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<document id="627428C0D83C182F76CD34E585780BB1" ID-CLB-Dataset="299054" ID-DOI="10.5281/zenodo.12117576" ID-GBIF-Dataset="9ec266f5-1dc7-4272-9fbc-67c7f2b0f5aa" ID-ISSN="0026-279X" ID-Zenodo-Dep="12117576" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1718741462746" checkinUser="felipe" docAuthor="Lobban, Christopher S." docDate="2015" docId="03826143FF908175FF34F9E710ACF8F8" docLanguage="en" docName="Micronesica.2015.02.1-49.pdf" docOrigin="Micronesica 2015 (2)" docStyle="DocumentStyle:73477DD41873F4E732E2765E250AF97D.1:Micronesica.2013-.journal_article.type1" docStyleId="73477DD41873F4E732E2765E250AF97D" docStyleName="Micronesica.2013-.journal_article.type1" docStyleVersion="1" docTitle="Psammodictyon pustulatum Lobban 2015, n. comb., n. stat." docType="treatment" docVersion="5" lastPageNumber="13" masterDocId="FFBB193BFF9B8179FFDCFF9E1460FD3F" masterDocTitle="Benthic marine diatom flora of Guam: new records, redescription of Psammodictyon pustulatum n. comb., n. stat., and three new species (Colliculoamphora gabgabensis, Lauderia excentrica, and Rhoiconeis pagoensis)" masterLastPageNumber="49" masterPageNumber="1" pageNumber="12" updateTime="1719541204275" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="89685B7DDE986CA74C8E5AB02CA91B04">Benthic marine diatom flora of Guam: new records, redescription of Psammodictyon pustulatum n. comb., n. stat., and three new species (Colliculoamphora gabgabensis, Lauderia excentrica, and Rhoiconeis pagoensis)</mods:title>
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<mods:namePart id="4C3C1EE5C33CF513AC0C14A6CD072420">Lobban, Christopher S.</mods:namePart>
<mods:affiliation id="0C9609620351BBC1E11118746AF42021">Division of Natural Sciences, University of Guam, Mangilao, GU 83823, USA e-mail: clobban @ guam. net</mods:affiliation>
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<treatment id="03826143FF908175FF34F9E710ACF8F8" ID-DOI="http://doi.org/10.5281/zenodo.12117582" ID-GBIF-Taxon="233814397" ID-Zenodo-Dep="12117582" LSID="urn:lsid:plazi:treatment:03826143FF908175FF34F9E710ACF8F8" httpUri="http://treatment.plazi.org/id/03826143FF908175FF34F9E710ACF8F8" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<subSubSection id="C33183DEFF908172FF34F9E71114FBAD" box="[232,1396,1657,1682]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8B94D055FF908172FF34F9E71114FBAD" blockId="11.[232,1403,1657,1886]" box="[232,1396,1657,1682]" pageId="11" pageNumber="12">
<heading id="D0DC6739FF908172FF34F9E71114FBAD" box="[232,1396,1657,1682]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
<emphasis id="B95F0C47FF908172FF34F9E71007FBAD" bold="true" box="[232,1127,1657,1682]" pageId="11" pageNumber="12">
<taxonomicName id="4C2BABD6FF908172FF34F9E717ECFBAD" authority="(Voigt ex Meister) Lobban" authorityName="Lobban" authorityYear="2015" baseAuthorityName="Voigt ex Meister" baseAuthorityYear="2015" box="[232,908,1657,1682]" class="Bacillariophyceae" family="Bacillariaceae" genus="Psammodictyon" higherTaxonomySource="GBIF,CoL" kingdom="Chromista" order="Bacillariales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="species" species="pustulatum" status="n. comb., n. stat.">
<emphasis id="B95F0C47FF908172FF34F9E71658FBAD" bold="true" box="[232,568,1657,1682]" italics="true" pageId="11" pageNumber="12">Psammodictyon pustulatum</emphasis>
(Voigt ex Meister) Lobban
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,
<taxonomicNameLabel id="A26CB13CFF908172FC46F9E71007FBAD" box="[922,1127,1657,1682]" pageId="11" pageNumber="12">n. comb., n. stat.</taxonomicNameLabel>
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<figureCitation id="1310CCD0FF908172FB0FF9E71114FBAD" box="[1235,1396,1657,1682]" captionStart-0="Figures 114126" captionStart-1="Figures 127138" captionStartId-0="43.[292,372,761,784]" captionStartId-1="45.[292,372,821,844]" captionTargetId-0="figure-14@44.[232,1402,376,1555]" captionTargetId-1="figure-14@46.[232,1402,232,1852]" captionTargetPageId-0="44" captionTargetPageId-1="46" captionText-0="Figures 114126. Figs 114117. Protokeelia cholnokyi. Fig. 114. Valve in LM (GU44I-1) (DIC). Fig. 115. External view of valve (GU44J-2 / ECT3569), SEM; image courtesy of Elizabeth Ruck. Fig. 116. Internal view of valve (GU44Z-15) (SEM). Fig. 117. External, dorsal view of whole frustule showing girdle bands and dorsal faces of valves (SEM); courtesy of Elizabeth Ruck. Figs 118126. Psammodictyon pustulatum. Figs 118, 119. Frustule at two focal planes (GU66F-7A) (DIC). Fig. 120. Valve from GU66A-3 (DIC). Fig. 121. Specimen from Nagasaki on Meisters slide 3509092. Fig. 122. Valve from Hustedt collection, slide W1/18, specimen from Vera Cruz, Mexico, image courtesy of Friedel Hinz. Fig. 123. External view of valve face (GU52Q-10a) (SEM). Fig. 124. Internal view of valve and part of the valvocopula. Arrows indicate where the valvocopula is broken. Inset shows full-resolution detail of valve margin and valvocopula, with the row of pores on the latter. Fig. 125. Frustule in oblique view showing the valves and girdle components. The ventral part of the valvocopula (vc) of the epivalve (ev) is visible at the front, and at the back the distal part of the valvocopula for the hypovalve (hv). Arrow indicates row of pores on the first pleura (p). Fig. 126. External detail of valve, showing the loculate character; arrow indicates row of pores on valvocopula. Scale bars: Figs 114117, 123, 124 = 5 µm, Figs 118122 = 10 µm, Figs 125, 126 = 2 µm." captionText-1="Figures 127138. Figs 125126. Psammodictyon pustulatum (GU66A-3) (SEM). Fig. 127. Oblique view of frustule showing linear pores on the dorsal valve face (arrow). In both Figs 127 and 128 the transition from the ventral to dorsal parts of the valvocopula is visible (arrowheads). Fig. 128. Apical view of frustule showing the distinctive topography of the ventral side of the raphe-keel, along with the linear pores on the dorsal side of the other valve (arrow) (two stacked images). Figs 129, 130. Pteroncola inane (SEM). Fig. 129. Frustule in girdle view (GU41D-1A). Fig. 130. Broken frustule showing internal and external valve faces, with the single row of internal pores (arrows) (Jordan collection, image by Misaki Ishizawa) (SEM). Figs. 131138. Rhoiconeis pagoensis. Figs 131, 132. Same live cell in valve and girdle view showing plastids (GU7X-8) (DIC). Figs 133, 134. Holotype (valve view) and a girdle view from the same slide (GU7X-7) (DIC). Fig. 135. Frustule in valve view (GU7X-7) (SEM). Fig. 136. Frustule in girdle view; note the very short striae at the central area on the concave valve (GU7X-2) (SEM). Fig. 137. Detail of a pair of frustules showing difference in striae lengths at central area on convex and concave valves (GU7X-2) (SEM). Fig. 138. Apex of a frustule in girdle view showing the segmented valvocopula (S1 and S2) and two pleural bands (P1 and P2) (GU7X-5) (SEM). Scale bars: Fig. 136 = 10 µm, Figs 127128, 131135 = 5 µm, Figs 129, 130, 137, 138 = 2 µm." figureDoi-0="http://doi.org/10.5281/zenodo.12575977" figureDoi-1="http://doi.org/10.5281/zenodo.12575981" httpUri-0="https://zenodo.org/record/12575977/files/figure.png" httpUri-1="https://zenodo.org/record/12575981/files/figure.png" pageId="11" pageNumber="12">Figs 118128</figureCitation>
</heading>
</paragraph>
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<subSubSection id="C33183DEFF908172FF34F9041069FBC8" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="8B94D055FF908172FF34F9041783FBEA" blockId="11.[232,1403,1657,1886]" pageId="11" pageNumber="12">
Basionym:
<taxonomicName id="4C2BABD6FF908172FEA2F90410A0FB8C" authority="Voigt ex Meister 1937" authorityName="Meister" authorityYear="1937" box="[382,1216,1690,1715]" class="Insecta" family="Menoponidae" genus="Nitzschia" kingdom="Animalia" order="Psocodea" pageId="11" pageNumber="12" phylum="Arthropoda" rank="variety" species="panduriformis" variety="pustulata">
<emphasis id="B95F0C47FF908172FEA2F90416C9FB8C" box="[382,681,1690,1715]" italics="true" pageId="11" pageNumber="12">Nitzschia panduriformis</emphasis>
var.
<emphasis id="B95F0C47FF908172FCDFF9041710FB8C" box="[771,880,1690,1715]" italics="true" pageId="11" pageNumber="12">pustulata</emphasis>
Voigt ex
<bibRefCitation id="EFBAADA4FF908172FBCFF90410A0FB8C" author="Meister, F." box="[1043,1216,1690,1715]" pageId="11" pageNumber="12" pagination="3 - 13" refId="ref16430" refString="Meister, F. 1937. Seltene und neue Kieselalgen. II. Berichte der Schweizerischen Botanischen Gesellschaft 47: 258 - 276; pls. 3 - 13." type="journal article" year="1937">Meister 1937</bibRefCitation>
</taxonomicName>
, Berichte der Schweizerischen Botanischen Gesellschaft 47: 270, pl. 11, fig. 9.
</paragraph>
<paragraph id="8B94D055FF908172FF34F9401069FBC8" blockId="11.[232,1403,1657,1886]" box="[232,1033,1758,1783]" pageId="11" pageNumber="12">
Samples:
<emphasis id="B95F0C47FF908172FE87F9401069FBC8" bold="true" box="[347,1033,1758,1783]" pageId="11" pageNumber="12">GU66A-1, GU66A-3, GU66F-2, GU66F-7A; GU52Q-10</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33183DEFF908175FF34F89C10ACF8F8" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
<paragraph id="8B94D055FF908172FF34F89C1100FA24" blockId="11.[232,1403,1657,1886]" box="[232,1376,1794,1820]" pageId="11" pageNumber="12">
Dimensions: Length 2233 µm, width 1317 µm; striae
<quantity id="4CD37DB0FF908172FCA2F89C1785FA24" box="[894,997,1794,1819]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" metricValueMax="6.096" metricValueMin="5.08" pageId="11" pageNumber="12" unit="in" value="22.0" valueMax="24.0" valueMin="20.0">2024 in</quantity>
10 µm, fibulae ca.
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10 µm.
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<paragraph id="8B94D055FF908172FF34F8BA1684FA61" blockId="11.[232,1403,1657,1886]" pageId="11" pageNumber="12">
Diagnostics: Differs from all other species of
<taxonomicName id="4C2BABD6FF908172FCD0F8BA17A7FA02" authorityName="D.G.Mann" authorityYear="1990" box="[780,967,1828,1853]" class="Bacillariophyceae" family="Bacillariaceae" genus="Psammodictyon" higherTaxonomySource="GBIF,CoL" kingdom="Chromista" order="Bacillariales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="genus">
<emphasis id="B95F0C47FF908172FCD0F8BA17A7FA02" box="[780,967,1828,1853]" italics="true" pageId="11" pageNumber="12">Psammodictyon</emphasis>
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in the presence of a pair of rounded elevations on the distal side of the sternum.
</paragraph>
<paragraph id="8B94D055FF978175FF34FF6E1500FFDE" blockId="12.[232,1404,240,1479]" pageId="12" pageNumber="13">
Description: Valves panduriform, highly sculptured, with a pair of rounded elevations on the distal side of the sternum and a pair of shallow depressions on the proximal side. Valves apiculate and loculate; areolae in quincunx arrangement, the striae interrupted by a hyaline sternum near the mid line (
<figureCitation id="1310CCD0FF978175FEFBFECB15B3FC51" box="[295,467,341,366]" captionStart="Figures 114126" captionStartId="43.[292,372,761,784]" captionTargetId="figure-14@44.[232,1402,376,1555]" captionTargetPageId="44" captionText="Figures 114126. Figs 114117. Protokeelia cholnokyi. Fig. 114. Valve in LM (GU44I-1) (DIC). Fig. 115. External view of valve (GU44J-2 / ECT3569), SEM; image courtesy of Elizabeth Ruck. Fig. 116. Internal view of valve (GU44Z-15) (SEM). Fig. 117. External, dorsal view of whole frustule showing girdle bands and dorsal faces of valves (SEM); courtesy of Elizabeth Ruck. Figs 118126. Psammodictyon pustulatum. Figs 118, 119. Frustule at two focal planes (GU66F-7A) (DIC). Fig. 120. Valve from GU66A-3 (DIC). Fig. 121. Specimen from Nagasaki on Meisters slide 3509092. Fig. 122. Valve from Hustedt collection, slide W1/18, specimen from Vera Cruz, Mexico, image courtesy of Friedel Hinz. Fig. 123. External view of valve face (GU52Q-10a) (SEM). Fig. 124. Internal view of valve and part of the valvocopula. Arrows indicate where the valvocopula is broken. Inset shows full-resolution detail of valve margin and valvocopula, with the row of pores on the latter. Fig. 125. Frustule in oblique view showing the valves and girdle components. The ventral part of the valvocopula (vc) of the epivalve (ev) is visible at the front, and at the back the distal part of the valvocopula for the hypovalve (hv). Arrow indicates row of pores on the first pleura (p). Fig. 126. External detail of valve, showing the loculate character; arrow indicates row of pores on valvocopula. Scale bars: Figs 114117, 123, 124 = 5 µm, Figs 118122 = 10 µm, Figs 125, 126 = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.12575977" httpUri="https://zenodo.org/record/12575977/files/figure.png" pageId="12" pageNumber="13">Figs 123, 124</figureCitation>
). (The part next to the raphe is considered proximal, and the side toward the mantle where the elevations occur is distal.) The raphe system is submarginal, dividing the valve into a wide ventral portion and a narrow dorsal band; pores on the dorsal portion consist of a longitudinal row of transapical slits (
<figureCitation id="1310CCD0FF978175FD46FE241721FCEC" box="[666,833,442,467]" captionStart="Figures 127138" captionStartId="45.[292,372,821,844]" captionTargetId="figure-14@46.[232,1402,232,1852]" captionTargetPageId="46" captionText="Figures 127138. Figs 125126. Psammodictyon pustulatum (GU66A-3) (SEM). Fig. 127. Oblique view of frustule showing linear pores on the dorsal valve face (arrow). In both Figs 127 and 128 the transition from the ventral to dorsal parts of the valvocopula is visible (arrowheads). Fig. 128. Apical view of frustule showing the distinctive topography of the ventral side of the raphe-keel, along with the linear pores on the dorsal side of the other valve (arrow) (two stacked images). Figs 129, 130. Pteroncola inane (SEM). Fig. 129. Frustule in girdle view (GU41D-1A). Fig. 130. Broken frustule showing internal and external valve faces, with the single row of internal pores (arrows) (Jordan collection, image by Misaki Ishizawa) (SEM). Figs. 131138. Rhoiconeis pagoensis. Figs 131, 132. Same live cell in valve and girdle view showing plastids (GU7X-8) (DIC). Figs 133, 134. Holotype (valve view) and a girdle view from the same slide (GU7X-7) (DIC). Fig. 135. Frustule in valve view (GU7X-7) (SEM). Fig. 136. Frustule in girdle view; note the very short striae at the central area on the concave valve (GU7X-2) (SEM). Fig. 137. Detail of a pair of frustules showing difference in striae lengths at central area on convex and concave valves (GU7X-2) (SEM). Fig. 138. Apex of a frustule in girdle view showing the segmented valvocopula (S1 and S2) and two pleural bands (P1 and P2) (GU7X-5) (SEM). Scale bars: Fig. 136 = 10 µm, Figs 127128, 131135 = 5 µm, Figs 129, 130, 137, 138 = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.12575981" httpUri="https://zenodo.org/record/12575981/files/figure.png" pageId="12" pageNumber="13">Figs 127, 128</figureCitation>
: arrows). The dorsal part of the valvocopula has a repand
<emphasis id="B95F0C47FF978175FE89FE421512FCCA" box="[341,370,476,501]" italics="true" pageId="12" pageNumber="13">ab</emphasis>
valvar edge (
<figureCitation id="1310CCD0FF978175FDD7FE4216D3FCCA" box="[523,691,476,501]" captionStart="Figures 114126" captionStartId="43.[292,372,761,784]" captionTargetId="figure-14@44.[232,1402,376,1555]" captionTargetPageId="44" captionText="Figures 114126. Figs 114117. Protokeelia cholnokyi. Fig. 114. Valve in LM (GU44I-1) (DIC). Fig. 115. External view of valve (GU44J-2 / ECT3569), SEM; image courtesy of Elizabeth Ruck. Fig. 116. Internal view of valve (GU44Z-15) (SEM). Fig. 117. External, dorsal view of whole frustule showing girdle bands and dorsal faces of valves (SEM); courtesy of Elizabeth Ruck. Figs 118126. Psammodictyon pustulatum. Figs 118, 119. Frustule at two focal planes (GU66F-7A) (DIC). Fig. 120. Valve from GU66A-3 (DIC). Fig. 121. Specimen from Nagasaki on Meisters slide 3509092. Fig. 122. Valve from Hustedt collection, slide W1/18, specimen from Vera Cruz, Mexico, image courtesy of Friedel Hinz. Fig. 123. External view of valve face (GU52Q-10a) (SEM). Fig. 124. Internal view of valve and part of the valvocopula. Arrows indicate where the valvocopula is broken. Inset shows full-resolution detail of valve margin and valvocopula, with the row of pores on the latter. Fig. 125. Frustule in oblique view showing the valves and girdle components. The ventral part of the valvocopula (vc) of the epivalve (ev) is visible at the front, and at the back the distal part of the valvocopula for the hypovalve (hv). Arrow indicates row of pores on the first pleura (p). Fig. 126. External detail of valve, showing the loculate character; arrow indicates row of pores on valvocopula. Scale bars: Figs 114117, 123, 124 = 5 µm, Figs 118122 = 10 µm, Figs 125, 126 = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.12575977" httpUri="https://zenodo.org/record/12575977/files/figure.png" pageId="12" pageNumber="13">Figs 124, 125</figureCitation>
). The distal part of the ventral valve face bears two rounded elevations; the proximal portion slopes down from the raphe-keel to the sternum in two shallow depressions (
<figureCitation id="1310CCD0FF978175FE5CFD811645FF07" box="[384,549,543,568]" captionStart="Figures 114126" captionStartId="43.[292,372,761,784]" captionTargetId="figure-14@44.[232,1402,376,1555]" captionTargetPageId="44" captionText="Figures 114126. Figs 114117. Protokeelia cholnokyi. Fig. 114. Valve in LM (GU44I-1) (DIC). Fig. 115. External view of valve (GU44J-2 / ECT3569), SEM; image courtesy of Elizabeth Ruck. Fig. 116. Internal view of valve (GU44Z-15) (SEM). Fig. 117. External, dorsal view of whole frustule showing girdle bands and dorsal faces of valves (SEM); courtesy of Elizabeth Ruck. Figs 118126. Psammodictyon pustulatum. Figs 118, 119. Frustule at two focal planes (GU66F-7A) (DIC). Fig. 120. Valve from GU66A-3 (DIC). Fig. 121. Specimen from Nagasaki on Meisters slide 3509092. Fig. 122. Valve from Hustedt collection, slide W1/18, specimen from Vera Cruz, Mexico, image courtesy of Friedel Hinz. Fig. 123. External view of valve face (GU52Q-10a) (SEM). Fig. 124. Internal view of valve and part of the valvocopula. Arrows indicate where the valvocopula is broken. Inset shows full-resolution detail of valve margin and valvocopula, with the row of pores on the latter. Fig. 125. Frustule in oblique view showing the valves and girdle components. The ventral part of the valvocopula (vc) of the epivalve (ev) is visible at the front, and at the back the distal part of the valvocopula for the hypovalve (hv). Arrow indicates row of pores on the first pleura (p). Fig. 126. External detail of valve, showing the loculate character; arrow indicates row of pores on valvocopula. Scale bars: Figs 114117, 123, 124 = 5 µm, Figs 118122 = 10 µm, Figs 125, 126 = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.12575977" httpUri="https://zenodo.org/record/12575977/files/figure.png" pageId="12" pageNumber="13">Figs 125, 126</figureCitation>
,
<figureCitation id="1310CCD0FF978175FDEDFD811601FF07" box="[561,609,543,568]" captionStart="Figures 127138" captionStartId="45.[292,372,821,844]" captionTargetId="figure-14@46.[232,1402,232,1852]" captionTargetPageId="46" captionText="Figures 127138. Figs 125126. Psammodictyon pustulatum (GU66A-3) (SEM). Fig. 127. Oblique view of frustule showing linear pores on the dorsal valve face (arrow). In both Figs 127 and 128 the transition from the ventral to dorsal parts of the valvocopula is visible (arrowheads). Fig. 128. Apical view of frustule showing the distinctive topography of the ventral side of the raphe-keel, along with the linear pores on the dorsal side of the other valve (arrow) (two stacked images). Figs 129, 130. Pteroncola inane (SEM). Fig. 129. Frustule in girdle view (GU41D-1A). Fig. 130. Broken frustule showing internal and external valve faces, with the single row of internal pores (arrows) (Jordan collection, image by Misaki Ishizawa) (SEM). Figs. 131138. Rhoiconeis pagoensis. Figs 131, 132. Same live cell in valve and girdle view showing plastids (GU7X-8) (DIC). Figs 133, 134. Holotype (valve view) and a girdle view from the same slide (GU7X-7) (DIC). Fig. 135. Frustule in valve view (GU7X-7) (SEM). Fig. 136. Frustule in girdle view; note the very short striae at the central area on the concave valve (GU7X-2) (SEM). Fig. 137. Detail of a pair of frustules showing difference in striae lengths at central area on convex and concave valves (GU7X-2) (SEM). Fig. 138. Apex of a frustule in girdle view showing the segmented valvocopula (S1 and S2) and two pleural bands (P1 and P2) (GU7X-5) (SEM). Scale bars: Fig. 136 = 10 µm, Figs 127128, 131135 = 5 µm, Figs 129, 130, 137, 138 = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.12575981" httpUri="https://zenodo.org/record/12575981/files/figure.png" pageId="12" pageNumber="13">128</figureCitation>
). Loculae are evident on the elevations (
<figureCitation id="1310CCD0FF978175FBE7FD8110BFFF07" box="[1083,1247,543,568]" captionStart="Figures 114126" captionStartId="43.[292,372,761,784]" captionTargetId="figure-14@44.[232,1402,376,1555]" captionTargetPageId="44" captionText="Figures 114126. Figs 114117. Protokeelia cholnokyi. Fig. 114. Valve in LM (GU44I-1) (DIC). Fig. 115. External view of valve (GU44J-2 / ECT3569), SEM; image courtesy of Elizabeth Ruck. Fig. 116. Internal view of valve (GU44Z-15) (SEM). Fig. 117. External, dorsal view of whole frustule showing girdle bands and dorsal faces of valves (SEM); courtesy of Elizabeth Ruck. Figs 118126. Psammodictyon pustulatum. Figs 118, 119. Frustule at two focal planes (GU66F-7A) (DIC). Fig. 120. Valve from GU66A-3 (DIC). Fig. 121. Specimen from Nagasaki on Meisters slide 3509092. Fig. 122. Valve from Hustedt collection, slide W1/18, specimen from Vera Cruz, Mexico, image courtesy of Friedel Hinz. Fig. 123. External view of valve face (GU52Q-10a) (SEM). Fig. 124. Internal view of valve and part of the valvocopula. Arrows indicate where the valvocopula is broken. Inset shows full-resolution detail of valve margin and valvocopula, with the row of pores on the latter. Fig. 125. Frustule in oblique view showing the valves and girdle components. The ventral part of the valvocopula (vc) of the epivalve (ev) is visible at the front, and at the back the distal part of the valvocopula for the hypovalve (hv). Arrow indicates row of pores on the first pleura (p). Fig. 126. External detail of valve, showing the loculate character; arrow indicates row of pores on valvocopula. Scale bars: Figs 114117, 123, 124 = 5 µm, Figs 118122 = 10 µm, Figs 125, 126 = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.12575977" httpUri="https://zenodo.org/record/12575977/files/figure.png" pageId="12" pageNumber="13">Figs 125, 126</figureCitation>
) but also can be seen on the proximal side of the sternum (
<figureCitation id="1310CCD0FF978175FD26FDDF1701FF65" box="[762,865,577,602]" captionStart="Figures 114126" captionStartId="43.[292,372,761,784]" captionTargetId="figure-14@44.[232,1402,376,1555]" captionTargetPageId="44" captionText="Figures 114126. Figs 114117. Protokeelia cholnokyi. Fig. 114. Valve in LM (GU44I-1) (DIC). Fig. 115. External view of valve (GU44J-2 / ECT3569), SEM; image courtesy of Elizabeth Ruck. Fig. 116. Internal view of valve (GU44Z-15) (SEM). Fig. 117. External, dorsal view of whole frustule showing girdle bands and dorsal faces of valves (SEM); courtesy of Elizabeth Ruck. Figs 118126. Psammodictyon pustulatum. Figs 118, 119. Frustule at two focal planes (GU66F-7A) (DIC). Fig. 120. Valve from GU66A-3 (DIC). Fig. 121. Specimen from Nagasaki on Meisters slide 3509092. Fig. 122. Valve from Hustedt collection, slide W1/18, specimen from Vera Cruz, Mexico, image courtesy of Friedel Hinz. Fig. 123. External view of valve face (GU52Q-10a) (SEM). Fig. 124. Internal view of valve and part of the valvocopula. Arrows indicate where the valvocopula is broken. Inset shows full-resolution detail of valve margin and valvocopula, with the row of pores on the latter. Fig. 125. Frustule in oblique view showing the valves and girdle components. The ventral part of the valvocopula (vc) of the epivalve (ev) is visible at the front, and at the back the distal part of the valvocopula for the hypovalve (hv). Arrow indicates row of pores on the first pleura (p). Fig. 126. External detail of valve, showing the loculate character; arrow indicates row of pores on valvocopula. Scale bars: Figs 114117, 123, 124 = 5 µm, Figs 118122 = 10 µm, Figs 125, 126 = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.12575977" httpUri="https://zenodo.org/record/12575977/files/figure.png" pageId="12" pageNumber="13">Fig. 125</figureCitation>
). Girdle bands lack pores, except for a single longitudinal row on the first pleura (
<figureCitation id="1310CCD0FF978175FD45FDFD1761FF43" box="[665,769,611,636]" captionStart="Figures 114126" captionStartId="43.[292,372,761,784]" captionTargetId="figure-14@44.[232,1402,376,1555]" captionTargetPageId="44" captionText="Figures 114126. Figs 114117. Protokeelia cholnokyi. Fig. 114. Valve in LM (GU44I-1) (DIC). Fig. 115. External view of valve (GU44J-2 / ECT3569), SEM; image courtesy of Elizabeth Ruck. Fig. 116. Internal view of valve (GU44Z-15) (SEM). Fig. 117. External, dorsal view of whole frustule showing girdle bands and dorsal faces of valves (SEM); courtesy of Elizabeth Ruck. Figs 118126. Psammodictyon pustulatum. Figs 118, 119. Frustule at two focal planes (GU66F-7A) (DIC). Fig. 120. Valve from GU66A-3 (DIC). Fig. 121. Specimen from Nagasaki on Meisters slide 3509092. Fig. 122. Valve from Hustedt collection, slide W1/18, specimen from Vera Cruz, Mexico, image courtesy of Friedel Hinz. Fig. 123. External view of valve face (GU52Q-10a) (SEM). Fig. 124. Internal view of valve and part of the valvocopula. Arrows indicate where the valvocopula is broken. Inset shows full-resolution detail of valve margin and valvocopula, with the row of pores on the latter. Fig. 125. Frustule in oblique view showing the valves and girdle components. The ventral part of the valvocopula (vc) of the epivalve (ev) is visible at the front, and at the back the distal part of the valvocopula for the hypovalve (hv). Arrow indicates row of pores on the first pleura (p). Fig. 126. External detail of valve, showing the loculate character; arrow indicates row of pores on valvocopula. Scale bars: Figs 114117, 123, 124 = 5 µm, Figs 118122 = 10 µm, Figs 125, 126 = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.12575977" httpUri="https://zenodo.org/record/12575977/files/figure.png" pageId="12" pageNumber="13">Fig. 125</figureCitation>
: arrow). The valvocopula delimits the ventral margin of the valve with a repand rim and there is a single row of pores at its base on the valve side (
<figureCitation id="1310CCD0FF978175FA9BFD1A1534FF80" captionStart="Figures 114126" captionStartId="43.[292,372,761,784]" captionTargetId="figure-14@44.[232,1402,376,1555]" captionTargetPageId="44" captionText="Figures 114126. Figs 114117. Protokeelia cholnokyi. Fig. 114. Valve in LM (GU44I-1) (DIC). Fig. 115. External view of valve (GU44J-2 / ECT3569), SEM; image courtesy of Elizabeth Ruck. Fig. 116. Internal view of valve (GU44Z-15) (SEM). Fig. 117. External, dorsal view of whole frustule showing girdle bands and dorsal faces of valves (SEM); courtesy of Elizabeth Ruck. Figs 118126. Psammodictyon pustulatum. Figs 118, 119. Frustule at two focal planes (GU66F-7A) (DIC). Fig. 120. Valve from GU66A-3 (DIC). Fig. 121. Specimen from Nagasaki on Meisters slide 3509092. Fig. 122. Valve from Hustedt collection, slide W1/18, specimen from Vera Cruz, Mexico, image courtesy of Friedel Hinz. Fig. 123. External view of valve face (GU52Q-10a) (SEM). Fig. 124. Internal view of valve and part of the valvocopula. Arrows indicate where the valvocopula is broken. Inset shows full-resolution detail of valve margin and valvocopula, with the row of pores on the latter. Fig. 125. Frustule in oblique view showing the valves and girdle components. The ventral part of the valvocopula (vc) of the epivalve (ev) is visible at the front, and at the back the distal part of the valvocopula for the hypovalve (hv). Arrow indicates row of pores on the first pleura (p). Fig. 126. External detail of valve, showing the loculate character; arrow indicates row of pores on valvocopula. Scale bars: Figs 114117, 123, 124 = 5 µm, Figs 118122 = 10 µm, Figs 125, 126 = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.12575977" httpUri="https://zenodo.org/record/12575977/files/figure.png" pageId="12" pageNumber="13">Figs 125, 126</figureCitation>
); however, along the dorsal portion of the valvocopula, the opposite edge is repand (
<figureCitation id="1310CCD0FF978175FA9BFD381533FFDE" captionStart="Figures 114126" captionStartId="43.[292,372,761,784]" captionTargetId="figure-14@44.[232,1402,376,1555]" captionTargetPageId="44" captionText="Figures 114126. Figs 114117. Protokeelia cholnokyi. Fig. 114. Valve in LM (GU44I-1) (DIC). Fig. 115. External view of valve (GU44J-2 / ECT3569), SEM; image courtesy of Elizabeth Ruck. Fig. 116. Internal view of valve (GU44Z-15) (SEM). Fig. 117. External, dorsal view of whole frustule showing girdle bands and dorsal faces of valves (SEM); courtesy of Elizabeth Ruck. Figs 118126. Psammodictyon pustulatum. Figs 118, 119. Frustule at two focal planes (GU66F-7A) (DIC). Fig. 120. Valve from GU66A-3 (DIC). Fig. 121. Specimen from Nagasaki on Meisters slide 3509092. Fig. 122. Valve from Hustedt collection, slide W1/18, specimen from Vera Cruz, Mexico, image courtesy of Friedel Hinz. Fig. 123. External view of valve face (GU52Q-10a) (SEM). Fig. 124. Internal view of valve and part of the valvocopula. Arrows indicate where the valvocopula is broken. Inset shows full-resolution detail of valve margin and valvocopula, with the row of pores on the latter. Fig. 125. Frustule in oblique view showing the valves and girdle components. The ventral part of the valvocopula (vc) of the epivalve (ev) is visible at the front, and at the back the distal part of the valvocopula for the hypovalve (hv). Arrow indicates row of pores on the first pleura (p). Fig. 126. External detail of valve, showing the loculate character; arrow indicates row of pores on valvocopula. Scale bars: Figs 114117, 123, 124 = 5 µm, Figs 118122 = 10 µm, Figs 125, 126 = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.12575977" httpUri="https://zenodo.org/record/12575977/files/figure.png" pageId="12" pageNumber="13">Figs 124, 125</figureCitation>
).
</paragraph>
<paragraph id="8B94D055FF978175FF34FD7410BCFEF2" blockId="12.[232,1404,240,1479]" pageId="12" pageNumber="13">
Comments: The loculate structure places this taxon in
<taxonomicName id="4C2BABD6FF978175FC4DFD74102CFE3C" authorityName="D.G.Mann" authorityYear="1990" box="[913,1100,746,771]" class="Bacillariophyceae" family="Bacillariaceae" genus="Psammodictyon" higherTaxonomySource="GBIF,CoL" kingdom="Chromista" order="Bacillariales" pageId="12" pageNumber="13" phylum="Ochrophyta" rank="genus">
<emphasis id="B95F0C47FF978175FC4DFD74102CFE3C" box="[913,1100,746,771]" italics="true" pageId="12" pageNumber="13">Psammodictyon</emphasis>
</taxonomicName>
rather than
<taxonomicName id="4C2BABD6FF978175FB35FD741136FE3C" authorityName="Baer" authorityYear="1826" box="[1257,1366,746,771]" class="Insecta" family="Menoponidae" genus="Nitzschia" kingdom="Animalia" order="Psocodea" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B95F0C47FF978175FB35FD741136FE3C" box="[1257,1366,746,771]" italics="true" pageId="12" pageNumber="13">Nitzschia</emphasis>
</taxonomicName>
or
<taxonomicName id="4C2BABD6FF978175FF34FC951519FE1B" authorityName="W.Smith" authorityYear="1853" box="[232,377,779,804]" class="Bacillariophyceae" family="Bacillariaceae" genus="Tryblionella" higherTaxonomySource="GBIF,CoL" kingdom="Chromista" order="Bacillariales" pageId="12" pageNumber="13" phylum="Ochrophyta" rank="genus">
<emphasis id="B95F0C47FF978175FF34FC951519FE1B" box="[232,377,779,804]" italics="true" pageId="12" pageNumber="13">Tryblionella</emphasis>
</taxonomicName>
; the character combinations distinguishing these genera are described by
<bibRefCitation id="EFBAADA4FF978175FB35FC95155AFE79" author="Round, F. E. &amp; R. M. Crawford &amp; D. G. Mann" pageId="12" pageNumber="13" refId="ref17569" refString="Round, F. E., R. M. Crawford &amp; D. G. Mann. 1990. The Diatoms: Biology and Morphology of the Genera. Cambridge University Press, Cambridge, U. K., 747 pp." type="book" year="1990">Round et al. (1990)</bibRefCitation>
. However, the valve face is sharply delimited by a vertical rim, which appears to be formed by the valvocopula, rather than merging imperceptibly into a shallow mantle on the ventral side, as described by
<bibRefCitation id="EFBAADA4FF978175FE5BFCEF160DFEB5" author="Round, F. E. &amp; R. M. Crawford &amp; D. G. Mann" box="[391,621,881,906]" pageId="12" pageNumber="13" refId="ref17569" refString="Round, F. E., R. M. Crawford &amp; D. G. Mann. 1990. The Diatoms: Biology and Morphology of the Genera. Cambridge University Press, Cambridge, U. K., 747 pp." type="book" year="1990">Round et al. (1990)</bibRefCitation>
for
<taxonomicName id="4C2BABD6FF978175FD7DFCEF173CFEB5" authorityName="D.G.Mann" authorityYear="1990" box="[673,860,881,906]" class="Bacillariophyceae" family="Bacillariaceae" genus="Psammodictyon" higherTaxonomySource="GBIF,CoL" kingdom="Chromista" order="Bacillariales" pageId="12" pageNumber="13" phylum="Ochrophyta" rank="genus">
<emphasis id="B95F0C47FF978175FD7DFCEF173CFEB5" box="[673,860,881,906]" italics="true" pageId="12" pageNumber="13">Psammodictyon</emphasis>
</taxonomicName>
. The possibility that this rim is simply due to collapse of the frustule during preparation is unlikely because of the consistency of its appearance in the
<collectionCode id="ED3A4890FF978175FEEDFC2A150DFEF2" box="[305,365,948,973]" pageId="12" pageNumber="13">SEM</collectionCode>
images and the way it goes down under the dorsal margin (
<figureCitation id="1310CCD0FF978175FBF5FC2A10AFFEF2" box="[1065,1231,948,973]" captionStart="Figures 127138" captionStartId="45.[292,372,821,844]" captionTargetId="figure-14@46.[232,1402,232,1852]" captionTargetPageId="46" captionText="Figures 127138. Figs 125126. Psammodictyon pustulatum (GU66A-3) (SEM). Fig. 127. Oblique view of frustule showing linear pores on the dorsal valve face (arrow). In both Figs 127 and 128 the transition from the ventral to dorsal parts of the valvocopula is visible (arrowheads). Fig. 128. Apical view of frustule showing the distinctive topography of the ventral side of the raphe-keel, along with the linear pores on the dorsal side of the other valve (arrow) (two stacked images). Figs 129, 130. Pteroncola inane (SEM). Fig. 129. Frustule in girdle view (GU41D-1A). Fig. 130. Broken frustule showing internal and external valve faces, with the single row of internal pores (arrows) (Jordan collection, image by Misaki Ishizawa) (SEM). Figs. 131138. Rhoiconeis pagoensis. Figs 131, 132. Same live cell in valve and girdle view showing plastids (GU7X-8) (DIC). Figs 133, 134. Holotype (valve view) and a girdle view from the same slide (GU7X-7) (DIC). Fig. 135. Frustule in valve view (GU7X-7) (SEM). Fig. 136. Frustule in girdle view; note the very short striae at the central area on the concave valve (GU7X-2) (SEM). Fig. 137. Detail of a pair of frustules showing difference in striae lengths at central area on convex and concave valves (GU7X-2) (SEM). Fig. 138. Apex of a frustule in girdle view showing the segmented valvocopula (S1 and S2) and two pleural bands (P1 and P2) (GU7X-5) (SEM). Scale bars: Fig. 136 = 10 µm, Figs 127128, 131135 = 5 µm, Figs 129, 130, 137, 138 = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.12575981" httpUri="https://zenodo.org/record/12575981/files/figure.png" pageId="12" pageNumber="13">Figs 127, 128</figureCitation>
).
</paragraph>
<paragraph id="8B94D055FF978175FE94FC4810CCF92E" blockId="12.[232,1404,240,1479]" pageId="12" pageNumber="13">
I examined Meisters slide 3509092, and found
<specimenCount id="9D2D1BDCFF978175FC59FC481047FED0" box="[901,1063,982,1007]" count="1" pageId="12" pageNumber="13" type="generic">one specimen</specimenCount>
—apparently not the
<typeStatus id="54906EF7FF978175FAF8FC481136FED0" box="[1316,1366,982,1007]" pageId="12" pageNumber="13">type</typeStatus>
he photographed, however, since it is partially overlain by another diatom (
<figureCitation id="1310CCD0FF978175FBE4FC6610FEF92E" box="[1080,1182,1016,1041]" captionStart="Figures 114126" captionStartId="43.[292,372,761,784]" captionTargetId="figure-14@44.[232,1402,376,1555]" captionTargetPageId="44" captionText="Figures 114126. Figs 114117. Protokeelia cholnokyi. Fig. 114. Valve in LM (GU44I-1) (DIC). Fig. 115. External view of valve (GU44J-2 / ECT3569), SEM; image courtesy of Elizabeth Ruck. Fig. 116. Internal view of valve (GU44Z-15) (SEM). Fig. 117. External, dorsal view of whole frustule showing girdle bands and dorsal faces of valves (SEM); courtesy of Elizabeth Ruck. Figs 118126. Psammodictyon pustulatum. Figs 118, 119. Frustule at two focal planes (GU66F-7A) (DIC). Fig. 120. Valve from GU66A-3 (DIC). Fig. 121. Specimen from Nagasaki on Meisters slide 3509092. Fig. 122. Valve from Hustedt collection, slide W1/18, specimen from Vera Cruz, Mexico, image courtesy of Friedel Hinz. Fig. 123. External view of valve face (GU52Q-10a) (SEM). Fig. 124. Internal view of valve and part of the valvocopula. Arrows indicate where the valvocopula is broken. Inset shows full-resolution detail of valve margin and valvocopula, with the row of pores on the latter. Fig. 125. Frustule in oblique view showing the valves and girdle components. The ventral part of the valvocopula (vc) of the epivalve (ev) is visible at the front, and at the back the distal part of the valvocopula for the hypovalve (hv). Arrow indicates row of pores on the first pleura (p). Fig. 126. External detail of valve, showing the loculate character; arrow indicates row of pores on valvocopula. Scale bars: Figs 114117, 123, 124 = 5 µm, Figs 118122 = 10 µm, Figs 125, 126 = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.12575977" httpUri="https://zenodo.org/record/12575977/files/figure.png" pageId="12" pageNumber="13">Fig. 121</figureCitation>
).
</paragraph>
<paragraph id="8B94D055FF978175FE94FB8417DDF949" blockId="12.[232,1404,240,1479]" pageId="12" pageNumber="13">
Although
<bibRefCitation id="EFBAADA4FF978175FE63FB84160FF90C" author="Meister, F." box="[447,623,1050,1075]" pageId="12" pageNumber="13" pagination="3 - 13" refId="ref16430" refString="Meister, F. 1937. Seltene und neue Kieselalgen. II. Berichte der Schweizerischen Botanischen Gesellschaft 47: 258 - 276; pls. 3 - 13." type="journal article" year="1937">Meister (1937)</bibRefCitation>
in the same paper also named
<taxonomicName id="4C2BABD6FF978175FC3CFB8410A3F90C" authorityName="Meister" authorityYear="1937" box="[992,1219,1050,1075]" class="Insecta" family="Menoponidae" genus="Nitzschia" kingdom="Animalia" order="Psocodea" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pustulata">
<emphasis id="B95F0C47FF978175FC3CFB8410A3F90C" box="[992,1219,1050,1075]" italics="true" pageId="12" pageNumber="13">Nitzschia pustulata</emphasis>
</taxonomicName>
, this is entirely different and certainly not a
<taxonomicName id="4C2BABD6FF978175FD9FFBA5169EF96B" authorityName="D.G.Mann" authorityYear="1990" box="[579,766,1083,1108]" class="Bacillariophyceae" family="Bacillariaceae" genus="Psammodictyon" higherTaxonomySource="GBIF,CoL" kingdom="Chromista" order="Bacillariales" pageId="12" pageNumber="13" phylum="Ochrophyta" rank="genus">
<emphasis id="B95F0C47FF978175FD9FFBA5169EF96B" box="[579,766,1083,1108]" italics="true" pageId="12" pageNumber="13">Psammodictyon</emphasis>
</taxonomicName>
, so I am confident there will be no future confusion from using
<taxonomicName id="4C2BABD6FF978175FEB3FBC31591F949" authorityName="Lobban" authorityYear="2015" baseAuthorityName="Voigt ex Meister" baseAuthorityYear="2015" box="[367,497,1117,1142]" class="Bacillariophyceae" family="Bacillariaceae" genus="Psammodictyon" higherTaxonomySource="GBIF,CoL" kingdom="Chromista" order="Bacillariales" pageId="12" pageNumber="13" phylum="Ochrophyta" rank="species" species="pustulatum">
<emphasis id="B95F0C47FF978175FEB3FBC31591F949" box="[367,497,1117,1142]" italics="true" pageId="12" pageNumber="13">pustulatum</emphasis>
</taxonomicName>
as the epithet in this new combination.
</paragraph>
<paragraph id="8B94D055FF978175FE94FBE116FAF820" blockId="12.[232,1404,240,1479]" pageId="12" pageNumber="13">
Specimens labeled as
<taxonomicName id="4C2BABD6FF978175FD88FBE117ACF9A7" authorityName="Voigt ex Meister" authorityYear="1937" box="[596,972,1151,1176]" class="Insecta" family="Menoponidae" genus="Nitzschia" kingdom="Animalia" order="Psocodea" pageId="12" pageNumber="13" phylum="Arthropoda" rank="variety" species="panduriforme" variety="pustulata">
<emphasis id="B95F0C47FF978175FD88FBE117ACF9A7" box="[596,972,1151,1176]" italics="true" pageId="12" pageNumber="13">N. panduriforme var. pustulata</emphasis>
</taxonomicName>
occur on two slides in the Hustedt collection (ZT2/46 and W1/18), kindly photographed by Friedel Hinz (one shown in
<figureCitation id="1310CCD0FF978175FB07FB3F1121F985" box="[1243,1345,1185,1210]" captionStart="Figures 114126" captionStartId="43.[292,372,761,784]" captionTargetId="figure-14@44.[232,1402,376,1555]" captionTargetPageId="44" captionText="Figures 114126. Figs 114117. Protokeelia cholnokyi. Fig. 114. Valve in LM (GU44I-1) (DIC). Fig. 115. External view of valve (GU44J-2 / ECT3569), SEM; image courtesy of Elizabeth Ruck. Fig. 116. Internal view of valve (GU44Z-15) (SEM). Fig. 117. External, dorsal view of whole frustule showing girdle bands and dorsal faces of valves (SEM); courtesy of Elizabeth Ruck. Figs 118126. Psammodictyon pustulatum. Figs 118, 119. Frustule at two focal planes (GU66F-7A) (DIC). Fig. 120. Valve from GU66A-3 (DIC). Fig. 121. Specimen from Nagasaki on Meisters slide 3509092. Fig. 122. Valve from Hustedt collection, slide W1/18, specimen from Vera Cruz, Mexico, image courtesy of Friedel Hinz. Fig. 123. External view of valve face (GU52Q-10a) (SEM). Fig. 124. Internal view of valve and part of the valvocopula. Arrows indicate where the valvocopula is broken. Inset shows full-resolution detail of valve margin and valvocopula, with the row of pores on the latter. Fig. 125. Frustule in oblique view showing the valves and girdle components. The ventral part of the valvocopula (vc) of the epivalve (ev) is visible at the front, and at the back the distal part of the valvocopula for the hypovalve (hv). Arrow indicates row of pores on the first pleura (p). Fig. 126. External detail of valve, showing the loculate character; arrow indicates row of pores on valvocopula. Scale bars: Figs 114117, 123, 124 = 5 µm, Figs 118122 = 10 µm, Figs 125, 126 = 2 µm." figureDoi="http://doi.org/10.5281/zenodo.12575977" httpUri="https://zenodo.org/record/12575977/files/figure.png" pageId="12" pageNumber="13">Fig. 122</figureCitation>
), are identical to my specimens; a similar specimen is illustrated in
<bibRefCitation id="EFBAADA4FF978175FC15FB5C10DDF9E4" author="Stidolph, S. R. &amp; F. A. S. Sterrenburg &amp; K. E. L. Smith &amp; A. Kraberg" box="[969,1213,1218,1243]" pageId="12" pageNumber="13" refId="ref17846" refString="Stidolph, S. R., F. A. S. Sterrenburg, K. E. L. Smith &amp; A. Kraberg. 2012. Stuart R. Stidolph Diatom Atlas: U. S. Geological Survey Open-File Report 2012 - 1163, available online at http: // pubs. usgs. gov / of / 2012 / 1163 / (accessed 25 July 2014)." type="url" year="2012">Stidolph et al. (2012</bibRefCitation>
, pl. 25, fig. 83). The known geographic range of
<taxonomicName id="4C2BABD6FF978175FD4EFB7A175DF9C2" authorityName="Lobban" authorityYear="2015" baseAuthorityName="Voigt ex Meister" baseAuthorityYear="2015" box="[658,829,1252,1277]" class="Bacillariophyceae" family="Bacillariaceae" genus="Psammodictyon" higherTaxonomySource="GBIF,CoL" kingdom="Chromista" order="Bacillariales" pageId="12" pageNumber="13" phylum="Ochrophyta" rank="species" species="pustulatum">
<emphasis id="B95F0C47FF978175FD4EFB7A175DF9C2" box="[658,829,1252,1277]" italics="true" pageId="12" pageNumber="13">P. pustulatum</emphasis>
</taxonomicName>
is thus
<collectingRegion id="49EF1EB7FF978175FC6FFB7A1040F9C2" box="[947,1056,1252,1277]" country="Japan" name="Nagasaki" pageId="12" pageNumber="13">Nagasaki</collectingRegion>
(
<collectingCountry id="F33C90C5FF978175FBE4FB7A10E0F9C2" box="[1080,1152,1252,1277]" name="Japan" pageId="12" pageNumber="13">Japan</collectingCountry>
),
<collectingCountry id="F33C90C5FF978175FB42FB7A1089F9C2" box="[1182,1257,1252,1277]" name="Guam" pageId="12" pageNumber="13">Guam</collectingCountry>
,
<collectingCountry id="F33C90C5FF978175FB21FB7A1117F9C2" box="[1277,1399,1252,1277]" name="Singapore" pageId="12" pageNumber="13">Singapore</collectingCountry>
, Galapagos, and Vera Cruz (
<collectingCountry id="F33C90C5FF978175FDF1FA9816EDF820" box="[557,653,1286,1311]" name="Mexico" pageId="12" pageNumber="13">Mexico</collectingCountry>
).
</paragraph>
<paragraph id="8B94D055FF978175FE94FAB610ACF8F8" blockId="12.[232,1404,240,1479]" pageId="12" pageNumber="13">
The size and outline of
<taxonomicName id="4C2BABD6FF978175FDB3FAB61776F87E" authorityName="Lobban" authorityYear="2015" baseAuthorityName="Voigt ex Meister" baseAuthorityYear="2015" box="[623,790,1320,1345]" class="Bacillariophyceae" family="Bacillariaceae" genus="Psammodictyon" higherTaxonomySource="GBIF,CoL" kingdom="Chromista" order="Bacillariales" pageId="12" pageNumber="13" phylum="Ochrophyta" rank="species" species="pustulatum">
<emphasis id="B95F0C47FF978175FDB3FAB61776F87E" box="[623,790,1320,1345]" italics="true" pageId="12" pageNumber="13">P. pustulatum</emphasis>
</taxonomicName>
are similar to
<taxonomicName id="4C2BABD6FF978175FC13FAB6101DF87E" box="[975,1149,1320,1345]" class="Bacillariophyceae" family="Bacillariaceae" genus="Psammodictyon" higherTaxonomySource="GBIF,CoL" kingdom="Chromista" order="Bacillariales" pageId="12" pageNumber="13" phylum="Ochrophyta" rank="species" species="constrictum">
<emphasis id="B95F0C47FF978175FC13FAB6101DF87E" box="[975,1149,1320,1345]" italics="true" pageId="12" pageNumber="13">P. constrictum</emphasis>
</taxonomicName>
in our flora, but the areolae, with single pores on both sides, are like those in
<emphasis id="B95F0C47FF978175FC1EFAD710F1F85D" box="[962,1169,1353,1378]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="4C2BABD6FF978175FC1EFAD710ECF85D" authorityName="W.Gregory" authorityYear="1857" box="[962,1164,1353,1378]" class="Insecta" family="Menoponidae" genus="Nitzschia" kingdom="Animalia" order="Psocodea" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="panduriforme">P. panduriforme</taxonomicName>
,
</emphasis>
unlike those in
<emphasis id="B95F0C47FF978175FABEFAD7151AF8BB" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="4C2BABD6FF978175FABEFAD71515F8BB" class="Bacillariophyceae" family="Bacillariaceae" genus="Psammodictyon" higherTaxonomySource="GBIF,CoL" kingdom="Chromista" order="Bacillariales" pageId="12" pageNumber="13" phylum="Ochrophyta" rank="species" species="constrictum">P. constrictum</taxonomicName>
,
</emphasis>
where there are complex external openings and clusters of 34 interior pores. Further differences in
<taxonomicName id="4C2BABD6FF978175FE4DFA13165CF899" box="[401,572,1421,1446]" class="Bacillariophyceae" family="Bacillariaceae" genus="Psammodictyon" higherTaxonomySource="GBIF,CoL" kingdom="Chromista" order="Bacillariales" pageId="12" pageNumber="13" phylum="Ochrophyta" rank="species" species="constrictum">
<emphasis id="B95F0C47FF978175FE4DFA13165CF899" box="[401,572,1421,1446]" italics="true" pageId="12" pageNumber="13">P. constrictum</emphasis>
</taxonomicName>
are that the areolae on the dorsal side of the raphe-keel are also large, and the pleura has transapical striae all along it (
<bibRefCitation id="EFBAADA4FF978175FCC2FA30179EF8F8" author="Lobban, C. S. &amp; M. Schefter &amp; R. W. Jordan &amp; Y. Arai &amp; A. Sasaki &amp; E. C. Theriot &amp; M. Ashworth &amp; E. C. Ruck &amp; C. Pennesi" box="[798,1022,1454,1479]" pageId="12" pageNumber="13" refId="ref16066" refString="Lobban, C. S., M. Schefter, R. W. Jordan, Y. Arai, A. Sasaki, E. C. Theriot, M. Ashworth, E. C. Ruck &amp; C. Pennesi. 2012. Coral-reef diatoms (Bacillariophyta) from Guam: new records and preliminary checklist, with emphasis on epiphytic species from farmer-fish territories. Micronesica 43: 237 - 479. [Errata on page 480 of the print edition and posted online at http: // micronesica. org / sites / default / files / 12 _ lobban _ et _ al _ 2012 _ errata. pdf]" type="url" year="2012">Lobban et al. 2012</bibRefCitation>
, pl. 59, figs 7, 8).
</paragraph>
</subSubSection>
</treatment>
</document>