553 lines
73 KiB
XML
553 lines
73 KiB
XML
<document id="BC23C5E1964601BC8C329C58B21B51CF" ID-CLB-Dataset="4796" ID-DOI="10.11646/zootaxa.3681.3.6" ID-GBIF-Dataset="a60dc5f9-4cdb-4689-ac94-b0d415592cee" ID-ISSN="1175-5326" ID-Zenodo-Dep="5981756" ID-ZooBank="F8E35CEC-5741-474F-B363-E7BC6DA678D3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629865992942" checkinUser="felipe" docAuthor="Pilato, Giovanni, D’Urso, Vera & Lisi, Oscar" docDate="2013" docId="039E87AA4C664530FF68F938E88D6956" docLanguage="en" docName="zt03681p280.pdf" docOrigin="Zootaxa 3681 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ramazzottius libycus Pilato & D’Urso & Lisi 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="279" masterDocId="FFA7FFD24C624539FFFFFF95EC626C79" masterDocTitle="Ramazzottius thulini (Pilato, 1970) bona species and description of Ramazzottius libycus sp. nov. (Eutardigrada, Ramazzottidae)" masterLastPageNumber="280" masterPageNumber="270" pageNumber="274" updateTime="1699144413453" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="54E67EF5D2FF71303E3CC0C2C45B35D2" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="E8DEE2790DD3DA09EBBC9D31D895A379">
|
||
<mods:title id="E4A34B9B5824C7455DF12C0E6D35BC93">Ramazzottius thulini (Pilato, 1970) bona species and description of Ramazzottius libycus sp. nov. (Eutardigrada, Ramazzottidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="6DC1DCE19D5615ED5728C4358D650035" type="personal">
|
||
<mods:role id="0BCCAA2E10DB531256AD2B3E89A69F59">
|
||
<mods:roleTerm id="CF45358CCD3A4BC4C1D90B266892CC3B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="BDC53099DFBE8B9DE872D7522B0CB81B">Pilato, Giovanni</mods:namePart>
|
||
<mods:nameIdentifier id="F2402A7E15A3C6A635B206F96C4CF772" type="email">pilato@unict.it</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="2F6A10C7E0CE33B8E2BC6D5FBB8550AD" type="personal">
|
||
<mods:role id="99852F919380AC98AA48628F4A1F8DF5">
|
||
<mods:roleTerm id="EC26ABD9038C017150CF61592EC2C6AE">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C7B748E35EED4D794F8AB9D78AABB99D">D’Urso, Vera</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="A1C1A8099CE642C893F8F9C253F83CF2" type="personal">
|
||
<mods:role id="7A4C18677FE853C310DBB64864FF7D93">
|
||
<mods:roleTerm id="B7DC7A2D8DF858A5D0C3753DB5D4C377">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="72CEAF97BE5BCFF955A58DCC527EB4E2">Lisi, Oscar</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="082D93D5DCA321165421FB08B5979ED7">text</mods:typeOfResource>
|
||
<mods:relatedItem id="0341764ECA8157F044584505AC581D0F" type="host">
|
||
<mods:titleInfo id="4C5FA720DAF1A9D1D58BD751CE359845">
|
||
<mods:title id="12CBB7C27A92B2CB0CE355FCDF9F75B5">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="16B3B1203A55C3F6BB0FC47D2C907F08">
|
||
<mods:date id="CF8AA6A51ACCCAC6FFA19C3304E7FE23">2013</mods:date>
|
||
<mods:detail id="6A51BBF5A7BD5219C456424409C8447E" type="pubDate">
|
||
<mods:number id="C29AE96A0CD0DDBC75B3A717E87FDA03">2013-06-21</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="4847897BCD73698EC19E3BBB749A5BA0" type="volume">
|
||
<mods:number id="3B58FFAB334D257BCFDCD3DDD03C25FD">3681</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="C64A97AAD7DA9FFA3BA87D3F76D9450D" type="issue">
|
||
<mods:number id="46D411B9E70770D010C0CFBB8BDDC566">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="84182F90D9F51E283198BB45C3F57097" unit="page">
|
||
<mods:start id="A874FB4672296664D7A3638B9CF731EE">270</mods:start>
|
||
<mods:end id="94A26FF8AED0512051CB2B0C5D9ADD56">280</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="88D1A23F4EF46A52C099A9D28384D654">journal article</mods:classification>
|
||
<mods:identifier id="2FD984A934AE5285529FEE32DB61213B" type="CLB-Dataset">4796</mods:identifier>
|
||
<mods:identifier id="CFC3EAC0A6BD5F8A6F2710267BFCBD1F" type="DOI">10.11646/zootaxa.3681.3.6</mods:identifier>
|
||
<mods:identifier id="C23A7B58FF1561345034D2354299C12B" type="GBIF-Dataset">a60dc5f9-4cdb-4689-ac94-b0d415592cee</mods:identifier>
|
||
<mods:identifier id="5EA7D932CE3EBDECC85DC4FC3C049C55" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="2F0B672EDCA9032C541724C99B1A029C" type="Zenodo-Dep">5981756</mods:identifier>
|
||
<mods:identifier id="9FD3B38BBC3EB639215E72D97AB38742" type="ZooBank">F8E35CEC-5741-474F-B363-E7BC6DA678D3</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="039E87AA4C664530FF68F938E88D6956" ID-DOI="http://doi.org/10.5281/zenodo.6159099" ID-GBIF-Taxon="184961065" ID-Zenodo-Dep="6159099" LSID="urn:lsid:plazi:treatment:039E87AA4C664530FF68F938E88D6956" httpUri="http://treatment.plazi.org/id/039E87AA4C664530FF68F938E88D6956" lastPageId="9" lastPageNumber="279" pageId="4" pageNumber="274">
|
||
<subSubSection id="C32D65374C66453DFF68F938ED8B6ABC" box="[151,489,1709,1734]" pageId="4" pageNumber="274" type="nomenclature">
|
||
<paragraph id="8B8836BC4C66453DFF68F938ED8B6ABC" blockId="4.[151,489,1709,1769]" box="[151,489,1709,1734]" pageId="4" pageNumber="274">
|
||
<heading id="D0C081D04C66453DFF68F938ED8B6ABC" bold="true" box="[151,489,1709,1734]" fontSize="10" level="2" pageId="4" pageNumber="274" reason="2">
|
||
<taxonomicName id="4C374D3F4C66453DFF68F938EDEB6ABF" authority="Pilato & D’Urso & Lisi, 2013" authorityName="Pilato & D’Urso & Lisi" authorityYear="2013" box="[151,393,1709,1734]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="274" phylum="Tardigrada" rank="species" species="libycus" status="sp. nov.">Ramazzottius libycus</taxonomicName>
|
||
<taxonomicNameLabel id="A27057D54C66453DFE6FF93BED8B6ABC" box="[400,489,1710,1734]" pageId="4" pageNumber="274" rank="species">
|
||
<emphasis id="B943EAAE4C66453DFE6FF93BED806ABF" bold="true" box="[400,482,1710,1734]" pageId="4" pageNumber="274">
|
||
<emphasis id="B943EAAE4C66453DFE6FF93BEDD36ABF" box="[400,433,1710,1734]" inLineHeading="true" pageId="4" pageNumber="274" reason="1">sp.</emphasis>
|
||
nov
|
||
</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32D65374C66453DFF68F944ED256A90" box="[151,327,1745,1769]" pageId="4" pageNumber="274" type="description">
|
||
<paragraph id="8B8836BC4C66453DFF68F944ED256A90" blockId="4.[151,489,1709,1769]" box="[151,327,1745,1769]" pageId="4" pageNumber="274">
|
||
(
|
||
<figureCitation id="130C2A394C66453DFF60F944ED416A90" box="[159,291,1745,1769]" captionStart="FIGURE 5" captionStartId="7.[151,250,1403,1426]" captionTargetBox="[159,1424,194,1372]" captionTargetId="figure-332@7.[151,1436,167,1400]" captionTargetPageId="7" captionText="FIGURE 5. A, Ramazzottius sp. with smooth cuticle from Sweden. Claws of the second pair of legs. B–D, Ramazzottius libycus sp. nov. B, Cuticle ornamentation (paratype). C, Buccal-pharyngeal apparatus of the holotype. D, Claws of the second pair of legs of the holotype. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981774" httpUri="https://zenodo.org/record/5981774/files/figure.png" pageId="4" pageNumber="274">
|
||
Figs. 5B
|
||
<emphasis id="B943EAAE4C66453DFF00F947ED6F6A90" box="[255,269,1746,1769]" italics="true" pageId="4" pageNumber="274">–</emphasis>
|
||
D
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="130C2A394C66453DFED0F944ED5F6A90" box="[303,317,1745,1769]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1966]" captionTargetBox="[206,1381,229,1881]" captionTargetId="figure-68@8.[178,1414,193,1919]" captionTargetPageId="8" captionText="FIGURE 6. A–D, Ramazzottius libycus sp. nov. A, claws of the hind legs of the holotype. Arrow indicates a visible lunule. B, Claws of the hind legs of a paratype. Arrows indicate visible lunules. C, D, Eggs. Arrows indicate some conical or truncoconical processes. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981778" httpUri="https://zenodo.org/record/5981778/files/figure.png" pageId="4" pageNumber="274">6</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32D65374C66453CFF68F88CEFF86B5D" lastPageId="5" lastPageNumber="275" pageId="4" pageNumber="274" type="materials_examined">
|
||
<paragraph id="8B8836BC4C66453DFF68F88CE8966B00" blockId="4.[151,1436,1817,1985]" pageId="4" pageNumber="274">
|
||
<emphasis id="B943EAAE4C66453DFF68F88CEDE66B4B" bold="true" box="[151,388,1817,1842]" inLineHeading="true" pageId="4" pageNumber="274" reason="1">Material examined.</emphasis>
|
||
<materialsCitation id="3B5F3CE14C66453DFE71F88CE8926B00" ID-GBIF-Occurrence="3338664301" country="Libya" latitude="32.488056" location="University of Catania" longLatPrecision="20" longitude="20.838612" pageId="4" pageNumber="274" specimenCount="8" specimenCount-egg="1" typeStatus="holotype">
|
||
<collectingCountry id="F320762C4C66453DFE71F88CEDB06B48" box="[398,466,1817,1841]" name="Libya" pageId="4" pageNumber="274">Libya</collectingCountry>
|
||
, Barce (
|
||
<geoCoordinate id="EE03507B4C66453DFDCAF88CEEDE6B48" box="[565,700,1817,1841]" degrees="32" direction="north" minutes="29" orientation="latitude" pageId="4" pageNumber="274" precision="15" seconds="17" value="32.488056">32°29'17"N</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE03507B4C66453DFD33F88CEF2D6B48" box="[716,847,1817,1841]" degrees="20" direction="east" minutes="50" orientation="longitude" pageId="4" pageNumber="274" precision="15" seconds="19" value="20.838612">20°50'19"E</geoCoordinate>
|
||
), in the neighbourhood of the town; moss sample:
|
||
<typeStatus id="548C881E4C66453DFF68F8A8EC9B6B2C" box="[151,249,1853,1877]" pageId="4" pageNumber="274" type="holotype">holotype</typeStatus>
|
||
(slide No. 2754),
|
||
<specimenCount id="9D31FD354C66453DFE2EF8ABEE366B2F" box="[465,596,1854,1878]" pageId="4" pageNumber="274" type="generic" typeStatus="paratypes">7 paratypes</specimenCount>
|
||
(slides No. 2753 and 2754), and
|
||
<specimenCount id="9D31FD354C66453DFC1FF8A8E8376B2F" box="[992,1109,1853,1878]" pageId="4" pageNumber="274" type="egg">three eggs</specimenCount>
|
||
(slides No. 2731 and 2752) collected in 1975 by the colleagues Francesco Furnari and Salvatore Brullo (
|
||
<location id="8EE860674C66453DFC0FF8F4E88B6B00" LSID="urn:lsid:plazi:treatment:039E87AA4C664530FF68F938E88D6956:8EE860674C66453DFC0FF8F4E88B6B00" box="[1008,1257,1889,1913]" country="Libya" latitude="32.488056" longLatPrecision="20" longitude="20.838612" name="University of Catania" pageId="4" pageNumber="274">University of Catania</location>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8836BC4C66453DFF38F813E8586BB8" blockId="4.[151,1436,1817,1985]" pageId="4" pageNumber="274">
|
||
<emphasis id="B943EAAE4C66453DFF38F813EDED6BE7" bold="true" box="[199,399,1925,1950]" inLineHeading="true" pageId="4" pageNumber="274" reason="1">Type repository.</emphasis>
|
||
<materialsCitation id="3B5F3CE14C66453DFE67F810E8546BB8" ID-GBIF-Occurrence="3338664302" collectorName="Binda & Pilato" country="Italy" county="University of Catania" location="Museum" municipality="Marcello La Greca" pageId="4" pageNumber="274" specimenCount="2" typeStatus="holotype">
|
||
<typeStatus id="548C881E4C66453DFE67F810EE626BE4" box="[408,512,1925,1949]" pageId="4" pageNumber="274" type="holotype">Holotype</typeStatus>
|
||
and
|
||
<typeStatus id="548C881E4C66453DFDBFF813EEC96BE7" box="[576,683,1926,1950]" pageId="4" pageNumber="274" type="paratype">paratypes</typeStatus>
|
||
are deposited in the
|
||
<collectorName id="26C2536A4C66453DFC55F810EF8F6BE4" box="[938,1005,1925,1949]" pageId="4" pageNumber="274">Binda</collectorName>
|
||
&
|
||
<collectorName id="26C2536A4C66453DFBE5F810E8396BE4" box="[1050,1115,1925,1949]" pageId="4" pageNumber="274">Pilato</collectorName>
|
||
collection (
|
||
<location id="8EE860674C66453DFB17F810E92E6BE4" LSID="urn:lsid:plazi:treatment:039E87AA4C664530FF68F938E88D6956:8EE860674C66453DFB17F810E92E6BE4" box="[1256,1356,1925,1949]" country="Italy" county="University of Catania" municipality="Marcello La Greca" name="Museum" pageId="4" pageNumber="274">Museum</location>
|
||
of the
|
||
<location id="8EE860674C66453DFF68F83FED976BB8" LSID="urn:lsid:plazi:treatment:039E87AA4C664530FF68F938E88D6956:8EE860674C66453DFF68F83FED976BB8" box="[151,501,1961,1985]" country="Italy" county="University of Catania" municipality="Marcello La Greca" name="Department of Animal Biology" pageId="4" pageNumber="274">Department of Animal Biology</location>
|
||
“
|
||
<collectingMunicipality id="6BECACC64C66453DFDF9F83CEE836BB8" box="[518,737,1961,1985]" pageId="4" pageNumber="274">Marcello La Greca</collectingMunicipality>
|
||
”,
|
||
<collectingCounty id="62E94E304C66453DFD09F83CEF886BB8" box="[758,1002,1961,1985]" pageId="4" pageNumber="274">University of Catania</collectingCounty>
|
||
,
|
||
<collectingCountry id="F320762C4C66453DFC09F83CE84C6BB8" box="[1014,1070,1961,1985]" name="Italy" pageId="4" pageNumber="274">Italy</collectingCountry>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF4866344C67453CFF68F994EF986A2C" ID-DOI="http://doi.org/10.5281/zenodo.5981766" ID-Zenodo-Dep="5981766" httpUri="https://zenodo.org/record/5981766/files/figure.png" pageId="5" pageNumber="275" startId="5.[151,250,1537,1560]" targetBox="[157,1432,210,1503]" targetPageId="5">
|
||
<paragraph id="8B8836BC4C67453CFF68F994EF986A2C" blockId="5.[151,1436,1537,1621]" pageId="5" pageNumber="275">
|
||
<emphasis id="B943EAAE4C67453CFF68F994ED746A61" bold="true" box="[151,278,1537,1560]" pageId="5" pageNumber="275">FIGURE 3.</emphasis>
|
||
A
|
||
<emphasis id="B943EAAE4C67453CFECCF996ED226A61" box="[307,320,1539,1560]" italics="true" pageId="5" pageNumber="275">–</emphasis>
|
||
E,
|
||
<emphasis id="B943EAAE4C67453CFEA0F996EDBD6A61" box="[351,479,1539,1560]" italics="true" pageId="5" pageNumber="275">Ramazzotius</emphasis>
|
||
specimens from the
|
||
<emphasis id="B943EAAE4C67453CFD3EF997EF2B6A61" box="[705,841,1538,1560]" italics="true" pageId="5" pageNumber="275">locus typicus</emphasis>
|
||
of
|
||
<taxonomicName id="4C374D3F4C67453CFC8DF996E8746A61" authorityName="Doyere" authorityYear="1840" box="[882,1046,1538,1560]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="275" phylum="Tardigrada" rank="species" species="oberhaeuseri">
|
||
<emphasis id="B943EAAE4C67453CFC8DF996E8746A61" box="[882,1046,1538,1560]" italics="true" pageId="5" pageNumber="275">R. oberhaeuseri</emphasis>
|
||
</taxonomicName>
|
||
. A, Buccal-pharyngeal apparatus. B, Claws of the first and second pairs of legs. C, Internal claw of the first pair of legs. D, Internal claw of the third pair of legs. E, Claws of the second pair of legs. Arrow indicates a visible lunule. Scale bar = 10 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8836BC4C67453CFF38F9E9EFF86B5D" blockId="5.[151,1436,1660,2009]" pageId="5" pageNumber="275">
|
||
<emphasis id="B943EAAE4C67453CFF38F9E9EDC66AEC" bold="true" box="[199,420,1660,1685]" inLineHeading="true" pageId="5" pageNumber="275" reason="1">Specific diagnosis.</emphasis>
|
||
Cuticle ornamentation and colour (5 longitudinal and 9 transverse pigmented bands) as usually in
|
||
<taxonomicName id="4C374D3F4C67453CFEE7F934EDB36AC0" authorityName="Doyere" authorityYear="1840" box="[280,465,1697,1721]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="275" phylum="Tardigrada" rank="species" species="oberhaeuseri">
|
||
<emphasis id="B943EAAE4C67453CFEE7F934EDB36AC0" box="[280,465,1697,1721]" italics="true" pageId="5" pageNumber="275">R. oberhaeuseri</emphasis>
|
||
</taxonomicName>
|
||
; eye spots absent; a pair of elliptical cephalic organs present; buccal-pharyngeal apparatus of
|
||
<taxonomicName id="4C374D3F4C67453CFED2F950EDFF6AA4" authorityName="Ehrenberg" authorityYear="1848" box="[301,413,1733,1757]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="275" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B943EAAE4C67453CFED2F950EDFF6AA4" box="[301,413,1733,1757]" italics="true" pageId="5" pageNumber="275">Hypsibius</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="548C881E4C67453CFE59F950EDB46AA4" box="[422,470,1733,1757]" pageId="5" pageNumber="275">type</typeStatus>
|
||
(
|
||
<taxonomicName id="4C374D3F4C67453CFE15F953EE1D6AA4" authorityName="Binda & Pilato" authorityYear="1987" box="[490,639,1734,1757]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="275" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B943EAAE4C67453CFE15F953EE1D6AA4" box="[490,639,1734,1757]" italics="true" pageId="5" pageNumber="275">Ramazzottius</emphasis>
|
||
</taxonomicName>
|
||
variant), pharyngeal bulb with two macroplacoids; microplacoid and septulum absent; claws of
|
||
<taxonomicName id="4C374D3F4C67453CFE22F97CEE106B79" authorityName="Binda & Pilato" authorityYear="1987" box="[477,626,1769,1792]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="275" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B943EAAE4C67453CFE22F97CEE106B79" box="[477,626,1769,1792]" italics="true" pageId="5" pageNumber="275">Ramazzottius</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="548C881E4C67453CFD7FF97CEED66B78" box="[640,692,1769,1793]" pageId="5" pageNumber="275">type</typeStatus>
|
||
; small lunules, difficult to see, present. Eggs, laid free, with hemispherical and, very rarely, trunco-conical and conical processes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32D65374C674530FF38F8A5EE5A6D65" lastPageId="9" lastPageNumber="279" pageId="5" pageNumber="275" type="description">
|
||
<paragraph id="8B8836BC4C67453EFF38F8A5EE536A95" blockId="5.[151,1436,1660,2009]" lastBlockId="7.[151,1436,1532,2024]" lastPageId="7" lastPageNumber="277" pageId="5" pageNumber="275">
|
||
<emphasis id="B943EAAE4C67453CFF38F8A5EE7F6B30" bold="true" box="[199,541,1840,1865]" inLineHeading="true" pageId="5" pageNumber="275" reason="1">
|
||
Description of the
|
||
<typeStatus id="548C881E4C67453CFE50F8A4EE7A6B30" box="[431,536,1841,1865]" pageId="5" pageNumber="275" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Body length 286 µm; colour (5 longitudinal and 9 transverse reddish bands separated by unpigmented bands) and cuticular ornamentation as usually in
|
||
<taxonomicName id="4C374D3F4C67453CFC0BF8C3E8C56B14" authorityName="Doyere" authorityYear="1840" box="[1012,1191,1877,1901]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="275" phylum="Tardigrada" rank="species" species="oberhaeuseri">
|
||
<emphasis id="B943EAAE4C67453CFC0BF8C3E8C56B14" box="[1012,1191,1877,1901]" italics="true" pageId="5" pageNumber="275">R. oberhaeuseri</emphasis>
|
||
</taxonomicName>
|
||
; polygonal tubercles, less visible in the anterior portion of the body and more visible in the posterior portion, are present having diameter up to 2.6 µm (
|
||
<figureCitation id="130C2A394C67453CFEBDF808EDFF6BCC" box="[322,413,1949,1973]" captionStart="FIGURE 5" captionStartId="7.[151,250,1403,1426]" captionTargetBox="[159,1424,194,1372]" captionTargetId="figure-332@7.[151,1436,167,1400]" captionTargetPageId="7" captionText="FIGURE 5. A, Ramazzottius sp. with smooth cuticle from Sweden. Claws of the second pair of legs. B–D, Ramazzottius libycus sp. nov. B, Cuticle ornamentation (paratype). C, Buccal-pharyngeal apparatus of the holotype. D, Claws of the second pair of legs of the holotype. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981774" httpUri="https://zenodo.org/record/5981774/files/figure.png" pageId="5" pageNumber="275">Fig. 5B</figureCitation>
|
||
); eye spots absent; a paired of cephalic, elliptical, organs present as ascertained in most species of the genus; buccal-pharyngeal apparatus of
|
||
<taxonomicName id="4C374D3F4C67453CFD13F854EF3E6BA0" authorityName="Ehrenberg" authorityYear="1848" box="[748,860,1985,2009]" class="Eutardigrada" family="Hypsibiidae" genus="Hypsibius" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="275" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B943EAAE4C67453CFD13F854EF3E6BA0" box="[748,860,1985,2009]" italics="true" pageId="5" pageNumber="275">Hypsibius</emphasis>
|
||
</taxonomicName>
|
||
type (
|
||
<taxonomicName id="4C374D3F4C67453CFC5BF854E85B6BA1" authorityName="Binda & Pilato" authorityYear="1987" box="[932,1081,1985,2008]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="275" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B943EAAE4C67453CFC5BF854E85B6BA1" box="[932,1081,1985,2008]" italics="true" pageId="5" pageNumber="275">Ramazzottius</emphasis>
|
||
</taxonomicName>
|
||
variant) i.e. with rigid buccal tube without ventral lamina, dorsal and ventral apophyses for the insertion of the stylet muscles in shape of blunt hooks, asymmetrical with respect to the frontal plane; peribuccal lamellae and peribuccal papulae absent (
|
||
<bibRefCitation id="EFA64B4D4C65453EFAC3F98AED446A25" author="Pilato, G. & Binda M. G." pageId="7" pageNumber="277" pagination="1 - 54" refId="ref5254" refString="Pilato, G. & Binda M. G. (2010) Definition of families, subfamilies. genera and subgenera of the Eutardigrada, and keys to their identification. Zootaxa, 2404, 1 - 54." type="journal article" year="2010">Pilato & Binda, 2010</bibRefCitation>
|
||
). Buccal tube, 29 µm long (with a slightly thicker tube wall immediately caudally to the insertion of the stylet supports) (
|
||
<figureCitation id="130C2A394C65453EFE81F9F2EDB46A06" box="[382,470,1639,1663]" captionStart="FIGURE 5" captionStartId="7.[151,250,1403,1426]" captionTargetBox="[159,1424,194,1372]" captionTargetId="figure-332@7.[151,1436,167,1400]" captionTargetPageId="7" captionText="FIGURE 5. A, Ramazzottius sp. with smooth cuticle from Sweden. Claws of the second pair of legs. B–D, Ramazzottius libycus sp. nov. B, Cuticle ornamentation (paratype). C, Buccal-pharyngeal apparatus of the holotype. D, Claws of the second pair of legs of the holotype. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981774" httpUri="https://zenodo.org/record/5981774/files/figure.png" pageId="7" pageNumber="277">Fig. 5C</figureCitation>
|
||
), and 1.5 µm wide internally (
|
||
<emphasis id="B943EAAE4C65453EFCD1F9FCEFEB6AF9" box="[814,905,1640,1664]" italics="true" pageId="7" pageNumber="277">pt = 5.2</emphasis>
|
||
). Stylet supports inserted on the buccal tube at 60 % of its length (
|
||
<emphasis id="B943EAAE4C65453EFE84F918EDF36ADD" box="[379,401,1677,1700]" italics="true" pageId="7" pageNumber="277">pt</emphasis>
|
||
=
|
||
<emphasis id="B943EAAE4C65453EFE4BF919ED876ADD" box="[436,485,1676,1700]" italics="true" pageId="7" pageNumber="277">60.0</emphasis>
|
||
). Pharyngeal bulb with apophyses and two short macroplacoids; microplacoid and septulum absent (
|
||
<figureCitation id="130C2A394C65453EFE9DF93AEDDF6ABE" box="[354,445,1711,1735]" captionStart="FIGURE 5" captionStartId="7.[151,250,1403,1426]" captionTargetBox="[159,1424,194,1372]" captionTargetId="figure-332@7.[151,1436,167,1400]" captionTargetPageId="7" captionText="FIGURE 5. A, Ramazzottius sp. with smooth cuticle from Sweden. Claws of the second pair of legs. B–D, Ramazzottius libycus sp. nov. B, Cuticle ornamentation (paratype). C, Buccal-pharyngeal apparatus of the holotype. D, Claws of the second pair of legs of the holotype. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981774" httpUri="https://zenodo.org/record/5981774/files/figure.png" pageId="7" pageNumber="277">Fig. 5C</figureCitation>
|
||
); first macroplacoid 3.9 µm long (
|
||
<emphasis id="B943EAAE4C65453EFCABF924EF086AB1" box="[852,874,1713,1736]" italics="true" pageId="7" pageNumber="277">pt</emphasis>
|
||
=
|
||
<emphasis id="B943EAAE4C65453EFC71F925EFDD6AB1" box="[910,959,1712,1736]" italics="true" pageId="7" pageNumber="277">13.4</emphasis>
|
||
), second 3.1 µm long (
|
||
<emphasis id="B943EAAE4C65453EFB2AF924E8896AB1" box="[1237,1259,1713,1736]" italics="true" pageId="7" pageNumber="277">pt</emphasis>
|
||
=
|
||
<emphasis id="B943EAAE4C65453EFAEFF925E9236AB1" box="[1296,1345,1712,1736]" italics="true" pageId="7" pageNumber="277">10.7</emphasis>
|
||
); entire placoid row 7.6 µm long (
|
||
<emphasis id="B943EAAE4C65453EFE43F940EDB06A95" box="[444,466,1749,1772]" italics="true" pageId="7" pageNumber="277">pt</emphasis>
|
||
= 26.2).
|
||
</paragraph>
|
||
<caption id="DF4866344C64453FFF68F855E81D6BAF" ID-DOI="http://doi.org/10.5281/zenodo.5981770" ID-Zenodo-Dep="5981770" box="[151,1151,1984,2007]" httpUri="https://zenodo.org/record/5981770/files/figure.png" pageId="6" pageNumber="276" startId="6.[151,250,1984,2007]" targetBox="[241,1351,210,1928]" targetPageId="6">
|
||
<paragraph id="8B8836BC4C64453FFF68F855E81D6BAF" blockId="6.[151,1151,1984,2007]" box="[151,1151,1984,2007]" pageId="6" pageNumber="276">
|
||
<emphasis id="B943EAAE4C64453FFF68F855ED706BAE" bold="true" box="[151,274,1984,2007]" pageId="6" pageNumber="276">FIGURE 4.</emphasis>
|
||
A
|
||
<emphasis id="B943EAAE4C64453FFED4F857ED556BAE" box="[299,311,1986,2007]" italics="true" pageId="6" pageNumber="276">–</emphasis>
|
||
B,
|
||
<taxonomicName id="4C374D3F4C64453FFEAAF857EDBE6BAE" authorityName="Binda & Pilato" authorityYear="1987" box="[341,476,1986,2007]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="276" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B943EAAE4C64453FFEAAF857EDBE6BAE" box="[341,476,1986,2007]" italics="true" pageId="6" pageNumber="276">Ramazzottius</emphasis>
|
||
</taxonomicName>
|
||
egg from the
|
||
<emphasis id="B943EAAE4C64453FFD95F854EE8C6BAE" box="[618,750,1985,2007]" italics="true" pageId="6" pageNumber="276">locus typicus</emphasis>
|
||
of
|
||
<taxonomicName id="4C374D3F4C64453FFCF0F857EFD36BAE" authorityName="Doyere" authorityYear="1840" box="[783,945,1985,2007]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="276" phylum="Tardigrada" rank="species" species="oberhaeuseri">
|
||
<emphasis id="B943EAAE4C64453FFCF0F857EFD36BAE" box="[783,945,1985,2007]" italics="true" pageId="6" pageNumber="276">R. oberhaeuseri</emphasis>
|
||
</taxonomicName>
|
||
. Scale bar = 10 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4866344C65453EFF68FAEEEE1469B6" ID-DOI="http://doi.org/10.5281/zenodo.5981774" ID-Zenodo-Dep="5981774" httpUri="https://zenodo.org/record/5981774/files/figure.png" pageId="7" pageNumber="277" startId="7.[151,250,1403,1426]" targetBox="[159,1424,194,1372]" targetPageId="7">
|
||
<paragraph id="8B8836BC4C65453EFF68FAEEEE1469B6" blockId="7.[151,1436,1403,1487]" pageId="7" pageNumber="277">
|
||
<emphasis id="B943EAAE4C65453EFF68FAEEED7469E8" bold="true" box="[151,278,1403,1426]" pageId="7" pageNumber="277">FIGURE 5.</emphasis>
|
||
A,
|
||
<taxonomicName id="4C374D3F4C65453EFEBDFAE9ED9269E8" box="[322,496,1404,1425]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="277" phylum="Tardigrada" rank="species" species="undetermined">
|
||
<emphasis id="B943EAAE4C65453EFEBDFAE9EDAB69E8" box="[322,457,1404,1425]" italics="true" pageId="7" pageNumber="277">Ramazzottius</emphasis>
|
||
<emphasis id="B943EAAE4C65453EFE2DFAE9ED9269E8" bold="true" box="[466,496,1404,1425]" pageId="7" pageNumber="277">sp.</emphasis>
|
||
</taxonomicName>
|
||
with smooth cuticle from Sweden. Claws of the second pair of legs. B
|
||
<emphasis id="B943EAAE4C65453EFB19FAE9E89069E8" box="[1254,1266,1404,1425]" italics="true" pageId="7" pageNumber="277">–</emphasis>
|
||
D,
|
||
<taxonomicName id="4C374D3F4C65453EFAEAFAE9ECBD69C9" authority="Pilato & D’Urso & Lisi, 2013" authorityName="Pilato & D’Urso & Lisi" authorityYear="2013" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="277" phylum="Tardigrada" rank="species" species="libycus" status="sp. nov.">
|
||
<emphasis id="B943EAAE4C65453EFAEAFAE9ECBD69C9" italics="true" pageId="7" pageNumber="277">Ramazzottius libycus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943EAAE4C65453EFF1AFA0EED5469C9" bold="true" box="[229,310,1435,1456]" pageId="7" pageNumber="277">
|
||
<taxonomicNameLabel id="A27057D54C65453EFF1AFA0EED5469C9" box="[229,310,1435,1456]" pageId="7" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
B, Cuticle ornamentation (paratype). C, Buccal-pharyngeal apparatus of the holotype. D, Claws of the second pair of legs of the holotype. Scale bar = 10 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8836BC4C65453EFF38F962EC846B91" blockId="7.[151,1436,1532,2024]" pageId="7" pageNumber="277">
|
||
Claws, of
|
||
<taxonomicName id="4C374D3F4C65453EFEC4F96DEDB26B76" authorityName="Binda & Pilato" authorityYear="1987" box="[315,464,1784,1807]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="277" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="B943EAAE4C65453EFEC4F96DEDB26B76" box="[315,464,1784,1807]" italics="true" pageId="7" pageNumber="277">Ramazzottius</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="548C881E4C65453EFE27F96DEE6A6B69" box="[472,520,1784,1808]" pageId="7" pageNumber="277">type</typeStatus>
|
||
(
|
||
<figureCitation id="130C2A394C65453EFDE7F962EEF36B76" box="[536,657,1783,1807]" captionStart="FIGURE 5" captionStartId="7.[151,250,1403,1426]" captionTargetBox="[159,1424,194,1372]" captionTargetId="figure-332@7.[151,1436,167,1400]" captionTargetPageId="7" captionText="FIGURE 5. A, Ramazzottius sp. with smooth cuticle from Sweden. Claws of the second pair of legs. B–D, Ramazzottius libycus sp. nov. B, Cuticle ornamentation (paratype). C, Buccal-pharyngeal apparatus of the holotype. D, Claws of the second pair of legs of the holotype. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981774" httpUri="https://zenodo.org/record/5981774/files/figure.png" pageId="7" pageNumber="277">Fig. 5C, D</figureCitation>
|
||
and
|
||
<figureCitation id="130C2A394C65453EFD35F962EF726B76" box="[714,784,1783,1807]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1966]" captionTargetBox="[206,1381,229,1881]" captionTargetId="figure-68@8.[178,1414,193,1919]" captionTargetPageId="8" captionText="FIGURE 6. A–D, Ramazzottius libycus sp. nov. A, claws of the hind legs of the holotype. Arrow indicates a visible lunule. B, Claws of the hind legs of a paratype. Arrows indicate visible lunules. C, D, Eggs. Arrows indicate some conical or truncoconical processes. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981778" httpUri="https://zenodo.org/record/5981778/files/figure.png" pageId="7" pageNumber="277">6A, B</figureCitation>
|
||
), well developed; length of external and internal claws of the third pair of legs 19.1 µm (
|
||
<emphasis id="B943EAAE4C65453EFE01F888EE766B4D" box="[510,532,1821,1844]" italics="true" pageId="7" pageNumber="277">pt</emphasis>
|
||
=
|
||
<emphasis id="B943EAAE4C65453EFDC9F889EE056B4D" box="[566,615,1820,1844]" italics="true" pageId="7" pageNumber="277">65.9</emphasis>
|
||
) and 11.2 µm (
|
||
<emphasis id="B943EAAE4C65453EFCE3F888EF506B4D" box="[796,818,1821,1844]" italics="true" pageId="7" pageNumber="277">pt</emphasis>
|
||
=
|
||
<emphasis id="B943EAAE4C65453EFCABF889EFE76B4D" box="[852,901,1820,1844]" italics="true" pageId="7" pageNumber="277">38.6</emphasis>
|
||
), respectively; length of posterior and anterior claws of the hind legs 24.6 µm (
|
||
<emphasis id="B943EAAE4C65453EFE00F8D4EE776B21" box="[511,533,1857,1880]" italics="true" pageId="7" pageNumber="277">pt</emphasis>
|
||
=
|
||
<emphasis id="B943EAAE4C65453EFDCCF8D5EE066B21" box="[563,612,1856,1880]" italics="true" pageId="7" pageNumber="277">84.8</emphasis>
|
||
) and 12.2 µm (
|
||
<emphasis id="B943EAAE4C65453EFCEDF8D4EF4A6B21" box="[786,808,1857,1880]" italics="true" pageId="7" pageNumber="277">pt</emphasis>
|
||
=
|
||
<emphasis id="B943EAAE4C65453EFCBAF8D5EF146B21" box="[837,886,1856,1880]" italics="true" pageId="7" pageNumber="277">42.1</emphasis>
|
||
), respectively. Due to the difficulty in measuring the total length of the external claws, we provided the separate lengths of the sclerified portions of the main branches and the basal portion + secondary branch in
|
||
<tableCitation id="C6B503074C65453EFD09F81DEF286BD9" box="[758,842,1928,1952]" captionStart="TABLE 1" captionStartId="3.[151,241,259,283]" captionText="TABLE 1. Length (in µm) and other parameters of selected structures (using, where possible, animals with similar body length), for two specimens of Ramazzottius thulini, two R. oberhaeuseri from the locus typicus, one Swedish Ramazzottius sp. (with smooth cuticle), and three Ramazzottius libycus sp. nov. Due to the variable orientation of the external claws, only the length of the sclerified portion of the main branch and basal portion + secondary branch of each claw have been measured. Pt ratios in italics." pageId="7" pageNumber="277">Table 1</tableCitation>
|
||
. Thin accessory points present on the main branches of all claws. Small, difficult to see, lunules present on all legs (
|
||
<figureCitation id="130C2A394C65453EFCA3F839EFB56BBD" box="[860,983,1964,1988]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1966]" captionTargetBox="[206,1381,229,1881]" captionTargetId="figure-68@8.[178,1414,193,1919]" captionTargetPageId="8" captionText="FIGURE 6. A–D, Ramazzottius libycus sp. nov. A, claws of the hind legs of the holotype. Arrow indicates a visible lunule. B, Claws of the hind legs of a paratype. Arrows indicate visible lunules. C, D, Eggs. Arrows indicate some conical or truncoconical processes. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981778" httpUri="https://zenodo.org/record/5981778/files/figure.png" pageId="7" pageNumber="277">Fig. 6A, B</figureCitation>
|
||
). Other cuticular thickening on the legs, absent.
|
||
</paragraph>
|
||
<caption id="DF4866344C6A4531FF68F802EE776B8B" ID-DOI="http://doi.org/10.5281/zenodo.5981778" ID-Zenodo-Dep="5981778" httpUri="https://zenodo.org/record/5981778/files/figure.png" pageId="8" pageNumber="278" startId="8.[151,250,1943,1966]" targetBox="[206,1381,229,1881]" targetPageId="8">
|
||
<paragraph id="8B8836BC4C6A4531FF68F802EE776B8B" blockId="8.[151,1437,1943,2034]" pageId="8" pageNumber="278">
|
||
<emphasis id="B943EAAE4C6A4531FF68F802ED716BD7" bold="true" box="[151,275,1943,1966]" pageId="8" pageNumber="278">FIGURE 6.</emphasis>
|
||
A
|
||
<emphasis id="B943EAAE4C6A4531FED3F80CED5A6BD7" box="[300,312,1945,1966]" italics="true" pageId="8" pageNumber="278">–</emphasis>
|
||
D,
|
||
<taxonomicName id="4C374D3F4C6A4531FEA8F80CEE4E6BD7" authority="Pilato & D’Urso & Lisi, 2013" authorityName="Pilato & D’Urso & Lisi" authorityYear="2013" box="[343,556,1944,1966]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="8" pageNumber="278" phylum="Tardigrada" rank="species" species="libycus" status="sp. nov.">
|
||
<emphasis id="B943EAAE4C6A4531FEA8F80CEE4E6BD7" box="[343,556,1944,1966]" italics="true" pageId="8" pageNumber="278">Ramazzottius libycus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B943EAAE4C6A4531FDCCF80DEEE16BD4" bold="true" box="[563,643,1944,1965]" pageId="8" pageNumber="278">
|
||
<taxonomicNameLabel id="A27057D54C6A4531FDCCF80DEEE16BD4" box="[563,643,1944,1965]" pageId="8" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A, claws of the hind legs of the holotype. Arrow indicates a visible lunule. B, Claws of the hind legs of a paratype. Arrows indicate visible lunules. C, D, Eggs. Arrows indicate some conical or truncoconical processes. Scale bar = 10 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8836BC4C6B4530FF38FF0DEE5A6D65" blockId="9.[150,1437,151,1328]" pageId="9" pageNumber="279">
|
||
Eggs laid freely (
|
||
<figureCitation id="130C2A394C6B4530FE6EFF02EE6A6CD6" box="[401,520,151,175]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1966]" captionTargetBox="[206,1381,229,1881]" captionTargetId="figure-68@8.[178,1414,193,1919]" captionTargetPageId="8" captionText="FIGURE 6. A–D, Ramazzottius libycus sp. nov. A, claws of the hind legs of the holotype. Arrow indicates a visible lunule. B, Claws of the hind legs of a paratype. Arrows indicate visible lunules. C, D, Eggs. Arrows indicate some conical or truncoconical processes. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981778" httpUri="https://zenodo.org/record/5981778/files/figure.png" pageId="9" pageNumber="279">Fig. 6C,D</figureCitation>
|
||
); we found
|
||
<specimenCount id="9D31FD354C6B4530FD69FF02EF6E6CC9" box="[662,780,151,176]" pageId="9" pageNumber="279" type="egg">three eggs</specimenCount>
|
||
but only one is measurable; its the diameter is 60.0 µm excluding and 64.0 µm including processes; there are 16 processes in the circumference and
|
||
<quantity id="4CCF9B594C6B4530FB55FF29E8816CAD" box="[1194,1251,188,212]" metricMagnitude="0" metricUnit="m" metricValue="1.2192" pageId="9" pageNumber="279" unit="in" value="48.0">48 in</quantity>
|
||
the hemisphere. Most processes are hemispherical, 3.0
|
||
<emphasis id="B943EAAE4C6B4530FDC0FF74EE2F6C81" box="[575,589,225,248]" italics="true" pageId="9" pageNumber="279">–</emphasis>
|
||
3.4 µm high, with the basal diameter 8.0
|
||
<emphasis id="B943EAAE4C6B4530FBF0FF74E87F6C81" box="[1039,1053,225,248]" italics="true" pageId="9" pageNumber="279">–</emphasis>
|
||
8.6 µm. Few processes are truncoconical or conical in shape (
|
||
<figureCitation id="130C2A394C6B4530FE2FFE91EE496D65" box="[464,555,260,284]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1966]" captionTargetBox="[206,1381,229,1881]" captionTargetId="figure-68@8.[178,1414,193,1919]" captionTargetPageId="8" captionText="FIGURE 6. A–D, Ramazzottius libycus sp. nov. A, claws of the hind legs of the holotype. Arrow indicates a visible lunule. B, Claws of the hind legs of a paratype. Arrows indicate visible lunules. C, D, Eggs. Arrows indicate some conical or truncoconical processes. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981778" httpUri="https://zenodo.org/record/5981778/files/figure.png" pageId="9" pageNumber="279">Fig. 6D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32D65374C6B4530FF38FEBDED926D1D" pageId="9" pageNumber="279" type="discussion">
|
||
<paragraph id="8B8836BC4C6B4530FF38FEBDED926D1D" blockId="9.[150,1437,151,1328]" pageId="9" pageNumber="279">
|
||
<emphasis id="B943EAAE4C6B4530FF38FEBDED596D39" bold="true" box="[199,315,296,320]" inLineHeading="true" pageId="9" pageNumber="279" reason="1">Remarks.</emphasis>
|
||
<typeStatus id="548C881E4C6B4530FEBCFEBDEDD26D39" box="[323,432,296,320]" pageId="9" pageNumber="279" type="paratype">Paratypes</typeStatus>
|
||
similar to the
|
||
<typeStatus id="548C881E4C6B4530FDAAFEB2EED56D46" box="[597,695,295,319]" pageId="9" pageNumber="279" type="holotype">holotype</typeStatus>
|
||
in both qualitative and metric characters: Measurements of some structures are given in
|
||
<tableCitation id="C6B503074C6B4530FE69FED9ED886D1D" box="[406,490,332,356]" captionStart="TABLE 1" captionStartId="3.[151,241,259,283]" captionText="TABLE 1. Length (in µm) and other parameters of selected structures (using, where possible, animals with similar body length), for two specimens of Ramazzottius thulini, two R. oberhaeuseri from the locus typicus, one Swedish Ramazzottius sp. (with smooth cuticle), and three Ramazzottius libycus sp. nov. Due to the variable orientation of the external claws, only the length of the sclerified portion of the main branch and basal portion + secondary branch of each claw have been measured. Pt ratios in italics." pageId="9" pageNumber="279">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32D65374C6B4530FF38FEE5E8C56DFE" box="[199,1191,367,392]" pageId="9" pageNumber="279" type="etymology">
|
||
<paragraph id="8B8836BC4C6B4530FF38FEE5E8C56DFE" blockId="9.[150,1437,151,1328]" box="[199,1191,367,392]" pageId="9" pageNumber="279">
|
||
<emphasis id="B943EAAE4C6B4530FF38FEE5ED2F6DF1" bold="true" box="[199,333,368,392]" inLineHeading="true" pageId="9" pageNumber="279" reason="1">Etymology.</emphasis>
|
||
The name refers to the geographic area (
|
||
<collectingCountry id="F320762C4C6B4530FCE7FEFAEF3E6DFE" box="[792,860,367,391]" name="Libya" pageId="9" pageNumber="279">Libya</collectingCountry>
|
||
) of the
|
||
<emphasis id="B943EAAE4C6B4530FC4CFEE5E8266DFE" box="[947,1092,368,392]" italics="true" pageId="9" pageNumber="279">locus typicus</emphasis>
|
||
(Barce).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32D65374C6B4530FF38FE06E88D6956" pageId="9" pageNumber="279" type="diagnosis">
|
||
<paragraph id="8B8836BC4C6B4530FF38FE06E8066EB5" blockId="9.[150,1437,151,1328]" pageId="9" pageNumber="279">
|
||
<emphasis id="B943EAAE4C6B4530FF38FE06EDAD6DD5" bold="true" box="[199,463,403,428]" inLineHeading="true" pageId="9" pageNumber="279" reason="1">Differential diagnosis.</emphasis>
|
||
For many characters of the adults and the eggs,
|
||
<taxonomicName id="4C374D3F4C6B4530FC07FE00E8866DD5" authority="Pilato & D’Urso & Lisi, 2013" authorityName="Pilato & D’Urso & Lisi" authorityYear="2013" box="[1016,1252,404,428]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="libycus" status="sp. nov.">
|
||
<emphasis id="B943EAAE4C6B4530FC07FE00E8866DD5" box="[1016,1252,404,428]" italics="true" pageId="9" pageNumber="279">Ramazzottius libycus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27057D54C6B4530FB12FE01E9256DD2" box="[1261,1351,404,428]" pageId="9" pageNumber="279" rank="species">
|
||
<emphasis id="B943EAAE4C6B4530FB12FE01E9226DD5" bold="true" box="[1261,1344,404,428]" pageId="9" pageNumber="279">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is very similar to
|
||
<taxonomicName id="4C374D3F4C6B4530FEF7FE2CEDD86DA9" authorityName="Doyere" authorityYear="1840" box="[264,442,440,464]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="oberhaeuseri">
|
||
<emphasis id="B943EAAE4C6B4530FEF7FE2CEDD86DA9" box="[264,442,440,464]" italics="true" pageId="9" pageNumber="279">R. oberhaeuseri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C374D3F4C6B4530FE0FFE2CEE3A6DA9" baseAuthorityName="Pilato" baseAuthorityYear="1970" box="[496,600,440,464]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="thulini">
|
||
<emphasis id="B943EAAE4C6B4530FE0FFE2CEE3A6DA9" box="[496,600,440,464]" italics="true" pageId="9" pageNumber="279">R. thulini</emphasis>
|
||
</taxonomicName>
|
||
. However, it differs from
|
||
<taxonomicName id="4C374D3F4C6B4530FC84FE2CE84F6DA9" authorityName="Doyere" authorityYear="1840" box="[891,1069,440,464]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="oberhaeuseri">
|
||
<emphasis id="B943EAAE4C6B4530FC84FE2CE84F6DA9" box="[891,1069,440,464]" italics="true" pageId="9" pageNumber="279">R. oberhaeuseri</emphasis>
|
||
</taxonomicName>
|
||
in having slightly shorter buccal tube with respect to the body length (% ratio = 10.0
|
||
<emphasis id="B943EAAE4C6B4530FD2CFE48EE836D8D" box="[723,737,477,500]" italics="true" pageId="9" pageNumber="279">–</emphasis>
|
||
<quantity id="4CCF9B594C6B4530FD1FFE49EF4C6D8D" box="[736,814,476,500]" metricMagnitude="-1" metricUnit="m" metricValue="2.5654" pageId="9" pageNumber="279" unit="in" value="10.1">10.1 in</quantity>
|
||
<taxonomicName id="4C374D3F4C6B4530FCCBFE48EFC06D8D" authority="Pilato & D’Urso & Lisi, 2013" authorityName="Pilato & D’Urso & Lisi" authorityYear="2013" box="[820,930,476,500]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="libycus" status="sp. nov.">
|
||
<emphasis id="B943EAAE4C6B4530FCCBFE48EFC06D8D" box="[820,930,476,500]" italics="true" pageId="9" pageNumber="279">R. libycus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27057D54C6B4530FC57FE49EF9C6D8D" box="[936,1022,476,500]" pageId="9" pageNumber="279" rank="species">
|
||
<emphasis id="B943EAAE4C6B4530FC57FE49EF986D8D" bold="true" box="[936,1018,476,500]" pageId="9" pageNumber="279">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
,11.5
|
||
<emphasis id="B943EAAE4C6B4530FBC7FE48E8246D8D" box="[1080,1094,477,500]" italics="true" pageId="9" pageNumber="279">–</emphasis>
|
||
12.0 in
|
||
<taxonomicName id="4C374D3F4C6B4530FB66FE48E9286D8D" authorityName="Doyere" authorityYear="1840" box="[1177,1354,476,500]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="oberhaeuseri">
|
||
<emphasis id="B943EAAE4C6B4530FB66FE48E9286D8D" box="[1177,1354,476,500]" italics="true" pageId="9" pageNumber="279">R. oberhaeuseri</emphasis>
|
||
</taxonomicName>
|
||
); more slender internal claws I
|
||
<emphasis id="B943EAAE4C6B4530FE5FFD94EDCC6E61" box="[416,430,513,536]" italics="true" pageId="9" pageNumber="279">–</emphasis>
|
||
III and anterior claws IV (
|
||
<figureCitation id="130C2A394C6B4530FD26FE6AEF5C6E6E" box="[729,830,511,535]" captionStart="FIGURE 5" captionStartId="7.[151,250,1403,1426]" captionTargetBox="[159,1424,194,1372]" captionTargetId="figure-332@7.[151,1436,167,1400]" captionTargetPageId="7" captionText="FIGURE 5. A, Ramazzottius sp. with smooth cuticle from Sweden. Claws of the second pair of legs. B–D, Ramazzottius libycus sp. nov. B, Cuticle ornamentation (paratype). C, Buccal-pharyngeal apparatus of the holotype. D, Claws of the second pair of legs of the holotype. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981774" httpUri="https://zenodo.org/record/5981774/files/figure.png" pageId="9" pageNumber="279">Figs. 5D</figureCitation>
|
||
,
|
||
<figureCitation id="130C2A394C6B4530FCB4FE6AEFF26E6E" box="[843,912,511,535]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1966]" captionTargetBox="[206,1381,229,1881]" captionTargetId="figure-68@8.[178,1414,193,1919]" captionTargetPageId="8" captionText="FIGURE 6. A–D, Ramazzottius libycus sp. nov. A, claws of the hind legs of the holotype. Arrow indicates a visible lunule. B, Claws of the hind legs of a paratype. Arrows indicate visible lunules. C, D, Eggs. Arrows indicate some conical or truncoconical processes. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981778" httpUri="https://zenodo.org/record/5981778/files/figure.png" pageId="9" pageNumber="279">6A, B</figureCitation>
|
||
and
|
||
<figureCitation id="130C2A394C6B4530FC36FE6AE8696E6E" box="[969,1035,511,536]" captionStart="FIGURE 3" captionStartId="5.[151,250,1537,1560]" captionTargetBox="[157,1432,210,1503]" captionTargetId="figure-237@5.[151,1436,195,1516]" captionTargetPageId="5" captionText="FIGURE 3. A–E, Ramazzotius specimens from the locus typicus of R. oberhaeuseri. A, Buccal-pharyngeal apparatus. B, Claws of the first and second pairs of legs. C, Internal claw of the first pair of legs. D, Internal claw of the third pair of legs. E, Claws of the second pair of legs. Arrow indicates a visible lunule. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981766" httpUri="https://zenodo.org/record/5981766/files/figure.png" pageId="9" pageNumber="279">
|
||
3C
|
||
<emphasis id="B943EAAE4C6B4530FC15FD94EF9A6E61" box="[1002,1016,513,536]" italics="true" pageId="9" pageNumber="279">–</emphasis>
|
||
E
|
||
</figureCitation>
|
||
) with higher values of the
|
||
<emphasis id="B943EAAE4C6B4530FAC0FD94E9376E61" box="[1343,1365,513,536]" italics="true" pageId="9" pageNumber="279">pt</emphasis>
|
||
index (
|
||
<tableCitation id="C6B503074C6B4530FF61FDB1EC906E45" box="[158,242,548,572]" captionStart="TABLE 1" captionStartId="3.[151,241,259,283]" captionText="TABLE 1. Length (in µm) and other parameters of selected structures (using, where possible, animals with similar body length), for two specimens of Ramazzottius thulini, two R. oberhaeuseri from the locus typicus, one Swedish Ramazzottius sp. (with smooth cuticle), and three Ramazzottius libycus sp. nov. Due to the variable orientation of the external claws, only the length of the sclerified portion of the main branch and basal portion + secondary branch of each claw have been measured. Pt ratios in italics." pageId="9" pageNumber="279">Table 1</tableCitation>
|
||
); higher
|
||
<emphasis id="B943EAAE4C6B4530FEA6FDB0ED0D6E45" box="[345,367,549,572]" italics="true" pageId="9" pageNumber="279">pt</emphasis>
|
||
values of the sclerified portion of both the main branches and basal portion + secondary branch in the external claws; the percent ratio between these portions of the claw is the same in the legs I
|
||
<emphasis id="B943EAAE4C6B4530FB23FDDCE8886E19" box="[1244,1258,585,608]" italics="true" pageId="9" pageNumber="279">–</emphasis>
|
||
III but higher in the hind legs (
|
||
<tableCitation id="C6B503074C6B4530FEC6FDF9EDEC6EFD" box="[313,398,620,644]" captionStart="TABLE 1" captionStartId="3.[151,241,259,283]" captionText="TABLE 1. Length (in µm) and other parameters of selected structures (using, where possible, animals with similar body length), for two specimens of Ramazzottius thulini, two R. oberhaeuseri from the locus typicus, one Swedish Ramazzottius sp. (with smooth cuticle), and three Ramazzottius libycus sp. nov. Due to the variable orientation of the external claws, only the length of the sclerified portion of the main branch and basal portion + secondary branch of each claw have been measured. Pt ratios in italics." pageId="9" pageNumber="279">Table 1</tableCitation>
|
||
). The eggs have a lower number of processes in the hemisphere (
|
||
<quantity id="4CCF9B594C6B4530FB81FDF9E8D86EFD" box="[1150,1210,620,644]" metricMagnitude="0" metricUnit="m" metricValue="1.2192" pageId="9" pageNumber="279" unit="in" value="48.0">48 in</quantity>
|
||
<taxonomicName id="4C374D3F4C6B4530FB3DFDF8E9536EFD" authority="Pilato & D’Urso & Lisi, 2013" authorityName="Pilato & D’Urso & Lisi" authorityYear="2013" box="[1218,1329,620,644]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="libycus" status="sp. nov.">
|
||
<emphasis id="B943EAAE4C6B4530FB3DFDF8E9536EFD" box="[1218,1329,620,644]" italics="true" pageId="9" pageNumber="279">R. libycus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27057D54C6B4530FAC4FDF9E9F76EFD" box="[1339,1429,620,644]" pageId="9" pageNumber="279" rank="species">
|
||
<emphasis id="B943EAAE4C6B4530FAC4FDF9E9EF6EFD" bold="true" box="[1339,1421,620,644]" pageId="9" pageNumber="279">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
,
|
||
<quantity id="4CCF9B594C6B4530FF68FD1AEC986EDE" box="[151,250,655,680]" metricMagnitude="0" metricUnit="m" metricValue="1.4859" metricValueMax="1.524" metricValueMin="1.4478" pageId="9" pageNumber="279" unit="in" value="58.5" valueMax="60.0" valueMin="57.0">
|
||
57
|
||
<emphasis id="B943EAAE4C6B4530FF4CFD04ECA36ED1" box="[179,193,657,680]" italics="true" pageId="9" pageNumber="279">–</emphasis>
|
||
60 in
|
||
</quantity>
|
||
<taxonomicName id="4C374D3F4C6B4530FEFDFD04EDD76ED1" authorityName="Doyere" authorityYear="1840" box="[258,437,656,680]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="oberhaeuseri">
|
||
<emphasis id="B943EAAE4C6B4530FEFDFD04EDD76ED1" box="[258,437,656,680]" italics="true" pageId="9" pageNumber="279">R. oberhaeuseri</emphasis>
|
||
</taxonomicName>
|
||
), but the number of eggs we examined is too low for us to know the range of individual variability of this character and therefore whether or not this is a significant difference.
|
||
</paragraph>
|
||
<paragraph id="8B8836BC4C6B4530FF38FD4CEF5A6805" blockId="9.[150,1437,151,1328]" pageId="9" pageNumber="279">
|
||
<taxonomicName id="4C374D3F4C6B4530FF38FD4CED556E89" authority="Pilato & D’Urso & Lisi, 2013" authorityName="Pilato & D’Urso & Lisi" authorityYear="2013" box="[199,311,728,752]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="libycus" status="sp. nov.">
|
||
<emphasis id="B943EAAE4C6B4530FF38FD4CED556E89" box="[199,311,728,752]" italics="true" pageId="9" pageNumber="279">R. libycus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27057D54C6B4530FEBFFD4DEDFB6E89" box="[320,409,728,752]" pageId="9" pageNumber="279" rank="species">
|
||
<emphasis id="B943EAAE4C6B4530FEBFFD4DEDF66E89" bold="true" box="[320,404,728,752]" pageId="9" pageNumber="279">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, differs from
|
||
<taxonomicName id="4C374D3F4C6B4530FDC5FD4CEEC66E89" baseAuthorityName="Pilato" baseAuthorityYear="1970" box="[570,676,728,752]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="thulini">
|
||
<emphasis id="B943EAAE4C6B4530FDC5FD4CEEC66E89" box="[570,676,728,752]" italics="true" pageId="9" pageNumber="279">R. thulini</emphasis>
|
||
</taxonomicName>
|
||
in having polygonal tubercles also in the anterior extremity of the body, slightly shorter buccal tube with respect to the body length (% ratio = 10.0
|
||
<emphasis id="B943EAAE4C6B4530FBE1FD68E84E6F6D" box="[1054,1068,765,788]" italics="true" pageId="9" pageNumber="279">–</emphasis>
|
||
<quantity id="4CCF9B594C6B4530FBD3FD69E8186F6D" box="[1068,1146,764,788]" metricMagnitude="-1" metricUnit="m" metricValue="2.5654" pageId="9" pageNumber="279" unit="in" value="10.1">10.1 in</quantity>
|
||
<taxonomicName id="4C374D3F4C6B4530FB7EFD68E88D6F6D" authority="Pilato & D’Urso & Lisi, 2013" authorityName="Pilato & D’Urso & Lisi" authorityYear="2013" box="[1153,1263,764,788]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="libycus" status="sp. nov.">
|
||
<emphasis id="B943EAAE4C6B4530FB7EFD68E88D6F6D" box="[1153,1263,764,788]" italics="true" pageId="9" pageNumber="279">R. libycus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27057D54C6B4530FB09FD69E92D6F6A" box="[1270,1359,764,788]" pageId="9" pageNumber="279" rank="species">
|
||
<emphasis id="B943EAAE4C6B4530FB09FD69E92B6F6D" bold="true" box="[1270,1353,764,788]" pageId="9" pageNumber="279">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, 11.6
|
||
<emphasis id="B943EAAE4C6B4530FA71FD68E9FE6F6D" box="[1422,1436,765,788]" italics="true" pageId="9" pageNumber="279">–</emphasis>
|
||
12.0 in
|
||
<taxonomicName id="4C374D3F4C6B4530FF13FCB4ED366F41" baseAuthorityName="Pilato" baseAuthorityYear="1970" box="[236,340,800,824]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="thulini">
|
||
<emphasis id="B943EAAE4C6B4530FF13FCB4ED366F41" box="[236,340,800,824]" italics="true" pageId="9" pageNumber="279">R. thulini</emphasis>
|
||
</taxonomicName>
|
||
); stylet supports inserted on the buccal tube in a slightly more caudal position (
|
||
<emphasis id="B943EAAE4C6B4530FB31FCB4E8866F41" box="[1230,1252,801,824]" italics="true" pageId="9" pageNumber="279">pt</emphasis>
|
||
=
|
||
<quantity id="4CCF9B594C6B4530FAFEFC8AE9FE6F4E" box="[1281,1436,799,823]" metricMagnitude="0" metricUnit="m" metricValue="1.54051" metricValueMax="1.56718" metricValueMin="1.51384" pageId="9" pageNumber="279" unit="in" value="60.65" valueMax="61.7" valueMin="59.6">59.6 – 61.7 in</quantity>
|
||
<taxonomicName id="4C374D3F4C6B4530FF68FCD0ED676F25" authority="Pilato & D’Urso & Lisi, 2013" authorityName="Pilato & D’Urso & Lisi" authorityYear="2013" box="[151,261,836,860]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="libycus" status="sp. nov.">
|
||
<emphasis id="B943EAAE4C6B4530FF68FCD0ED676F25" box="[151,261,836,860]" italics="true" pageId="9" pageNumber="279">R. libycus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27057D54C6B4530FEF3FCD1ED066F25" box="[268,356,836,860]" pageId="9" pageNumber="279" rank="species">
|
||
<emphasis id="B943EAAE4C6B4530FEF3FCD1ED3F6F25" bold="true" box="[268,349,836,860]" pageId="9" pageNumber="279">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
,
|
||
<quantity id="4CCF9B594C6B4530FE8DFCD1EE6F6F25" box="[370,525,836,860]" metricMagnitude="0" metricUnit="m" metricValue="1.46304" metricValueMax="1.47066" metricValueMin="1.45542" pageId="9" pageNumber="279" unit="in" value="57.599999999999994" valueMax="57.9" valueMin="57.3">57.3 – 57.9 in</quantity>
|
||
<taxonomicName id="4C374D3F4C6B4530FDECFCD0EE196F25" baseAuthorityName="Pilato" baseAuthorityYear="1970" box="[531,635,836,860]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="thulini">
|
||
<emphasis id="B943EAAE4C6B4530FDECFCD0EE196F25" box="[531,635,836,860]" italics="true" pageId="9" pageNumber="279">R. thulini</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<tableCitation id="C6B503074C6B4530FD75FCD1EEB96F25" box="[650,731,836,860]" captionStart="TABLE 1" captionStartId="3.[151,241,259,283]" captionText="TABLE 1. Length (in µm) and other parameters of selected structures (using, where possible, animals with similar body length), for two specimens of Ramazzottius thulini, two R. oberhaeuseri from the locus typicus, one Swedish Ramazzottius sp. (with smooth cuticle), and three Ramazzottius libycus sp. nov. Due to the variable orientation of the external claws, only the length of the sclerified portion of the main branch and basal portion + secondary branch of each claw have been measured. Pt ratios in italics." pageId="9" pageNumber="279">Table 1</tableCitation>
|
||
); different shape of the internal claws: the main branches of
|
||
<taxonomicName id="4C374D3F4C6B4530FA7BFCD0EC846FF9" authority="Pilato & D’Urso & Lisi, 2013" authorityName="Pilato & D’Urso & Lisi" authorityYear="2013" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="libycus" status="sp. nov.">
|
||
<emphasis id="B943EAAE4C6B4530FA7BFCD0EC846FF9" italics="true" pageId="9" pageNumber="279">R. libycus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27057D54C6B4530FF12FCFDED276F06" box="[237,325,872,896]" pageId="9" pageNumber="279" rank="species">
|
||
<emphasis id="B943EAAE4C6B4530FF12FCFDED5D6FF9" bold="true" box="[237,319,872,896]" pageId="9" pageNumber="279">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
are longer and more slender, and the secondary branches are longer and taper gradually, whereas in
|
||
<taxonomicName id="4C374D3F4C6B4530FF69FC18ED606FDD" baseAuthorityName="Pilato" baseAuthorityYear="1970" box="[150,258,908,932]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="thulini">
|
||
<emphasis id="B943EAAE4C6B4530FF69FC18ED606FDD" box="[150,258,908,932]" italics="true" pageId="9" pageNumber="279">R. thulini</emphasis>
|
||
</taxonomicName>
|
||
the secondary branches taper more abruptly (
|
||
<figureCitation id="130C2A394C6B4530FCE4FC19EFE06FDD" box="[795,898,908,932]" captionStart="FIGURE 5" captionStartId="7.[151,250,1403,1426]" captionTargetBox="[159,1424,194,1372]" captionTargetId="figure-332@7.[151,1436,167,1400]" captionTargetPageId="7" captionText="FIGURE 5. A, Ramazzottius sp. with smooth cuticle from Sweden. Claws of the second pair of legs. B–D, Ramazzottius libycus sp. nov. B, Cuticle ornamentation (paratype). C, Buccal-pharyngeal apparatus of the holotype. D, Claws of the second pair of legs of the holotype. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981774" httpUri="https://zenodo.org/record/5981774/files/figure.png" pageId="9" pageNumber="279">Figs. 5D</figureCitation>
|
||
,
|
||
<figureCitation id="130C2A394C6B4530FC6EFC19EFBA6FDA" box="[913,984,908,932]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1966]" captionTargetBox="[206,1381,229,1881]" captionTargetId="figure-68@8.[178,1414,193,1919]" captionTargetPageId="8" captionText="FIGURE 6. A–D, Ramazzottius libycus sp. nov. A, claws of the hind legs of the holotype. Arrow indicates a visible lunule. B, Claws of the hind legs of a paratype. Arrows indicate visible lunules. C, D, Eggs. Arrows indicate some conical or truncoconical processes. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981778" httpUri="https://zenodo.org/record/5981778/files/figure.png" pageId="9" pageNumber="279">6A, B</figureCitation>
|
||
and
|
||
<figureCitation id="130C2A394C6B4530FBEAFC19E8E36FDD" box="[1045,1153,908,932]" captionStart="FIGURE 2" captionStartId="4.[151,250,1575,1598]" captionTargetBox="[157,1427,223,1532]" captionTargetId="figure-150@4.[151,1436,195,1551]" captionTargetPageId="4" captionText="FIGURE 2. A–E, Ramazzottius thulini (holotype). A, Buccal-pharyngeal apparatus. B, Claws of the second and third pairs of legs. C, Claws of the second pair of legs. D, Claws of the third pair of legs. E, Claws of the hind legs. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981762" httpUri="https://zenodo.org/record/5981762/files/figure.png" pageId="9" pageNumber="279">2C, D, E</figureCitation>
|
||
); higher
|
||
<emphasis id="B943EAAE4C6B4530FB14FC18E9636FDD" box="[1259,1281,909,932]" italics="true" pageId="9" pageNumber="279">pt</emphasis>
|
||
index values relative to internal claws I
|
||
<emphasis id="B943EAAE4C6B4530FE36FC24EDB56FB1" box="[457,471,945,968]" italics="true" pageId="9" pageNumber="279">–</emphasis>
|
||
III and anterior claws IV (
|
||
<tableCitation id="C6B503074C6B4530FCF3FC3AEF016FBE" box="[780,867,943,967]" captionStart="TABLE 1" captionStartId="3.[151,241,259,283]" captionText="TABLE 1. Length (in µm) and other parameters of selected structures (using, where possible, animals with similar body length), for two specimens of Ramazzottius thulini, two R. oberhaeuseri from the locus typicus, one Swedish Ramazzottius sp. (with smooth cuticle), and three Ramazzottius libycus sp. nov. Due to the variable orientation of the external claws, only the length of the sclerified portion of the main branch and basal portion + secondary branch of each claw have been measured. Pt ratios in italics." pageId="9" pageNumber="279">Table 1</tableCitation>
|
||
); in the external claws the
|
||
<emphasis id="B943EAAE4C6B4530FB56FC24E8DD6FB1" box="[1193,1215,945,968]" italics="true" pageId="9" pageNumber="279">pt</emphasis>
|
||
values of both the sclerified portion of the main branches and the basal portion + secondary branches length are higher; in legs I
|
||
<emphasis id="B943EAAE4C6B4530FA8DFC40E9E26F95" box="[1394,1408,981,1004]" italics="true" pageId="9" pageNumber="279">–</emphasis>
|
||
III the percent ratio between the length of these structures is the same but it is clearly higher in the hind legs (
|
||
<tableCitation id="C6B503074C6B4530FAC8FC62E9E96876" box="[1335,1419,1015,1039]" captionStart="TABLE 1" captionStartId="3.[151,241,259,283]" captionText="TABLE 1. Length (in µm) and other parameters of selected structures (using, where possible, animals with similar body length), for two specimens of Ramazzottius thulini, two R. oberhaeuseri from the locus typicus, one Swedish Ramazzottius sp. (with smooth cuticle), and three Ramazzottius libycus sp. nov. Due to the variable orientation of the external claws, only the length of the sclerified portion of the main branch and basal portion + secondary branch of each claw have been measured. Pt ratios in italics." pageId="9" pageNumber="279">Table 1</tableCitation>
|
||
). The eggs have a lower number of processes in the hemisphere (about
|
||
<quantity id="4CCF9B594C6B4530FC5FFB89EFBB684D" box="[928,985,1052,1076]" metricMagnitude="0" metricUnit="m" metricValue="1.2192" pageId="9" pageNumber="279" unit="in" value="48.0">48 in</quantity>
|
||
<taxonomicName id="4C374D3F4C6B4530FC20FB88E82F684D" authority="Pilato & D’Urso & Lisi, 2013" authorityName="Pilato & D’Urso & Lisi" authorityYear="2013" box="[991,1101,1052,1076]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="libycus" status="sp. nov.">
|
||
<emphasis id="B943EAAE4C6B4530FC20FB88E82F684D" box="[991,1101,1052,1076]" italics="true" pageId="9" pageNumber="279">R. libycus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27057D54C6B4530FBABFB89E8CF684D" box="[1108,1197,1052,1076]" pageId="9" pageNumber="279" rank="species">
|
||
<emphasis id="B943EAAE4C6B4530FBABFB89E8C4684D" bold="true" box="[1108,1190,1052,1076]" pageId="9" pageNumber="279">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
,
|
||
<quantity id="4CCF9B594C6B4530FB45FB89E97F684D" box="[1210,1309,1052,1076]" metricMagnitude="0" metricUnit="m" metricValue="1.4859" metricValueMax="1.524" metricValueMin="1.4478" pageId="9" pageNumber="279" unit="in" value="58.5" valueMax="60.0" valueMin="57.0">
|
||
57
|
||
<emphasis id="B943EAAE4C6B4530FB29FB88E886684D" box="[1238,1252,1053,1076]" italics="true" pageId="9" pageNumber="279">–</emphasis>
|
||
60 in
|
||
</quantity>
|
||
<taxonomicName id="4C374D3F4C6B4530FADBFB88E9EE684D" baseAuthorityName="Pilato" baseAuthorityYear="1970" box="[1316,1420,1052,1076]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="thulini">
|
||
<emphasis id="B943EAAE4C6B4530FADBFB88E9EE684D" box="[1316,1420,1052,1076]" italics="true" pageId="9" pageNumber="279">R. thulini</emphasis>
|
||
</taxonomicName>
|
||
), but the limited number of eggs we examined for
|
||
<taxonomicName id="4C374D3F4C6B4530FD35FBD4EF586821" authority="Pilato & D’Urso & Lisi, 2013" authorityName="Pilato & D’Urso & Lisi" authorityYear="2013" box="[714,826,1088,1112]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="libycus" status="sp. nov.">
|
||
<emphasis id="B943EAAE4C6B4530FD35FBD4EF586821" box="[714,826,1088,1112]" italics="true" pageId="9" pageNumber="279">R. libycus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27057D54C6B4530FCBCFBD5EFFF6821" box="[835,925,1088,1112]" pageId="9" pageNumber="279" rank="species">
|
||
<emphasis id="B943EAAE4C6B4530FCBCFBD5EFF56821" bold="true" box="[835,919,1088,1112]" pageId="9" pageNumber="279">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
has not provided the range of the individual variability and we cannot say this is a significant difference.
|
||
</paragraph>
|
||
<paragraph id="8B8836BC4C6B4530FF38FB1DE88D6956" blockId="9.[150,1437,151,1328]" pageId="9" pageNumber="279">
|
||
<taxonomicName id="4C374D3F4C6B4530FF38FB1DEDD168D9" authority="Pilato & D’Urso & Lisi, 2013" authorityName="Pilato & D’Urso & Lisi" authorityYear="2013" box="[199,435,1160,1184]" class="Eutardigrada" family="Hypsibiidae" genus="Ramazzottius" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="libycus" status="sp. nov.">
|
||
<emphasis id="B943EAAE4C6B4530FF38FB1DEDD168D9" box="[199,435,1160,1184]" italics="true" pageId="9" pageNumber="279">Ramazzottius libycus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27057D54C6B4530FE44FB1DEE7668E6" box="[443,532,1160,1184]" pageId="9" pageNumber="279" rank="species">
|
||
<emphasis id="B943EAAE4C6B4530FE44FB1DEE6C68D9" bold="true" box="[443,526,1160,1184]" pageId="9" pageNumber="279">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is clearly different from the Swedish smooth specimens, which
|
||
<bibRefCitation id="EFA64B4D4C6B4530FB16FB1DED5C68BD" author="Durante Pasa, M. V. & Maucci, W." pageId="9" pageNumber="279" pagination="47 - 85" refId="ref5052" refString="Durante Pasa, M. V. & Maucci, W. (1979) Moss Tardigrada from the Scandinavian Peninsula. Zeszyty Naukowe Uniwersytetu Jagiello n skiego, Prace Zoologiczne, 25, 47 - 85." type="journal article" year="1979">Durante Pasa & Maucci (1979)</bibRefCitation>
|
||
attributed to
|
||
<taxonomicName id="4C374D3F4C6B4530FE26FB38EEEF68BD" authorityName="Doyere" authorityYear="1840" box="[473,653,1196,1220]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="279" phylum="Tardigrada" rank="species" species="oberhaeuseri">
|
||
<emphasis id="B943EAAE4C6B4530FE26FB38EEEF68BD" box="[473,653,1196,1220]" italics="true" pageId="9" pageNumber="279">R. oberhaeuseri</emphasis>
|
||
</taxonomicName>
|
||
, in having ornamented cuticle; higher
|
||
<emphasis id="B943EAAE4C6B4530FBB6FB38E83D68BD" box="[1097,1119,1197,1220]" italics="true" pageId="9" pageNumber="279">pt</emphasis>
|
||
index values relative to the internal claws (
|
||
<tableCitation id="C6B503074C6B4530FEBEFB5AEDF7689E" box="[321,405,1231,1255]" captionStart="TABLE 1" captionStartId="3.[151,241,259,283]" captionText="TABLE 1. Length (in µm) and other parameters of selected structures (using, where possible, animals with similar body length), for two specimens of Ramazzottius thulini, two R. oberhaeuseri from the locus typicus, one Swedish Ramazzottius sp. (with smooth cuticle), and three Ramazzottius libycus sp. nov. Due to the variable orientation of the external claws, only the length of the sclerified portion of the main branch and basal portion + secondary branch of each claw have been measured. Pt ratios in italics." pageId="9" pageNumber="279">Table 1</tableCitation>
|
||
); external claws with larger
|
||
<emphasis id="B943EAAE4C6B4530FD2AFB44EE896891" box="[725,747,1233,1256]" italics="true" pageId="9" pageNumber="279">pt</emphasis>
|
||
values relative to the basal portion + secondary branch (
|
||
<figureCitation id="130C2A394C6B4530FA9CFB5AECD86975" captionStart="FIGURE 5" captionStartId="7.[151,250,1403,1426]" captionTargetBox="[159,1424,194,1372]" captionTargetId="figure-332@7.[151,1436,167,1400]" captionTargetPageId="7" captionText="FIGURE 5. A, Ramazzottius sp. with smooth cuticle from Sweden. Claws of the second pair of legs. B–D, Ramazzottius libycus sp. nov. B, Cuticle ornamentation (paratype). C, Buccal-pharyngeal apparatus of the holotype. D, Claws of the second pair of legs of the holotype. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981774" httpUri="https://zenodo.org/record/5981774/files/figure.png" pageId="9" pageNumber="279">Figs. 5C</figureCitation>
|
||
,
|
||
<figureCitation id="130C2A394C6B4530FF39FB61ED6B6972" box="[198,265,1268,1292]" captionStart="FIGURE 6" captionStartId="8.[151,250,1943,1966]" captionTargetBox="[206,1381,229,1881]" captionTargetId="figure-68@8.[178,1414,193,1919]" captionTargetPageId="8" captionText="FIGURE 6. A–D, Ramazzottius libycus sp. nov. A, claws of the hind legs of the holotype. Arrow indicates a visible lunule. B, Claws of the hind legs of a paratype. Arrows indicate visible lunules. C, D, Eggs. Arrows indicate some conical or truncoconical processes. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981778" httpUri="https://zenodo.org/record/5981778/files/figure.png" pageId="9" pageNumber="279">6A, B</figureCitation>
|
||
and
|
||
<figureCitation id="130C2A394C6B4530FEBFFB61ED076975" box="[320,357,1268,1292]" captionStart="FIGURE 5" captionStartId="7.[151,250,1403,1426]" captionTargetBox="[159,1424,194,1372]" captionTargetId="figure-332@7.[151,1436,167,1400]" captionTargetPageId="7" captionText="FIGURE 5. A, Ramazzottius sp. with smooth cuticle from Sweden. Claws of the second pair of legs. B–D, Ramazzottius libycus sp. nov. B, Cuticle ornamentation (paratype). C, Buccal-pharyngeal apparatus of the holotype. D, Claws of the second pair of legs of the holotype. Scale bar = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5981774" httpUri="https://zenodo.org/record/5981774/files/figure.png" pageId="9" pageNumber="279">5A</figureCitation>
|
||
;
|
||
<tableCitation id="C6B503074C6B4530FE8DFB61EDA16975" box="[370,451,1268,1292]" captionStart="TABLE 1" captionStartId="3.[151,241,259,283]" captionText="TABLE 1. Length (in µm) and other parameters of selected structures (using, where possible, animals with similar body length), for two specimens of Ramazzottius thulini, two R. oberhaeuseri from the locus typicus, one Swedish Ramazzottius sp. (with smooth cuticle), and three Ramazzottius libycus sp. nov. Due to the variable orientation of the external claws, only the length of the sclerified portion of the main branch and basal portion + secondary branch of each claw have been measured. Pt ratios in italics." pageId="9" pageNumber="279">Table 1</tableCitation>
|
||
) and different percent ratio between the main branch and the basal portion + secondary branch (
|
||
<tableCitation id="C6B503074C6B4530FF0EFA82ED276956" box="[241,325,1303,1327]" captionStart="TABLE 1" captionStartId="3.[151,241,259,283]" captionText="TABLE 1. Length (in µm) and other parameters of selected structures (using, where possible, animals with similar body length), for two specimens of Ramazzottius thulini, two R. oberhaeuseri from the locus typicus, one Swedish Ramazzottius sp. (with smooth cuticle), and three Ramazzottius libycus sp. nov. Due to the variable orientation of the external claws, only the length of the sclerified portion of the main branch and basal portion + secondary branch of each claw have been measured. Pt ratios in italics." pageId="9" pageNumber="279">Table 1</tableCitation>
|
||
). We have no eggs produced by the specimens with smooth cuticle for comparison.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |