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<document id="95B49B9347CC6E16F94FA6847D80C747" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFC8E5385098819B615DFB15" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) exasperatus SCHUBERT 1908" docType="treatment" docVersion="11" lastPageNumber="127" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="126" updateTime="1699471407760" updateUser="ExternalLinkService">
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<mods:title id="4F3D2B33FDF40034C35ED87C68ED453A">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:namePart id="9D99CA5B5D415EB167233BC408837A42">Assing, Volker</mods:namePart>
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<mods:title id="E88B9820A5AF0AAAA3E6620B03C65FF5">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="A2BBEAF8CEF551490D7BD5C610FBF8D3">
<mods:date id="692E08C9C5BF0F6AB0183711BE036AC8">2017</mods:date>
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<mods:number id="144F060989D010F7D519814B41B18735">2017-06-30</mods:number>
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<mods:number id="6EA3913A71669D47B4AEEDEF692D775E">67</mods:number>
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<mods:identifier id="4A4865BD11C85BF373248796E8AAB1F9" type="CLB-Dataset">2472</mods:identifier>
<mods:identifier id="AE5C29D7B171B507F21C8AD2185D3130" type="DOI">10.21248/contrib.entomol.67.1.117-192</mods:identifier>
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<treatment id="03A787BAFFC8E5385098819B615DFB15" ID-DOI="http://doi.org/10.5281/zenodo.5888455" ID-GBIF-Taxon="192498156" ID-Zenodo-Dep="5888455" LSID="urn:lsid:plazi:treatment:03A787BAFFC8E5385098819B615DFB15" httpUri="http://treatment.plazi.org/id/03A787BAFFC8E5385098819B615DFB15" lastPageId="10" lastPageNumber="127" pageId="9" pageNumber="126">
<subSubSection id="C3146527FFC8E53B5098819B6167F91A" box="[189,620,1683,1710]" pageId="9" pageNumber="126" type="nomenclature">
<paragraph id="8BB136ACFFC8E53B5098819B6167F91A" blockId="9.[189,620,1683,1743]" box="[189,620,1683,1710]" pageId="9" pageNumber="126">
<heading id="D0F981C0FFC8E53B5098819B6167F91A" box="[189,620,1683,1710]" centered="true" fontSize="12" level="2" pageId="9" pageNumber="126" reason="2">
<taxonomicName id="4C0E4D2FFFC8E53B5098819B6167F91A" ID-CoL="5DDBZ" authority="SCHUBERT, 1908" authorityName="SCHUBERT" authorityYear="1908" box="[189,620,1683,1710]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="126" phylum="Arthropoda" rank="species" species="exasperatus" subGenus="Zyras">
<emphasis id="B97AEABEFFC8E53B5098819B63F8F91A" box="[189,243,1683,1710]" italics="true" pageId="9" pageNumber="126">Zyras</emphasis>
(
<emphasis id="B97AEABEFFC8E53B5124819B623CF91A" box="[257,311,1683,1710]" italics="true" pageId="9" pageNumber="126">Zyras</emphasis>
)
<emphasis id="B97AEABEFFC8E53B5160819B62CBF91A" box="[325,448,1683,1710]" italics="true" pageId="9" pageNumber="126">exasperatus</emphasis>
<bibRefCitation id="EF9F4B5DFFC8E53B51E3819B6167F91A" author="SCHUBERT, K." box="[454,620,1683,1710]" pageId="9" pageNumber="126" pagination="609 - 625" refId="ref59207" refString="SCHUBERT, K. 1908: Beitrag zur Staphylinidenfauna Ostindiens (West-Himalaya). (Col.). - Deutsche Entomologische Zeitschrift 1908: 609 - 625." type="journal article" year="1908">SCHUBERT, 1908</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFC8E53B509881B3629DF97B" box="[189,406,1723,1743]" pageId="9" pageNumber="126" type="description">
<paragraph id="8BB136ACFFC8E53B509881B3629DF97B" blockId="9.[189,620,1683,1743]" box="[189,406,1723,1743]" pageId="9" pageNumber="126">
(
<figureCitation id="13352A29FFC8E53B50E481B363FBF97B" box="[193,240,1723,1743]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="9" pageNumber="126">Figs 2</figureCitation>
,
<figureCitation id="13352A29FFC8E53B50DD81B36206F97B" box="[248,269,1723,1743]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="9" pageNumber="126">43</figureCitation>
,
<figureCitation id="13352A29FFC8E53B513181B36252F97B" box="[276,345,1723,1743]" captionStart="Figs 117–133" captionStartId="55.[151,191,2014,2036]" captionTargetBox="[159,1393,236,1954]" captionTargetId="figure-108@55.[151,1398,233,1972]" captionText="Figs 117–133: Zyras kraatzi (117–120), Z. exasperatus (121–124), Z. pindarae (125–131), and Z. perforatus (132–133): median lobe of aedeagus in lateral and in ventral view (117–118, 121–122, 125–128, 132–133); male tergite VIII (119, 123, 130); male sternite VIII (120, 124, 131); paramere (129). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888364" httpUri="https://zenodo.org/record/5888364/files/figure.png" pageId="9" pageNumber="126">121–124</figureCitation>
,
<figureCitation id="13352A29FFC8E53B517A81B36284F97B" box="[351,399,1723,1743]" captionStart="Map 1" captionStartId="64.[189,234,822,843]" captionTargetBox="[189,1436,232,813]" captionTargetId="figure-76@64.[189,1436,232,813]" captionTargetPageId="64" captionText="Map 1: Revised distributions of Zyras kraatzi (black and white circles; white circle: type locality), Z. exasperatus (white circle), Z. iniquus (triangles), Z. pallipes (diamonds), and Z. championi (white stars) in the Himalaya." figureDoi="http://doi.org/10.5281/zenodo.5742393" httpUri="https://zenodo.org/record/5742393/files/figure.png" pageId="9" pageNumber="126">Map 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFC8E53B50B281E4612BF8B5" box="[151,544,1771,1793]" pageId="9" pageNumber="126" type="reference_group">
<paragraph id="8BB136ACFFC8E53B50B281E4612BF8B5" blockId="9.[151,544,1771,1793]" box="[151,544,1771,1793]" pageId="9" pageNumber="126">
<treatmentCitationGroup id="AB1E1182FFC8E53B50B281E4612BF8B5" box="[151,544,1771,1793]" pageId="9" pageNumber="126">
<treatmentCitation id="0AAF10BDFFC8E53B50B281E46105F8B5" author="SCHUBERT, K." box="[151,526,1771,1793]" page="610" pageId="9" pageNumber="126" year="1908">
<taxonomicName id="4C0E4D2FFFC8E53B50B281E46105F8B5" ID-CoL="5DDBZ" authority="SCHUBERT, 1908: 610" authorityName="SCHUBERT" authorityPageNumber="610" authorityYear="1908" box="[151,526,1771,1793]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="126" phylum="Arthropoda" rank="species" species="exasperatus">
<emphasis id="B97AEABEFFC8E53B50B281E46234F8B5" box="[151,319,1772,1793]" italics="true" pageId="9" pageNumber="126">Zyras exasperatus</emphasis>
<bibRefCitation id="EF9F4B5DFFC8E53B516181E36105F8B5" author="SCHUBERT, K." box="[324,526,1771,1793]" pageId="9" pageNumber="126" pagination="609 - 625" refId="ref59207" refString="SCHUBERT, K. 1908: Beitrag zur Staphylinidenfauna Ostindiens (West-Himalaya). (Col.). - Deutsche Entomologische Zeitschrift 1908: 609 - 625." type="journal article" year="1908">
SCHUBERT, 1908:
<specimenCount id="9D08FD25FFC8E53B51CE81E36105F8B5" box="[491,526,1771,1793]" pageId="9" pageNumber="126" type="generic">610</specimenCount>
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
f.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFC8E53B50B2802163E6F7B5" pageId="9" pageNumber="126" type="materials_examined">
<paragraph id="8BB136ACFFC8E53B50B2802163E6F7B5" blockId="9.[151,756,1833,2049]" pageId="9" pageNumber="126">
<emphasis id="B97AEABEFFC8E53B50B2802162A2F8F5" bold="true" box="[151,425,1833,1857]" pageId="9" pageNumber="126">Type material examined</emphasis>
:
<materialsCitation id="3B663CF1FFC8E53B5192802263E6F7B5" ID-GBIF-Occurrence="3450162391" collectionCode="MNB" collectorName="V. Assing" determinerName="V. Assing" location="Kulu" municipality="Kulu" pageId="9" pageNumber="126" specimenCount="1" typeStatus="lectotype">
<typeStatus id="54B5880EFFC8E53B519280226129F8F6" box="[439,546,1834,1858]" pageId="9" pageNumber="126" type="lectotype">Lectotype</typeStatus>
<specimenCount id="9D08FD25FFC8E53B520E8022614BF8F5" box="[555,576,1834,1857]" pageId="9" pageNumber="126" type="generic">♂</specimenCount>
[teneral]: “
<collectingMunicipality id="6BD5ACD6FFC8E53B529B802161F8F8F5" box="[702,755,1833,1857]" pageId="9" pageNumber="126">Kulu</collectingMunicipality>
10 /
<taxonomicName id="4C0E4D2FFFC8E53B50E2804162BEF8D5" box="[199,437,1865,1890]" class="Insecta" family="Staphylinidae" genus="Myrmedonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="126" phylum="Arthropoda" rank="subGenus" subGenus="Zyras">
<emphasis id="B97AEABEFFC8E53B50E28041625AF8D5" box="[199,337,1865,1889]" italics="true" pageId="9" pageNumber="126">Myrmedonia</emphasis>
sg.
<emphasis id="B97AEABEFFC8E53B515F804262BEF8D5" box="[378,437,1866,1889]" italics="true" pageId="9" pageNumber="126">Zyras</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0E4D2FFFC8E53B51E780426134F8D5" authorityName="SCHUBERT" authorityYear="1908" box="[450,575,1866,1889]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="126" phylum="Arthropoda" rank="species" species="exasperatus">
<emphasis id="B97AEABEFFC8E53B51E780426134F8D5" box="[450,575,1866,1889]" italics="true" pageId="9" pageNumber="126">exasperatus</emphasis>
</taxonomicName>
m. type. / Typus /
<taxonomicName id="4C0E4D2FFFC8E53B508880626221F835" authorityName="SCHUBERT" authorityYear="1908" box="[173,298,1898,1921]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="126" phylum="Arthropoda" rank="species" species="exasperatus">
<emphasis id="B97AEABEFFC8E53B508880626221F835" box="[173,298,1898,1921]" italics="true" pageId="9" pageNumber="126">exasperatus</emphasis>
</taxonomicName>
m. / Holotypus [sic]
<taxonomicName id="4C0E4D2FFFC8E53B520A80626248F815" authority="Schubert, 1908" authorityName="SCHUBERT" authorityYear="1908" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="126" phylum="Arthropoda" rank="species" species="exasperatus">
<emphasis id="B97AEABEFFC8E53B520A806261FFF835" box="[559,756,1898,1921]" italics="true" pageId="9" pageNumber="126">Zyras exasperatus</emphasis>
<bibRefCitation id="EF9F4B5DFFC8E53B50B280816248F815" author="SCHUBERT, K." box="[151,323,1929,1953]" pageId="9" pageNumber="126" pagination="609 - 625" refId="ref59207" refString="SCHUBERT, K. 1908: Beitrag zur Staphylinidenfauna Ostindiens (West-Himalaya). (Col.). - Deutsche Entomologische Zeitschrift 1908: 609 - 625." type="journal article" year="1908">Schubert, 1908</bibRefCitation>
</taxonomicName>
, labelled by MNHUB 2016 / Lectotypus ♂
<taxonomicName id="4C0E4D2FFFC8E53B50DA80A2613FF875" authority="Schubert" authorityName="SCHUBERT" authorityYear="1908" box="[255,564,1961,1985]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="126" phylum="Arthropoda" rank="species" species="exasperatus">
<emphasis id="B97AEABEFFC8E53B50DA80A262CFF875" box="[255,452,1962,1985]" italics="true" pageId="9" pageNumber="126">Zyras exasperatus</emphasis>
Schubert
</taxonomicName>
, desig. V. Assing 2016 /
<taxonomicName id="4C0E4D2FFFC8E53B50C680C26106F855" authority="Schubert" authorityName="SCHUBERT" authorityYear="1908" box="[227,525,1993,2017]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="126" phylum="Arthropoda" rank="species" species="exasperatus">
<emphasis id="B97AEABEFFC8E53B50C680C262A9F855" box="[227,418,1994,2017]" italics="true" pageId="9" pageNumber="126">Zyras exasperatus</emphasis>
Schubert
</taxonomicName>
, det.
<determinerName id="7E6DDA68FFC8E53B526D80C261A6F855" box="[584,685,1993,2017]" pageId="9" pageNumber="126">V. Assing</determinerName>
2016” (
<collectionCode id="ED1FAE69FFC8E53B50BA80E163EAF7B5" box="[159,225,2025,2049]" collectionName="MNB" pageId="9" pageNumber="126">MNB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFC8E53B533F87E16637FE35" pageId="9" pageNumber="126" type="discussion">
<paragraph id="8BB136ACFFC8E53B533F87E16637FE35" blockId="9.[794,1399,233,385]" pageId="9" pageNumber="126">
<emphasis id="B97AEABEFFC8E53B533F87E16082FEB5" bold="true" box="[794,905,233,257]" pageId="9" pageNumber="126">Comment</emphasis>
: The original description is based on an unspecified number of
<typeStatus id="54B5880EFFC8E53B540886026782FE96" box="[1069,1161,266,290]" pageId="9" pageNumber="126" type="syntype">syntypes</typeStatus>
from “Kulu, Himalaya, ca.
<quantity id="4CF69B49FFC8E53B5353862160CFFEF5" box="[886,964,297,321]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="126" unit="m" value="3000.0">3000 m</quantity>
” (
<bibRefCitation id="EF9F4B5DFFC8E53B53FB86216786FEF5" author="SCHUBERT, K." box="[990,1165,297,321]" pageId="9" pageNumber="126" pagination="609 - 625" refId="ref59207" refString="SCHUBERT, K. 1908: Beitrag zur Staphylinidenfauna Ostindiens (West-Himalaya). (Col.). - Deutsche Entomologische Zeitschrift 1908: 609 - 625." type="journal article" year="1908">SCHUBERT 1908</bibRefCitation>
). Only
<specimenCount id="9D08FD25FFC8E53B54F886226654FEF6" box="[1245,1375,298,322]" pageId="9" pageNumber="126" type="generic" typeStatus="syntype">one syntype</specimenCount>
, a slightly teneral male, was located in the collections of the MNB. This specimen is designated as the
<typeStatus id="54B5880EFFC8E53B54F186616633FE35" box="[1236,1336,361,385]" pageId="9" pageNumber="126" type="lectotype">lectotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFC8E538533F86A1627BFBB5" lastPageId="10" lastPageNumber="127" pageId="9" pageNumber="126" type="description">
<paragraph id="8BB136ACFFC8E53B533F86A1670AFCB5" blockId="9.[794,1399,425,2081]" pageId="9" pageNumber="126">
<emphasis id="B97AEABEFFC8E53B533F86A160BCFE75" bold="true" box="[794,951,425,449]" pageId="9" pageNumber="126">Redescription</emphasis>
: Body length 7.0 mm; length of forebody
<quantity id="4CF69B49FFC8E53B533F86C16064FE55" box="[794,879,457,481]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="9" pageNumber="126" unit="mm" value="3.3">3.3 mm</quantity>
. Coloration (note that the
<typeStatus id="54B5880EFFC8E53B548486C1660BFE55" box="[1185,1280,457,481]" pageId="9" pageNumber="126" type="holotype">holotype</typeStatus>
is teneral; mature specimens are most likely of significantly darker coloration) (
<figureCitation id="13352A29FFC8E53B5382850160E0FD95" box="[935,1003,521,545]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="9" pageNumber="126">Figs 2</figureCitation>
,
<figureCitation id="13352A29FFC8E53B53D985016711FD95" box="[1020,1050,521,545]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="9" pageNumber="126">43</figureCitation>
): head dark-brown; pronotum and elytra brown, with the postero-lateral portions of the elytra slightly and diffusely darker; abdomen: tergites II–VI brown with paler posterior margins and paratergites, tergites VII–X reddish; legs yellowishbrown; antennae dark-brown with antennomere XI reddish-brown; maxillary palpi brown, with the apical palpomere yellowish.
</paragraph>
<paragraph id="8BB136ACFFC8E53B533F840160EEFB95" blockId="9.[794,1399,425,2081]" pageId="9" pageNumber="126">
Head (
<figureCitation id="13352A29FFC8E53B534C840160B8FC95" box="[873,947,777,801]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="9" pageNumber="126">Fig. 43</figureCitation>
) distinctly transverse, broadly impunctate along middle; punctation in lateral dorsal portions moderately fine and moderately sparse. Eyes slightly longer than postocular region in dorsal view. Antenna (
<figureCitation id="13352A29FFC8E53B530484816054FC15" box="[801,863,905,929]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="9" pageNumber="126">Fig. 2</figureCitation>
)
<quantity id="4CF69B49FFC8E53B5357848160C8FC15" box="[882,963,905,929]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="126" unit="mm" value="2.6">2.6 mm</quantity>
long and slender; antennomeres IV–V weakly oblong, VI–VII approximately as long as broad, VIII–X weakly transverse, X much less than 1.5 times as broad as long, and XI slightly longer than the combined length of IX and X.
</paragraph>
<paragraph id="8BB136ACFFC8E53B533F832267DAFA95" blockId="9.[794,1399,425,2081]" pageId="9" pageNumber="126">
Pronotum (
<figureCitation id="13352A29FFC8E53B53B0832160D4FBF5" box="[917,991,1065,1089]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="9" pageNumber="126">Fig. 43</figureCitation>
) weakly transverse, 1.1 times as broad as long and 1.37 times as broad as head, broadest near anterior angles, distinctly tapering posteriad; lateral margins straight in posterior half (dorsal view); punctation moderately sparse and irregularly distributed, in antero-lateral portions with rather extensive impunctate patch on either side; midline broadly impunctate; lateral margins each with four long brown setae.
</paragraph>
<paragraph id="8BB136ACFFC8E53B533F82216725FA55" blockId="9.[794,1399,425,2081]" pageId="9" pageNumber="126">
Elytra (
<figureCitation id="13352A29FFC8E53B534C822160BFFAF5" box="[873,948,1321,1345]" captionStart="Figs 42–54" captionStartId="46.[189,229,2012,2034]" captionTargetBox="[191,1436,232,1943]" captionTargetId="figure-109@46.[189,1436,232,1971]" captionTargetPageId="46" captionText="Figs 42–54: Forebody of Zyras spp.: kraatzi (42), exasperatus (43), pindarae (44), perforatus (45), championi (46), pallipes (47), condignus, holotype (48), condignus, Nepal (49), nigroaeneus (50), morvani (51), truncatus (52), longilobatus (53), hastatus (54). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="9" pageNumber="126">Fig. 43</figureCitation>
) 0.84 times as long as pronotum; punctation coarse, somewhat asperate or granulose in anterior half, rather dense near anterior margins and scutellum, moderately dense elsewhere. Hind wings probably present. Metatarsomere I approximately as long as the combined length of II–IV.
</paragraph>
<paragraph id="8BB136ACFFC8E538533F82E16189FEF5" blockId="9.[794,1399,425,2081]" lastBlockId="10.[189,794,233,481]" lastPageId="10" lastPageNumber="127" pageId="9" pageNumber="126">
Abdomen slightly narrower than elytra, with rather shallow anterior impressions on tergites III–V; anterior impressions of tergites III–V each with few scattered fine non-setiferous punctures in the middle, laterally impunctate; tergites III–IV with a lateral setiferous puncture on either side and with four setiferous punctures at posterior margins; tergite V with a lateral setiferous puncture on either side and with six setiferous punctures at posterior margin; tergite VI with a narrow transverse row of approximately ten non-setiferous punctures confined to the middle of anterior portion (laterally impunctate), with a lateral setiferous puncture on either side, and with six setiferous punctures at posterior margin; tergite VII with a narrow transverse band of non-setiferous punctures anteriorly, with a median pair of setiferous punctures at posterior fourth, and with a transverse series of six setiferous punctures near posterior margin, posterior margin with palisade fringe; tergite VIII (
<figureCitation id="13352A29FFC8E53B55328F016664F795" box="[1303,1391,2057,2081]" captionStart="Figs 117–133" captionStartId="55.[151,191,2014,2036]" captionTargetBox="[159,1393,236,1954]" captionTargetId="figure-108@55.[151,1398,233,1972]" captionText="Figs 117–133: Zyras kraatzi (117–120), Z. exasperatus (121–124), Z. pindarae (125–131), and Z. perforatus (132–133): median lobe of aedeagus in lateral and in ventral view (117–118, 121–122, 125–128, 132–133); male tergite VIII (119, 123, 130); male sternite VIII (120, 124, 131); paramere (129). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888364" httpUri="https://zenodo.org/record/5888364/files/figure.png" pageId="9" pageNumber="126">Fig. 123</figureCitation>
) with 14 long black setae near posterior margin, posterior margin concavely excised in the middle, on either side of this concavity with a triangular projection.
</paragraph>
<paragraph id="8BB136ACFFCBE53850988642600AFE75" blockId="10.[189,794,233,481]" pageId="10" pageNumber="127">
♂: sternite VIII (
<figureCitation id="13352A29FFCBE5385158864162DFFED5" box="[381,468,329,353]" captionStart="Figs 117–133" captionStartId="55.[151,191,2014,2036]" captionTargetBox="[159,1393,236,1954]" captionTargetId="figure-108@55.[151,1398,233,1972]" captionText="Figs 117–133: Zyras kraatzi (117–120), Z. exasperatus (121–124), Z. pindarae (125–131), and Z. perforatus (132–133): median lobe of aedeagus in lateral and in ventral view (117–118, 121–122, 125–128, 132–133); male tergite VIII (119, 123, 130); male sternite VIII (120, 124, 131); paramere (129). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888364" httpUri="https://zenodo.org/record/5888364/files/figure.png" pageId="10" pageNumber="127">Fig. 124</figureCitation>
) with strongly convex posterior margin; median lobe of aedeagus
<quantity id="4CF69B49FFCBE5385245866161B9FE35" box="[608,690,361,385]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="127" unit="mm" value="0.9">0.9 mm</quantity>
long and shaped as in
<figureCitation id="13352A29FFCBE5385177868162EBFE15" box="[338,480,393,417]" captionStart="Figs 117–133" captionStartId="55.[151,191,2014,2036]" captionTargetBox="[159,1393,236,1954]" captionTargetId="figure-108@55.[151,1398,233,1972]" captionText="Figs 117–133: Zyras kraatzi (117–120), Z. exasperatus (121–124), Z. pindarae (125–131), and Z. perforatus (132–133): median lobe of aedeagus in lateral and in ventral view (117–118, 121–122, 125–128, 132–133); male tergite VIII (119, 123, 130); male sternite VIII (120, 124, 131); paramere (129). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888364" httpUri="https://zenodo.org/record/5888364/files/figure.png" pageId="10" pageNumber="127">Figs 121–122</figureCitation>
; paramere nearly as long as median lobe, with rather short and slender apical lobe.
</paragraph>
<paragraph id="8BB136ACFFCBE538509886C26240FE55" blockId="10.[189,794,233,481]" box="[189,331,457,481]" pageId="10" pageNumber="127">♀: unknown.</paragraph>
<paragraph id="8BB136ACFFCBE53850988501617DFCD5" blockId="10.[189,794,521,1025]" pageId="10" pageNumber="127">
<emphasis id="B97AEABEFFCBE538509885016299FD95" bold="true" box="[189,402,521,545]" pageId="10" pageNumber="127">Comparative notes</emphasis>
: Based on the external and sexual characters,
<taxonomicName id="4C0E4D2FFFCBE538511C852262C4FDF5" authorityName="SCHUBERT" authorityYear="1908" box="[313,463,554,577]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="127" phylum="Arthropoda" rank="species" species="exasperatus">
<emphasis id="B97AEABEFFCBE538511C852262C4FDF5" box="[313,463,554,577]" italics="true" pageId="10" pageNumber="127">Z. exasperatus</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="4C0E4D2FFFCBE538529185226012FDF5" authorityName="SCHUBERT" authorityYear="1908" box="[692,793,553,577]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="127" phylum="Arthropoda" rank="species" species="kraatzi">
<emphasis id="B97AEABEFFCBE538529185226012FDF5" box="[692,793,553,577]" italics="true" pageId="10" pageNumber="127">Z. kraatzi</emphasis>
</taxonomicName>
and allied species. It differs from the sympatric
<taxonomicName id="4C0E4D2FFFCBE538529185426012FDD5" authorityName="SCHUBERT" authorityYear="1908" box="[692,793,585,609]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="127" phylum="Arthropoda" rank="species" species="kraatzi">
<emphasis id="B97AEABEFFCBE538529185426012FDD5" box="[692,793,585,609]" italics="true" pageId="10" pageNumber="127">Z. kraatzi</emphasis>
</taxonomicName>
by larger body size, paler coloration (but note that the
<typeStatus id="54B5880EFFCBE538509885816215FD15" box="[189,286,649,673]" pageId="10" pageNumber="127" type="lectotype">lectotype</typeStatus>
is teneral), longer and more slender antennae, a less convex pronotum (cross-section) with less sparse punctation, denser, coarser, and somewhat asperate or granulose punctation of the elytra, even fewer nonsetiferous punctures on the abdomen, and by the larger aedeagus with a slightly broader crista apicalis and a ventral process of slightly different shape.
</paragraph>
<paragraph id="8BB136ACFFCBE53850988461627BFBB5" blockId="10.[189,794,521,1025]" pageId="10" pageNumber="127">
The possibility that
<taxonomicName id="4C0E4D2FFFCBE53851AC846262F8FC35" authorityName="ASSING" authorityYear="2016" box="[393,499,874,897]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="127" phylum="Arthropoda" rank="species" species="iniquus">
<emphasis id="B97AEABEFFCBE53851AC846262F8FC35" box="[393,499,874,897]" italics="true" pageId="10" pageNumber="127">Z. iniquus</emphasis>
</taxonomicName>
from
<collectingCountry id="F319763CFFCBE538520A84616181FC35" box="[559,650,873,897]" name="Pakistan" pageId="10" pageNumber="127">Pakistan</collectingCountry>
and
<collectingCountry id="F319763CFFCBE538529F846163E2FC15" name="Afghanistan" pageId="10" pageNumber="127">Afghanistan</collectingCountry>
is conspecific with
<taxonomicName id="4C0E4D2FFFCBE53851EF8482616BFC15" authorityName="SCHUBERT" authorityYear="1908" box="[458,608,906,929]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="127" phylum="Arthropoda" rank="species" species="exasperatus">
<emphasis id="B97AEABEFFCBE53851EF8482616BFC15" box="[458,608,906,929]" italics="true" pageId="10" pageNumber="127">Z. exasperatus</emphasis>
</taxonomicName>
cannot be ruled out completely. Mature males of
<taxonomicName id="4C0E4D2FFFCBE538523E84A261BAFC75" authorityName="SCHUBERT" authorityYear="1908" box="[539,689,938,961]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="127" phylum="Arthropoda" rank="species" species="exasperatus">
<emphasis id="B97AEABEFFCBE538523E84A261BAFC75" box="[539,689,938,961]" italics="true" pageId="10" pageNumber="127">Z. exasperatus</emphasis>
</taxonomicName>
would be needed to assess intraspecific variation of coloration and other characters.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFCBE53850988321615DFB15" pageId="10" pageNumber="127" type="distribution">
<paragraph id="8BB136ACFFCBE53850988321615DFB15" blockId="10.[189,794,1065,1185]" pageId="10" pageNumber="127">
<emphasis id="B97AEABEFFCBE53850988321624DFBF5" bold="true" box="[189,326,1065,1089]" pageId="10" pageNumber="127">Distribution</emphasis>
: This species is currently known only from the
<typeStatus id="54B5880EFFCBE53850C083426218FBD6" box="[229,275,1098,1122]" pageId="10" pageNumber="127">type</typeStatus>
locality in Himalachal Pradesh, Northwest
<collectingCountry id="F319763CFFCBE53852C783416011FBD5" box="[738,794,1097,1121]" name="India" pageId="10" pageNumber="127">India</collectingCountry>
(
<figureCitation id="13352A29FFCBE53850E183616202FB35" box="[196,265,1129,1153]" captionStart="Map 1" captionStartId="64.[189,234,822,843]" captionTargetBox="[189,1436,232,813]" captionTargetId="figure-76@64.[189,1436,232,813]" captionTargetPageId="64" captionText="Map 1: Revised distributions of Zyras kraatzi (black and white circles; white circle: type locality), Z. exasperatus (white circle), Z. iniquus (triangles), Z. pallipes (diamonds), and Z. championi (white stars) in the Himalaya." figureDoi="http://doi.org/10.5281/zenodo.5742393" httpUri="https://zenodo.org/record/5742393/files/figure.png" pageId="10" pageNumber="127">Map 1</figureCitation>
). The previous record from
<collectingRegion id="49CAF84EFFCBE5385210836161B6FB35" box="[565,701,1129,1153]" country="India" name="Uttarakhand" pageId="10" pageNumber="127">Uttarakhand</collectingRegion>
(
<bibRefCitation id="EF9F4B5DFFCBE53852EE836163F5FB15" author="ASSING, V." pageId="10" pageNumber="127" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING 2016a</bibRefCitation>
) is based on a misidentification.
</paragraph>
</subSubSection>
</treatment>
</document>
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