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<document id="4FCF64B09053F5BD060A777741DAE2D2" ID-DOI="10.3853/j.2201-4349.75.2023.1827" ID-ISSN="2201-4349" ID-Zenodo-Dep="10414024" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1703108455257" checkinUser="felipe" docAuthor="Blake, James A." docDate="2023" docId="03A787E9FFE09746FC21F9FB5456FA03" docLanguage="en" docName="RecAustMus.75.3.271-298.pdf" docOrigin="Records of the Australian Museum (Rec. Aust. Mus.) 75 (3)" docSource="http://dx.doi.org/10.3853/j.2201-4349.75.2023.1827" docStyle="DocumentStyle:997BA24B490466A2F99594A367FA147F.4:RecAustMus.2019-.journal_article.0cover" docStyleId="997BA24B490466A2F99594A367FA147F" docStyleName="RecAustMus.2019-.journal_article.0cover" docStyleVersion="4" docTitle="Axiokebuita australis Blake 2023, sp. nov." docType="treatment" docUuid="C93CC779-84C1-4FC0-8252-FEE01BBA2968" docUuidSource="ZooBank" docVersion="6" lastPageNumber="279" masterDocId="FF9EFF91FFE6974EFF86FFEA511AFFC0" masterDocTitle="New Species of Scalibregmatidae (Annelida) from Slope and Abyssal Depths off Eastern Australia" masterLastPageNumber="298" masterPageNumber="271" pageNumber="277" updateTime="1713474973228" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="B7D8F80C9566095B0469CC7264093E0A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B604BB1B118F829F26F4DA5BC36CA736">
<mods:title id="24BCC0CF2C2AA98F1DFE45A002C30713">New Species of Scalibregmatidae (Annelida) from Slope and Abyssal Depths off Eastern Australia</mods:title>
</mods:titleInfo>
<mods:name id="2283129554E23E706008D1167F1FCA3C" type="personal">
<mods:role id="6B6D03C2FB0A7DC24B91E945A8D7A3DB">
<mods:roleTerm id="035027FF4D61C40B4926A611BE1DF00E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5F4253A676BED1F7246BDF3BF0D2F517">Blake, James A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="AF51324E28587894292929B6E1573DC4">text</mods:typeOfResource>
<mods:relatedItem id="63918D01E697E4B96F95B4CE00886AC8" type="host">
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<mods:title id="2C0DB3D216FADBF6AF3F8FE088F044D0">Records of the Australian Museum</mods:title>
</mods:titleInfo>
<mods:part id="6838D3985656E7068515CE927B62FE22">
<mods:date id="978FE5DBC6CBF29788C3ABD8AD3B6376">2023</mods:date>
<mods:detail id="79D434D8075FC23E065A2A19C80DCE28" type="series">
<mods:title id="063DAE7A98E773C87A513D702A3C8CED">Rec. Aust. Mus.</mods:title>
</mods:detail>
<mods:detail id="34A46A7F3C167AD560CE07BAD9A99509" type="pubDate">
<mods:number id="CC16491992A358F28F3A1A5157064CBA">2023-05-17</mods:number>
</mods:detail>
<mods:detail id="063002C2E6BFBA693BADA7D2F62411BE" type="volume">
<mods:number id="A0E69F3570469F6E03E7C181D7D7D224">75</mods:number>
</mods:detail>
<mods:detail id="CA984ED52B62E052FBECD52FE234E8D9" type="issue">
<mods:number id="2E970EF25890AAE70A6BA1AB1C467285">3</mods:number>
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<mods:extent id="4A7B56A50A368E419F86C129A2887ABD" unit="page">
<mods:start id="1EEE4747B842497520BCBA3501B3B49C">271</mods:start>
<mods:end id="40C880B1AE83D9133AE440FE25B9ED41">298</mods:end>
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<mods:location id="49A7C6C24EA62E67A997F42E32F909CD">
<mods:url id="A9621AF38B1692D5E970698B90002DD0">http://dx.doi.org/10.3853/j.2201-4349.75.2023.1827</mods:url>
</mods:location>
<mods:classification id="E3F4C9D8B423F8C4B8FCD0121E699A56">journal article</mods:classification>
<mods:identifier id="59C031E1AD3163004247C65821B1F0F2" type="DOI">10.3853/j.2201-4349.75.2023.1827</mods:identifier>
<mods:identifier id="D263B5CCE3CF1A7F455845589B07C978" type="ISSN">2201-4349</mods:identifier>
<mods:identifier id="2459627AEEFDA1E9A213A8F6F17BF454" type="Zenodo-Dep">10414024</mods:identifier>
<mods:identifier id="7E2B4D30910235612E4CF0E296095697" type="ZooBank">7D3BDF25-010F-41A4-AD15-763C3F067D8A</mods:identifier>
</mods:mods>
<treatment id="03A787E9FFE09746FC21F9FB5456FA03" ID-DOI="http://doi.org/10.5281/zenodo.10995052" ID-Zenodo-Dep="10995052" LSID="urn:lsid:zoobank.org:act:C93CC779-84C1-4FC0-8252-FEE01BBA2968" httpUri="http://treatment.plazi.org/id/03A787E9FFE09746FC21F9FB5456FA03" lastPageId="8" lastPageNumber="279" pageId="6" pageNumber="277">
<subSubSection id="C3146574FFE09748FC21F9FB54B6F994" pageId="6" pageNumber="277" type="nomenclature">
<paragraph id="8BB136FFFFE09748FC21F9FB5440F9F1" blockId="6.[935,1370,1553,1585]" box="[935,1370,1553,1585]" pageId="6" pageNumber="277">
<heading id="D0F98193FFE09748FC21F9FB5440F9F1" bold="true" box="[935,1370,1553,1585]" centered="true" fontSize="12" level="2" pageId="6" pageNumber="277" reason="2">
<emphasis id="B97AEAEDFFE09748FC21F9FB5440F9F1" bold="true" box="[935,1370,1553,1585]" pageId="6" pageNumber="277">
<taxonomicName id="4C0E4D7CFFE09748FC21F9FB55FAF9F0" ID-CoL="BDS5B" authority="Blake, 2023" authorityName="Blake" authorityYear="2023" box="[935,1248,1553,1584]" class="Polychaeta" family="Scalibregmatidae" genus="Axiokebuita" kingdom="Animalia" pageId="6" pageNumber="277" phylum="Annelida" rank="species" species="australis" status="sp. nov.">
<emphasis id="B97AEAEDFFE09748FC21F9FB55FAF9F0" bold="true" box="[935,1248,1553,1584]" italics="true" pageId="6" pageNumber="277">Axiokebuita australis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2495796FFE09748FB6FF9F85440F9F1" box="[1257,1370,1554,1585]" pageId="6" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB136FFFFE09748FCD3F9A854B6F994" blockId="6.[853,1452,1602,1620]" box="[853,1452,1602,1620]" pageId="6" pageNumber="277">
<uri id="FF9F3AFDFFE09748FCD3F9A854B6F994" box="[853,1452,1602,1620]" pageId="6" pageNumber="277">
urn:lsid:zoobank.org:act:
<uuid id="FFA80C2AFFE09748FBA6F9A854B6F994" box="[1056,1452,1602,1620]" pageId="6" pageNumber="277">C93CC779-84C1-4FC0-8252-FEE01BBA2968</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3146574FFE09748FBCBF99B55AFF94C" box="[1101,1205,1649,1676]" pageId="6" pageNumber="277" type="description">
<paragraph id="8BB136FFFFE09748FBCBF99B55AFF94C" blockId="6.[1101,1205,1649,1676]" box="[1101,1205,1649,1676]" pageId="6" pageNumber="277">
<figureCitation id="13352A7AFFE09748FBCBF99B55AFF94C" box="[1101,1205,1649,1676]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStartId-0="7.[121,189,1657,1679]" captionStartId-1="8.[166,233,983,1005]" captionTargetBox-0="[121,1421,151,1645]" captionTargetBox-1="[184,1444,161,957]" captionTargetId-0="figure-94@7.[121,1421,151,1645]" captionTargetId-1="figure-6@8.[167,1467,148,983]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figure 3. Axiokebuita australis sp. nov. (A) Anterior two thirds of holotype, lateral view; (B) detail of pre-setiger region of same; (C) entire paratype in dorsolateral view; (D) posterior end, right lateral view. A, B, holotype (AM W.52681); C, D, paratype (AM W.53989). All stained with Shirlastain A." captionText-1="Figure 4. Axiokebuita australis sp.nov.(A) anterior end, dorsal view; (B) setiger 9, anterior view.A, paratype (AM W.53989); B, holotype (AM W.52681)." figureDoi-0="http://doi.org/10.5281/zenodo.10414030" figureDoi-1="http://doi.org/10.5281/zenodo.10414032" httpUri-0="https://zenodo.org/record/10414030/files/figure.png" httpUri-1="https://zenodo.org/record/10414032/files/figure.png" pageId="6" pageNumber="277">Figs 3–4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3146574FFE09748FCC1F94B55A1F863" pageId="6" pageNumber="277" type="materials_examined">
<paragraph id="8BB136FFFFE09748FCC1F94B55A1F863" blockId="6.[839,1467,1697,1955]" pageId="6" pageNumber="277">
<materialsCitation id="3B663CA2FFE09748FCC1F94B52AEF8D0" collectingDate="2017-05-19" collectionCode="AM" country="Australia" elevation="2793" latitude="-41.721" location="Freycinet Marine Park" longLatPrecision="69" longitude="149.125" pageId="6" pageNumber="277" specimenCount="1" stateProvince="Tasmania" typeStatus="holotype">
<emphasis id="B97AEAEDFFE09748FCC1F94B52B4F979" bold="true" box="[839,942,1697,1721]" pageId="6" pageNumber="277">
<typeStatus id="54B5885DFFE09748FCC1F94B52B4F979" box="[839,942,1697,1721]" pageId="6" pageNumber="277" type="holotype">Holotype</typeStatus>
</emphasis>
: eastern
<collectingCountry id="F319766FFFE09748FB8FF94B5577F979" box="[1033,1133,1697,1721]" name="Australia" pageId="6" pageNumber="277">Australia</collectingCountry>
, continental slope off Tasmania,
<location id="8ED16024FFE09748FCC1F9545527F916" LSID="urn:lsid:plazi:treatment:03A787E9FFE09746FC21F9FB5456FA03:8ED16024FFE09748FCC1F9545527F916" box="[839,1085,1726,1750]" country="Australia" latitude="-41.721" longLatPrecision="69" longitude="149.125" name="Freycinet Marine Park" pageId="6" pageNumber="277" stateProvince="Tasmania">Freycinet Marine Park</location>
, RV
<taxonomicName id="4C0E4D7CFFE09748FBF5F9555436F916" authority=", Sta." authorityName="Sta." box="[1139,1324,1726,1750]" class="Actinopterygii" family="Nemichthyidae" genus="Investigator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="277" phylum="Chordata" rank="genus">
<emphasis id="B97AEAEDFFE09748FBF5F95555EEF916" box="[1139,1268,1727,1750]" italics="true" pageId="6" pageNumber="277">Investigator</emphasis>
, Sta.
</taxonomicName>
011, coll.
<date id="FFB0103FFFE09748FA26F95452A0F933" pageId="6" pageNumber="277" value="2017-05-19">
<collectingDate id="EFF4E9D7FFE09748FA26F95452A0F933" pageId="6" pageNumber="277" value="2017-05-19">19 May 2017</collectingDate>
</date>
, box core,
<geoCoordinate id="EE3A5038FFE09748FBBFF9315586F933" box="[1081,1180,1755,1779]" degrees="41.721" direction="south" orientation="latitude" pageId="6" pageNumber="277" precision="55" value="-41.721">41.721°S</geoCoordinate>
<geoCoordinate id="EE3A5038FFE09748FB20F9315400F933" box="[1190,1306,1755,1779]" degrees="149.125" direction="east" orientation="longitude" pageId="6" pageNumber="277" precision="55" value="149.125">149.125°E</geoCoordinate>
,
<quantity id="4CF69B1AFFE09748FAAEF9315467F933" box="[1320,1405,1755,1779]" metricMagnitude="3" metricUnit="m" metricValue="2.793" pageId="6" pageNumber="277" unit="m" value="2793.0">
<elevation id="0023D1CCFFE09748FAAEF9315467F933" box="[1320,1405,1755,1779]" metricMagnitude="3" metricUnit="m" metricValue="2.793" pageId="6" pageNumber="277" unit="m" value="2793.0">2793 m</elevation>
</quantity>
(
<collectionCode id="ED1FAE3AFFE09748FA08F93154A1F933" box="[1422,1467,1755,1779]" country="Australia" name="Australian Museum" pageId="6" pageNumber="277" type="Museum">AM</collectionCode>
W.52681)
</materialsCitation>
.
<materialsCitation id="3B663CA2FFE09748FC46F91352A1F8EE" collectingDate="2017-05-24" collectionCode="AM" country="Australia" elevation="4064" latitude="-41.721" location="East Gippsland Marine Park" longLatPrecision="69" longitude="149.125" pageId="6" pageNumber="277" specimenCount="1" stateProvince="Victoria" typeStatus="paratype">
<emphasis id="B97AEAEDFFE09748FC46F9135528F8D1" bold="true" box="[960,1074,1785,1809]" pageId="6" pageNumber="277">
<typeStatus id="54B5885DFFE09748FC46F9135528F8D1" box="[960,1074,1785,1809]" pageId="6" pageNumber="277" type="paratype">Paratypes</typeStatus>
</emphasis>
(2): same data as holotype (1,
<collectionCode id="ED1FAE3AFFE09748FA16F91354A1F8D1" box="[1424,1467,1785,1809]" country="Australia" name="Australian Museum" pageId="6" pageNumber="277" type="Museum">AM</collectionCode>
W.53989);
</materialsCitation>
<materialsCitation id="3B663CA2FFE09748FC40F8FC55ADF863" collectingDate="2017-05-24" collectionCode="AM" country="Australia" elevation="4064" latitude="-41.721" location="East Gippsland Marine Park" longLatPrecision="69" longitude="149.125" pageId="6" pageNumber="277" specimenCode="W.52695" specimenCount="1" stateProvince="Victoria" typeStatus="paratype">
abyssal plain off southeastern
<collectingRegion id="49CAF81DFFE09748FAA7F8FC5467F8EE" box="[1313,1405,1814,1838]" country="Australia" name="Victoria" pageId="6" pageNumber="277">Victoria</collectingRegion>
,
<location id="8ED16024FFE09748FA0DF8FC5554F88B" LSID="urn:lsid:plazi:treatment:03A787E9FFE09746FC21F9FB5456FA03:8ED16024FFE09748FA0DF8FC5554F88B" country="Australia" latitude="-41.721" longLatPrecision="69" longitude="149.125" name="East Gippsland Marine Park" pageId="6" pageNumber="277" stateProvince="Victoria">East Gippsland Marine Park</location>
, RV
<taxonomicName id="4C0E4D7CFFE09748FB0FF8DE545DF88B" authority=", Sta." authorityName="Sta." box="[1161,1351,1843,1867]" class="Actinopterygii" family="Nemichthyidae" genus="Investigator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="277" phylum="Chordata" rank="genus">
<emphasis id="B97AEAEDFFE09748FB0FF8DE5416F88B" box="[1161,1292,1844,1867]" italics="true" pageId="6" pageNumber="277">Investigator</emphasis>
, Sta.
</taxonomicName>
033, coll.
<date id="FFB0103FFFE09748FCC1F8BB52F9F8A9" box="[839,995,1873,1897]" pageId="6" pageNumber="277" value="2017-05-24">
<collectingDate id="EFF4E9D7FFE09748FCC1F8BB52F9F8A9" box="[839,995,1873,1897]" pageId="6" pageNumber="277" value="2017-05-24">24 May 2017</collectingDate>
</date>
, Brenke Sled, distance
<quantity id="4CF69B1AFFE09748FB75F8BB545FF8A9" box="[1267,1349,1873,1897]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="6" pageNumber="277" unit="km" value="2.6">2.6 km</quantity>
, 38.521°S 153.213°E 4107 m to
<geoCoordinate id="EE3A5038FFE09748FBA1F8845592F846" box="[1063,1160,1902,1926]" degrees="38.498" direction="south" orientation="latitude" pageId="6" pageNumber="277" precision="55" value="-38.498">38.498°S</geoCoordinate>
<geoCoordinate id="EE3A5038FFE09748FB0BF88455E7F846" box="[1165,1277,1902,1926]" degrees="150.207" direction="east" orientation="longitude" pageId="6" pageNumber="277" precision="55" value="150.207">150.207°E</geoCoordinate>
<quantity id="4CF69B1AFFE09748FA87F8845448F845" box="[1281,1362,1902,1926]" metricMagnitude="3" metricUnit="m" metricValue="4.064" pageId="6" pageNumber="277" unit="m" value="4064.0">
<elevation id="0023D1CCFFE09748FA87F8845448F845" box="[1281,1362,1902,1926]" metricMagnitude="3" metricUnit="m" metricValue="4.064" pageId="6" pageNumber="277" unit="m" value="4064.0">4064 m</elevation>
</quantity>
, dissected in part for DNA (1,
<collectionCode id="ED1FAE3AFFE09748FB9FF861555EF863" box="[1049,1092,1931,1955]" country="Australia" name="Australian Museum" pageId="6" pageNumber="277" type="Museum">AM</collectionCode>
<specimenCode id="DBA89E84FFE09748FBCCF86155B5F863" box="[1098,1199,1931,1955]" pageId="6" pageNumber="277">W.52695</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146574FFE09746FCC1F8525305F916" lastPageId="8" lastPageNumber="279" pageId="6" pageNumber="277" type="description">
<paragraph id="8BB136FFFFE09749FCC1F852543AF813" blockId="6.[839,1466,1976,2001]" lastBlockId="7.[793,1422,1773,2003]" lastPageId="7" lastPageNumber="278" pageId="6" pageNumber="277">
<emphasis id="B97AEAEDFFE09748FCC1F85252D0F810" bold="true" box="[839,970,1976,2000]" pageId="6" pageNumber="277">Description</emphasis>
. All
<specimenCount id="9D08FD76FFE09748FB86F85355B1F811" box="[1024,1195,1977,2001]" count="3" pageId="6" pageNumber="277" type="generic">three specimens</specimenCount>
complete:
<typeStatus id="54B5885DFFE09748FAA5F853549AF811" box="[1315,1408,1977,2001]" pageId="6" pageNumber="277" type="holotype">holotype</typeStatus>
(AM W.52681) with 22 setigers,
<quantity id="4CF69B1AFFE19749FE16F90750E7F8C5" box="[400,509,1773,1797]" metricMagnitude="-2" metricUnit="m" metricValue="1.055" pageId="7" pageNumber="278" unit="mm" value="10.55">10. 55 mm</quantity>
long and about
<quantity id="4CF69B1AFFE19749FD18F90753F6F8C5" box="[670,748,1773,1797]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="278" unit="mm" value="1.8">1.8 mm</quantity>
wide;
<typeStatus id="54B5885DFFE19749FF31F8E1500BF8E3" box="[183,273,1803,1827]" pageId="7" pageNumber="278" type="paratype">paratype</typeStatus>
from Sta. 011 (AM W.53989) with 19 setigers,
<quantity id="4CF69B1AFFE19749FFFFF8C251A9F8FF" box="[121,179,1832,1856]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="278" unit="mm" value="8.0">8 mm</quantity>
long and
<quantity id="4CF69B1AFFE19749FE95F8C25054F8FF" box="[275,334,1832,1856]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="278" unit="mm" value="2.0">2 mm</quantity>
wide;
<typeStatus id="54B5885DFFE19749FE16F8C250F3F880" box="[400,489,1832,1856]" pageId="7" pageNumber="278" type="paratype">paratype</typeStatus>
(AM W.52695) from Sta. 033 complete, but damaged and in two parts, with 19 setigers, about
<quantity id="4CF69B1AFFE19749FF38F889501AF8BA" box="[190,256,1891,1915]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="278" unit="mm" value="6.0">6 mm</quantity>
long, about
<quantity id="4CF69B1AFFE19749FE16F88950FCF8BA" box="[400,486,1891,1915]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="278" unit="mm" value="1.2">1.2 mm</quantity>
wide (posterior section removed for DNA). Body thick, sausage-shaped; more or less rectangular in cross section (
<figureCitation id="13352A7AFFE19749FE2AF877533BF875" box="[428,545,1949,1973]" captionStart="Figure 3" captionStartId="7.[121,189,1657,1679]" captionTargetBox="[121,1421,151,1645]" captionTargetId="figure-94@7.[121,1421,151,1645]" captionTargetPageId="7" captionText="Figure 3. Axiokebuita australis sp. nov. (A) Anterior two thirds of holotype, lateral view; (B) detail of pre-setiger region of same; (C) entire paratype in dorsolateral view; (D) posterior end, right lateral view. A, B, holotype (AM W.52681); C, D, paratype (AM W.53989). All stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.10414030" httpUri="https://zenodo.org/record/10414030/files/figure.png" pageId="7" pageNumber="278">Fig. 3A, C</figureCitation>
), without dorsal or ventral grooves or ridges along body. Segments with one or two rows of elevated pads across dorsum; setigers 1–4 with a biannulate pattern (
<figureCitation id="13352A7AFFE19749FC76F8E1555EF8E3" box="[1008,1092,1803,1827]" captionStart="Figure 4" captionStartId="8.[166,233,983,1005]" captionTargetBox="[184,1444,161,957]" captionTargetId="figure-6@8.[167,1467,148,983]" captionTargetPageId="8" captionText="Figure 4. Axiokebuita australis sp.nov.(A) anterior end, dorsal view; (B) setiger 9, anterior view.A, paratype (AM W.53989); B, holotype (AM W.52681)." figureDoi="http://doi.org/10.5281/zenodo.10414032" httpUri="https://zenodo.org/record/10414032/files/figure.png" pageId="7" pageNumber="278">Fig. 4A</figureCitation>
); most of body with uniannulate segments; ventrally, each anterior and middle segment with one to four annulated rows, these narrow, with separate raised pads indistinct, appearing more or less as single elevated ridge; posteriorly ventral surfaces with a single row of raised pads more or less continuous with dorsal rows (
<figureCitation id="13352A7AFFE19749FA88F877549AF875" box="[1294,1408,1949,1973]" captionStart="Figure 3" captionStartId="7.[121,189,1657,1679]" captionTargetBox="[121,1421,151,1645]" captionTargetId="figure-94@7.[121,1421,151,1645]" captionTargetPageId="7" captionText="Figure 3. Axiokebuita australis sp. nov. (A) Anterior two thirds of holotype, lateral view; (B) detail of pre-setiger region of same; (C) entire paratype in dorsolateral view; (D) posterior end, right lateral view. A, B, holotype (AM W.52681); C, D, paratype (AM W.53989). All stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.10414030" httpUri="https://zenodo.org/record/10414030/files/figure.png" pageId="7" pageNumber="278">Fig. 3C, D</figureCitation>
). Colour in alcohol light tan with no pigmentation.
</paragraph>
<caption id="DF716677FFE19749FFFFF9935086F904" ID-DOI="http://doi.org/10.5281/zenodo.10414030" ID-Zenodo-Dep="10414030" httpUri="https://zenodo.org/record/10414030/files/figure.png" pageId="7" pageNumber="278" startId="7.[121,189,1657,1679]" targetBox="[121,1421,151,1645]" targetPageId="7" targetType="figure">
<paragraph id="8BB136FFFFE19749FFFFF9935086F904" blockId="7.[121,1422,1657,1733]" pageId="7" pageNumber="278">
<emphasis id="B97AEAEDFFE19749FFFFF99351D5F94E" bold="true" box="[121,207,1657,1679]" pageId="7" pageNumber="278">Figure 3</emphasis>
.
<taxonomicName id="4C0E4D7CFFE19749FF65F99050AAF94F" authority="Blake, 2023" authorityName="Blake" authorityYear="2023" box="[227,432,1658,1679]" class="Polychaeta" family="Scalibregmatidae" genus="Axiokebuita" kingdom="Animalia" pageId="7" pageNumber="278" phylum="Annelida" rank="species" species="australis" status="sp. nov.">
<emphasis id="B97AEAEDFFE19749FF65F99050AAF94F" box="[227,432,1658,1679]" italics="true" pageId="7" pageNumber="278">Axiokebuita australis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2495796FFE19749FE31F9905318F94F" box="[439,514,1658,1679]" pageId="7" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="B97AEAEDFFE19749FD96F9905305F94F" box="[528,543,1658,1679]" italics="true" pageId="7" pageNumber="278">A</emphasis>
) Anterior two thirds of holotype, lateral view; (
<emphasis id="B97AEAEDFFE19749FC68F99052E7F94F" box="[1006,1021,1658,1679]" italics="true" pageId="7" pageNumber="278">B</emphasis>
) detail of pre-setiger region of same; (
<emphasis id="B97AEAEDFFE19749FAF3F990549FF94F" box="[1397,1413,1658,1679]" italics="true" pageId="7" pageNumber="278">C</emphasis>
) entire paratype in dorsolateral view; (
<emphasis id="B97AEAEDFFE19749FE66F97F50EBF96A" box="[480,497,1685,1706]" italics="true" pageId="7" pageNumber="278">D</emphasis>
) posterior end, right lateral view. A, B, holotype (AM W.52681); C, D, paratype (AM W.53989). All stained with Shirlastain A.
</paragraph>
</caption>
<caption id="DF716677FFEE9746FF20FC3D505AFBC8" ID-DOI="http://doi.org/10.5281/zenodo.10414032" ID-Zenodo-Dep="10414032" httpUri="https://zenodo.org/record/10414032/files/figure.png" pageId="8" pageNumber="279" startId="8.[166,233,983,1005]" targetBox="[184,1444,161,957]" targetPageId="8" targetType="figure">
<paragraph id="8BB136FFFFEE9746FF20FC3D505AFBC8" blockId="8.[166,1467,983,1032]" pageId="8" pageNumber="279">
<emphasis id="B97AEAEDFFEE9746FF20FC3D51E0FC2C" bold="true" box="[166,250,983,1005]" pageId="8" pageNumber="279">Figure 4</emphasis>
.
<taxonomicName id="4C0E4D7CFFEE9746FE81FC3D50C8FC2C" authority="Blake, 2023" authorityName="Blake" authorityYear="2023" box="[263,466,983,1004]" class="Polychaeta" family="Scalibregmatidae" genus="Axiokebuita" kingdom="Animalia" pageId="8" pageNumber="279" phylum="Annelida" rank="species" species="australis" status="sp. nov.">
<emphasis id="B97AEAEDFFEE9746FE81FC3D50C8FC2C" box="[263,466,983,1004]" italics="true" pageId="8" pageNumber="279">Axiokebuita australis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2495796FFEE9746FE5EFC32533BFC2D" box="[472,545,984,1005]" pageId="8" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
(
<emphasis id="B97AEAEDFFEE9746FDAAFC325321FC2D" box="[556,571,984,1005]" italics="true" pageId="8" pageNumber="279">A</emphasis>
) anterior end, dorsal view; (
<emphasis id="B97AEAEDFFEE9746FCC5FC325248FC2D" box="[835,850,984,1005]" italics="true" pageId="8" pageNumber="279">B</emphasis>
) setiger 9, anterior view.A, paratype (AM W.53989); B, holotype (AM W.52681).
</paragraph>
</caption>
<paragraph id="8BB136FFFFEE9746FF43FBD2535BFA52" blockId="8.[166,795,1080,1750]" pageId="8" pageNumber="279">
Prostomium pentagonal in shape (
<figureCitation id="13352A7AFFEE9746FD99FBD25395FB90" box="[543,655,1080,1104]" captionStart="Figure 3" captionStartId="7.[121,189,1657,1679]" captionTargetBox="[121,1421,151,1645]" captionTargetId="figure-94@7.[121,1421,151,1645]" captionTargetPageId="7" captionText="Figure 3. Axiokebuita australis sp. nov. (A) Anterior two thirds of holotype, lateral view; (B) detail of pre-setiger region of same; (C) entire paratype in dorsolateral view; (D) posterior end, right lateral view. A, B, holotype (AM W.52681); C, D, paratype (AM W.53989). All stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.10414030" httpUri="https://zenodo.org/record/10414030/files/figure.png" pageId="8" pageNumber="279">Figs 3A, B</figureCitation>
,
<figureCitation id="13352A7AFFEE9746FD1FFBD253A6FB90" box="[665,700,1080,1104]" captionStart="Figure 4" captionStartId="8.[166,233,983,1005]" captionTargetBox="[184,1444,161,957]" captionTargetId="figure-6@8.[167,1467,148,983]" captionTargetPageId="8" captionText="Figure 4. Axiokebuita australis sp.nov.(A) anterior end, dorsal view; (B) setiger 9, anterior view.A, paratype (AM W.53989); B, holotype (AM W.52681)." figureDoi="http://doi.org/10.5281/zenodo.10414032" httpUri="https://zenodo.org/record/10414032/files/figure.png" pageId="8" pageNumber="279">4A</figureCitation>
), straight across anterior end with two thick lateral horns arising from narrow subterminal sides (
<figureCitation id="13352A7AFFEE9746FE3EFB995331FB4A" box="[440,555,1139,1163]" captionStart="Figure 3" captionStartId="7.[121,189,1657,1679]" captionTargetBox="[121,1421,151,1645]" captionTargetId="figure-94@7.[121,1421,151,1645]" captionTargetPageId="7" captionText="Figure 3. Axiokebuita australis sp. nov. (A) Anterior two thirds of holotype, lateral view; (B) detail of pre-setiger region of same; (C) entire paratype in dorsolateral view; (D) posterior end, right lateral view. A, B, holotype (AM W.52681); C, D, paratype (AM W.53989). All stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.10414030" httpUri="https://zenodo.org/record/10414030/files/figure.png" pageId="8" pageNumber="279">Figs 3A, B</figureCitation>
,
<figureCitation id="13352A7AFFEE9746FDB2FB995342FB4B" box="[564,600,1139,1163]" captionStart="Figure 4" captionStartId="8.[166,233,983,1005]" captionTargetBox="[184,1444,161,957]" captionTargetId="figure-6@8.[167,1467,148,983]" captionTargetPageId="8" captionText="Figure 4. Axiokebuita australis sp.nov.(A) anterior end, dorsal view; (B) setiger 9, anterior view.A, paratype (AM W.53989); B, holotype (AM W.52681)." figureDoi="http://doi.org/10.5281/zenodo.10414032" httpUri="https://zenodo.org/record/10414032/files/figure.png" pageId="8" pageNumber="279">4A</figureCitation>
), followed by long curving borders meeting mid-dorsally; eyes absent, nuchal organs not observed. Peristomium irregularly encompassing prostomium dorsolaterally (
<figureCitation id="13352A7AFFEE9746FE40FB215323FB22" box="[454,569,1227,1251]" captionStart="Figure 3" captionStartId="7.[121,189,1657,1679]" captionTargetBox="[121,1421,151,1645]" captionTargetId="figure-94@7.[121,1421,151,1645]" captionTargetPageId="7" captionText="Figure 3. Axiokebuita australis sp. nov. (A) Anterior two thirds of holotype, lateral view; (B) detail of pre-setiger region of same; (C) entire paratype in dorsolateral view; (D) posterior end, right lateral view. A, B, holotype (AM W.52681); C, D, paratype (AM W.53989). All stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.10414030" httpUri="https://zenodo.org/record/10414030/files/figure.png" pageId="8" pageNumber="279">Figs 3A, B</figureCitation>
,
<figureCitation id="13352A7AFFEE9746FDC4FB215379FB23" box="[578,611,1227,1251]" captionStart="Figure 4" captionStartId="8.[166,233,983,1005]" captionTargetBox="[184,1444,161,957]" captionTargetId="figure-6@8.[167,1467,148,983]" captionTargetPageId="8" captionText="Figure 4. Axiokebuita australis sp.nov.(A) anterior end, dorsal view; (B) setiger 9, anterior view.A, paratype (AM W.53989); B, holotype (AM W.52681)." figureDoi="http://doi.org/10.5281/zenodo.10414032" httpUri="https://zenodo.org/record/10414032/files/figure.png" pageId="8" pageNumber="279">4A</figureCitation>
) and surrounding mouth ventrally; proboscis emergent, with numerous elongate lobes surrounding oral opening (
<figureCitation id="13352A7AFFEE9746FDD3FAEC53B0FADE" box="[597,682,1286,1310]" captionStart="Figure 3" captionStartId="7.[121,189,1657,1679]" captionTargetBox="[121,1421,151,1645]" captionTargetId="figure-94@7.[121,1421,151,1645]" captionTargetPageId="7" captionText="Figure 3. Axiokebuita australis sp. nov. (A) Anterior two thirds of holotype, lateral view; (B) detail of pre-setiger region of same; (C) entire paratype in dorsolateral view; (D) posterior end, right lateral view. A, B, holotype (AM W.52681); C, D, paratype (AM W.53989). All stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.10414030" httpUri="https://zenodo.org/record/10414030/files/figure.png" pageId="8" pageNumber="279">Figs 3B</figureCitation>
,
<figureCitation id="13352A7AFFEE9746FD35FAEC53C2FADE" box="[691,728,1286,1310]" captionStart="Figure 4" captionStartId="8.[166,233,983,1005]" captionTargetBox="[184,1444,161,957]" captionTargetId="figure-6@8.[167,1467,148,983]" captionTargetPageId="8" captionText="Figure 4. Axiokebuita australis sp.nov.(A) anterior end, dorsal view; (B) setiger 9, anterior view.A, paratype (AM W.53989); B, holotype (AM W.52681)." figureDoi="http://doi.org/10.5281/zenodo.10414032" httpUri="https://zenodo.org/record/10414032/files/figure.png" pageId="8" pageNumber="279">4A</figureCitation>
); neck organs observed on
<typeStatus id="54B5885DFFEE9746FE03FAC950FBFAFB" box="[389,481,1315,1339]" pageId="8" pageNumber="279" type="paratype">paratype</typeStatus>
(AM W.52695) as a pair of swellings ventrolateral on enlarged oral area; these obscured on
<typeStatus id="54B5885DFFEE9746FF40FAB45039FAB6" box="[198,291,1374,1398]" pageId="8" pageNumber="279" type="holotype">holotype</typeStatus>
and other
<typeStatus id="54B5885DFFEE9746FE14FAB450F4FAB6" box="[402,494,1374,1398]" pageId="8" pageNumber="279" type="paratype">paratype</typeStatus>
due to ventral curvature and compression of entire pre-setiger area.
</paragraph>
<paragraph id="8BB136FFFFEE9746FF43FA7250B4F95B" blockId="8.[166,795,1080,1750]" pageId="8" pageNumber="279">
Parapodia reduced to blister-like swellings throughout, with noto- and neurosetae arising from notch on posterior sides. Notopodia with single short, oval to rounded postsetal lamella, this less conspicuous in posterior setigers; neuropodia with an elongate, fingerlike lamella; both lamellae from setiger 2 or 3 continuing to posterior end (
<figureCitation id="13352A7AFFEE9746FF28F9A2501CF9A0" box="[174,262,1608,1632]" captionStart="Figure 4" captionStartId="8.[166,233,983,1005]" captionTargetBox="[184,1444,161,957]" captionTargetId="figure-6@8.[167,1467,148,983]" captionTargetPageId="8" captionText="Figure 4. Axiokebuita australis sp.nov.(A) anterior end, dorsal view; (B) setiger 9, anterior view.A, paratype (AM W.53989); B, holotype (AM W.52681)." figureDoi="http://doi.org/10.5281/zenodo.10414032" httpUri="https://zenodo.org/record/10414032/files/figure.png" pageId="8" pageNumber="279">Fig. 4B</figureCitation>
). Setae all thin capillaries in spreading fascicles throughout; furcate setae not observed. Notosetae long, with up to 35–45 per fascicle.
</paragraph>
<paragraph id="8BB136FFFFEE9746FF43F94A5305F916" blockId="8.[166,795,1080,1750]" pageId="8" pageNumber="279">
Pygidium a simple plate composed of 6–8 thickened lobes surrounded anal opening (
<figureCitation id="13352A7AFFEE9746FE3AF9545308F916" box="[444,530,1726,1750]" captionStart="Figure 3" captionStartId="7.[121,189,1657,1679]" captionTargetBox="[121,1421,151,1645]" captionTargetId="figure-94@7.[121,1421,151,1645]" captionTargetPageId="7" captionText="Figure 3. Axiokebuita australis sp. nov. (A) Anterior two thirds of holotype, lateral view; (B) detail of pre-setiger region of same; (C) entire paratype in dorsolateral view; (D) posterior end, right lateral view. A, B, holotype (AM W.52681); C, D, paratype (AM W.53989). All stained with Shirlastain A." figureDoi="http://doi.org/10.5281/zenodo.10414030" httpUri="https://zenodo.org/record/10414030/files/figure.png" pageId="8" pageNumber="279">Fig. 3D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3146574FFEE9746FF20F901545EFB68" pageId="8" pageNumber="279" type="discussion">
<paragraph id="8BB136FFFFEE9746FF20F901545EFB68" blockId="8.[166,794,1771,2000]" lastBlockId="8.[839,1467,1080,1192]" pageId="8" pageNumber="279">
<emphasis id="B97AEAEDFFEE9746FF20F9015016F8C3" bold="true" box="[166,268,1771,1795]" pageId="8" pageNumber="279">Remarks</emphasis>
.
<taxonomicName id="4C0E4D7CFFEE9746FE9EF90150ECF8C3" authority="Blake, 2023" authorityName="Blake" authorityYear="2023" box="[280,502,1771,1795]" class="Polychaeta" family="Scalibregmatidae" genus="Axiokebuita" kingdom="Animalia" pageId="8" pageNumber="279" phylum="Annelida" rank="species" species="australis" status="sp. nov.">
<emphasis id="B97AEAEDFFEE9746FE9EF90150ECF8C3" box="[280,502,1771,1795]" italics="true" pageId="8" pageNumber="279">Axiokebuita australis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2495796FFEE9746FE7CF9015350F8C2" box="[506,586,1771,1794]" pageId="8" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to
<taxonomicName id="4C0E4D7CFFEE9746FC85F90151FBF8E0" authorityName="Pocklington &amp; Fournier" authorityYear="1987" class="Polychaeta" family="Scalibregmatidae" genus="Axiokebuita" kingdom="Animalia" pageId="8" pageNumber="279" phylum="Annelida" rank="species" species="millsi">
<emphasis id="B97AEAEDFFEE9746FC85F90151FBF8E0" italics="true" pageId="8" pageNumber="279">A. millsi</emphasis>
</taxonomicName>
, the type-species, in lacking papillated pygidial lobes. Specimens of
<taxonomicName id="4C0E4D7CFFEE9746FEC2F8CC5085F8FE" authorityName="Pocklington &amp; Fournier" authorityYear="1987" box="[324,415,1830,1854]" class="Polychaeta" family="Scalibregmatidae" genus="Axiokebuita" kingdom="Animalia" pageId="8" pageNumber="279" phylum="Annelida" rank="species" species="millsi">
<emphasis id="B97AEAEDFFEE9746FEC2F8CC5085F8FE" box="[324,415,1830,1854]" italics="true" pageId="8" pageNumber="279">A. millsi</emphasis>
</taxonomicName>
from off New
<collectingCountry id="F319766FFFEE9746FDCCF8CC53B9F8FE" box="[586,675,1830,1854]" name="United Kingdom" pageId="8" pageNumber="279">England</collectingCountry>
in bathyal depths were observed as part of a separate study (Blake in preparation) to have a fusiform-shaped body, two large non-papillated lobes around the anal opening, and 2–3 rows of annulated pads along the body; these observations agree with the published observations of
<bibRefCitation id="EF9F4B0EFFEE9746FD89F8525297FB90" author="Pocklington, P. &amp; J. A. Fournier" box="[527,909,1080,2000]" pageId="8" pageNumber="279" pagination="108 - 113" refId="ref17927" refString="Pocklington, P., and J. A. Fournier. 1987. Axiokebuita millsi, new genus, new species (Polychaeta: Scalibregmatidae) from eastern Canada. Bulletin of the Biological Society of Washington 7: 108 - 113." type="journal article" year="1987">Pocklington and Fournier (1987)</bibRefCitation>
from off Nova Scotia. In contrast,
<taxonomicName id="4C0E4D7CFFEE9746FB69FBD3547DFB90" authority="Blake, 2023" authorityName="Blake" authorityYear="2023" box="[1263,1383,1080,1104]" class="Polychaeta" family="Scalibregmatidae" genus="Axiokebuita" kingdom="Animalia" pageId="8" pageNumber="279" phylum="Annelida" rank="species" species="australis" status="sp. nov.">
<emphasis id="B97AEAEDFFEE9746FB69FBD3547DFB90" box="[1263,1383,1080,1104]" italics="true" pageId="8" pageNumber="279">A. australis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2495796FFEE9746FAEDFBD354A1FB90" box="[1387,1467,1081,1104]" pageId="8" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
has a sausage or grub-shaped body, 6–8 small lobes around the anus, and two annulated rows of pads on setigers 1–4 but only a single row along the rest of the body.
</paragraph>
</subSubSection>
<subSubSection id="C3146574FFEE9746FCC1FB545559FAD0" pageId="8" pageNumber="279" type="etymology">
<paragraph id="8BB136FFFFEE9746FCC1FB545559FAD0" blockId="8.[839,1466,1214,1296]" pageId="8" pageNumber="279">
<emphasis id="B97AEAEDFFEE9746FCC1FB5452DCFB16" bold="true" box="[839,966,1214,1238]" pageId="8" pageNumber="279">Etymology</emphasis>
. The epithet is from the Latin
<taxonomicName id="4C0E4D7CFFEE9746FAB5FB54548EFB16" authorityName="Blake" authorityYear="2023" box="[1331,1428,1214,1238]" class="Polychaeta" family="Scalibregmatidae" genus="Axiokebuita" kingdom="Animalia" pageId="8" pageNumber="279" phylum="Annelida" rank="species" species="australis">
<emphasis id="B97AEAEDFFEE9746FAB5FB54548EFB16" box="[1331,1428,1214,1238]" italics="true" pageId="8" pageNumber="279">australis</emphasis>
</taxonomicName>
, in reference to the Australian and Southern Hemisphere location of this species.
</paragraph>
</subSubSection>
<subSubSection id="C3146574FFEE9746FCC1FACF5552FAB8" pageId="8" pageNumber="279" type="biology_ecology">
<paragraph id="8BB136FFFFEE9746FCC1FACF5552FAB8" blockId="8.[839,1467,1317,1401]" pageId="8" pageNumber="279">
<emphasis id="B97AEAEDFFEE9746FCC1FACF52B8FAFD" bold="true" box="[839,930,1317,1341]" pageId="8" pageNumber="279">Biology</emphasis>
.
<typeStatus id="54B5885DFFEE9746FC33FACC5504FAFE" box="[949,1054,1318,1342]" pageId="8" pageNumber="279" type="holotype">Holotype</typeStatus>
(AM W.52681) and
<typeStatus id="54B5885DFFEE9746FA9EFACC5463FAFE" box="[1304,1401,1318,1342]" pageId="8" pageNumber="279" type="paratype">paratype</typeStatus>
(AM W.53989) with numerous oocytes in coelom measuring 80–100 µm in diameter.
</paragraph>
</subSubSection>
<subSubSection id="C3146574FFEE9746FCC1FA675456FA03" pageId="8" pageNumber="279" type="distribution">
<paragraph id="8BB136FFFFEE9746FCC1FA675456FA03" blockId="8.[839,1466,1421,1475]" pageId="8" pageNumber="279">
<emphasis id="B97AEAEDFFEE9746FCC1FA6752C8FA65" bold="true" box="[839,978,1421,1445]" pageId="8" pageNumber="279">Distribution</emphasis>
. Continental slope and abyssal plain off
<collectingRegion id="49CAF81DFFEE9746FA1DFA6452B6FA03" country="Australia" name="Western Australia" pageId="8" pageNumber="279">SE Australia</collectingRegion>
,
<collectingRegion id="49CAF81DFFEE9746FC30FA415506FA03" box="[950,1052,1451,1475]" country="Australia" name="Tasmania" pageId="8" pageNumber="279">Tasmania</collectingRegion>
and
<collectingRegion id="49CAF81DFFEE9746FBC9FA4155B2FA03" box="[1103,1192,1451,1475]" country="Australia" name="Victoria" pageId="8" pageNumber="279">Victoria</collectingRegion>
,
<quantity id="4CF69B1AFFEE9746FB34FA415453FA03" box="[1202,1353,1451,1475]" metricMagnitude="3" metricUnit="m" metricValue="3.4285" metricValueMax="4.064" metricValueMin="2.793" pageId="8" pageNumber="279" unit="m" value="3428.5" valueMax="4064.0" valueMin="2793.0">2793–4064 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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