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<document id="0028C53D924DA0C73D632FBBC0D27B24" ID-CLB-Dataset="28431" ID-DOI="10.11646/zootaxa.4564.1.2" ID-GBIF-Dataset="d3a56aac-92aa-48f2-b9fe-9576a4c2a0ec" ID-ISSN="1175-5326" ID-Zenodo-Dep="2588621" ID-ZooBank="27F70B5F-24DD-4C9E-BBA7-2B87476BD49C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552162070752" checkinUser="plazi" docAuthor="Gunter, Nicole L. &amp; Weir, Thomas A." docDate="2019" docId="03832713FF8A8D50A3DAFC1451A79336" docLanguage="en" docName="zootaxa.4564.1.2.pdf" docOrigin="Zootaxa 4564 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepanus pungalina Gunter &amp; Weir 2019, new species" docType="treatment" docUuid="688F650E-BD39-48CC-BDFD-312550D1C7BE" docUuidSource="ZooBank" docVersion="9" lastPageNumber="75" masterDocId="FFBA5F6BFFAB8D72A34DFFB65569910A" masterDocTitle="Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group" masterLastPageNumber="80" masterPageNumber="41" pageNumber="74" updateTime="1698700752634" updateUser="ExternalLinkService">
<mods:mods id="52D8145B298367A7FADC40D756FFC562" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FE73CFE9CFFF1197583DA43E7CCB0F5B">
<mods:title id="766DDF10955D12945639B905D383F7D3">Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group</mods:title>
</mods:titleInfo>
<mods:name id="C3AA314E0475ED063FEF2C37430E4A0E" type="personal">
<mods:role id="60EE68D3E495CF154EA6A40F4E72B22E">
<mods:roleTerm id="F0C8E80DB152BA29786567CCC6C401B6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F98B02C1BEB3F0B576BA289F33B1119E">Gunter, Nicole L.</mods:namePart>
</mods:name>
<mods:name id="FF60FD5C3CB2FACCB2305E741A62E776" type="personal">
<mods:role id="A77E775CF24E6E2C064C819B047B2379">
<mods:roleTerm id="0B02D7E495A0CAB8410CC893CA56E593">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="030E55FD1BB72885D83CF7742CDC47BC">Weir, Thomas A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="03143FFE382F29028E273CBF3C86442A">text</mods:typeOfResource>
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<mods:titleInfo id="4A26365C9E227464DB1171A231A6AD2B">
<mods:title id="790D41B40D6B8D6578F4350D17A5F7B6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="07B1FE116924E38FEA66058A8561689D">
<mods:date id="39AA5D07A20EDF807D77952E95EF0379">2019</mods:date>
<mods:detail id="B14FFAFDD39E0288632B7C8F8A556A23" type="pubDate">
<mods:number id="F7EBF3F21A124AA0BBC9BB8602CAB302">2019-03-05</mods:number>
</mods:detail>
<mods:detail id="906C459D35137E0903DC666F9EE508BF" type="volume">
<mods:number id="56C426A83E1DCE325CAD43DA45BA8C3C">4564</mods:number>
</mods:detail>
<mods:detail id="495B1546A721C5E4FA33DCFAD7DC79E4" type="issue">
<mods:number id="5D3DC17D0B2D42E277DBC94B76BDE394">1</mods:number>
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<mods:start id="05277296FCD30C5D9E30B1A9840F36C4">41</mods:start>
<mods:end id="3D905D1857B780D9C26AB50E5AE542EC">80</mods:end>
</mods:extent>
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<mods:classification id="0FB70076907510A59511A3DD0E412171">journal article</mods:classification>
<mods:identifier id="8093A75FECB38EC497ABDBCB9400130A" type="CLB-Dataset">28431</mods:identifier>
<mods:identifier id="851F82716EB35DDFF80DBA7FB7A3E482" type="DOI">10.11646/zootaxa.4564.1.2</mods:identifier>
<mods:identifier id="954D5C11BD28D49049C713040DA7031E" type="GBIF-Dataset">d3a56aac-92aa-48f2-b9fe-9576a4c2a0ec</mods:identifier>
<mods:identifier id="8414DC03E7D227A8B5A1C3906403B940" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E2655B6AD9E070E819C39454565ADECF" type="Zenodo-Dep">2588621</mods:identifier>
<mods:identifier id="4650FC09B04A04F09C441E5525C26937" type="ZooBank">27F70B5F-24DD-4C9E-BBA7-2B87476BD49C</mods:identifier>
</mods:mods>
<treatment id="03832713FF8A8D50A3DAFC1451A79336" ID-DOI="http://doi.org/10.5281/zenodo.5922830" ID-GBIF-Taxon="154366362" ID-Zenodo-Dep="5922830" LSID="urn:lsid:zoobank.org:act:688F650E-BD39-48CC-BDFD-312550D1C7BE" httpUri="http://treatment.plazi.org/id/03832713FF8A8D50A3DAFC1451A79336" lastPageId="34" lastPageNumber="75" pageId="33" pageNumber="74">
<subSubSection id="C330C58EFF8A8D53A3DAFC14578C92B6" box="[151,741,930,956]" pageId="33" pageNumber="74" type="nomenclature">
<paragraph id="8B959605FF8A8D53A3DAFC14578C92B6" blockId="33.[151,741,930,990]" box="[151,741,930,956]" pageId="33" pageNumber="74">
<heading id="D0DD2169FF8A8D53A3DAFC14578C92B6" bold="true" box="[151,741,930,956]" fontSize="11" level="1" pageId="33" pageNumber="74" reason="1">
<emphasis id="B95E4A17FF8A8D53A3DAFC14578C92B6" bold="true" box="[151,741,930,956]" pageId="33" pageNumber="74">
<taxonomicName id="4C2AED86FF8A8D53A3DAFC14572092B6" ID-CoL="3T8VJ" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[151,585,930,956]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="74" phylum="Arthropoda" rank="species" species="pungalina" status="sp. nov.">
<emphasis id="B95E4A17FF8A8D53A3DAFC1454EB92B6" bold="true" box="[151,386,930,956]" italics="true" pageId="33" pageNumber="74">Lepanus pungalina</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A26DF76CFF8A8D53A118FC14578C92B6" box="[597,741,930,956]" pageId="33" pageNumber="74" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF8A8D53A3DAFC73548192D4" box="[151,488,965,990]" pageId="33" pageNumber="74" type="description">
<paragraph id="8B959605FF8A8D53A3DAFC73548192D4" blockId="33.[151,741,930,990]" box="[151,488,965,990]" pageId="33" pageNumber="74">
(
<figureCitation id="13118A80FF8A8D53A3D2FC73558592D4" box="[159,236,965,990]" captionStart="FIGURE 4" captionStartId="10.[151,250,1773,1795]" captionTargetBox="[153,1436,509,1747]" captionTargetId="figure@10.[151,1436,481,1759]" captionTargetPageId="10" captionText="FIGURE 4. Some diagnostic characters of the Lepanus pygmaeus species group. A, Head of Lepanus guthrieae featuring medially interrupted basal margin (white arrow) and reticulations on clypeal area only (black arrow); B, head of Lepanus lentil featuring angulate and complete basal margin (white arrow); C, pterothorax of Lepanus andersonorum showing fine, short carina extending backwards from the meso-metaventral suture (white arrow); D, posterior aspect of metatibia of Lepanus pecki; E, anterior aspect of metatibia of Lepanus pecki; F, anterior aspect of metatibia of Lepanus guthrieae; G, female protibia of Lepanus pygmaeus; H, male protibia of Lepanus pygmaeus; I, female protibia of Lepanus pecki; J, male protibia of Lepanus pecki; K, male protibia of Lepanus crenidens; L, male protibia of Lepanus pungalina; M, inner laterodorsal aspect male protibia of Lepanus pungalina showing bent apical digit (black arrow)." httpUri="https://zenodo.org/record/2588629/files/figure.png" pageId="33" pageNumber="74">Figs. 4</figureCitation>
L–K, 6F, 9C, 12B, 14)
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF8A8D53A3DAFBB054E695FD" pageId="33" pageNumber="74" type="materials_examined">
<paragraph id="8B959605FF8A8D53A3DAFBB054E695FD" blockId="33.[151,1437,1030,2027]" pageId="33" pageNumber="74">
<emphasis id="B95E4A17FF8A8D53A3DAFBB054CA9515" bold="true" box="[151,419,1030,1055]" pageId="33" pageNumber="74">
Type series.
<materialsCitation id="3B429C58FF8A8D53A260FBB054CA9515" ID-GBIF-Occurrence="2012745326" box="[301,419,1030,1055]" collectingDate="2012-06-26" collectingDateMax="2012-07-08" collectingDateMin="2012-06-26" collectionCode="ANIC" collectorName="N. Gunter Pitfall" county="Karns Crk" determinerName="T. A. Weir" latitude="-16.791666" location="Pungalina" longLatPrecision="21" longitude="137.4575" municipality="Pungalina" pageId="33" pageNumber="74" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="549128A7FF8A8D53A260FBB054F79515" box="[301,414,1030,1055]" pageId="33" pageNumber="74" type="holotype">Holotype</typeStatus>
:
</materialsCitation>
</emphasis>
<specimenCount id="9D2C5D8CFF8A8D53A2E6FBB154A99514" box="[427,448,1031,1054]" pageId="33" pageNumber="74" type="male">♂</specimenCount>
“
<geoCoordinate id="EE1EF0C2FF8A8D53A299FBB0573D9515" box="[468,596,1030,1055]" direction="south" orientation="latitude" pageId="33" pageNumber="74" precision="15" value="-16.791666">16°47'30&quot;S</geoCoordinate>
<geoCoordinate id="EE1EF0C2FF8A8D53A110FBB157879515" box="[605,750,1031,1055]" direction="east" orientation="longitude" pageId="33" pageNumber="74" precision="15" value="137.4575">137°27'27&quot;E</geoCoordinate>
, NT:
<location id="8EF5C0DEFF8A8D53A07DFBB056CA9515" LSID="urn:lsid:plazi:treatment:03832713FF8A8D50A3DAFC1451A79336:8EF5C0DEFF8A8D53A07DFBB056CA9515" box="[816,931,1030,1055]" county="Karns Crk" latitude="-16.791666" longLatPrecision="21" longitude="137.4575" municipality="Pungalina" name="Pungalina" pageId="33" pageNumber="74">Pungalina</location>
,
<quantity id="4CD23BE0FF8A8D53A0E2FBB0569E9515" box="[943,1015,1030,1055]" metricMagnitude="3" metricUnit="m" metricValue="9.6" pageId="33" pageNumber="74" unit="km" value="9.6">9.6km</quantity>
SEbyS
<location id="8EF5C0DEFF8A8D53A719FBB050739515" LSID="urn:lsid:plazi:treatment:03832713FF8A8D50A3DAFC1451A79336:8EF5C0DEFF8A8D53A719FBB050739515" box="[1108,1306,1030,1055]" county="Karns Crk" latitude="-16.791666" longLatPrecision="21" longitude="137.4575" municipality="Pungalina" name="Pungalina Hmstd" pageId="33" pageNumber="74">
<collectingMunicipality id="6BF10C7FFF8A8D53A719FBB051AD9515" box="[1108,1220,1030,1055]" pageId="33" pageNumber="74">Pungalina</collectingMunicipality>
Hmstd
</location>
,
<collectingCounty id="62F4EE89FF8A8D53A66AFBB150F59515" box="[1319,1436,1030,1055]" pageId="33" pageNumber="74">Karns Crk</collectingCounty>
<date id="FF94B0C5FF8A8D53A3DAFB9C540F9549" box="[151,358,1066,1091]" pageId="33" pageNumber="74" value="2012-06-26" valueMax="2012-07-08" valueMin="2012-06-26">
<collectingDate id="EFD0492DFF8A8D53A3DAFB9C540F9549" box="[151,358,1066,1091]" pageId="33" pageNumber="74" value="2012-06-26" valueMax="2012-07-08" valueMin="2012-06-26">26Jun – 8Jul 2012</collectingDate>
</date>
,
<collectorName id="26DFF3D3FF8A8D53A23DFB9D57449549" box="[368,557,1066,1091]" pageId="33" pageNumber="74">N. Gunter Pitfall</collectorName>
” / “mushroom and human dung bait” / “COL2443” / “ANIC Database No. 25- 057708” / “
<taxonomicName id="4C2AED86FF8A8D53A26DFBF954E8956D" authorityName=", Matthews" authorityYear="1974" box="[288,385,1103,1127]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="79" phylum="Arthropoda" rank="genus">Lepanus</taxonomicName>
NT5 det.
<determinerName id="7E497AC1FF8A8D53A2B2FBF8570F956D" box="[511,614,1102,1127]" pageId="33" pageNumber="74">T.A.Weir</determinerName>
2013” / “ANIC Image” in ANIC, also as DNA extraction COL2443.
<materialsCitation id="3B429C58FF8A8D53A3DAFBC454E295FD" ID-GBIF-Occurrence="2012745330" collectingDate="2012-05-23" collectingDateMax="2012-07-08" collectingDateMin="2012-05-23" collectionCode="ANIC" collectorName="N. L. Gunter &amp; Bush Blitz Site B" county="Fish River Station Reserve" latitude="-13.826944" location="Pungalina" longLatPrecision="21" longitude="130.71611" municipality="Karns Creek" pageId="33" pageNumber="74" specimenCount="1" typeStatus="paratype">
<emphasis id="B95E4A17FF8A8D53A3DAFBC454719581" bold="true" box="[151,280,1138,1163]" pageId="33" pageNumber="74">
<typeStatus id="549128A7FF8A8D53A3DAFBC4547A9581" box="[151,275,1138,1163]" pageId="33" pageNumber="74" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<location id="8EF5C0DEFF8A8D53A265FBC454F29581" LSID="urn:lsid:plazi:treatment:03832713FF8A8D50A3DAFC1451A79336:8EF5C0DEFF8A8D53A265FBC454F29581" box="[296,411,1138,1163]" county="Fish River Station Reserve" latitude="-13.826944" longLatPrecision="21" longitude="130.71611" municipality="Karns Creek" name="Pungalina" pageId="33" pageNumber="74">Pungalina</location>
,
<locationDeviation id="9A278F27FF8A8D53A2E2FBC4563E9581" box="[431,855,1138,1163]" location="Pungalina Homestead" pageId="33" pageNumber="74">
<quantity id="4CD23BE0FF8A8D53A2E2FBC4576F9581" box="[431,518,1138,1163]" metricMagnitude="3" metricUnit="m" metricValue="9.6" pageId="33" pageNumber="74" unit="km" value="9.6">9.6 km</quantity>
SSE Pungalina Homestead
</locationDeviation>
,
<collectingMunicipality id="6BF10C7FFF8A8D53A021FBC5516C9581" box="[876,1029,1138,1163]" pageId="33" pageNumber="74">Karns Creek</collectingMunicipality>
,
<geoCoordinate id="EE1EF0C2FF8A8D53A754FBC451F39581" box="[1049,1178,1138,1163]" direction="south" orientation="latitude" pageId="33" pageNumber="74" precision="15" value="-16.791666">16°47'30&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FF8A8D53A7E2FBC550299581" box="[1199,1344,1139,1163]" direction="east" orientation="longitude" pageId="33" pageNumber="74" precision="15" value="137.4575">137°27'27&quot;E</geoCoordinate>
,
<date id="FF94B0C5FF8A8D53A618FBC4546295A5" pageId="33" pageNumber="74" value="2012-06-26" valueMax="2012-07-08" valueMin="2012-06-26">
<collectingDate id="EFD0492DFF8A8D53A618FBC4546295A5" pageId="33" pageNumber="74" value="2012-06-26" valueMax="2012-07-08" valueMin="2012-06-26">26.vi– 8.vii.2012</collectingDate>
</date>
,
<collectorName id="26DFF3D3FF8A8D53A25BFB2154F795A5" box="[278,414,1175,1199]" pageId="33" pageNumber="74">N.L. Gunter</collectorName>
(25-057709
<date id="FF94B0C5FF8A8D53A163FB21572295A5" box="[558,587,1175,1199]" pageId="33" pageNumber="74">
<collectingDate id="EFD0492DFF8A8D53A163FB21572295A5" box="[558,587,1175,1199]" pageId="33" pageNumber="74">x1</collectingDate>
</date>
♀ in
<collectionCode id="ED3B0EC0FF8A8D53A1C7FB2157A795A5" box="[650,718,1175,1199]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="33" pageNumber="74">ANIC</collectionCode>
also as DNA extraction COL2444);
<collectingCounty id="62F4EE89FF8A8D53A724FB2050F195A4" box="[1129,1432,1174,1199]" pageId="33" pageNumber="74">Fish River Station Reserve</collectingCounty>
,
<collectorName id="26DFF3D3FF8A8D53A3DAFB0C543795D9" box="[151,350,1210,1235]" pageId="33" pageNumber="74">Bush Blitz Site B</collectorName>
3,
<geoCoordinate id="EE1EF0C2FF8A8D53A235FB0D549095D9" box="[376,505,1211,1235]" direction="south" orientation="latitude" pageId="33" pageNumber="74" precision="15" value="-13.826944">13°49'37&quot;S</geoCoordinate>
,
<geoCoordinate id="EE1EF0C2FF8A8D53A148FB0C57FE95D9" box="[517,663,1210,1235]" direction="east" orientation="longitude" pageId="33" pageNumber="74" precision="15" value="130.71611">130°42'58&quot;E</geoCoordinate>
,
<date id="FF94B0C5FF8A8D53A1EEFB0C565795D9" box="[675,830,1210,1235]" pageId="33" pageNumber="74" value="2012-05-23" valueMax="2012-05-26" valueMin="2012-05-23">
<collectingDate id="EFD0492DFF8A8D53A1EEFB0C565795D9" box="[675,830,1210,1235]" pageId="33" pageNumber="74" value="2012-05-23" valueMax="2012-05-26" valueMin="2012-05-23">23–26.v.2012</collectingDate>
</date>
,
<collectorName id="26DFF3D3FF8A8D53A004FB0D56BB95D9" box="[841,978,1211,1235]" pageId="33" pageNumber="74">N.L. Gunter</collectorName>
(25-057609
<date id="FF94B0C5FF8A8D53A72EFB0D51E995D9" box="[1123,1152,1211,1235]" pageId="33" pageNumber="74">
<collectingDate id="EFD0492DFF8A8D53A72EFB0D51E995D9" box="[1123,1152,1211,1235]" pageId="33" pageNumber="74">x1</collectingDate>
</date>
♀ in
<collectionCode id="ED3B0EC0FF8A8D53A78DFB0D506C95D9" box="[1216,1285,1211,1235]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="33" pageNumber="74">ANIC</collectionCode>
also as DNA extraction COL2438)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF8A8D50A38AFAB4505F91BA" lastPageId="34" lastPageNumber="75" pageId="33" pageNumber="74" type="description">
<paragraph id="8B959605FF8A8D53A38AFAB456CF9411" blockId="33.[151,1437,1030,2027]" box="[199,934,1282,1307]" pageId="33" pageNumber="74">
<emphasis id="B95E4A17FF8A8D53A38AFAB454309411" bold="true" box="[199,345,1282,1307]" pageId="33" pageNumber="74">Description.</emphasis>
Uniform dark brown in colour. Antennal clubs grey.
</paragraph>
<paragraph id="8B959605FF8A8D53A38AFA9051A89434" blockId="33.[151,1437,1030,2027]" box="[199,1217,1318,1343]" pageId="33" pageNumber="74">
Total length:
<quantity id="4CD23BE0FF8A8D53A211FA9154DC9434" box="[348,437,1319,1343]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="33" pageNumber="74" unit="mm" value="2.3">2.3 mm</quantity>
.
<typeStatus id="549128A7FF8A8D53A2F2FA90574E9435" box="[447,551,1318,1343]" pageId="33" pageNumber="74" type="holotype">Holotype</typeStatus>
measurements: Total length
<quantity id="4CD23BE0FF8A8D53A026FA9156AA9434" box="[875,963,1319,1343]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="33" pageNumber="74" unit="mm" value="2.3">2.3 mm</quantity>
, elytral width
<quantity id="4CD23BE0FF8A8D53A729FA9051D49434" box="[1124,1213,1318,1343]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="33" pageNumber="74" unit="mm" value="1.5">1.5 mm</quantity>
.
</paragraph>
<paragraph id="8B959605FF8A8D53A38AFAFC563F971D" blockId="33.[151,1437,1030,2027]" pageId="33" pageNumber="74">Male: Head: U-shaped between clypeal teeth, which are short and upturned. Margin slightly angulate at clypeogenal suture. Genal angle weakly defined. Basal carina present but interrupted medially. Surface of head reticulate between clypeal suture and apical margin. Dorsal part of eyes narrow in size, separated by interocular space approximately 16–17 times eye width (33: 2); eye canthus nearly dividing eye. Head 1.3 × wider than long (41:32). Prothorax: Hypomeral surface finely reticulate with light punctation. Hypomeral striae about half the length of hypomeron. Pronotum 1.6 × wider than long (66:41).</paragraph>
<paragraph id="8B959605FF8A8D53A38AF99456FD9755" blockId="33.[151,1437,1030,2027]" pageId="33" pageNumber="74">Elytra: Striae 1–5 narrow and double, with very fine punctate edges. All striae on disc not equal width, striae 6–7 are weaker and narrower. Elytra 1.15 × wider than long (68:80).</paragraph>
<paragraph id="8B959605FF8A8D53A38AF9DD56D997C1" blockId="33.[151,1437,1030,2027]" pageId="33" pageNumber="74">Legs: Apical tooth of protibiae elongate and bent downwards. Front edge of protibiae deeply angulate; apical digit somewhat elongate and sharp; two teeth on outer edge. Inner edge of metatibiae almost straight. Metatarsi with 2nd tarsomere subequal to the 5th. Tarsal claws small and simple.</paragraph>
<paragraph id="8B959605FF8A8D53A38AF96057E59650" blockId="33.[151,1437,1030,2027]" pageId="33" pageNumber="74">Abdomen: Pygidium with a secondary basal line across base of depression, pygidial depression tear-drop shaped, containing an inverted V-shaped tubercle that positioned in upper third. Depression occupies at least half size of disc and just extends under the virtually straight upper edge of pygidium. Abdominal ventrites reticulated right across. Segment 6 very finely punctate.</paragraph>
<paragraph id="8B959605FF8A8D53A38AF8D054BB96E1" blockId="33.[151,1437,1030,2027]" pageId="33" pageNumber="74">Pterothorax: Medial lobe of metaventrite finely punctate and broadly margined between mesocoxae. Lateral lobe of metaventrite smooth and punctate with setae. Meso-metaventral suture very slightly arched. Mesoventrite with slight notch anteriorly. Mesoventrite with discernible punctures at least at sides. Mesepimeron reticulate. Metanepisternum reticulate.</paragraph>
<paragraph id="8B959605FF898D50A38AFF21505F91BA" blockId="34.[151,1437,151,573]" box="[199,1334,151,176]" pageId="34" pageNumber="75">
Female: Apical tooth on protibia similar to the male, otherwise as per the
<taxonomicName id="4C2AED86FF898D50A74CFF2F51E791BA" authorityName="in Matthews" authorityYear="1974" box="[1025,1166,152,176]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="75" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FF898D50A74CFF2F517991BA" box="[1025,1040,153,176]" italics="true" pageId="34" pageNumber="75">L</emphasis>
.
<emphasis id="B95E4A17FF898D50A753FF2F51E791BA" box="[1054,1166,153,176]" italics="true" pageId="34" pageNumber="75">pygmaeus</emphasis>
</taxonomicName>
species group.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF898D50A38AFF0A566591DF" box="[199,780,188,213]" pageId="34" pageNumber="75" type="etymology">
<paragraph id="8B959605FF898D50A38AFF0A566591DF" blockId="34.[151,1437,151,573]" box="[199,780,188,213]" pageId="34" pageNumber="75">
<emphasis id="B95E4A17FF898D50A38AFF0A542791DF" bold="true" box="[199,334,188,213]" pageId="34" pageNumber="75">Etymology.</emphasis>
Named for the
<typeStatus id="549128A7FF898D50A2B0FF0B574491DF" box="[509,557,189,213]" pageId="34" pageNumber="75">type</typeStatus>
locality, Pungalina.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF898D50A38AFF695663904A" pageId="34" pageNumber="75" type="distribution">
<paragraph id="8B959605FF898D50A38AFF695663904A" blockId="34.[151,1437,151,573]" pageId="34" pageNumber="75">
<emphasis id="B95E4A17FF898D50A38AFF69540991F2" bold="true" box="[199,352,223,248]" pageId="34" pageNumber="75">Distribution.</emphasis>
Known only from three specimens collected in open forests of the
<collectingRegion id="49EE58E7FF898D50A731FF69502791F2" box="[1148,1358,223,248]" country="Australia" name="Northern Territory" pageId="34" pageNumber="75">Northern Territory</collectingRegion>
IBRA regions GUC and DAC (
<figureCitation id="13118A80FF898D50A2E3FEB2576B9016" box="[430,514,260,285]" captionStart="FIGURE 14" captionStartId="31.[151,250,1220,1242]" captionTargetBox="[151,1436,375,1198]" captionTargetId="figure@31.[151,1436,375,1198]" captionTargetPageId="31" captionText="FIGURE 14. Map of known records for 10 species within the Lepanus pygmaeus species group overlaid with predicted distribution boundary of L. pygmaeus (dashed line) taken from Fig. 13 map." httpUri="https://zenodo.org/record/2588649/files/figure.png" pageId="34" pageNumber="75">Fig. 14</figureCitation>
). Given the disjunct distribution of these two IBRA regions, it is likely it is likely
<taxonomicName id="4C2AED86FF898D50A3DAFE9E544D904A" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[151,292,295,320]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="75" phylum="Arthropoda" rank="species" species="pungalina">L. pungalina</taxonomicName>
also occurs in the adjoining IBRA regions.
</paragraph>
</subSubSection>
<subSubSection id="C330C58EFF898D50A38AFEFA51A79336" pageId="34" pageNumber="75" type="discussion">
<paragraph id="8B959605FF898D50A38AFEFA51A79336" blockId="34.[151,1437,151,573]" pageId="34" pageNumber="75">
<emphasis id="B95E4A17FF898D50A38AFEFA5426906F" bold="true" box="[199,335,332,357]" pageId="34" pageNumber="75">Comments.</emphasis>
<taxonomicName id="4C2AED86FF898D50A214FEFB575D906F" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[345,564,332,357]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="75" phylum="Arthropoda" rank="species" species="pungalina">
<emphasis id="B95E4A17FF898D50A214FEFB575D906F" box="[345,564,332,357]" italics="true" pageId="34" pageNumber="75">Lepanus pungalina</emphasis>
</taxonomicName>
was unknown at the time when the informal nomenclature for undescribed species of
<taxonomicName id="4C2AED86FF898D50A240FEC754059082" authorityName=", Matthews" authorityYear="1974" box="[269,364,369,392]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FF898D50A240FEC754059082" box="[269,364,369,392]" italics="true" pageId="34" pageNumber="75">Lepanus</emphasis>
</taxonomicName>
was reported (
<bibRefCitation id="EFBBEBF4FF898D50A15CFEC657B59082" author="Yeates, D. &amp; Gunter, N. L. &amp; Weir, T. &amp; Cameron, S. L." box="[529,732,367,392]" pageId="34" pageNumber="75" pagination="92 - 99" refId="ref30201" refString="Yeates, D. K, Gunter, N. L., Weir, T. &amp; Cameron, S. L. (2011) Developing a benchmark invertebrate group for biodiversity modelling. In: West, J. (Ed.), 21 st Century Taxonomy: accelerating research and discovery of Australia's biodiversity, Taxonomy Research and Identification Network Final Report- March, 2011, pp. 92 - 99. Available from: https: // www. environment. gov. au / system / files / pages / 56 c 08 f 6 f- 7 bf 7 - 4594 - 8175 - b 4 c 143110156 / files / hub-taxonomy. pdf (accessed 12 December 2018)" type="journal article" year="2011">
Yeates
<emphasis id="B95E4A17FF898D50A129FEC757F59082" box="[612,668,367,392]" italics="true" pageId="34" pageNumber="75">et al.</emphasis>
2011
</bibRefCitation>
) but was designated the informal name “
<taxonomicName id="4C2AED86FF898D50A7EAFEC7506F9082" authorityName=", Matthews" authorityYear="1974" box="[1191,1286,369,392]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FF898D50A7EAFEC7506F9082" box="[1191,1286,369,392]" italics="true" pageId="34" pageNumber="75">Lepanus</emphasis>
</taxonomicName>
NT5” within this revision, which was followed in the phylogenetic study of
<bibRefCitation id="EFBBEBF4FF898D50A019FE22515C90A6" author="Gunter, N. L. &amp; Monteith, G. B. &amp; Cameron, S. L. &amp; Weir, T." box="[852,1077,404,429]" pageId="34" pageNumber="75" pagination="1 - 27" refId="ref29280" refString="Gunter, N. L., Monteith, G. B., Cameron, S. L. &amp; Weir, T. A (2018) Evidence from Australian mesic zone dung beetles support Gondwanan vicariance and Mesozoic origin of the Scarabaeinae (Coleoptera: Scarabaeidae). Invertebrate Systematics and Evolution, 1 - 27. [Advance online publication] https: // doi. org / 10.1163 / 1876312 X- 00002171" type="book chapter" year="2018">
Gunter
<emphasis id="B95E4A17FF898D50A0E4FE23568890A7" box="[937,993,404,429]" italics="true" pageId="34" pageNumber="75">et al.</emphasis>
(2018)
</bibRefCitation>
. The
<typeStatus id="549128A7FF898D50A73FFE2251BD90A7" box="[1138,1236,404,429]" pageId="34" pageNumber="75" type="holotype">holotype</typeStatus>
<taxonomicName id="4C2AED86FF898D50A796FE23500390A7" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[1243,1386,404,429]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="75" phylum="Arthropoda" rank="species" species="pungalina">
<emphasis id="B95E4A17FF898D50A796FE23500390A7" box="[1243,1386,404,429]" italics="true" pageId="34" pageNumber="75">L. pungalina</emphasis>
</taxonomicName>
was sequenced in the study of
<bibRefCitation id="EFBBEBF4FF898D50A297FE0E57A290DA" author="Gunter, N. L. &amp; Monteith, G. B. &amp; Cameron, S. L. &amp; Weir, T." box="[474,715,439,464]" pageId="34" pageNumber="75" pagination="1 - 27" refId="ref29280" refString="Gunter, N. L., Monteith, G. B., Cameron, S. L. &amp; Weir, T. A (2018) Evidence from Australian mesic zone dung beetles support Gondwanan vicariance and Mesozoic origin of the Scarabaeinae (Coleoptera: Scarabaeidae). Invertebrate Systematics and Evolution, 1 - 27. [Advance online publication] https: // doi. org / 10.1163 / 1876312 X- 00002171" type="book chapter" year="2018">
Gunter
<emphasis id="B95E4A17FF898D50A178FE0F571D90DA" box="[565,628,439,464]" italics="true" pageId="34" pageNumber="75">et al.</emphasis>
(2018)
</bibRefCitation>
(ANIC 25-057708) and was recovered as a member of the monsoonal clade of
<taxonomicName id="4C2AED86FF898D50A2CDFE6B54B690FE" authorityName=", Matthews" authorityYear="1974" box="[384,479,477,500]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="B95E4A17FF898D50A2CDFE6B54B690FE" box="[384,479,477,500]" italics="true" pageId="34" pageNumber="75">Lepanus</emphasis>
</taxonomicName>
, which contains species in the
<taxonomicName id="4C2AED86FF898D50A00AFE6B56BE90FE" authorityName="in Matthews" authorityYear="1974" box="[839,983,477,500]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="75" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B95E4A17FF898D50A00AFE6B56BE90FE" box="[839,983,477,500]" italics="true" pageId="34" pageNumber="75">L. pygmaeus</emphasis>
</taxonomicName>
species group, sister to
<taxonomicName id="4C2AED86FF898D50A7A2FE6B501390FF" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[1263,1402,476,501]" class="Insecta" family="Scarabaeidae" genus="Lepanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="75" phylum="Arthropoda" rank="species" species="guthrieae">
<emphasis id="B95E4A17FF898D50A7A2FE6B501390FF" box="[1263,1402,476,501]" italics="true" pageId="34" pageNumber="75">L. guthrieae</emphasis>
</taxonomicName>
. It should be noted that as the
<typeStatus id="549128A7FF898D50A2BEFE49573C9312" box="[499,597,511,536]" pageId="34" pageNumber="75" type="holotype">holotype</typeStatus>
has been sampled for DNA, it is missing a mesothoracic leg and a metathoracic leg. All specimens were collected in mushroom or human dung baited pitfall traps.
</paragraph>
</subSubSection>
</treatment>
</document>
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