242 lines
32 KiB
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242 lines
32 KiB
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<document id="E9115CED7305D075B445E191DA10A5A1" ID-CLB-Dataset="7383" ID-DOI="10.11646/zootaxa.4951.3.7" ID-GBIF-Dataset="a6ee038f-4cda-4f38-8fde-8ad0efec8f0f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4668453" ID-ZooBank="EB876346-4282-4B4E-8585-ED50F1A616E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617782094822" checkinUser="plazi" docAuthor="Martens, Jochen, Maghradze, Eter & Barjadze, Shalva" docDate="2021" docId="03A787A4FFFDFFD43584F3981FAAFC1D" docLanguage="en" docName="zootaxa.4951.3.7.pdf" docOrigin="Zootaxa 4951 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nemaspela Silhavy 1966" docType="treatment" docVersion="8" lastPageNumber="543" masterDocId="FF9EFFDCFFFFFFD63513F30F1D1AFFA4" masterDocTitle="Two new species of the genus Nemaspela Šilhavý from caves in Georgia (Opiliones Nemastomatidae)" masterLastPageNumber="558" masterPageNumber="541" pageNumber="543" updateTime="1698936134096" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="CA2C708E1BC0AEA182A42C11F558B44D">
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<mods:title id="9EF23971B7640618929A439BFE6518EA">Two new species of the genus Nemaspela Šilhavý from caves in Georgia (Opiliones Nemastomatidae)</mods:title>
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<mods:name id="FC86881F209930D736FE3288EB17FC79" type="personal">
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<mods:namePart id="9A2A31B845B67A934546472D4463A17C">Martens, Jochen</mods:namePart>
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<mods:affiliation id="44F94C0611FDC3DCC54D7C2BBBF2519B">Johannes Gutenberg-Universität, Institut für Organismische und Molekulare Evolutionsbiologie (iomE), D- 55099 Mainz, Germany; Senckenberg Research Institute, Arachnology, D- 60325 Frankfurt am Main, Germany</mods:affiliation>
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<mods:namePart id="B66C392487A3FE27FADCD830DB7FA314">Maghradze, Eter</mods:namePart>
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<mods:affiliation id="9B17D9DBF10AF37C1B81BE2D09F98A45">Institute of Zoology, Ilia State University, Giorgi Tsereteli 3, 0162, Tbilisi, Georgia</mods:affiliation>
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<mods:namePart id="4FAF79C289A46DCEE3C7C0DBF9FDABC2">Barjadze, Shalva</mods:namePart>
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<mods:affiliation id="0F61187E61A50F8462425D4992B7E130">Institute of Zoology, Ilia State University, Giorgi Tsereteli 3, 0162, Tbilisi, Georgia</mods:affiliation>
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<mods:title id="D0771597DD75ABB0182CF95E12ED5EC7">Zootaxa</mods:title>
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<mods:date id="B3B2E5B764C88C25F04F98272CAC6B56">2021</mods:date>
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<mods:number id="1AA8406157EF297A474F83E5FB36E8F6">2021-04-07</mods:number>
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<treatment id="03A787A4FFFDFFD43584F3981FAAFC1D" ID-DOI="http://doi.org/10.5281/zenodo.4685055" ID-GBIF-Taxon="181003512" ID-Zenodo-Dep="4685055" LSID="urn:lsid:plazi:treatment:03A787A4FFFDFFD43584F3981FAAFC1D" httpUri="http://treatment.plazi.org/id/03A787A4FFFDFFD43584F3981FAAFC1D" lastPageNumber="543" pageId="2" pageNumber="543">
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<subSubSection id="C3146539FFFDFFD43584F3981F48FF15" box="[151,594,151,180]" pageId="2" pageNumber="543" type="nomenclature">
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<paragraph id="8BB136B2FFFDFFD43584F3981F48FF15" blockId="2.[151,594,151,180]" box="[151,594,151,180]" pageId="2" pageNumber="543">
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<heading id="D0F981DEFFFDFFD43584F3981F48FF15" bold="true" box="[151,594,151,180]" fontSize="11" level="1" pageId="2" pageNumber="543" reason="1">
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<emphasis id="B97AEAA0FFFDFFD43584F3981F48FF15" bold="true" box="[151,594,151,180]" pageId="2" pageNumber="543">
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The genus
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<taxonomicName id="4C0E4D31FFFDFFD4340DF3981F48FF15" ID-CoL="7PPVQ" authority="Silhavy, 1966" authorityName="Silhavy" authorityYear="1966" box="[286,594,151,180]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus">
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<emphasis id="B97AEAA0FFFDFFD4340DF3981CBEFF16" bold="true" box="[286,420,151,178]" italics="true" pageId="2" pageNumber="543">Nemaspela</emphasis>
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<bibRefCitation id="EF9F4B43FFFDFFD434B8F3971F48FF15" author="Silhavy, V." box="[427,594,151,180]" pageId="2" pageNumber="543" pagination="67 - 72" refId="ref10547" refString="Silhavy, V. (1966) Uber die Genitalmorphologie der Nemastomatidae (Arach., Opiliones). Senckenbergiana biologica, 47 (1), 67 - 72." type="journal article" year="1966">Šilhavý, 1966</bibRefCitation>
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</emphasis>
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<paragraph id="8BB136B2FFFDFFD43584F3D31FE5FF50" blockId="2.[151,1437,219,305]" box="[151,767,219,246]" pageId="2" pageNumber="543">
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<taxonomicName id="4C0E4D31FFFDFFD43584F3D31CB9FF57" ID-CoL="7PPVQ" authority="Silhavy, 1966" authorityName="Silhavy" authorityYear="1966" box="[151,419,219,246]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus">
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<emphasis id="B97AEAA0FFFDFFD43584F3D31C12FF50" box="[151,264,220,244]" italics="true" pageId="2" pageNumber="543">Nemaspela</emphasis>
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<bibRefCitation id="EF9F4B43FFFDFFD43402F3D21CB9FF57" author="Silhavy, V." box="[273,419,219,246]" pageId="2" pageNumber="543" pagination="67 - 72" refId="ref10547" refString="Silhavy, V. (1966) Uber die Genitalmorphologie der Nemastomatidae (Arach., Opiliones). Senckenbergiana biologica, 47 (1), 67 - 72." type="journal article" year="1966">Šilhavý, 1966</bibRefCitation>
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, Senckenbergia biologica 47: 72.
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<typeStatus id="54B58810FFFDFFD43614F3D31E23FF50" box="[775,825,220,244]" pageId="2" pageNumber="543">Type</typeStatus>
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species:
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<taxonomicName id="4C0E4D31FFFDFFD4368FF3D31DE0FEB6" ID-CoL="9X6CS" authority="Ljovuschkin & Starobogatov, 1963" authorityName="Ljovuschkin & Starobogatov" authorityYear="1963" class="Arachnida" family="Nemastomatidae" genus="Buresiolla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="sokolovi">
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<emphasis id="B97AEAA0FFFDFFD4368FF3D31978FF50" box="[924,1122,220,244]" italics="true" pageId="2" pageNumber="543">Buresiolla sokolovi</emphasis>
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<bibRefCitation id="EF9F4B43FFFDFFD4317FF3D31DE0FEB6" author="Ljovuschkin, S. T. & Starobogatov, Y. I." pageId="2" pageNumber="543" pagination="41 - 51" refId="ref9622" refString="Ljovuschkin, S. T. & Starobogatov, Y. I. (1963) The cavernicolous Opilionidea in the Crimea and Caucasus. Biospeologica Sovietica, XVIII. Byulletin Moskovskogo Obshchestva Ispytatelei Prirody, Otdel Biologicheskii, 68 (1), 41 - 51. [in Russian]" type="journal article" year="1963">Ljovuschkin & Starobogatov, 1963</bibRefCitation>
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(by original designation).
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</paragraph>
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<paragraph id="8BB136B2FFFDFFD4371AF3F51E93FE95" blockId="2.[151,1437,219,305]" pageId="2" pageNumber="543">
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—
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<bibRefCitation id="EF9F4B43FFFDFFD43737F3F51FF5FEB6" author="Martens, J." box="[548,751,250,274]" pageId="2" pageNumber="543" pagination="145 - 210" refId="ref9859" refString="Martens, J. (2006) Weberknechte aus dem Kaukasus (Arachnida, Opiliones, Nemastomatidae). Senckenbergiana biologica, 86 (2), 145 - 210." type="journal article" year="2006">Martens 2006: 169</bibRefCitation>
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;
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<bibRefCitation id="EF9F4B43FFFDFFD437EEF3F51ECFFEB6" author="Chemeris, A." box="[765,981,250,274]" pageId="2" pageNumber="543" pagination="286 - 377" refId="ref9136" refString="Chemeris, A. (2009) New data on the harvestman genus Nemaspela S ilhavy, 1966 (Arachnida: Opiliones). Bulletin of the British Arachnological Society, 14 (7), 286 - 377. https: // doi. org / 10.13156 / arac. 2009.14.7.286" type="journal article" year="2009">Chemeris 2009: 287</bibRefCitation>
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;
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<bibRefCitation id="EF9F4B43FFFDFFD436F0F3F519ACFEB6" author="Tchemeris, A. N." box="[995,1206,250,274]" pageId="2" pageNumber="543" pagination="41 - 46" refId="ref10734" refString="Tchemeris, A. N. (2013) Two new harvestman species (Arachnida: Opiliones) from the collection of the Siberian Zoological Museum. Arthropoda selecta, 22, 41 - 46. https: // doi. org / 10.15298 / arthsel. 22.1.03" type="journal article" year="2013">Tchemeris 2013: 41</bibRefCitation>
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;
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<collectingRegion id="49CAF850FFFDFFD431D7F3F4183BFEB6" box="[1220,1313,251,274]" country="Turkey" name="Karaman" pageId="2" pageNumber="543">Karaman</collectingRegion>
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2013: 241
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;
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<bibRefCitation id="EF9F4B43FFFDFFD435D4F2161C84FE95" author="Schonhofer, A. L." box="[199,414,281,305]" pageId="2" pageNumber="543" pagination="1 - 68" refId="ref10322" refString="Schonhofer, A. L. (2013) A taxonomic catalogue of the Dyspnoi Hansen and Sorensen, 1904 (Arachnida: Opiliones). Zootaxa, 3679 (1), 1 - 68. https: // doi. org / 10.11646 / zootaxa. 3679.1.1" type="journal article" year="2013">Schönhofer 2013: 35</bibRefCitation>
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<bibRefCitation id="EF9F4B43FFFDFFD434BAF2161FABFE95" author="Turbanov, I. S. & Demidov, P. E. & Kolesnikov, V. B. & Turbanova, A. A." box="[425,689,281,305]" pageId="2" pageNumber="543" pagination="360 - 368" refId="ref10892" refString="Turbanov, I. S., Demidov, P. E., Kolesnikov, V. B. & Turbanova, A. A. (2018) Preliminary results of investigation of invertebrate animals study in the Veryovkina Cave (The Western Caucasus, Abkhazia). In: Gunko, A. A. (Eds.), Speleology and Spelestology. Collection of materials of IX International scientific conference. Nizhny Novgorod Pedagogical University Press, Nizhny Novgorod, pp. 360 - 368." type="book chapter" year="2018">Turbanov et al. 2018: 363</bibRefCitation>
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;
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<treatmentCitation id="0AAF10A3FFFDFFD437AFF2161E9FFE95" author="Kozel, P. & Delic, T. & Novak, T." box="[700,901,281,305]" page="4" pageId="2" pageNumber="543" year="2020">
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<bibRefCitation id="EF9F4B43FFFDFFD437AFF2161E9FFE95" author="Kozel, P. & Delic, T. & Novak, T." box="[700,901,281,305]" pageId="2" pageNumber="543" pagination="90 - 107" refId="ref9487" refString="Kozel, P., Delic, T. & Novak, T. (2020) Nemaspela borkoae sp. nov. (Opiliones: Nemastomatidae), the second species of the genus from the Dinaric Karst. European Journal of Taxonomy, 717 (1), 90 - 107. https: // doi. org / 10.5852 / ejt. 2020.717.1103" type="journal article" year="2020">Kozel et al. 2020: 4</bibRefCitation>
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</treatmentCitation>
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.
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<subSubSection id="C3146539FFFDFFD43584F2501FAAFC1D" pageId="2" pageNumber="543" type="reference_group">
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<paragraph id="8BB136B2FFFDFFD43584F2501FAAFC1D" blockId="2.[151,1437,351,954]" pageId="2" pageNumber="543">
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The taxonomic history of
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<taxonomicName id="4C0E4D31FFFDFFD434ADF2501F20FEDD" ID-CoL="7PPVQ" authorityName="Silhavy" authorityYear="1966" box="[446,570,351,377]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus">
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<emphasis id="B97AEAA0FFFDFFD434ADF2501F20FEDD" box="[446,570,351,377]" italics="true" pageId="2" pageNumber="543">Nemaspela</emphasis>
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</taxonomicName>
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is relatively short. The first troglobiont nemastomatid species was placed in
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<taxonomicName id="4C0E4D31FFFDFFD43584F28B1C01FE39" ID-CoL="7PLQD" box="[151,283,388,413]" class="Arachnida" family="Nemastomatidae" genus="Nemastoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus">
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<emphasis id="B97AEAA0FFFDFFD43584F28B1C01FE39" box="[151,283,388,413]" italics="true" pageId="2" pageNumber="543">Nemastoma</emphasis>
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</taxonomicName>
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by
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<bibRefCitation id="EF9F4B43FFFDFFD43455F28C1CCDFE39" author="Grese, N." box="[326,471,387,413]" pageId="2" pageNumber="543" pagination="108" refId="ref9373" refString="Grese, N. (1911) Uber eine blinde Nemastoma - Art aus einer Hohle in der Krim (Nemastoma caecum nov. sp.). Zoologischer Anzeiger, 37, 108." type="journal article" year="1911">Grese (1911)</bibRefCitation>
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from caves in the
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<collectingRegion id="49CAF850FFFDFFD437BBF28C1FE7FE39" box="[680,765,387,413]" country="Ukraine" name="Crimea" pageId="2" pageNumber="543">Crimea</collectingRegion>
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,
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<collectingCountry id="F3197622FFFDFFD4361BF28C1E7FFE39" box="[776,869,387,413]" name="Ukraine" pageId="2" pageNumber="543">Ukraine</collectingCountry>
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, then the genus name covering most species of the family. Later
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<taxonomicName id="4C0E4D31FFFDFFD43423F2A81F70FE66" ID-CoL="9WZ4D" authority="Kratochvil, 1958" authorityName="Kratochvil" authorityYear="1958" box="[304,618,423,450]" class="Arachnida" family="Nemastomatidae" genus="Buresiolla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus">
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<emphasis id="B97AEAA0FFFDFFD43423F2A81CBEFE65" box="[304,420,423,449]" italics="true" pageId="2" pageNumber="543">Buresiolla</emphasis>
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<bibRefCitation id="EF9F4B43FFFDFFD434B8F2A81F70FE66" author="Kratochvil, J." box="[427,618,423,450]" pageId="2" pageNumber="543" pagination="523 - 576" refId="ref9553" refString="Kratochvil, J. (1958) Hohlenweberknechte Bulgariens (Palpatores-Nemastomatidae). Prace Brnenske Zakladny Ceskoslovenske Akademie Ved, 30 (12), 523 - 576." type="journal article" year="1958">Kratochvíl, 1958</bibRefCitation>
|
|
</taxonomicName>
|
|
was erected for a troglobiotic species from Bulgarian caves, but its
|
|
<typeStatus id="54B58810FFFDFFD4307FF2A71886FE66" box="[1388,1436,424,450]" pageId="2" pageNumber="543">type</typeStatus>
|
|
species did not belong to the later
|
|
<taxonomicName id="4C0E4D31FFFDFFD4373AF2C41FBFFE41" ID-CoL="7PPVQ" authorityName="Silhavy" authorityYear="1966" box="[553,677,459,485]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B97AEAA0FFFDFFD4373AF2C41FBFFE41" box="[553,677,459,485]" italics="true" pageId="2" pageNumber="543">Nemaspela</emphasis>
|
|
</taxonomicName>
|
|
and the genus
|
|
<taxonomicName id="4C0E4D31FFFDFFD4364BF2C41ED6FE41" ID-CoL="9WZ4D" authorityName="Kratochvil" authorityYear="1958" box="[856,972,459,485]" class="Arachnida" family="Nemastomatidae" genus="Buresiolla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B97AEAA0FFFDFFD4364BF2C41ED6FE41" box="[856,972,459,485]" italics="true" pageId="2" pageNumber="543">Buresiolla</emphasis>
|
|
</taxonomicName>
|
|
was synonymized with
|
|
<taxonomicName id="4C0E4D31FFFDFFD431F6F2C31C7DFDAD" ID-CoL="7PS37" authority="Redikorzew, 1936" authorityName="Redikorzew" authorityYear="1936" class="Arachnida" family="Nemastomatidae" genus="Paranemastoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B97AEAA0FFFDFFD431F6F2C31887FE41" box="[1253,1437,460,485]" italics="true" pageId="2" pageNumber="543">Paranemastoma</emphasis>
|
|
Redikorzew, 1936
|
|
</taxonomicName>
|
|
. But
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|
<bibRefCitation id="EF9F4B43FFFDFFD434B6F2E01E5BFDAE" author="Ljovuschkin, S. T. & Starobogatov, Y. I." box="[421,833,495,522]" pageId="2" pageNumber="543" pagination="41 - 51" refId="ref9622" refString="Ljovuschkin, S. T. & Starobogatov, Y. I. (1963) The cavernicolous Opilionidea in the Crimea and Caucasus. Biospeologica Sovietica, XVIII. Byulletin Moskovskogo Obshchestva Ispytatelei Prirody, Otdel Biologicheskii, 68 (1), 41 - 51. [in Russian]" type="journal article" year="1963">Ljovuschkin & Starobogatov (1963)</bibRefCitation>
|
|
followed the
|
|
<taxonomicName id="4C0E4D31FFFDFFD436F2F2E0194FFDAD" ID-CoL="9WZ4D" authorityName="Kratochvil" authorityYear="1958" box="[993,1109,495,521]" class="Arachnida" family="Nemastomatidae" genus="Buresiolla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B97AEAA0FFFDFFD436F2F2E0194FFDAD" box="[993,1109,495,521]" italics="true" pageId="2" pageNumber="543">Buresiolla</emphasis>
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|
</taxonomicName>
|
|
affiliation of troglobiont nemastomatids, when they named the first Caucasian troglobiont nemastomatids (
|
|
<taxonomicName id="4C0E4D31FFFDFFD4313FF11C1C8AFDF5" ID-CoL="9X6CS" authority="Ljovuschkin & Starobogatov, 1963" authorityName="Ljovuschkin & Starobogatov" authorityYear="1963" class="Arachnida" family="Nemastomatidae" genus="Buresiolla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="sokolovi">
|
|
<emphasis id="B97AEAA0FFFDFFD4313FF11C181CFD89" box="[1068,1286,531,557]" italics="true" pageId="2" pageNumber="543">Buresiolla sokolovi</emphasis>
|
|
<bibRefCitation id="EF9F4B43FFFDFFD43003F11C1C8AFDF5" author="Ljovuschkin, S. T. & Starobogatov, Y. I." pageId="2" pageNumber="543" pagination="41 - 51" refId="ref9622" refString="Ljovuschkin, S. T. & Starobogatov, Y. I. (1963) The cavernicolous Opilionidea in the Crimea and Caucasus. Biospeologica Sovietica, XVIII. Byulletin Moskovskogo Obshchestva Ispytatelei Prirody, Otdel Biologicheskii, 68 (1), 41 - 51. [in Russian]" type="journal article" year="1963">Ljovuschkin & Starobogatov, 1963</bibRefCitation>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C0E4D31FFFDFFD434DAF1381955FDF6" ID-CoL="9X6L2" authority="Ljovuschkin & Starobogatov, 1963" authorityName="Ljovuschkin & Starobogatov" authorityYear="1963" box="[457,1103,567,594]" class="Arachnida" family="Nemastomatidae" genus="Buresiolla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="abchasica">
|
|
<emphasis id="B97AEAA0FFFDFFD434DAF1381FADFDF5" box="[457,695,567,593]" italics="true" pageId="2" pageNumber="543">Buresiolla abchasica</emphasis>
|
|
<bibRefCitation id="EF9F4B43FFFDFFD437ADF1381955FDF6" author="Ljovuschkin, S. T. & Starobogatov, Y. I." box="[702,1103,567,594]" pageId="2" pageNumber="543" pagination="41 - 51" refId="ref9622" refString="Ljovuschkin, S. T. & Starobogatov, Y. I. (1963) The cavernicolous Opilionidea in the Crimea and Caucasus. Biospeologica Sovietica, XVIII. Byulletin Moskovskogo Obshchestva Ispytatelei Prirody, Otdel Biologicheskii, 68 (1), 41 - 51. [in Russian]" type="journal article" year="1963">Ljovuschkin & Starobogatov, 1963</bibRefCitation>
|
|
</taxonomicName>
|
|
).
|
|
<bibRefCitation id="EF9F4B43FFFDFFD43177F1361816FDF6" author="Silhavy, V." box="[1124,1292,567,596]" pageId="2" pageNumber="543" pagination="67 - 72" refId="ref10547" refString="Silhavy, V. (1966) Uber die Genitalmorphologie der Nemastomatidae (Arach., Opiliones). Senckenbergiana biologica, 47 (1), 67 - 72." type="journal article" year="1966">Šilhavý (1966)</bibRefCitation>
|
|
clarified the situation and erected the genus
|
|
<taxonomicName id="4C0E4D31FFFDFFD434EBF1541F6EFDD1" ID-CoL="7PPVQ" authorityName="Silhavy" authorityYear="1966" box="[504,628,603,629]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B97AEAA0FFFDFFD434EBF1541F6EFDD1" box="[504,628,603,629]" italics="true" pageId="2" pageNumber="543">Nemaspela</emphasis>
|
|
</taxonomicName>
|
|
for the Caucasian troglobiont nemastomatid species. After a long hiatus of about 34 years,
|
|
<bibRefCitation id="EF9F4B43FFFDFFD43477F18F1F0BFD3E" author="Martens, J." box="[356,529,639,666]" pageId="2" pageNumber="543" pagination="145 - 210" refId="ref9859" refString="Martens, J. (2006) Weberknechte aus dem Kaukasus (Arachnida, Opiliones, Nemastomatidae). Senckenbergiana biologica, 86 (2), 145 - 210." type="journal article" year="2006">Martens (2006)</bibRefCitation>
|
|
described
|
|
<taxonomicName id="4C0E4D31FFFDFFD4379AF170194CFD3D" ID-CoL="7XFN5" authority="Martens, 2006" authorityName="Martens" authorityYear="2006" box="[649,1110,639,666]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="femorecurvata">
|
|
<emphasis id="B97AEAA0FFFDFFD4379AF1701EB4FD3D" box="[649,942,639,665]" italics="true" pageId="2" pageNumber="543">Nemaspela femorecurvata</emphasis>
|
|
<bibRefCitation id="EF9F4B43FFFDFFD436A7F18F194CFD3D" author="Martens, J." box="[948,1110,639,666]" pageId="2" pageNumber="543" pagination="145 - 210" refId="ref9859" refString="Martens, J. (2006) Weberknechte aus dem Kaukasus (Arachnida, Opiliones, Nemastomatidae). Senckenbergiana biologica, 86 (2), 145 - 210." type="journal article" year="2006">Martens, 2006</bibRefCitation>
|
|
</taxonomicName>
|
|
from a cave in Racha region, western
|
|
<collectingCountry id="F3197622FFFDFFD435E5F1AC1C49FD19" box="[246,339,675,701]" name="Georgia" pageId="2" pageNumber="543">Georgia</collectingCountry>
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|
, then the easternmost record of the genus. The last novelties in the Caucasian fauna were
|
|
<taxonomicName id="4C0E4D31FFFDFFD43054F1AB1CD6FD45" ID-CoL="7XFN7" authority="Chemeris, 2009" authorityName="Chemeris" authorityYear="2009" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="kovali">
|
|
<emphasis id="B97AEAA0FFFDFFD43054F1AB1C09FD45" italics="true" pageId="2" pageNumber="543">Nemaspela kovali</emphasis>
|
|
<bibRefCitation id="EF9F4B43FFFDFFD43408F1C81CD6FD45" author="Chemeris, A." box="[283,460,711,737]" pageId="2" pageNumber="543" pagination="286 - 377" refId="ref9136" refString="Chemeris, A. (2009) New data on the harvestman genus Nemaspela S ilhavy, 1966 (Arachnida: Opiliones). Bulletin of the British Arachnological Society, 14 (7), 286 - 377. https: // doi. org / 10.13156 / arac. 2009.14.7.286" type="journal article" year="2009">Chemeris, 2009</bibRefCitation>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C0E4D31FFFDFFD43717F1C81EB9FD45" ID-CoL="7XFN6" authority="Tchemeris, 2013" authorityName="Tchemeris" authorityYear="2013" box="[516,931,711,737]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="gagrica">
|
|
<emphasis id="B97AEAA0FFFDFFD43717F1C81FC5FD45" box="[516,735,711,737]" italics="true" pageId="2" pageNumber="543">Nemaspela gagrica</emphasis>
|
|
<bibRefCitation id="EF9F4B43FFFDFFD437F5F1C81EB9FD45" author="Tchemeris, A. N." box="[742,931,711,737]" pageId="2" pageNumber="543" pagination="41 - 46" refId="ref10734" refString="Tchemeris, A. N. (2013) Two new harvestman species (Arachnida: Opiliones) from the collection of the Siberian Zoological Museum. Arthropoda selecta, 22, 41 - 46. https: // doi. org / 10.15298 / arthsel. 22.1.03" type="journal article" year="2013">Tchemeris, 2013</bibRefCitation>
|
|
</taxonomicName>
|
|
, the latter from
|
|
<collectingCountry id="F3197622FFFDFFD43144F1C819ABFD45" box="[1111,1201,711,737]" name="Georgia" pageId="2" pageNumber="543">Georgia</collectingCountry>
|
|
and the former from
|
|
<collectingRegion id="49CAF850FFFDFFD43584F1E41C62FCA1" box="[151,376,747,773]" country="Russia" name="Kabardino-Balkariya" pageId="2" pageNumber="543">Kabardino-Balkaria</collectingRegion>
|
|
,
|
|
<collectingCountry id="F3197622FFFDFFD43497F1E41CC8FCA1" box="[388,466,747,773]" name="Russia" pageId="2" pageNumber="543">Russia</collectingCountry>
|
|
. This rather strict concentration of
|
|
<taxonomicName id="4C0E4D31FFFDFFD4364FF1E41EC2FCA1" ID-CoL="7PPVQ" authorityName="Silhavy" authorityYear="1966" box="[860,984,747,773]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B97AEAA0FFFDFFD4364FF1E41EC2FCA1" box="[860,984,747,773]" italics="true" pageId="2" pageNumber="543">Nemaspela</emphasis>
|
|
</taxonomicName>
|
|
species in the Caucasus and the Crimea was greatly enlarged by the discovery of two species in a Dinaric cave in
|
|
<collectingCountry id="F3197622FFFDFFD436F2F00019DDFC8D" box="[993,1223,783,809]" name="Bosnia and Herzegovina" pageId="2" pageNumber="543">Bosnia-Herzegovina</collectingCountry>
|
|
(
|
|
<taxonomicName id="4C0E4D31FFFDFFD431CAF0001C52FCE9" ID-CoL="7XFN8" authority="Karaman, 2013" authorityName="Karaman" authorityYear="2013" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="ladae">
|
|
<emphasis id="B97AEAA0FFFDFFD431CAF0001886FC8D" box="[1241,1436,783,809]" italics="true" pageId="2" pageNumber="543">Nemaspela ladae</emphasis>
|
|
<bibRefCitation id="EF9F4B43FFFDFFD43584F03B1C52FCE9" author="Karaman, I." box="[151,328,819,846]" pageId="2" pageNumber="543" pagination="240 - 248" refId="ref9434" refString="Karaman, I. (2013) Nemaspela ladae sp. n., a new troglobitic nemastomatid (Opiliones, Dyspnoi, Nemastomatidae) from a Dinaric cave. Zootaxa, 3694 (3), 240 - 248. https: // doi. org / 10.11646 / zootaxa. 3694.3.4" type="journal article" year="2013">Karaman, 2013</bibRefCitation>
|
|
</taxonomicName>
|
|
) and in
|
|
<collectingCountry id="F3197622FFFDFFD434BAF03B1F28FCEA" box="[425,562,820,846]" name="Serbia" pageId="2" pageNumber="543">Montenegro</collectingCountry>
|
|
in the south-western Balkan Peninsula (
|
|
<taxonomicName id="4C0E4D31FFFDFFD43110F03C1C34FCD5" ID-CoL="9X5JX" authority="Kozel, Delic & Novak, 2020" authorityName="Kozel, Delic & Novak" authorityYear="2020" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="species" species="borkoae">
|
|
<emphasis id="B97AEAA0FFFDFFD43110F03C19FFFCE9" box="[1027,1253,819,845]" italics="true" pageId="2" pageNumber="543">Nemaspela borkoae</emphasis>
|
|
<bibRefCitation id="EF9F4B43FFFDFFD431FDF03C1C34FCD5" author="Kozel, P. & Delic, T. & Novak, T." pageId="2" pageNumber="543" pagination="90 - 107" refId="ref9487" refString="Kozel, P., Delic, T. & Novak, T. (2020) Nemaspela borkoae sp. nov. (Opiliones: Nemastomatidae), the second species of the genus from the Dinaric Karst. European Journal of Taxonomy, 717 (1), 90 - 107. https: // doi. org / 10.5852 / ejt. 2020.717.1103" type="journal article" year="2020">Kozel, Delić & Novak, 2020</bibRefCitation>
|
|
</taxonomicName>
|
|
). These authors called them the western species group, these species lacking cheliceral apophysis in males. The additional discovery of two further
|
|
<taxonomicName id="4C0E4D31FFFDFFD437B5F0741E38FC31" ID-CoL="7PPVQ" authorityName="Silhavy" authorityYear="1966" box="[678,802,891,917]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B97AEAA0FFFDFFD437B5F0741E38FC31" box="[678,802,891,917]" italics="true" pageId="2" pageNumber="543">Nemaspela</emphasis>
|
|
</taxonomicName>
|
|
species as proposed here expands the Caucasian species set to eight and
|
|
<taxonomicName id="4C0E4D31FFFDFFD4345BF0901CDEFC1D" ID-CoL="7PPVQ" authorityName="Silhavy" authorityYear="1966" box="[328,452,927,953]" class="Arachnida" family="Nemastomatidae" genus="Nemaspela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="543" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B97AEAA0FFFDFFD4345BF0901CDEFC1D" box="[328,452,927,953]" italics="true" pageId="2" pageNumber="543">Nemaspela</emphasis>
|
|
</taxonomicName>
|
|
in general to eleven.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |