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<document id="E8DF1B1B48E9A0DFF2D95919E3D70659" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFE8E518509884DB601AFD35" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) matangensis Cameron 1943" docType="treatment" docVersion="12" lastPageNumber="159" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="158" updateTime="1699471407760" updateUser="ExternalLinkService">
<mods:mods id="EB380A431253DB3EA2DFBDC0432AED00" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="ACC75CFFABDA7CA7B1622B0CDC9EF945">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
</mods:titleInfo>
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<mods:role id="81CE4B70C4A65BB352017995F00CCFB8">
<mods:roleTerm id="6D9A378C1C8AEAF1A8EAB9A8297E8B95">Author</mods:roleTerm>
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<mods:namePart id="97EA73F8BCE6CF769C901591DD9FD586">Assing, Volker</mods:namePart>
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<mods:title id="9212F742AE0ADEEEFF4DD74D5BC6266C">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="DD51AC90547DFB808A2DE6C251FA2F24">
<mods:date id="42CB5EB8384F278D87CD0EFF13BEE168">2017</mods:date>
<mods:detail id="A79F441D922B30837DD4323CEE99FB15" type="pubDate">
<mods:number id="E0936497A9B66269C3616032A741300C">2017-06-30</mods:number>
</mods:detail>
<mods:detail id="00AFB0F11D71CB785C275AC2B5964013" type="volume">
<mods:number id="645AD1F9D438FA7CC60D831807655494">67</mods:number>
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<mods:number id="E39FA8350F793D0952731631C9C705E5">1</mods:number>
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<mods:classification id="794910C123D1062BE730ECB0218FEFE1">journal article</mods:classification>
<mods:identifier id="9D4B088688DC9A0810D42EFE35779889" type="CLB-Dataset">2472</mods:identifier>
<mods:identifier id="E917EB51C271D2357C2DF627F96A4367" type="DOI">10.21248/contrib.entomol.67.1.117-192</mods:identifier>
<mods:identifier id="6533D6A316160662DCC4C819EC8F6205" type="GBIF-Dataset">39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43</mods:identifier>
<mods:identifier id="0985381A11000ECA1765C1095D3D1C61" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="1ED6DB6CBDB722C37C0B6A00EE825F9D" type="Zenodo-Dep">5742363</mods:identifier>
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</mods:mods>
<treatment id="03A787BAFFE8E518509884DB601AFD35" ID-DOI="http://doi.org/10.5281/zenodo.5888489" ID-GBIF-Taxon="192498196" ID-Zenodo-Dep="5888489" LSID="urn:lsid:plazi:treatment:03A787BAFFE8E518509884DB601AFD35" httpUri="http://treatment.plazi.org/id/03A787BAFFE8E518509884DB601AFD35" lastPageId="42" lastPageNumber="159" pageId="41" pageNumber="158">
<subSubSection id="C3146527FFE8E51B509884DB6178FC5A" box="[189,627,979,1006]" pageId="41" pageNumber="158" type="nomenclature">
<paragraph id="8BB136ACFFE8E51B509884DB6178FC5A" blockId="41.[189,627,979,1039]" box="[189,627,979,1006]" pageId="41" pageNumber="158">
<heading id="D0F981C0FFE8E51B509884DB6178FC5A" box="[189,627,979,1006]" centered="true" fontSize="12" level="2" pageId="41" pageNumber="158" reason="2">
<taxonomicName id="4C0E4D2FFFE8E51B509884DB6178FC5A" ID-CoL="5DDDR" authority="CAMERON, 1943" authorityName="Cameron" authorityYear="1943" box="[189,627,979,1006]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="158" phylum="Arthropoda" rank="species" species="matangensis" subGenus="Zyras">
<emphasis id="B97AEABEFFE8E51B509884DB63F8FC5A" box="[189,243,979,1006]" italics="true" pageId="41" pageNumber="158">Zyras</emphasis>
(
<emphasis id="B97AEABEFFE8E51B512484DB623CFC5A" box="[257,311,979,1006]" italics="true" pageId="41" pageNumber="158">Zyras</emphasis>
)
<emphasis id="B97AEABEFFE8E51B516084DB62C2FC5A" box="[325,457,979,1006]" italics="true" pageId="41" pageNumber="158">matangensis</emphasis>
<bibRefCitation id="EF9F4B5DFFE8E51B51EA84DB6178FC5A" author="CAMERON, M." box="[463,627,979,1006]" pageId="41" pageNumber="158" pagination="139 - 143" refId="ref57563" refString="CAMERON, M. 1943: New species of Staphylinidae (Col.) from Borneo. - Entomologist's Monthly Magazine 79: 39 - 42, 139 - 143, 183." type="journal article" year="1943">CAMERON, 1943</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFE8E51B509884F362A4FBBB" box="[189,431,1019,1039]" pageId="41" pageNumber="158" type="description">
<paragraph id="8BB136ACFFE8E51B509884F362A4FBBB" blockId="41.[189,627,979,1039]" box="[189,431,1019,1039]" pageId="41" pageNumber="158">
(
<figureCitation id="13352A29FFE8E51B50E484F363F1FBBB" box="[193,250,1019,1039]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="41" pageNumber="158">Figs 35</figureCitation>
,
<figureCitation id="13352A29FFE8E51B512484F3621DFBBB" box="[257,278,1019,1039]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="41" pageNumber="158">73</figureCitation>
,
<figureCitation id="13352A29FFE8E51B513884F36230FBBB" box="[285,315,1019,1039]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="41" pageNumber="158">108</figureCitation>
,
<figureCitation id="13352A29FFE8E51B516784F362A0FBBB" box="[322,427,1019,1039]" captionStart="Figs 227–244" captionStartId="61.[151,191,1983,2005]" captionTargetBox="[163,1397,240,1949]" captionTargetId="figure-119@61.[151,1398,231,1970]" captionText="Figs 227–244: Zyras parahirtus (227–228, 231–232), Z. matangensis (229–230, 233), Z. gratellus (Sulawesi) (234–238), and Z. densissimus (239–244): median lobe of aedeagus in lateral and in ventral view (227–230, 234–235, 239–241); paramere (231, 236, 242); male tergite VIII (232, 237, 243); male sternite VIII (233, 238, 244). Scale bars: 227–233, 237–238, 243–244: 0.5 mm; 234–236, 239–242: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888378" httpUri="https://zenodo.org/record/5888378/files/figure.png" pageId="41" pageNumber="158">229–230, 233</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFE8E51B50B283216143FB75" pageId="41" pageNumber="158" type="materials_examined">
<paragraph id="8BB136ACFFE8E51B50B283216143FB75" blockId="41.[151,757,1065,1217]" pageId="41" pageNumber="158">
<emphasis id="B97AEABEFFE8E51B50B2832162BBFBF5" bold="true" box="[151,432,1065,1089]" pageId="41" pageNumber="158">Type material examined</emphasis>
:
<materialsCitation id="3B663CF1FFE8E51B51E183216143FB75" ID-GBIF-Occurrence="3450162383" collectingDate="2014-01-19" collectionCode="BMNH" collectorName="G. E. Bryant. &amp; V. Assing" country="Malaysia" elevation="610" location="Mt. Matang" pageId="41" pageNumber="158" specimenCount="1" typeStatus="holotype">
<typeStatus id="54B5880EFFE8E51B51E18321612CFBF5" box="[452,551,1065,1089]" pageId="41" pageNumber="158" type="holotype">Holotype</typeStatus>
♂: “
<location id="8ED16077FFE8E51B524C832161FBFBF6" LSID="urn:lsid:plazi:treatment:03A787BAFFE8E518509884DB601AFD35:8ED16077FFE8E51B524C832161FBFBF6" box="[617,752,1065,1090]" country="Malaysia" name="Mt. Matang" pageId="41" pageNumber="158">Mt. Matang</location>
, W.
<collectingCountry id="F319763CFFE8E51B509C8341621EFBD5" box="[185,277,1097,1121]" name="Malaysia" pageId="41" pageNumber="158">Sarawak</collectingCountry>
.
<collectorName id="26FB537AFFE8E51B510D834162A7FBD6" box="[296,428,1097,1122]" pageId="41" pageNumber="158">G.E. Bryant.</collectorName>
<date id="FFB0106CFFE8E51B519E8341610DFBD5" box="[443,518,1097,1121]" pageId="41" pageNumber="158" value="2014-01-19">
<collectingDate id="EFF4E984FFE8E51B519E8341610DFBD5" box="[443,518,1097,1121]" pageId="41" pageNumber="158" value="2014-01-19">19.I.14</collectingDate>
</date>
,
<quantity id="4CF69B49FFE8E51B523F83416165FBD5" box="[538,622,1097,1121]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="41" pageNumber="158" unit="ft" value="2000.0">
<elevation id="0023D19FFFE8E51B523F83416165FBD5" box="[538,622,1097,1121]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="41" pageNumber="158" unit="ft" value="2000.0">2000 ft</elevation>
</quantity>
/ G. 23 /
<taxonomicName id="4C0E4D2FFFE8E51B50B283626270FB35" authority="Cam." authorityName="Cameron" authorityYear="1943" box="[151,379,1129,1153]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="158" phylum="Arthropoda" rank="species" species="matangensis">
<emphasis id="B97AEABEFFE8E51B50B28362623FFB35" box="[151,308,1130,1153]" italics="true" pageId="41" pageNumber="158">Z. matangensis</emphasis>
Cam.
</taxonomicName>
Type / Holotype / M. Cameron. Bequest. B.M. 1955-147 / Holotypus
<taxonomicName id="4C0E4D2FFFE8E51B5215838263F6FB75" authority="Cameron" authorityName="Cameron" authorityYear="1943" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="158" phylum="Arthropoda" rank="species" species="matangensis">
<emphasis id="B97AEABEFFE8E51B5215838261FFFB15" box="[560,756,1162,1185]" italics="true" pageId="41" pageNumber="158">Zyras matangensis</emphasis>
Cameron
</taxonomicName>
, rev.
<collectorName id="26FB537AFFE8E51B511683A2629EFB75" box="[307,405,1193,1217]" pageId="41" pageNumber="158">V. Assing</collectorName>
2017” (
<collectionCode id="ED1FAE69FFE8E51B51C283A16136FB75" box="[487,573,1193,1217]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="41" pageNumber="158">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFE8E51B50B283E16112F9B5" pageId="41" pageNumber="158" type="discussion">
<paragraph id="8BB136ACFFE8E51B50B283E163D1FAF5" blockId="41.[151,756,1257,1345]" pageId="41" pageNumber="158">
<emphasis id="B97AEABEFFE8E51B50B283E1620DFAB5" bold="true" box="[151,262,1257,1281]" pageId="41" pageNumber="158">Comment</emphasis>
: The original description is based on a unique
<typeStatus id="54B5880EFFE8E51B50B2820163FEFA95" box="[151,245,1289,1313]" pageId="41" pageNumber="158" type="holotype">holotype</typeStatus>
from “W.
<collectingCountry id="F319763CFFE8E51B5152820162D8FA95" box="[375,467,1289,1313]" name="Malaysia" pageId="41" pageNumber="158">Sarawak</collectingCountry>
: Mt. Matang” (
<bibRefCitation id="EF9F4B5DFFE8E51B52A3820163C5FAF5" author="CAMERON, M." pageId="41" pageNumber="158" pagination="139 - 143" refId="ref57563" refString="CAMERON, M. 1943: New species of Staphylinidae (Col.) from Borneo. - Entomologist's Monthly Magazine 79: 39 - 42, 139 - 143, 183." type="journal article" year="1943">CAMERON 1943</bibRefCitation>
).
</paragraph>
<paragraph id="8BB136ACFFE8E51B50B282616112F9B5" blockId="41.[151,756,1385,1537]" pageId="41" pageNumber="158">
<emphasis id="B97AEABEFFE8E51B50B2826162E3FA35" bold="true" box="[151,488,1385,1409]" pageId="41" pageNumber="158">Additional material examined</emphasis>
:
<emphasis id="B97AEABEFFE8E51B51DC82616141FA35" bold="true" box="[505,586,1385,1409]" pageId="41" pageNumber="158">Borneo</emphasis>
:
<materialsCitation id="3B663CF1FFE8E51B527F82616159FA75" ID-GBIF-Occurrence="3450162456" collectingDate="1997-03" collectingMethod="flight interception trap" collectionCode="BMNH" collectorName="S Lokan &amp; Chung" country="Malaysia" location="Sandakan" pageId="41" pageNumber="158" specimenCount="1" stateProvince="Sabah">
1 ♂,
<collectingCountry id="F319763CFFE8E51B52B5826161FBFA35" box="[656,752,1385,1409]" name="Malaysia" pageId="41" pageNumber="158">Malaysia</collectingCountry>
,
<collectingRegion id="49CAF84EFFE8E51B50B2828163D2FA15" box="[151,217,1417,1441]" country="Malaysia" name="Sabah" pageId="41" pageNumber="158">Sabah</collectingRegion>
,
<location id="8ED16077FFE8E51B50C182816245FA15" LSID="urn:lsid:plazi:treatment:03A787BAFFE8E518509884DB601AFD35:8ED16077FFE8E51B50C182816245FA15" box="[228,334,1417,1441]" country="Malaysia" name="Sandakan" pageId="41" pageNumber="158" stateProvince="Sabah">Sandakan</location>
,
<collectorName id="26FB537AFFE8E51B517C828162B8FA15" box="[345,435,1417,1441]" pageId="41" pageNumber="158">S Lokan</collectorName>
, logged forest,
<collectingMethod id="524F4EBBFFE8E51B5271828163F2FA75" pageId="41" pageNumber="158">flight interception trap</collectingMethod>
,
<date id="FFB0106CFFE8E51B512382A16257FA75" box="[262,348,1449,1473]" pageId="41" pageNumber="158" value="1997-03">
<collectingDate id="EFF4E984FFE8E51B512382A16257FA75" box="[262,348,1449,1473]" pageId="41" pageNumber="158" value="1997-03">III.1997</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFE8E51B51BD82A162EAFA75" box="[408,481,1449,1473]" pageId="41" pageNumber="158">Chung</collectorName>
(
<collectionCode id="ED1FAE69FFE8E51B51D482A1614DFA75" box="[497,582,1449,1473]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="41" pageNumber="158">BMNH</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B663CF1FFE8E51B527E82A16112F9B5" ID-GBIF-Occurrence="3450162367" collectingDate="1996-10" collectingMethod="flight interception trap" collectionCode="BMNH" collectorName="Chung" country="Malaysia" location="Sepilok" municipality="Sandakan" pageId="41" pageNumber="158" specimenCount="1" stateProvince="Sabah">
1 ♀,
<collectingCountry id="F319763CFFE8E51B52B582A161FBFA75" box="[656,752,1449,1473]" name="Malaysia" pageId="41" pageNumber="158">Malaysia</collectingCountry>
,
<collectingRegion id="49CAF84EFFE8E51B50B282C163D2FA55" box="[151,217,1481,1505]" country="Malaysia" name="Sabah" pageId="41" pageNumber="158">Sabah</collectingRegion>
,
<collectingMunicipality id="6BD5ACD6FFE8E51B50C682C16246FA55" box="[227,333,1481,1505]" pageId="41" pageNumber="158">Sandakan</collectingMunicipality>
,
<location id="8ED16077FFE8E51B517282C162A3FA55" LSID="urn:lsid:plazi:treatment:03A787BAFFE8E518509884DB601AFD35:8ED16077FFE8E51B517282C162A3FA55" box="[343,424,1481,1505]" country="Malaysia" municipality="Sandakan" name="Sepilok" pageId="41" pageNumber="158" stateProvince="Sabah">Sepilok</location>
, primary forest,
<collectingMethod id="524F4EBBFFE8E51B527082C163FDF9B5" pageId="41" pageNumber="158">flight interception trap</collectingMethod>
,
<date id="FFB0106CFFE8E51B50DA82E16247F9B5" box="[255,332,1513,1537]" pageId="41" pageNumber="158" value="1996-10">
<collectingDate id="EFF4E984FFE8E51B50DA82E16247F9B5" box="[255,332,1513,1537]" pageId="41" pageNumber="158" value="1996-10">X.1996</collectingDate>
</date>
, leg.
<collectorName id="26FB537AFFE8E51B51A482E162C1F9B5" box="[385,458,1513,1537]" pageId="41" pageNumber="158">Chung</collectorName>
(cAss).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFE8E51850B281216131FE55" lastPageId="42" lastPageNumber="159" pageId="41" pageNumber="158" type="description">
<paragraph id="8BB136ACFFE8E51B50B281216188F895" blockId="41.[151,756,1577,2081]" pageId="41" pageNumber="158">
<emphasis id="B97AEABEFFE8E51B50B281216226F9F5" bold="true" box="[151,301,1577,1601]" pageId="41" pageNumber="158">Redescription</emphasis>
: Body length
<quantity id="4CF69B49FFE8E51B51EF81216143F9F5" box="[458,584,1577,1601]" metricMagnitude="-3" metricUnit="m" metricValue="6.35" metricValueMax="7.5" metricValueMin="5.2" pageId="41" pageNumber="158" unit="mm" value="6.35" valueMax="7.5" valueMin="5.2">5.2–7.5 mm</quantity>
; length of forebody
<quantity id="4CF69B49FFE8E51B50F681416259F9D5" box="[211,338,1609,1633]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.2" metricValueMin="2.6" pageId="41" pageNumber="158" unit="mm" value="2.9" valueMax="3.2" valueMin="2.6">2.6–3.2 mm</quantity>
. Coloration (
<figureCitation id="13352A29FFE8E51B51F981416122F9D5" box="[476,553,1609,1633]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="41" pageNumber="158">Figs 35</figureCitation>
,
<figureCitation id="13352A29FFE8E51B521181416144F9D5" box="[564,591,1609,1633]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="41" pageNumber="158">73</figureCitation>
,
<figureCitation id="13352A29FFE8E51B527E8141618FF9D5" box="[603,644,1609,1633]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="41" pageNumber="158">108</figureCitation>
): forebody reddish to castaneous-brown, elytra of uniform coloration or with the postero-lateral portions slightly paler; legs dark-yellowish; antennae reddish-brown to brown with the basal 2–3 antennomeres reddish and the apex of antennomere XI sometimes slightly paler brown; maxillary palpi reddish with the terminal palpomere yellowish.
</paragraph>
<paragraph id="8BB136ACFFE8E51B50B2802167BFFE95" blockId="41.[151,756,1577,2081]" lastBlockId="41.[794,1399,233,1505]" pageId="41" pageNumber="158">
Head (
<figureCitation id="13352A29FFE8E51B50C780216226F8F5" box="[226,301,1833,1857]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="41" pageNumber="158">Fig. 73</figureCitation>
) moderately transverse, broadly impunctate along middle; lateral dorsal portions with sparse to moderately dense coarse punctation. Eyes moderately large, as long as, or slightly longer than postocular region in dorsal view. Antenna (
<figureCitation id="13352A29FFE8E51B51E980A1611DF875" box="[460,534,1961,1985]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="41" pageNumber="158">Fig. 35</figureCitation>
) 1.9–2.0 mm long; antennomeres IV approximately as long as broad or weakly transverse, V–X of gradually increasing width and increasingly transverse, X approximately twice as broad as long, and XI short and of conical shape, slightly shorter than the combined length of IX and X.
</paragraph>
<paragraph id="8BB136ACFFE8E51B533F86226790FDF5" blockId="41.[794,1399,233,1505]" pageId="41" pageNumber="158">
Pronotum (
<figureCitation id="13352A29FFE8E51B53BE862160EDFEF5" box="[923,998,297,321]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="41" pageNumber="158">Fig. 73</figureCitation>
) weakly transverse, 1.03–1.10 times as broad as long and 1.20–1.25 times as broad as head, broadest in anterior half, distinctly tapering posteriad; lateral margins straight or weakly sinuate in posterior two-thirds (dorsal view); punctation moderately coarse, moderately dense to dense, and slightly irregularly distributed, laterally with impunctate patches; midline broadly impunctate; lateral and anterior margins with numerous long and erect dark setae.
</paragraph>
<paragraph id="8BB136ACFFE8E51B533F85416787FCB5" blockId="41.[794,1399,233,1505]" pageId="41" pageNumber="158">
Elytra (
<figureCitation id="13352A29FFE8E51B534D854160B8FDD5" box="[872,947,585,609]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="41" pageNumber="158">Fig. 73</figureCitation>
) approximately 0.75–0.80 times as long as pronotum; punctation fine, moderately sparse to dense; pubescence long, suberect, and pale; lateral margins with numerous stouter, dark and erect setae. Hind wings fully developed. Metatarsomere I slightly to distinctly shorter than the combined length of II–IV.
</paragraph>
<paragraph id="8BB136ACFFE8E51B533F840160C2FAD5" blockId="41.[794,1399,233,1505]" pageId="41" pageNumber="158">
Abdomen (
<figureCitation id="13352A29FFE8E51B53BC840160F8FC95" box="[921,1011,777,801]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="41" pageNumber="158">Fig. 108</figureCitation>
) broad, nearly as broad as elytra, with moderately deep anterior impressions on tergites III–V, these impressions with or without sparse nonsetiferous punctures; tergites III–V with some lateral setiferous punctures on either side, with few scattered non-setiferous punctures on disc, and with 6–8 setiferous punctures at posterior margin, disc without micropunctation; tergite VI with a transverse band of rather coarse and dense non-setiferous punctures anteriorly, with scattered non-setiferous punctures on disc, and with some setiferous punctures laterally and at posterior margin; tergite VII with a transverse band of rather coarse and dense non-setiferous punctures situated in an anterior transverse impression, with scattered non-setiferous punctures on disc, and with two transverse series of setiferous punctures posteriorly, posterior margin with palisade fringe; tergite VIII with long dark setae in posterior third, posterior margin broadly and weakly concave.
</paragraph>
<paragraph id="8BB136ACFFE8E51B533F8262664FFA55" blockId="41.[794,1399,233,1505]" pageId="41" pageNumber="158">
♂: posterior margin of sternite VIII (
<figureCitation id="13352A29FFE8E51B54B3826167E6FA35" box="[1174,1261,1385,1409]" captionStart="Figs 227–244" captionStartId="61.[151,191,1983,2005]" captionTargetBox="[163,1397,240,1949]" captionTargetId="figure-119@61.[151,1398,231,1970]" captionText="Figs 227–244: Zyras parahirtus (227–228, 231–232), Z. matangensis (229–230, 233), Z. gratellus (Sulawesi) (234–238), and Z. densissimus (239–244): median lobe of aedeagus in lateral and in ventral view (227–230, 234–235, 239–241); paramere (231, 236, 242); male tergite VIII (232, 237, 243); male sternite VIII (233, 238, 244). Scale bars: 227–233, 237–238, 243–244: 0.5 mm; 234–236, 239–242: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888378" httpUri="https://zenodo.org/record/5888378/files/figure.png" pageId="41" pageNumber="158">Fig. 233</figureCitation>
) with convex posterior margin; median lobe of aedeagus
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long and shaped as in
<figureCitation id="13352A29FFE8E51B540982A167B3FA75" box="[1068,1208,1449,1473]" captionStart="Figs 227–244" captionStartId="61.[151,191,1983,2005]" captionTargetBox="[163,1397,240,1949]" captionTargetId="figure-119@61.[151,1398,231,1970]" captionText="Figs 227–244: Zyras parahirtus (227–228, 231–232), Z. matangensis (229–230, 233), Z. gratellus (Sulawesi) (234–238), and Z. densissimus (239–244): median lobe of aedeagus in lateral and in ventral view (227–230, 234–235, 239–241); paramere (231, 236, 242); male tergite VIII (232, 237, 243); male sternite VIII (233, 238, 244). Scale bars: 227–233, 237–238, 243–244: 0.5 mm; 234–236, 239–242: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888378" httpUri="https://zenodo.org/record/5888378/files/figure.png" pageId="41" pageNumber="158">Figs 229–230</figureCitation>
; paramere
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long and with moderately short apical lobe.
</paragraph>
<paragraph id="8BB136ACFFE8E51B533F8101663EF8D5" blockId="41.[794,1399,1545,1889]" pageId="41" pageNumber="158">
<emphasis id="B97AEABEFFE8E51B533F81016719F995" bold="true" box="[794,1042,1545,1569]" pageId="41" pageNumber="158">Intraspecific variation</emphasis>
: The aedeagus of the non-type male is distinguished from that of the
<typeStatus id="54B5880EFFE8E51B548D8121660DF9F5" box="[1192,1286,1577,1601]" pageId="41" pageNumber="158" type="holotype">holotype</typeStatus>
by slightly larger size of the median lobe, a crista apicalis of slightly different shape, and somewhat larger parameres. In the non-type female, the non-setiferous punctation of tergite VI and VII is much finer and less extensive than in the
<specimenCount id="9D08FD25FFE8E51B533F81C26083F955" box="[794,904,1737,1761]" pageId="41" pageNumber="158" type="male">two males</specimenCount>
examined. Since no additional evidence was found suggesting that the
<specimenCount id="9D08FD25FFE8E51B547581E1660EF8B5" box="[1104,1285,1769,1793]" pageId="41" pageNumber="158" type="generic">three specimens</specimenCount>
represent different species, and in view of the often pronounced variability of
<taxonomicName id="4C0E4D2FFFE8E51B539480226739F8F5" authority="sensu" authorityName="SenSU" box="[945,1074,1834,1857]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="186" phylum="Arthropoda" rank="genus">
<emphasis id="B97AEABEFFE8E51B5394802260E0F8F5" box="[945,1003,1834,1857]" italics="true" pageId="41" pageNumber="158">Zyras</emphasis>
sensu
</taxonomicName>
strictu species, the observed differences are attributed to intraspecific variation.
</paragraph>
<paragraph id="8BB136ACFFE8E518533F80816131FE55" blockId="41.[794,1399,1929,2081]" lastBlockId="42.[189,794,233,481]" lastPageId="42" lastPageNumber="159" pageId="41" pageNumber="158">
<emphasis id="B97AEABEFFE8E51B533F808160E5F815" bold="true" box="[794,1006,1929,1953]" pageId="41" pageNumber="158">Comparative notes</emphasis>
: In general appearance (coloration, size, habitus),
<taxonomicName id="4C0E4D2FFFE8E51B539780A26744F875" authorityName="Cameron" authorityYear="1943" box="[946,1103,1962,1985]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="158" phylum="Arthropoda" rank="species" species="matangensis">
<emphasis id="B97AEABEFFE8E51B539780A26744F875" box="[946,1103,1962,1985]" italics="true" pageId="41" pageNumber="158">Z. matangensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C0E4D2FFFE8E51B54FE80A2663EF875" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[1243,1333,1961,1985]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="158" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B97AEABEFFE8E51B54FE80A2663EF875" box="[1243,1333,1961,1985]" italics="true" pageId="41" pageNumber="158">Z. hirtus</emphasis>
</taxonomicName>
, from which it is reliably distinguished only by the completely different shape of the aedeagus. In habitus, coloration, and other external characters, the species is also highly similar to
<taxonomicName id="4C0E4D2FFFEBE51850FD87E2611AFEB5" authority="CAMERON, 1939" authorityName="CAMERON" authorityYear="1939" box="[216,529,233,257]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="159" phylum="Arthropoda" rank="species" species="flavorufus">
<emphasis id="B97AEABEFFEBE51850FD87E26254FEB5" box="[216,351,233,257]" italics="true" pageId="42" pageNumber="159">Z. flavorufus</emphasis>
CAMERON, 1939
</taxonomicName>
from
<collectingRegion id="49CAF84EFFEBE518527787E2618BFEB5" box="[594,640,234,257]" country="Indonesia" name="Jawa Timur" pageId="42" pageNumber="159">Java</collectingRegion>
, from which it differs by darker coloration, a less transverse pronotum with less distinctly sinuate lateral margins, shorter antennae with more transverse antennomeres IV-X, shorter elytra, the punctation pattern of the abdomen (particularly much coarser and denser non-setiferous punctation on tergites VI and VII, and by the differently shaped and smaller median lobe of the adeagus.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFEBE51850988501601AFD35" pageId="42" pageNumber="159" type="distribution">
<paragraph id="8BB136ACFFEBE51850988501601AFD35" blockId="42.[189,794,521,641]" pageId="42" pageNumber="159">
<emphasis id="B97AEABEFFEBE51850988501612FFD95" bold="true" box="[189,548,521,545]" pageId="42" pageNumber="159">Distribution and natural history</emphasis>
: At present,
<taxonomicName id="4C0E4D2FFFEBE5185297850263F2FDF5" authorityName="Cameron" authorityYear="1943" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="159" phylum="Arthropoda" rank="species" species="matangensis">
<emphasis id="B97AEABEFFEBE5185297850263F2FDF5" italics="true" pageId="42" pageNumber="159">Z. matangensis</emphasis>
</taxonomicName>
is known from three localities in
<collectingCountry id="F319763CFFEBE51852A8852161EEFDF5" box="[653,741,553,577]" name="Malaysia" pageId="42" pageNumber="159">Sarawak</collectingCountry>
and
<collectingRegion id="49CAF84EFFEBE5185098854163F5FDD5" box="[189,254,585,609]" country="Malaysia" name="Sabah" pageId="42" pageNumber="159">Sabah</collectingRegion>
, Borneo. The additional material was collected with flight interception traps in a primary and a logged forest.
</paragraph>
</subSubSection>
</treatment>
</document>
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