fanagoriyskaya Cave (ZMUM). A. Paratype ♀, habitus, lateral view. B. Paratype ♀, anterior part of body, lateral view. C. Paratype ♂, posterior part of body, lateral view. D. Holotype ♂, body ring 7, lateral view. E. Paratype ♀, midbody rings, lateral view. Arrow indicated metazonal seta. Scale bars: 1 mm." captionText-1="Fig. 17. Leucogeorgia borealis sp. nov., paratype ♂ from Fanagoriyskaya Cave (ZMUM). A. Leg-pair 1, anterior view. B. Right leg 1, lateral view. C. Head, ventral view. D. Left gonopods, mesal view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: A–B = 0.05 mm; C = 0.2 mm; D = 0.1 mm." captionText-2="Fig. 18. Leucogeorgia borealis sp. nov., holotype ♂ from Fanagoriyskaya Cave (ZMUM). A. Right gonopods, mesal view. B. Right gonopods, lateral view. C. Penes, posterior view. D. Leg-pair 1, anterior view. E. Right leg 1, lateral view. Scale bar: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4020786" figureDoi-1="http://doi.org/10.5281/zenodo.4020788" figureDoi-2="http://doi.org/10.5281/zenodo.4020790" httpUri-0="https://zenodo.org/record/4020786/files/figure.png" httpUri-1="https://zenodo.org/record/4020788/files/figure.png" httpUri-2="https://zenodo.org/record/4020790/files/figure.png" pageId="27" pageNumber="28">Figs 16–18</figureCitation>
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,
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<figureCitation id="133BCEF12846FFB62F1A1DE30C80FA16" box="[830,904,1501,1528]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="96.[189,232,1921,1947]" captionStartId-1="97.[189,232,1991,2017]" captionTargetBox-0="[189,1399,979,1893]" captionTargetBox-1="[189,1399,890,1963]" captionTargetId-0="figure@96.[189,1399,979,1893]" captionTargetId-1="figure@97.[189,1399,890,1963]" captionTargetPageId-0="96" captionTargetPageId-1="97" captionText-0="Fig. 56. General distribution of the Caucasian leucogeorgiinines. The white dashed line hypothetically indicates the boundary between the northwestern and southeastern groups of species of the genus Leucogeorgia Verhoeff, 1930." captionText-1="Fig. 57. Distribution of the northwestern group of the genus Leucogeorgia Verhoeff, 1930." figureDoi-0="http://doi.org/10.5281/zenodo.4020873" figureDoi-1="http://doi.org/10.5281/zenodo.4020875" httpUri-0="https://zenodo.org/record/4020873/files/figure.png" httpUri-1="https://zenodo.org/record/4020875/files/figure.png" pageId="27" pageNumber="28">56–57</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A81FF2846FFB62C991E180BBEF888" pageId="27" pageNumber="28" type="diagnosis">
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<paragraph id="8BBFD2742846FFB62C991E180E3EF9AE" blockId="27.[189,310,1574,1600]" box="[189,310,1574,1600]" pageId="27" pageNumber="28">
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<heading id="D0F765182846FFB62C991E180E3EF9AE" bold="true" box="[189,310,1574,1600]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
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<emphasis id="B9740E662846FFB62C991E180E3EF9AE" bold="true" box="[189,310,1574,1600]" pageId="27" pageNumber="28">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBFD2742846FFB62C991E6B0BBEF888" blockId="27.[189,1399,1621,1894]" pageId="27" pageNumber="28">
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This species belongs to the group of
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<taxonomicName id="4C00A9F72846FFB62E491E680C07F981" authorityName="Verhoeff" authorityYear="1930" box="[621,783,1622,1647]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
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<emphasis id="B9740E662846FFB62E491E680C07F981" box="[621,783,1622,1647]" italics="true" pageId="27" pageNumber="28">Leucogeorgia</emphasis>
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</taxonomicName>
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spp. without modified mouthparts and with teeth on the mesomeral claw (vs absent in
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<taxonomicName id="4C00A9F72846FFB62E681E470DEFF97C" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[588,743,1656,1682]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
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<emphasis id="B9740E662846FFB62E681E470DEFF97C" box="[588,743,1656,1682]" italics="true" pageId="27" pageNumber="28">L. golovatchi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2846FFB62ECA1E470C43F97D" box="[750,843,1657,1683]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
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,
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<taxonomicName id="4C00A9F72846FFB62F731E470CC9F97C" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[855,961,1656,1682]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
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<emphasis id="B9740E662846FFB62F731E470CC9F97C" box="[855,961,1656,1682]" italics="true" pageId="27" pageNumber="28">L. lobata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2846FFB62FED1E470B2CF97D" box="[969,1060,1657,1683]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
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and
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<taxonomicName id="4C00A9F72846FFB628791E470A0CF97C" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[1117,1284,1656,1682]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="prometheus" status="sp. nov.">
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<emphasis id="B9740E662846FFB628791E470A0CF97C" box="[1117,1284,1656,1682]" italics="true" pageId="27" pageNumber="28">L. prometheus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2846FFB6292F1E470A62F97D" box="[1291,1386,1657,1683]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
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).
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<taxonomicName id="4C00A9F72846FFB62C991EA20EC0F95B" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[189,456,1691,1717]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
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<emphasis id="B9740E662846FFB62C991EA20EC0F95B" box="[189,456,1691,1717]" italics="true" pageId="27" pageNumber="28">Leucogeorgia borealis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2846FFB62DF61EA20D38F958" box="[466,560,1692,1718]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
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differs from
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<taxonomicName id="4C00A9F72846FFB62EF71EA20C64F95B" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[723,876,1691,1717]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="abchasica">
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<emphasis id="B9740E662846FFB62EF71EA20C64F95B" box="[723,876,1691,1717]" italics="true" pageId="27" pageNumber="28">L. abchasica</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C00A9F72846FFB62F591EA20CF6F95B" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[893,1022,1692,1717]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="satunini">
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<emphasis id="B9740E662846FFB62F591EA20CF6F95B" box="[893,1022,1692,1717]" italics="true" pageId="27" pageNumber="28">L. satunini</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C00A9F72846FFB628181EA20B84F95B" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[1084,1164,1692,1717]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="gioi" status="sp. nov.">
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<emphasis id="B9740E662846FFB628181EA20B84F95B" box="[1084,1164,1692,1717]" italics="true" pageId="27" pageNumber="28">L. gioi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2846FFB628B21EA20BFDF95B" box="[1174,1269,1692,1718]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
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by having a stout mesomeral claw (vs elongate and slender in
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<taxonomicName id="4C00A9F72846FFB62F641E810CD2F937" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[832,986,1727,1753]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="abchasica">
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<emphasis id="B9740E662846FFB62F641E810CD2F937" box="[832,986,1727,1753]" italics="true" pageId="27" pageNumber="28">L. abchasica</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C00A9F72846FFB62FC91E810B67F936" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[1005,1135,1727,1752]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="satunini">
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<emphasis id="B9740E662846FFB62FC91E810B67F936" box="[1005,1135,1727,1752]" italics="true" pageId="27" pageNumber="28">L. satunini</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C00A9F72846FFB628941E810A09F936" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[1200,1281,1727,1752]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="gioi" status="sp. nov.">
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<emphasis id="B9740E662846FFB628941E810A09F936" box="[1200,1281,1727,1752]" italics="true" pageId="27" pageNumber="28">L. gioi</emphasis>
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</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2846FFB629281E810A62F937" box="[1292,1386,1727,1753]" pageId="27" pageNumber="28" rank="species">sp. nov</taxonomicNameLabel>
|
||
).
|
||
<taxonomicName id="4C00A9F72846FFB62C991EDD0ECEF912" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[189,454,1762,1788]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
|
||
<emphasis id="B9740E662846FFB62C991EDD0ECEF912" box="[189,454,1762,1788]" italics="true" pageId="27" pageNumber="28">Leucogeorgia borealis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2846FFB62DE91EDC0D21F912" box="[461,553,1762,1788]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C00A9F72846FFB62EE11EDD0C34F912" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[709,828,1762,1788]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="oculata" status="sp. nov.">
|
||
<emphasis id="B9740E662846FFB62EE11EDD0C34F912" box="[709,828,1762,1788]" italics="true" pageId="27" pageNumber="28">L. oculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2846FFB62F601EDC0C97F912" box="[836,927,1762,1788]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by having a strongly serrate mesomeral lamella without a posterior rise (vs lamella non-serrate and with a posterior rise in
|
||
<taxonomicName id="4C00A9F72846FFB628A81F380A0BF8F1" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[1164,1283,1797,1823]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="oculata" status="sp. nov.">
|
||
<emphasis id="B9740E662846FFB628A81F380A0BF8F1" box="[1164,1283,1797,1823]" italics="true" pageId="27" pageNumber="28">L. oculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2846FFB6292F1F3B0A61F8F1" box="[1291,1385,1797,1823]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
), and in the absence both of pigmentation and ommatidia, and the presence of metazonal setae (vs a pigmented body, ommatidia present and metazonal setae absent in
|
||
<taxonomicName id="4C00A9F72846FFB62FE81F720B4BF88B" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[972,1091,1867,1893]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="oculata" status="sp. nov.">
|
||
<emphasis id="B9740E662846FFB62FE81F720B4BF88B" box="[972,1091,1867,1893]" italics="true" pageId="27" pageNumber="28">L. oculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2846FFB6286F1F720BA1F888" box="[1099,1193,1868,1894]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A81FF2846FFB62C991FAB0A7FF833" box="[189,1399,1941,2014]" pageId="27" pageNumber="28" type="etymology">
|
||
<paragraph id="8BBFD2742846FFB62C991FAB0E4CF841" blockId="27.[189,324,1941,1967]" box="[189,324,1941,1967]" pageId="27" pageNumber="28">
|
||
<heading id="D0F765182846FFB62C991FAB0E4CF841" bold="true" box="[189,324,1941,1967]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
|
||
<emphasis id="B9740E662846FFB62C991FAB0E4CF841" bold="true" box="[189,324,1941,1967]" pageId="27" pageNumber="28">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742846FFB62C991FFD0A7FF833" blockId="27.[189,1399,1987,2014]" box="[189,1399,1987,2014]" pageId="27" pageNumber="28">
|
||
From the Latin ʻ
|
||
<emphasis id="B9740E662846FFB62D531FFD0ECEF833" box="[375,454,1987,2013]" italics="true" pageId="27" pageNumber="28">boreas</emphasis>
|
||
ʼ (= ʻnorthʼ), reflecting its having the northernmost species in the genus.Adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7F82FC2841FFB12C991F580C1AF805" ID-DOI="http://doi.org/10.5281/zenodo.4020786" ID-Zenodo-Dep="4020786" httpUri="https://zenodo.org/record/4020786/files/figure.png" pageId="28" pageNumber="29" startId="28.[189,232,1894,1920]" targetBox="[281,1306,265,1866]" targetPageId="28">
|
||
<paragraph id="8BBFD2742841FFB12C991F580C1AF805" blockId="28.[189,1399,1894,2027]" pageId="28" pageNumber="29">
|
||
<emphasis id="B9740E662841FFB12C991F580E14F86E" bold="true" box="[189,284,1894,1920]" pageId="28" pageNumber="29">Fig. 16.</emphasis>
|
||
<taxonomicName id="4C00A9F72841FFB12D021F590D3AF86E" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[294,562,1894,1920]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
|
||
<emphasis id="B9740E662841FFB12D021F590D3AF86E" box="[294,562,1894,1920]" italics="true" pageId="28" pageNumber="29">Leucogeorgia borealis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2841FFB12E181F590D94F86F" box="[572,668,1895,1921]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, types from Fanagoriyskaya Cave (ZMUM).
|
||
<emphasis id="B9740E662841FFB128E51F580BDEF86E" bold="true" box="[1217,1238,1894,1920]" pageId="28" pageNumber="29">A</emphasis>
|
||
. Paratype ♀, habitus, lateral view.
|
||
<emphasis id="B9740E662841FFB12D931FB40EC3F84A" bold="true" box="[439,459,1930,1956]" pageId="28" pageNumber="29">B</emphasis>
|
||
. Paratype ♀, anterior part of body, lateral view.
|
||
<emphasis id="B9740E662841FFB128261FB70B1FF84D" bold="true" box="[1026,1047,1929,1955]" pageId="28" pageNumber="29">C</emphasis>
|
||
. Paratype ♂, posterior part of body, lateral view.
|
||
<emphasis id="B9740E662841FFB12DBD1F930EA6F829" bold="true" box="[409,430,1965,1991]" pageId="28" pageNumber="29">D</emphasis>
|
||
. Holotype ♂, body ring 7, lateral view.
|
||
<emphasis id="B9740E662841FFB12F591F930C99F829" bold="true" box="[893,913,1965,1991]" pageId="28" pageNumber="29">E</emphasis>
|
||
. Paratype ♀, midbody rings, lateral view. Arrow indicated metazonal seta. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31A81FF2840FFB02C9919350CE8FD87" pageId="29" pageNumber="30" type="materials_examined">
|
||
<paragraph id="8BBFD2742840FFB02C9919350E64FECB" blockId="29.[189,364,267,293]" box="[189,364,267,293]" pageId="29" pageNumber="30">
|
||
<heading id="D0F765182840FFB02C9919350E64FECB" bold="true" box="[189,364,267,293]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
|
||
<emphasis id="B9740E662840FFB02C9919350E64FECB" bold="true" box="[189,364,267,293]" pageId="29" pageNumber="30">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<materialsCitation id="3B68D8292840FFB02CCE19040ED1FE50" ID-GBIF-Occurrence="2858644315" collectingDate="1943-11-03" collectionCode="ZMUM" collectorName="J. Birstein" country="Russia" county="Greater Caucasus" latitude="44.47" location="Fanagoriyskaya (= Bolshaya Fanagoriyskaya, = Psekupskaya) Cave" longLatPrecision="682" longitude="38.98" municipality="Fanagoriyskaya" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Krasnodar Province" typeStatus="holotype">
|
||
<paragraph id="8BBFD2742840FFB02CCE19040E54FEBA" blockId="29.[189,1399,314,446]" box="[234,348,314,340]" pageId="29" pageNumber="30">
|
||
<emphasis id="B9740E662840FFB02CCE19040E54FEBA" bold="true" box="[234,348,314,340]" pageId="29" pageNumber="30">
|
||
<typeStatus id="54BB6CD62840FFB02CCE19040E54FEBA" box="[234,348,314,340]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742840FFB02C9919630ED1FE50" blockId="29.[189,1399,314,446]" pageId="29" pageNumber="30">
|
||
<collectingCountry id="F31792E42840FFB02C9919630E2CFE99" box="[189,292,349,375]" name="Russia" pageId="29" pageNumber="30">RUSSIA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D0619FD2840FFB02D6E19650E69FE99" box="[330,353,347,375]" pageId="29" pageNumber="30" type="male">♂</specimenCount>
|
||
;
|
||
<collectingRegion id="49C41C962840FFB02D5219620D61FE98" box="[374,617,348,374]" country="Russia" name="Krasnodar" pageId="29" pageNumber="30">Krasnodar Province</collectingRegion>
|
||
,
|
||
<locationDeviation id="9A0DCB562840FFB02E5F19630A7AFE99" box="[635,1394,348,375]" pageId="29" pageNumber="30">near Goryachiy Klyuch, NW karst area of Greater Caucasus</locationDeviation>
|
||
,
|
||
<location id="8EDF84AF2840FFB02C9919BE0CD4FE74" LSID="urn:lsid:plazi:treatment:03A963622846FFB22E461DA90C44FE2A:8EDF84AF2840FFB02C9919BE0CD4FE74" box="[189,988,384,411]" country="Russia" county="Greater Caucasus" latitude="44.47" longLatPrecision="682" longitude="38.98" municipality="Fanagoriyskaya" name="Fanagoriyskaya (= Bolshaya Fanagoriyskaya, = Psekupskaya) Cave" pageId="29" pageNumber="30" stateProvince="Krasnodar Province">Fanagoriyskaya (= Bolshaya Fanagoriyskaya, = Psekupskaya) Cave</location>
|
||
;
|
||
<geoCoordinate id="EE34B4B32840FFB02FC819BE0B52FE74" box="[1004,1114,384,410]" degrees="44.47" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="555" value="44.47">44.47° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE34B4B32840FFB0284219BE0BD9FE74" box="[1126,1233,384,410]" degrees="38.98" direction="east" orientation="longitude" pageId="29" pageNumber="30" precision="555" value="38.98">38.98° E</geoCoordinate>
|
||
;
|
||
<date id="FFBEF4B42840FFB028F919BE0A7AFE74" box="[1245,1394,384,410]" pageId="29" pageNumber="30" value="1943-11-03">
|
||
<collectingDate id="EFFA0D5C2840FFB028F919BE0A7AFE74" box="[1245,1394,384,410]" pageId="29" pageNumber="30" value="1943-11-03">3 Nov. 1943</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F5B7A22840FFB02C99199A0E3CFE53" box="[189,308,419,446]" pageId="29" pageNumber="30">J. Birstein</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED114AB12840FFB02D52199A0EDDFE50" box="[374,469,420,446]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="29" pageNumber="30">ZMUM</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="8BBFD2742840FFB02CCE19DB0E6FFE11" blockId="29.[189,1398,485,617]" box="[234,359,485,511]" pageId="29" pageNumber="30">
|
||
<emphasis id="B9740E662840FFB02CCE19DB0E6FFE11" bold="true" box="[234,359,485,511]" pageId="29" pageNumber="30">
|
||
<typeStatus id="54BB6CD62840FFB02CCE19DB0E6FFE11" box="[234,359,485,511]" pageId="29" pageNumber="30" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742840FFB02C991A360CE8FD87" blockId="29.[189,1398,485,617]" pageId="29" pageNumber="30">
|
||
<materialsCitation id="3B68D8292840FFB02C991A360E10FDAB" ID-GBIF-Occurrence="2858644376" collectingDate="1961-07-08" collectionCode="ZMUM" collectorName="S. I. Ljovuschkin" country="Russia" county="Greater Caucasus" latitude="44.47" location="RUSSIA" longLatPrecision="682" longitude="38.98" municipality="Fanagoriyskaya" pageId="29" pageNumber="30" specimenCount="4" specimenCount-female="1" specimenCount-male="1" stateProvince="Krasnodar" typeStatus="paratype">
|
||
<collectingCountry id="F31792E42840FFB02C991A360E2CFDCC" box="[189,292,520,546]" name="Russia" pageId="29" pageNumber="30">RUSSIA</collectingCountry>
|
||
•
|
||
<specimenCount id="9D0619FD2840FFB02D191A360E61FDCC" box="[317,361,518,546]" pageId="29" pageNumber="30" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2840FFB02D5C1A360EACFDCC" box="[376,420,520,546]" pageId="29" pageNumber="30" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2840FFB02D971A360EF7FDCC" box="[435,511,520,546]" pageId="29" pageNumber="30">2 juvs</specimenCount>
|
||
; same collection data as for holotype but
|
||
<date id="FFBEF4B42840FFB02FC31A360B64FDCF" box="[999,1132,519,546]" pageId="29" pageNumber="30" value="1961-07-08">
|
||
<collectingDate id="EFFA0D5C2840FFB02FC31A360B64FDCF" box="[999,1132,519,546]" pageId="29" pageNumber="30" value="1961-07-08">8 Jul. 1961</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F5B7A22840FFB0285D1A360A34FDCC" box="[1145,1340,520,546]" pageId="29" pageNumber="30">S.I. Ljovuschkin</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED114AB12840FFB02C991A150E10FDAB" box="[189,280,555,581]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="29" pageNumber="30">ZMUM</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B68D8292840FFB02D101A150A6BFDAB" ID-GBIF-Occurrence="2858644362" box="[308,1379,555,581]" collectingDate="1983-05-19" collectionCode="ZMUM" collectorName="S. I. Golovatch" country="Russia" county="Greater Caucasus" latitude="44.47" location="Cave" longLatPrecision="682" longitude="38.98" municipality="Fanagoriyskaya" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Krasnodar" typeStatus="paratype">
|
||
<specimenCount id="9D0619FD2840FFB02D101A150E71FDAB" box="[308,377,555,581]" pageId="29" pageNumber="30" type="generic">1 juv.</specimenCount>
|
||
; same collection data as for holotype but
|
||
<date id="FFBEF4B42840FFB02F4D1A150B03FDAB" box="[873,1035,555,581]" pageId="29" pageNumber="30" value="1983-05-19">
|
||
<collectingDate id="EFFA0D5C2840FFB02F4D1A150B03FDAB" box="[873,1035,555,581]" pageId="29" pageNumber="30" value="1983-05-19">19 May 1983</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F5B7A22840FFB0283D1A150BCCFDAB" box="[1049,1220,555,581]" pageId="29" pageNumber="30">S.I. Golovatch</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED114AB12840FFB0292C1A150A6BFDAB" box="[1288,1379,555,581]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="29" pageNumber="30">ZMUM</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B68D8292840FFB02C991A700CD4FD87" ID-GBIF-Occurrence="2858644373" box="[189,988,590,617]" collectingDate="1968-09-20" collectionCode="ZMUM" collectorName="Goryachiy Klyuch, NW & J. Birstein" country="Russia" county="Greater Caucasus" latitude="44.47" location="Cave" longLatPrecision="682" longitude="38.98" municipality="Fanagoriyskaya" pageId="29" pageNumber="30" specimenCount="1" specimenCount-female="1" stateProvince="Krasnodar" typeStatus="paratype">
|
||
<specimenCount id="9D0619FD2840FFB02C991A700FE2FD87" box="[189,234,590,617]" pageId="29" pageNumber="30" type="female">1 ♀</specimenCount>
|
||
; same collection data as for holotype but
|
||
<date id="FFBEF4B42840FFB02EF71A700C79FD86" box="[723,881,590,616]" pageId="29" pageNumber="30" value="1968-09-20">
|
||
<collectingDate id="EFFA0D5C2840FFB02EF71A700C79FD86" box="[723,881,590,616]" pageId="29" pageNumber="30" value="1968-09-20">20 Sep. 1968</collectingDate>
|
||
</date>
|
||
;
|
||
<collectionCode id="ED114AB12840FFB02F591A710CD4FD87" box="[893,988,591,617]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="29" pageNumber="30">ZMUM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A81FF2840FFB32C991AB10D76FEA7" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="description">
|
||
<paragraph id="8BBFD2742840FFB02C991AB10E46FD47" blockId="29.[189,334,655,681]" box="[189,334,655,681]" pageId="29" pageNumber="30">
|
||
<heading id="D0F765182840FFB02C991AB10E46FD47" bold="true" box="[189,334,655,681]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
|
||
<emphasis id="B9740E662840FFB02C991AB10E46FD47" bold="true" box="[189,334,655,681]" pageId="29" pageNumber="30">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742840FFB02C991A800EF8FC8B" blockId="29.[189,1399,702,869]" pageId="29" pageNumber="30">
|
||
SIZE AND NUMBER OF BODY RINGS.
|
||
<typeStatus id="54BB6CD62840FFB02E181A800DA1FD36" box="[572,681,702,728]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
|
||
male
|
||
<quantity id="4CF87F912840FFB02ED71A800C4EFD37" box="[755,838,702,729]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="29" pageNumber="30" unit="mm" value="24.0">24 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F912840FFB029761A800FE6FD12" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="mm" value="1.5">1.5 mm</quantity>
|
||
, body with 43 podous rings + 1 apodous ring + telson.
|
||
<typeStatus id="54BB6CD62840FFB02F4E1ADC0CD9FD12" box="[874,977,738,764]" pageId="29" pageNumber="30" type="paratype">Paratype</typeStatus>
|
||
male
|
||
<quantity id="4CF87F912840FFB028331ADF0B61FD12" box="[1047,1129,737,764]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="29" pageNumber="30" unit="mm" value="28.0">28 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F912840FFB02D881B3A0D10FCF1" box="[428,536,772,799]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="29" pageNumber="30" unit="mm" value="1.55">1.55 mm</quantity>
|
||
, body with 49 podous rings + 2 apodous rings + telson.
|
||
<typeStatus id="54BB6CD62840FFB028891B3B0A1CFCF1" box="[1197,1300,773,799]" pageId="29" pageNumber="30" type="paratype">Paratype</typeStatus>
|
||
females 21 and
|
||
<quantity id="4CF87F912840FFB02D361B160E6CFCAC" box="[274,356,808,834]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="29" pageNumber="30" unit="mm" value="24.0">24 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F912840FFB02F471B190CFBFCAC" box="[867,1011,807,834]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="29" pageNumber="30" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4–1.5 mm</quantity>
|
||
, body with 37 or 44 podous rings + 1 apodous ring + telson.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742840FFB02C991BB20DDDFC48" blockId="29.[189,725,908,935]" box="[189,725,908,935]" pageId="29" pageNumber="30">
|
||
COLOUR (
|
||
<figureCitation id="133BCEF12840FFB02D0C1BB20E77FC49" box="[296,383,908,935]" captionStart="Fig" captionStartId="28.[189,232,1894,1920]" captionTargetBox="[281,1306,265,1866]" captionTargetId="figure@28.[281,1306,265,1866]" captionTargetPageId="28" captionText="Fig. 16. Leucogeorgia borealis sp. nov., types from Fanagoriyskaya Cave (ZMUM). A. Paratype ♀, habitus, lateral view. B. Paratype ♀, anterior part of body, lateral view. C. Paratype ♂, posterior part of body, lateral view. D. Holotype ♂, body ring 7, lateral view. E. Paratype ♀, midbody rings, lateral view. Arrow indicated metazonal seta. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020786" httpUri="https://zenodo.org/record/4020786/files/figure.png" pageId="29" pageNumber="30">Fig. 16</figureCitation>
|
||
). Yellowish white in alcohol.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742840FFB02C991BF00A25FB30" blockId="29.[189,1399,974,1247]" pageId="29" pageNumber="30">
|
||
HEAD (
|
||
<figureCitation id="133BCEF12840FFB02D2B1BF00E74FC06" box="[271,380,974,1000]" captionStart="Fig" captionStartId="28.[189,232,1894,1920]" captionTargetBox="[281,1306,265,1866]" captionTargetId="figure@28.[281,1306,265,1866]" captionTargetPageId="28" captionText="Fig. 16. Leucogeorgia borealis sp. nov., types from Fanagoriyskaya Cave (ZMUM). A. Paratype ♀, habitus, lateral view. B. Paratype ♀, anterior part of body, lateral view. C. Paratype ♂, posterior part of body, lateral view. D. Holotype ♂, body ring 7, lateral view. E. Paratype ♀, midbody rings, lateral view. Arrow indicated metazonal seta. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020786" httpUri="https://zenodo.org/record/4020786/files/figure.png" pageId="29" pageNumber="30">Figs 16B</figureCitation>
|
||
,
|
||
<figureCitation id="133BCEF12840FFB02DA81BF00EC9FC06" box="[396,449,974,1000]" captionStart="Fig" captionStartId="30.[189,232,1832,1858]" captionTargetBox="[189,1399,370,1804]" captionTargetId="figure@30.[189,1399,370,1804]" captionTargetPageId="30" captionText="Fig. 17. Leucogeorgia borealis sp. nov., paratype ♂ from Fanagoriyskaya Cave (ZMUM). A. Leg-pair 1, anterior view. B. Right leg 1, lateral view. C. Head, ventral view. D. Left gonopods, mesal view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: A–B = 0.05 mm; C = 0.2 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020788" httpUri="https://zenodo.org/record/4020788/files/figure.png" pageId="29" pageNumber="30">17C</figureCitation>
|
||
). Without ommatidia. Frontal setae absent. Labrum with three labral teeth, four supralabral setae and 17 labral setae in
|
||
<typeStatus id="54BB6CD62840FFB02EB71BCF0DF0FBE5" box="[659,760,1009,1035]" pageId="29" pageNumber="30" type="paratype">paratype</typeStatus>
|
||
male. Gnathochilarium with a rhomboid promentum; lamellae linguales with 4+4 setae, stipites with 3+3 distolateral and 5+6 medial setae. Antennae
|
||
<quantity id="4CF87F912840FFB029391C2A0A7FFBC1" box="[1309,1399,1044,1071]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="29" pageNumber="30" unit="mm" value="1.9">1.9 mm</quantity>
|
||
long in smaller
|
||
<typeStatus id="54BB6CD62840FFB02D501C060ED1FBBC" box="[372,473,1080,1106]" pageId="29" pageNumber="30" type="paratype">paratype</typeStatus>
|
||
female, their length ca 140% of vertical diameter of largest body ring. Lengths of antennomeres I–VIII (in mm): 0.13 (I), 0.45 (II), 0.36 (III), 0.30 (IV), 0.40 (V), 0.18 (VI), 0.06 (VII) and 0.02 (VIII). Length/width ratio of antennomeres I–VII: 0.8 (I), 3 (II), 2.4 (III), 2 (IV), 2.7 (V), 1 (VI) and 0.5 (VII). Antennomeres V and VI each with a terminal corolla of large sensilla basiconica bacilliformia; antennomere VII with a terminal corolla of small sensilla basiconica bacilliformia.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742840FFB02C991D380D54FAAD" blockId="29.[189,1399,1285,1347]" pageId="29" pageNumber="30">
|
||
BODY RINGS (
|
||
<figureCitation id="133BCEF12840FFB02D7E1D3B0ECFFACE" box="[346,455,1285,1312]" captionStart="Fig" captionStartId="28.[189,232,1894,1920]" captionTargetBox="[281,1306,265,1866]" captionTargetId="figure@28.[281,1306,265,1866]" captionTargetPageId="28" captionText="Fig. 16. Leucogeorgia borealis sp. nov., types from Fanagoriyskaya Cave (ZMUM). A. Paratype ♀, habitus, lateral view. B. Paratype ♀, anterior part of body, lateral view. C. Paratype ♂, posterior part of body, lateral view. D. Holotype ♂, body ring 7, lateral view. E. Paratype ♀, midbody rings, lateral view. Arrow indicated metazonal seta. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020786" httpUri="https://zenodo.org/record/4020786/files/figure.png" pageId="29" pageNumber="30">Fig. 16E</figureCitation>
|
||
). Entire metazonal area with longitudinal striations. Length of midbody setae ca 5% of vertical diameter of rings.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742840FFB02C991D550CCCFA49" blockId="29.[189,1399,1386,1448]" pageId="29" pageNumber="30">
|
||
TELSON (
|
||
<figureCitation id="133BCEF12840FFB02D051D540E82FA6B" box="[289,394,1386,1413]" captionStart="Fig" captionStartId="28.[189,232,1894,1920]" captionTargetBox="[281,1306,265,1866]" captionTargetId="figure@28.[281,1306,265,1866]" captionTargetPageId="28" captionText="Fig. 16. Leucogeorgia borealis sp. nov., types from Fanagoriyskaya Cave (ZMUM). A. Paratype ♀, habitus, lateral view. B. Paratype ♀, anterior part of body, lateral view. C. Paratype ♂, posterior part of body, lateral view. D. Holotype ♂, body ring 7, lateral view. E. Paratype ♀, midbody rings, lateral view. Arrow indicated metazonal seta. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020786" httpUri="https://zenodo.org/record/4020786/files/figure.png" pageId="29" pageNumber="30">Fig. 16C</figureCitation>
|
||
). Epiproct with a short and blunt preanal process, covered with dorsal and lateral setae. Paraprocts rounded, setose. Hypoproct without any modifications.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742840FFB02C991DF10DD3F99B" blockId="29.[189,1399,1487,1653]" pageId="29" pageNumber="30">
|
||
LEGS IN MALES. First pair of legs modified, hook-shaped (
|
||
<figureCitation id="133BCEF12840FFB02F561DF10CC5FA07" box="[882,973,1487,1513]" captionStart="Fig" captionStartId="30.[189,232,1832,1858]" captionTargetBox="[189,1399,370,1804]" captionTargetId="figure@30.[189,1399,370,1804]" captionTargetPageId="30" captionText="Fig. 17. Leucogeorgia borealis sp. nov., paratype ♂ from Fanagoriyskaya Cave (ZMUM). A. Leg-pair 1, anterior view. B. Right leg 1, lateral view. C. Head, ventral view. D. Left gonopods, mesal view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: A–B = 0.05 mm; C = 0.2 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020788" httpUri="https://zenodo.org/record/4020788/files/figure.png" pageId="29" pageNumber="30">Figs 17</figureCitation>
|
||
A–B, 18D–E), with three complete podomeres; coxa with two setae; prefemur with four setae; femur, postfemur and tibiotarsus coalesced; femur with two setae; postfemur with one seta; tibiotarsus with a prominent distal lobe (tarsal remnant). Tip slightly tuberculated. Postfemoral and tibial ventral pads well developed on anterior legs, then gradually disappearing towards posterior legs.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742840FFB02C991EA00A26F959" blockId="29.[189,1326,1693,1720]" box="[189,1326,1693,1720]" pageId="29" pageNumber="30">
|
||
VENTRAL MARGIN OF MALE BODY RING 7 (
|
||
<figureCitation id="133BCEF12840FFB02EA71EA30DE7F956" box="[643,751,1693,1720]" captionStart="Fig" captionStartId="28.[189,232,1894,1920]" captionTargetBox="[281,1306,265,1866]" captionTargetId="figure@28.[281,1306,265,1866]" captionTargetPageId="28" captionText="Fig. 16. Leucogeorgia borealis sp. nov., types from Fanagoriyskaya Cave (ZMUM). A. Paratype ♀, habitus, lateral view. B. Paratype ♀, anterior part of body, lateral view. C. Paratype ♂, posterior part of body, lateral view. D. Holotype ♂, body ring 7, lateral view. E. Paratype ♀, midbody rings, lateral view. Arrow indicated metazonal seta. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020786" httpUri="https://zenodo.org/record/4020786/files/figure.png" pageId="29" pageNumber="30">Fig. 16D</figureCitation>
|
||
). Poorly developed, low, rounded in lateral view.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742840FFB02C991EE10BEDF916" blockId="29.[189,1253,1758,1785]" box="[189,1253,1758,1785]" pageId="29" pageNumber="30">
|
||
PENES (
|
||
<figureCitation id="133BCEF12840FFB02D361EE00E74F917" box="[274,380,1758,1785]" captionStart="Fig" captionStartId="31.[189,232,1929,1955]" captionTargetBox="[202,1379,487,1902]" captionTargetId="figure@31.[189,1399,471,1904]" captionTargetPageId="31" captionText="Fig. 18. Leucogeorgia borealis sp. nov., holotype ♂ from Fanagoriyskaya Cave (ZMUM). A. Right gonopods, mesal view. B. Right gonopods, lateral view. C. Penes, posterior view. D. Leg-pair 1, anterior view. E. Right leg 1, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020790" httpUri="https://zenodo.org/record/4020790/files/figure.png" pageId="29" pageNumber="30">Fig. 18C</figureCitation>
|
||
). In form of a short trapezoid, apically with two small subtriangular lobes.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742840FFB02C991F1E0A7FF829" blockId="29.[189,1399,1824,1991]" pageId="29" pageNumber="30">
|
||
GONOPODS (
|
||
<figureCitation id="133BCEF12840FFB02D601F1E0EB9F8D4" box="[324,433,1824,1850]" captionStart="Fig" captionStartId="30.[189,232,1832,1858]" captionTargetBox="[189,1399,370,1804]" captionTargetId="figure@30.[189,1399,370,1804]" captionTargetPageId="30" captionText="Fig. 17. Leucogeorgia borealis sp. nov., paratype ♂ from Fanagoriyskaya Cave (ZMUM). A. Leg-pair 1, anterior view. B. Right leg 1, lateral view. C. Head, ventral view. D. Left gonopods, mesal view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: A–B = 0.05 mm; C = 0.2 mm; D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020788" httpUri="https://zenodo.org/record/4020788/files/figure.png" pageId="29" pageNumber="30">Figs 17D</figureCitation>
|
||
,
|
||
<figureCitation id="133BCEF12840FFB02DE41F1E0ED5F8D4" box="[448,477,1824,1850]" captionStart="Fig" captionStartId="31.[189,232,1929,1955]" captionTargetBox="[202,1379,487,1902]" captionTargetId="figure@31.[189,1399,471,1904]" captionTargetPageId="31" captionText="Fig. 18. Leucogeorgia borealis sp. nov., holotype ♂ from Fanagoriyskaya Cave (ZMUM). A. Right gonopods, mesal view. B. Right gonopods, lateral view. C. Penes, posterior view. D. Leg-pair 1, anterior view. E. Right leg 1, lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020790" httpUri="https://zenodo.org/record/4020790/files/figure.png" pageId="29" pageNumber="30">18</figureCitation>
|
||
A–B). Promere (p) long and slender, with a flagellum (f); apical part spatulate, with denticulate margins; basal half with two developed ridges. Mesomere (m) with a well-developed, robust and denticulate mesomeral claw (mc); mesomeral lamella (ml) convex in central part, distal margin serrate, posterior part finely fimbriate. Opisthomere (o) bipartite. Anterior branch of o with a solenomere (s), with a medium-sized tip, and a well-developed and fimbriate velum (v). Posterior branch
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742843FFB32C9919350D76FEA7" blockId="30.[189,1398,267,329]" pageId="30" pageNumber="31">of o in form of a shield-like protective lamella (pl). Mesomere and opisthomere connected basally through an accessory membrane (am).</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7F82FC2843FFB32C991F160C4BF83E" ID-DOI="http://doi.org/10.5281/zenodo.4020788" ID-Zenodo-Dep="4020788" httpUri="https://zenodo.org/record/4020788/files/figure.png" pageId="30" pageNumber="31" startId="30.[189,232,1832,1858]" targetBox="[189,1399,370,1804]" targetPageId="30">
|
||
<paragraph id="8BBFD2742843FFB32C991F160C4BF83E" blockId="30.[189,1399,1831,2000]" pageId="30" pageNumber="31">
|
||
<emphasis id="B9740E662843FFB32C991F160E12F8AD" bold="true" box="[189,282,1832,1859]" pageId="30" pageNumber="31">Fig. 17.</emphasis>
|
||
<taxonomicName id="4C00A9F72843FFB32D061F140D23F8AD" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[290,555,1833,1859]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
|
||
<emphasis id="B9740E662843FFB32D061F140D23F8AD" box="[290,555,1833,1859]" italics="true" pageId="30" pageNumber="31">Leucogeorgia borealis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2843FFB32E171F170D98F8AD" box="[563,656,1833,1859]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, paratype ♂ from Fanagoriyskaya Cave (ZMUM).
|
||
<emphasis id="B9740E662843FFB328C91F160A0AF8AC" bold="true" box="[1261,1282,1832,1858]" pageId="30" pageNumber="31">A</emphasis>
|
||
. Leg-pair 1, anterior view.
|
||
<emphasis id="B9740E662843FFB32DA81F720EA8F888" bold="true" box="[396,416,1868,1894]" pageId="30" pageNumber="31">B</emphasis>
|
||
. Right leg 1, lateral view.
|
||
<emphasis id="B9740E662843FFB32ECC1F720DF5F888" bold="true" box="[744,765,1868,1894]" pageId="30" pageNumber="31">C</emphasis>
|
||
. Head, ventral view.
|
||
<emphasis id="B9740E662843FFB328271F720B10F888" bold="true" box="[1027,1048,1868,1894]" pageId="30" pageNumber="31">D</emphasis>
|
||
. Left gonopods, mesal view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: A–B = 0.05 mm; C = 0.2 mm; D = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31A81FF2842FFB22C9919350C6BFEBA" box="[189,867,267,340]" pageId="31" pageNumber="32" type="distribution">
|
||
<paragraph id="8BBFD2742842FFB22C9919350E5EFECB" blockId="31.[189,342,267,293]" box="[189,342,267,293]" pageId="31" pageNumber="32">
|
||
<heading id="D0F765182842FFB22C9919350E5EFECB" bold="true" box="[189,342,267,293]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
|
||
<emphasis id="B9740E662842FFB22C9919350E5EFECB" bold="true" box="[189,342,267,293]" pageId="31" pageNumber="32">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742842FFB22C9919040C6BFEBA" blockId="31.[189,867,313,340]" box="[189,867,313,340]" pageId="31" pageNumber="32">
|
||
Known only from its
|
||
<typeStatus id="54BB6CD62842FFB22D9E19040EE5FEBA" box="[442,493,314,340]" pageId="31" pageNumber="32">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="133BCEF12842FFB22E7F19070DBAFEBA" box="[603,690,313,340]" captionStart="Fig" captionStartId="97.[189,232,1991,2017]" captionTargetBox="[189,1399,890,1963]" captionTargetId="figure@97.[189,1399,890,1963]" captionTargetPageId="97" captionText="Fig. 57. Distribution of the northwestern group of the genus Leucogeorgia Verhoeff, 1930." figureDoi="http://doi.org/10.5281/zenodo.4020875" httpUri="https://zenodo.org/record/4020875/files/figure.png" pageId="31" pageNumber="32">Fig. 57</figureCitation>
|
||
, blue triangle).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A81FF2842FFB22C9919450C44FE2A" box="[189,844,379,452]" pageId="31" pageNumber="32" type="discussion">
|
||
<paragraph id="8BBFD2742842FFB22C9919450E27FE7B" blockId="31.[189,303,379,405]" box="[189,303,379,405]" pageId="31" pageNumber="32">
|
||
<heading id="D0F765182842FFB22C9919450E27FE7B" bold="true" box="[189,303,379,405]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
|
||
<emphasis id="B9740E662842FFB22C9919450E27FE7B" bold="true" box="[189,303,379,405]" pageId="31" pageNumber="32">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742842FFB22C9919970C44FE2A" blockId="31.[189,844,425,452]" box="[189,844,425,452]" pageId="31" pageNumber="32">This is the northwesternmost record of the entire genus.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |