151 lines
19 KiB
XML
151 lines
19 KiB
XML
<document id="44A37C65C28C5B9280F9FC98B47D685D" ID-DOI="10.1206/0003-0090(2005)291<0001:SBOTGA>2.0.CO;2" ID-ISSN="0003-0090" ID-Zenodo-Dep="5362016" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630237249567" checkinUser="felipe" docAuthor="MILLER, KELLY B. & WHEELER, QUENTIN D." docDate="2005" docId="038387B33721B969FD495445FD300817" docLanguage="en" docName="B291.pdf" docOrigin="Bulletin of the American Museum of Natural History 2005 (291)" docSource="http://www.bioone.org/perlserv/?request=get-abstract&doi=10.1206%2F0003-0090(2005)291%3C0001%3ASBOTGA%3E2.0.CO%3B2" docStyle="DocumentStyle:2B34016C1EBFE58B07368F9517618743.3:BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleId="2B34016C1EBFE58B07368F9517618743" docStyleName="BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleVersion="3" docTitle="Agathidium tribulograndum Miller and Wheeler 2005, new species" docType="treatment" docVersion="3" lastPageNumber="110" masterDocId="FFBAFFCB374DB904FF9F5642FFD3095A" masterDocTitle="Slime-Mold Beetles Of The Genus Agathidium Panzer In North And Central America, Part Ii. Coleoptera: Leiodidae" masterLastPageNumber="167" masterPageNumber="1" pageNumber="109" updateTime="1699194605127" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="311A2FA83DE2EA01762CD2027B0640B6">Slime-Mold Beetles Of The Genus Agathidium Panzer In North And Central America, Part Ii. Coleoptera: Leiodidae</mods:title>
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<mods:namePart id="438F800993D227EDD0E8EC0C3B65FE9D">MILLER, KELLY B.</mods:namePart>
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<mods:namePart id="DADEDBF7CEA808E0DB36715CA246C230">WHEELER, QUENTIN D.</mods:namePart>
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<mods:typeOfResource id="55698EBBEEC56D80678ABDCAD223E09D">text</mods:typeOfResource>
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<mods:title id="9D7093F3C3C96071577AF6202B629767">Bulletin of the American Museum of Natural History</mods:title>
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<mods:part id="9989C7D7B1034C190A81EAE92C33C350">
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<mods:date id="03CF3C7CA5E3CD6EBC1608B8D92FB0B5">2005</mods:date>
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<mods:number id="EF8AF2B2293B4B8CE93BDE6C889C5B56">2005-03-24</mods:number>
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<mods:number id="76B8D78D9D2C4E8E8C4A990913B4DA03">2005</mods:number>
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<mods:number id="28C69E1448AF67B2E83E9C4C3BACE592">291</mods:number>
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<mods:classification id="04188C7E673F23B24363872E0207F897">journal article</mods:classification>
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<mods:identifier id="F74965E0B9D3E47C53D24F0BA1B66ED9" type="DOI">10.1206/0003-0090(2005)291<0001:SBOTGA>2.0.CO;2</mods:identifier>
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<treatment id="038387B33721B969FD495445FD300817" LSID="urn:lsid:plazi:treatment:038387B33721B969FD495445FD300817" httpUri="http://treatment.plazi.org/id/038387B33721B969FD495445FD300817" lastPageId="109" lastPageNumber="110" pageId="108" pageNumber="109">
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<subSubSection id="C330652E3721B968FD495445FB500B03" pageId="108" pageNumber="109" type="nomenclature">
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<paragraph id="8B9536A53721B968FD495445FB500B03" blockId="108.[726,1202,519,601]" pageId="108" pageNumber="109">
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<heading id="D0DD81C93721B968FD495445FB500B03" centered="true" fontSize="9" level="2" pageId="108" pageNumber="109" reason="2">
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<taxonomicName id="4C2A4D263721B968FD495445FC700B66" ID-CoL="662RL" authority="Miller and Wheeler" authorityName="Miller and Wheeler" authorityYear="2005" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="tribulograndum" status="sp. nov.">
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<emphasis id="B95EEAB73721B968FD495445FBFE0B45" bold="true" box="[726,1069,519,543]" italics="true" pageId="108" pageNumber="109">Agathidium tribulograndum</emphasis>
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Miller and Wheeler
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26D57CC3721B968FC2E5467FBEC0B66" box="[945,1087,549,572]" pageId="108" pageNumber="109" rank="species">new species</taxonomicNameLabel>
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<figureCitation id="13112A203721B968FC9A5400FC470B03" box="[773,916,578,601]" captionStart="Figs" captionStartId="72.[150,198,991,1012]" captionTargetBox="[323,1037,187,970]" captionTargetId="figure-16@72.[317,1037,187,970]" captionTargetPageId="72" captionText="Figs. 113–117. Agathidium oniscoidesgroup species, sterna and legs: 113, A. gomezae, metasternum, arrow indicating medial metasternal lobe. 114, A. skoliosternum, mesosternum, arrow indicating anterior excavation of mesosternal margin. 115, A. tribulosum, Ƌ left proleg, arrow indicating spine on Ƌ left protrochanter. 116, A. tribulograndum, Ƌ left proleg, arrow indicating spine on Ƌ left procoxa. 117, A. framea, Ƌ left metaleg, arrow indicating broad lobe along anterior margin of Ƌ left metacoxa." figureDoi="http://doi.org/10.5281/zenodo.5362137" httpUri="https://zenodo.org/record/5362137/files/figure.png" pageId="108" pageNumber="109">Figures 116</figureCitation>
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,
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<figureCitation id="13112A203721B968FC3C5400FC020B03" box="[931,977,578,601]" captionStart="Figs" captionStartId="73.[149,197,1560,1581]" captionTargetBox="[181,1166,187,1535]" captionTargetId="figure-13@73.[181,1168,187,1539]" captionTargetPageId="73" captionText="Figs. 118–144. Agathidium concinnum and A. oniscoidesgroup species, left male metafemur, ventral aspect: 118, A. concinnum. 119, A. kimberlae. 120, A. vaderi. 121, A. bituberculum. 122, A. oedema. 123, A. gomezae. 124, A. hidalgoense. 125, A. skoliosternum. 126, A. erythromelas. 127, A. rhamphastes. 128, A. megoniscoides. 129, A. grumum. 130, A. triangularum. 131, A. lobosternum. 132, A. potosii. 133, A. popocatepetlae. 134, A. hyle. 135, A. stenomma. 136, A. tribulosum. 137, A. tribulograndum. 138, A. invisitatum. 139, A. multidentatum. 140, A. sejunctum. 141, A. grandidentatum. 142, A. andersoni. 143, A. disgregum. 144, A. oaxacaense. Bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5362145" httpUri="https://zenodo.org/record/5362145/files/figure.png" pageId="108" pageNumber="109">137</figureCitation>
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,
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<figureCitation id="13112A203721B968FC7F5400FB9A0B03" box="[992,1097,578,601]" captionStart="Figs" captionStartId="79.[149,197,1615,1636]" captionTargetBox="[153,1201,191,1587]" captionTargetId="figure-13@79.[148,1201,187,1594]" captionTargetPageId="79" captionText="Figs. 232–260. Agathidium oniscoidesgroup species, aedeagus: 232–235, A. tribulosum: 232, ventral; 233, median lobe, apex, ventral; 234, median lobe, lateral; 235, left lateral lobe, lateral. 236–239, A. tribulograndum: 236, ventral; 237, median lobe, apex, ventral; 238, median lobe, lateral; 239, left lateral lobe, lateral. 240–243, A. invisitatum: 240, ventral; 241, median lobe, apex, ventral; 242, median lobe, lateral; 243, left lateral lobe, lateral. 244–247, A. multidentatum: 244, ventral; 245, median lobe," figureDoi="http://doi.org/10.5281/zenodo.5362159" httpUri="https://zenodo.org/record/5362159/files/figure.png" pageId="108" pageNumber="109">236–239</figureCitation>
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,
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<figureCitation id="13112A203721B968FBC85400FB500B03" box="[1111,1155,578,601]" captionStart="Figs" captionStartId="103.[149,197,1634,1655]" captionTargetBox="[170,1180,187,1612]" captionTargetId="figure-14@103.[169,1180,187,1613]" captionTargetPageId="103" captionText="Figs. 363, 364. Geographic distribution of Agathidium oniscoidesgroup species: 363, A. popocatepetlae = -; A. potosii = •; A. stenomma = *; A. tribulosum = ^. 364, A. hyle = *; A. grandidentatum = O; A. multidentatum = •; A. sejunctum = ^; A. tribulograndum = •." figureDoi="http://doi.org/10.5281/zenodo.5362176" httpUri="https://zenodo.org/record/5362176/files/figure.png" pageId="108" pageNumber="109">364</figureCitation>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C330652E3721B968FD465434FCB20ACF" pageId="108" pageNumber="109" type="materials_examined">
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<paragraph id="8B9536A53721B968FD465434FC9F0A00" blockId="108.[700,1229,630,1741]" pageId="108" pageNumber="109">
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TYPE
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<collectionCode id="ED3BAE603721B968FCBD5434FC440BD6" box="[802,919,630,653]" pageId="108" pageNumber="109">MATERIAL</collectionCode>
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:
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<typeStatus id="549188073721B968FC325434FBCF0BD7" box="[941,1052,630,653]" pageId="108" pageNumber="109" type="holotype">Holotype</typeStatus>
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, Ƌ in
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<collectionCode id="ED3BAE603721B968FBE75434FB1F0BD7" box="[1144,1228,630,653]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="108" pageNumber="109" type="Museum">FMNH</collectionCode>
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labeled ‘‘
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<collectingCountry id="F33D76353721B968FCB554D1FC740BF0" box="[810,935,659,682]" name="Mexico" pageId="108" pageNumber="109">MEXICO</collectingCountry>
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:
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<collectingRegion id="49EEF8473721B968FC2654D1FBC40BF0" box="[953,1047,659,682]" country="Mexico" name="Hidalgo" pageId="108" pageNumber="109">Hidalgo</collectingRegion>
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<quantity id="4CD29B403721B968FBBA54D1FB7A0BF0" box="[1061,1193,659,682]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" metricValueMax="5.632704" metricValueMin="4.02336" pageId="108" pageNumber="109" unit="mi" value="3.0" valueMax="3.5" valueMin="2.5">2.5–3.5 mi</quantity>
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N Tlanchinol,
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<quantity id="4CD29B403721B968FCC754F2FC350B9D" box="[856,998,688,711]" metricMagnitude="2" metricUnit="m" metricValue="8.001" metricValueMax="15.8496" metricValueMin="0.1524" pageId="108" pageNumber="109" unit="ft" value="2625.0" valueMax="5200.0" valueMin="50.0">50–5200 ft.</quantity>
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vii.6–11.1973/fungusy wood chips A. Newton/
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<typeStatus id="549188073721B968FBAC548CFB1E0BBF" box="[1075,1229,718,741]" pageId="108" pageNumber="109" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="4C2A4D263721B968FD2354A9FCB50A45" authority="Miller and Wheeler, 2003" authorityName="Miller and Wheeler" authorityYear="2003" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="tribulograndum">
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<emphasis id="B95EEAB73721B968FD2354A9FBFA0A58" box="[700,1065,747,770]" italics="true" pageId="108" pageNumber="109">Agathidium tribulograndum</emphasis>
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Miller and Wheeler, 2003
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</taxonomicName>
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[red label with black line bor der]’’. Only a single specimen is known of this species.
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</paragraph>
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<paragraph id="8B9536A53721B968FD465522FCB20ACF" blockId="108.[700,1229,630,1741]" pageId="108" pageNumber="109">
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<collectionCode id="ED3BAE603721B968FD465522FC5C0A2D" box="[729,911,864,887]" pageId="108" pageNumber="109">
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<typeStatus id="549188073721B968FD465522FCC00A2D" box="[729,787,864,887]" pageId="108" pageNumber="109">TYPE</typeStatus>
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LOCALITY
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</collectionCode>
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:
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<collectingCountry id="F33D76353721B968FC3D5522FC2D0A2D" box="[930,1022,864,887]" name="Mexico" pageId="108" pageNumber="109">Mexico</collectingCountry>
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,
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<collectingRegion id="49EEF8473721B968FB915522FBBC0A2D" box="[1038,1135,864,887]" country="Mexico" name="Hidalgo" pageId="108" pageNumber="109">Hidalgo</collectingRegion>
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,
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<quantity id="4CD29B403721B968FBE15522FB1E0A2D" box="[1150,1229,864,887]" metricMagnitude="3" metricUnit="m" metricValue="4.02336" pageId="108" pageNumber="109" unit="mi" value="2.5">2.5 mi</quantity>
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N Tlanchinol.
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</paragraph>
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</subSubSection>
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<subSubSection id="C330652E3721B968FD4655D9FB8E0DAF" pageId="108" pageNumber="109" type="diagnosis">
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<paragraph id="8B9536A53721B968FD4655D9FB8E0DAF" blockId="108.[700,1229,630,1741]" pageId="108" pageNumber="109">
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DIAGNOSIS: Males of this species are immediately recognizable by the combination of the very large, flattened tooth extending from the ventral surface of the procoxa (fig. 116), the characteristic metafemur with two prominent teeth and a relatively strongly concave surface between them (fig. 137), and the relatively broad metasternum with the fovea located anterad of the middle. The male genitalia, while unique in shape (figs. 236– 239), are also similar to many of the other members of the
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<taxonomicName id="4C2A4D263721B968FC1F529CFC0C0DAF" box="[896,991,1246,1269]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="aztec">
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<emphasis id="B95EEAB73721B968FC1F529CFC0C0DAF" box="[896,991,1246,1269]" italics="true" pageId="108" pageNumber="109">A. aztec</emphasis>
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</taxonomicName>
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subgroup.
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</paragraph>
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</subSubSection>
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<subSubSection id="C330652E3721B969FD4652B9FE4C0F0F" lastPageId="109" lastPageNumber="110" pageId="108" pageNumber="109" type="description">
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<paragraph id="8B9536A53721B968FD4652B9FC250C17" blockId="108.[700,1229,630,1741]" pageId="108" pageNumber="109">
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DESCRIPTION: Body moderate in size (TBL =
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<quantity id="4CD29B403721B968FD7E535AFC870C75" box="[737,852,1304,1327]" metricMagnitude="-3" metricUnit="m" metricValue="3.44" pageId="108" pageNumber="109" unit="mm" value="3.44">3.44 mm</quantity>
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), broad, robust (PNW/TBL = 0.48), strongly contractile.
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</paragraph>
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<paragraph id="8B9536A53721B968FD465311FC980C99" blockId="108.[700,1229,630,1741]" pageId="108" pageNumber="109">Head and pronotum red; elytra red, with blue iridescence laterally; venter yellow, dark brown on metasternum; antennae, palpi, and legs yellow.</paragraph>
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<paragraph id="8B9536A53721B969FD465388FE4A0BC2" blockId="108.[700,1229,630,1741]" lastBlockId="109.[123,652,193,1741]" lastPageId="109" lastPageNumber="110" pageId="108" pageNumber="109">Head broad (MDL/OHW = 0.54), dorsal surface flattened, dorsoventrally compressed; with very fine punctures, each with a short, very fine seta, surface between punctures shiny, smooth; frontoclypeal suture obsolete medially; eyes prominent, not strongly compressed; gula flat; antennomere ratios: length I:II:III = 1.6:1.0:1.6, width VII:VIII:IX = 1.0:1.0:2.1. Pronotum very large, broad (PNL/PNW = 0.76), strongly convex, anterolateral lobes strongly produced, lateral margin broadly curved, not angulate; with very fine, sparse punctures, each with a short, very fine seta, surface between punctures smooth. Elytra broad, lateral margins strongly rounded, apically rounded (SEL/ELW = 1.00); punctation similar to pronotum; sutural stria present in apical onehalf. Flight wings fully developed. Mesosternum broadly convex; medial carina prominent. Metasternum narrow medially (MTL/MTW = 0.18), slightly sloping dorsad anteriorly, flattened, smooth; oblique femoral carinae low, not strongly developed, especially laterally, medially low and obsolete.</paragraph>
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<paragraph id="8B9536A53720B969FF0754DDFF220F6D" blockId="109.[123,652,193,1741]" pageId="109" pageNumber="110">Male tarsi 554; pro and mesobasotarsomeres distinctly laterally expanded, protarsomeres more so and with large field of spatulate setae ventrally; mandibles not modified; metafemur moderately broad, prominent, with distinct tooth about onethird distance from apex, smaller tooth about onethird distance from base, with concave area between, apical margin rounded (fig. 137); metasternal fovea large, anteromedial, transversely oval, with dense brush of long, fine setae; procoxa with large ventral, broad flattened tooth (fig. 116). Median lobe in lateral aspect long, slender, evenly curved, expanded subapically along dorsal margin, apical portion directed dorsad at angle, slender, straight, apically slightly expanded and curved ventrad, apex narrowly rounded (fig. 238); in ventral aspect slender, evenly constricted medially, expand ed basal to apical portion with a distinct carina which forms, together with dorsal carina, a sulcus in which fits the lateral lobe, apical portion narrowed to elongate, slender apex (figs. 236, 237); operculum elongate, parallelsided, slightly expanded apically, apex rounded with small medial emargination (fig. 236); lateral lobes long, slender, subapically expanded, apex evenly expand ed, rounded, sinuate apically, apex abruptly narrowed, with a single stout subapical seta (fig. 239).</paragraph>
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<paragraph id="8B9536A53720B969FF07507CFE4C0F0F" blockId="109.[123,652,193,1741]" box="[152,415,1598,1621]" pageId="109" pageNumber="110">Female not examined.</paragraph>
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</subSubSection>
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<subSubSection id="C330652E3720B969FF07501EFD980F97" pageId="109" pageNumber="110" type="etymology">
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<paragraph id="8B9536A53720B969FF07501EFD980F97" blockId="109.[123,652,193,1741]" pageId="109" pageNumber="110">
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ETYMOLOGY: This species is named from the Latin words
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<emphasis id="B95EEAB73720B969FED85038FE7A0FCB" box="[327,425,1658,1681]" italics="true" pageId="109" pageNumber="110">tribulus,</emphasis>
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meaning ‘‘thorn’’, and
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<emphasis id="B95EEAB73720B969FF3150DAFEDD0FF5" box="[174,270,1688,1711]" italics="true" pageId="109" pageNumber="110">grandis,</emphasis>
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meaning ‘‘large’’, for the large, thornlike process on the male procoxa.
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</paragraph>
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</subSubSection>
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<subSubSection id="C330652E3720B969FD475683FB1509AF" pageId="109" pageNumber="110" type="distribution">
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<paragraph id="8B9536A53720B969FD475683FB1509AF" blockId="109.[699,1228,193,333]" pageId="109" pageNumber="110">
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DISTRIBUTION: This species is known only from the
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<typeStatus id="549188073720B969FCB4569CFC8E09AF" box="[811,861,222,245]" pageId="109" pageNumber="110">type</typeStatus>
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locality in
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<collectingRegion id="49EEF8473720B969FC77569CFB9509AF" box="[1000,1094,222,245]" country="Mexico" name="Hidalgo" pageId="109" pageNumber="110">Hidalgo</collectingRegion>
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(fig. 364).
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</paragraph>
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</subSubSection>
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<subSubSection id="C330652E3720B969FD4756BEFD300817" pageId="109" pageNumber="110" type="discussion">
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||
<paragraph id="8B9536A53720B969FD4756BEFD300817" blockId="109.[699,1228,193,333]" pageId="109" pageNumber="110">
|
||
DISCUSSION: The
|
||
<typeStatus id="549188073720B969FC2E56BEFBC00849" box="[945,1043,252,275]" pageId="109" pageNumber="110" type="holotype">holotype</typeStatus>
|
||
was collected from ‘‘fungusy wood chips’’ at ‘‘5000–5200 ft’’.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |