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<document id="87240D37E2B7742902DC9E76C5B5AEE2" ID-CLB-Dataset="95354" ID-DOI="10.11646/palaeoentomology.5.3.6" ID-GBIF-Dataset="ba07339a-df37-474e-81b2-d676d1eb7da5" ID-ISSN="2624-2834" ID-Zenodo-Dep="6820841" ID-ZooBank="21855A12-EBBD-4125-A47B-4D159FDDC9F5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657610399486" checkinUser="plazi" docAuthor="HUANG, DI-YING, CORENTIN, JOUAULT &amp; NEL, ANDRÉ" docDate="2022" docId="03824721FFE5FF8BFCD0FC321153FCEF" docLanguage="en" docName="palaeoentomology.5.3.6.pdf" docOrigin="Palaeoentomology 5 (3)" docSource="http://dx.doi.org/10.11646/palaeoentomology.5.3.6" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Rudiaeschna jarzembowskii HUANG &amp; CORENTIN &amp; NEL 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="241" masterDocId="FFBB3F59FFE4FF89FFFAFF801558FFB2" masterDocTitle="The second species of Rudiaeschna (Odonata, Rudiaeschnidae) discovered in the Lower Cretaceous of Inner Mongolia, Northeast China" masterLastPageNumber="245" masterPageNumber="240" pageNumber="241" updateTime="1699358429918" updateUser="ExternalLinkService">
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<mods:title id="2D241B77CE0E4F0BD9A690BF4A036FC1">The second species of Rudiaeschna (Odonata, Rudiaeschnidae) discovered in the Lower Cretaceous of Inner Mongolia, Northeast China</mods:title>
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<mods:namePart id="B85CAF12677D15F1135F796D6A8AA16F">HUANG, DI-YING</mods:namePart>
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<mods:namePart id="8780E5B732BE90118E6EA77BF564AA4A">CORENTIN, JOUAULT</mods:namePart>
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<mods:namePart id="0FF0E8822292DC6A436F3FDD4743B697">NEL, ANDRÉ</mods:namePart>
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<mods:title id="31DE1EC9ACBC3AF2568E0A8892C17044">Palaeoentomology</mods:title>
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<mods:date id="32378A688FACA3405AD5D43D07301366">2022</mods:date>
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<mods:number id="1B0E06AF5F8DE7C02460FAB0FD0722AD">2022-06-27</mods:number>
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<mods:identifier id="154580D7F8D17D6434FD297F432E80C0" type="CLB-Dataset">95354</mods:identifier>
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<treatment id="03824721FFE5FF8BFCD0FC321153FCEF" ID-DOI="http://doi.org/10.5281/zenodo.6828727" ID-GBIF-Taxon="197516827" ID-Zenodo-Dep="6828727" LSID="urn:lsid:plazi:treatment:03824721FFE5FF8BFCD0FC321153FCEF" httpUri="http://treatment.plazi.org/id/03824721FFE5FF8BFCD0FC321153FCEF" lastPageId="2" lastPageNumber="241" pageId="1" pageNumber="241">
<subSubSection id="C331A5BCFFE5FF88FCD0FC321195FC7F" box="[810,1229,946,973]" pageId="1" pageNumber="241" type="nomenclature">
<paragraph id="8B94F637FFE5FF88FCD0FC321195FC7F" blockId="1.[810,1452,946,1082]" box="[810,1229,946,973]" pageId="1" pageNumber="241">
<heading id="D0DC415BFFE5FF88FCD0FC321195FC7F" bold="true" box="[810,1229,946,973]" fontSize="10" level="2" pageId="1" pageNumber="241" reason="3">
<emphasis id="B95F2A25FFE5FF88FCD0FC321195FC7F" bold="true" box="[810,1229,946,973]" pageId="1" pageNumber="241">
<taxonomicName id="4C2B8DB4FFE5FF88FCD0FC321135FC7F" authority="HUANG &amp; CORENTIN &amp; NEL, 2022" authorityName="HUANG &amp; CORENTIN &amp; NEL" authorityYear="2022" box="[810,1133,946,973]" class="Insecta" family="Rudiaeschnidae" genus="Rudiaeschna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="241" phylum="Arthropoda" rank="species" species="jarzembowskii" status="sp. nov.">
<emphasis id="B95F2A25FFE5FF88FCD0FC321135FC7F" bold="true" box="[810,1133,946,973]" italics="true" pageId="1" pageNumber="241">Rudiaeschna jarzembowskii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C975EFFE5FF88FB8EFC331195FC7F" box="[1140,1229,947,973]" pageId="1" pageNumber="241" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C331A5BCFFE5FF88FCD0FC5716B9FB88" pageId="1" pageNumber="241" type="description">
<paragraph id="8B94F637FFE5FF88FCD0FC5716B9FB88" blockId="1.[810,1452,946,1082]" pageId="1" pageNumber="241">
(
<figureCitation id="1310EAB2FFE5FF88FCC8FC571622FC43" box="[818,890,983,1009]" captionStart="FIGURE 2" captionStartId="2.[136,235,648,672]" captionTargetBox="[151,1436,189,624]" captionTargetId="figure-618@2.[151,1436,189,624]" captionTargetPageId="2" captionText="FIGURE 2. Rudiaeschna jarzembowskii sp. nov., Early Cretaceous, holotype NIGP180347, Photograph of forewing. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.6820845" httpUri="https://zenodo.org/record/6820845/files/figure.png" pageId="1" pageNumber="241">Figs 2</figureCitation>
,
<figureCitation id="1310EAB2FFE5FF88FC7CFC5716CCFC43" box="[902,916,983,1009]" captionStart="FIGURE 3" captionStartId="3.[136,235,1617,1641]" captionTargetBox="[425,1162,189,1594]" captionTargetId="figure-210@3.[425,1162,189,1594]" captionTargetPageId="3" captionText="FIGURE 3. Details of Rudiaeschna jarzembowskii sp. nov., Early Cretaceous, holotype NIGP180347. A, Wing base. B, Nodus area. C, Wing apex. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6820847" httpUri="https://zenodo.org/record/6820847/files/figure.png" pageId="1" pageNumber="241">3</figureCitation>
)
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urn:lsid:zoobank.org:act:
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<subSubSection id="C331A5BCFFE5FF88FCD0FBE911E0FB7E" pageId="1" pageNumber="241" type="materials_examined">
<paragraph id="8B94F637FFE5FF88FCD0FBE911E0FB7E" blockId="1.[810,1452,1129,2070]" pageId="1" pageNumber="241">
<emphasis id="B95F2A25FFE5FF88FCD0FBE916C1FB31" bold="true" box="[810,921,1129,1155]" pageId="1" pageNumber="241">Material.</emphasis>
<materialsCitation id="3B43FC6AFFE5FF88FC5BFBE911E0FB7E" ID-GBIF-Occurrence="3862611302" collectionCode="NIGP" country="China" county="Nanjing" location="Nanjing Institute of Geology" municipality="Palaeontology" pageId="1" pageNumber="241" specimenCode="NIGP180347" specimenCount="1" typeStatus="holotype">
<typeStatus id="54904895FFE5FF88FC5BFBE91151FB31" box="[929,1033,1129,1155]" pageId="1" pageNumber="241" type="holotype">Holotype</typeStatus>
<specimenCode id="DB8D5E4CFFE5FF88FBEBFBE911FDFB31" box="[1041,1189,1129,1155]" collectionCode="NIGP" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="1" pageNumber="241">NIGP180347</specimenCode>
(a complete imprint of a forewing), housed in the
<location id="8EF4A0ECFFE5FF88FB98FB0E10F3FB1A" LSID="urn:lsid:plazi:treatment:03824721FFE5FF8BFCD0FC321153FCEF:8EF4A0ECFFE5FF88FB98FB0E10F3FB1A" box="[1122,1451,1166,1192]" country="China" county="Nanjing" municipality="Palaeontology" name="Nanjing Institute of Geology" pageId="1" pageNumber="241">Nanjing Institute of Geology</location>
and
<collectingMunicipality id="6BF06C4DFFE5FF88FCA3FB3216A2FB7E" box="[857,1018,1202,1228]" pageId="1" pageNumber="241">Palaeontology</collectingMunicipality>
,
<collectingCounty id="62F58EBBFFE5FF88FBFCFB32113BFB7E" box="[1030,1123,1202,1228]" pageId="1" pageNumber="241">Nanjing</collectingCounty>
,
<collectingCountry id="F33CB6A7FFE5FF88FB94FB3211EDFB7E" box="[1134,1205,1202,1228]" name="China" pageId="1" pageNumber="241">China</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C331A5BCFFE5FF88FCA5FB571057FA89" pageId="1" pageNumber="241" type="etymology">
<paragraph id="8B94F637FFE5FF88FCA5FB571057FA89" blockId="1.[810,1452,1129,2070]" pageId="1" pageNumber="241">
<emphasis id="B95F2A25FFE5FF88FCA5FB5716BEFB43" bold="true" box="[863,998,1239,1265]" pageId="1" pageNumber="241">Etymology.</emphasis>
Named in honor of our friend and colleague Edmund A. Jarzembowski. The specific epithet is to be treated as a noun in a genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C331A5BCFFE5FF88FCA5FAC4168CF9ED" pageId="1" pageNumber="241" type="diagnosis">
<paragraph id="8B94F637FFE5FF88FCA5FAC4168CF9ED" blockId="1.[810,1452,1129,2070]" pageId="1" pageNumber="241">
<emphasis id="B95F2A25FFE5FF88FCA5FAC41681FAEC" bold="true" box="[863,985,1348,1374]" pageId="1" pageNumber="241">Diagnosis.</emphasis>
Forewing characters only. Eleven postnodal crossveins; discoidal triangle divided into four cells; three cells in subdiscoidal triangle; four cells between RP3/4 and MAa along wing margin; four crossveins in space between base of IR2, RP1/2, RP2 and first oblique crossvein; five rows of cells between MAa and MP; presence of a distal fork of RP2, below distal end of pterostigma.
</paragraph>
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<subSubSection id="C331A5BCFFE5FF88FCA5F9E911FCF943" pageId="1" pageNumber="241" type="materials_examined">
<paragraph id="8B94F637FFE5FF88FCA5F9E911FCF943" blockId="1.[810,1452,1129,2070]" pageId="1" pageNumber="241">
<materialsCitation id="3B43FC6AFFE5FF88FCA5F9E911F8F943" ID-GBIF-Occurrence="3862611301" country="China" county="Ningcheng County" location="The" municipality="Lower Cretaceous Yixian Formation" pageId="1" pageNumber="241" specimenCount="1" stateProvince="Nei Mongol" typeStatus="holotype">
<emphasis id="B95F2A25FFE5FF88FCA5F9E911FFF931" bold="true" box="[863,1191,1641,1667]" pageId="1" pageNumber="241">
<typeStatus id="54904895FFE5FF88FCA5F9E916C2F931" box="[863,922,1641,1667]" pageId="1" pageNumber="241">Type</typeStatus>
locality and horizon.
</emphasis>
<location id="8EF4A0ECFFE5FF88FB49F9E91187F931" LSID="urn:lsid:plazi:treatment:03824721FFE5FF8BFCD0FC321153FCEF:8EF4A0ECFFE5FF88FB49F9E91187F931" box="[1203,1247,1641,1667]" country="China" county="Ningcheng County" municipality="Lower Cretaceous Yixian Formation" name="The" pageId="1" pageNumber="241" stateProvince="Nei Mongol">The</location>
locality near the
<location id="8EF4A0ECFFE5FF88FCD0F90E16A3F91A" LSID="urn:lsid:plazi:treatment:03824721FFE5FF8BFCD0FC321153FCEF:8EF4A0ECFFE5FF88FCD0F90E16A3F91A" box="[810,1019,1678,1704]" country="China" county="Ningcheng County" municipality="Lower Cretaceous Yixian Formation" name="Liutiaogou village" pageId="1" pageNumber="241" stateProvince="Nei Mongol">Liutiaogou village</location>
,
<location id="8EF4A0ECFFE5FF88FBFCF90E107DF91A" LSID="urn:lsid:plazi:treatment:03824721FFE5FF8BFCD0FC321153FCEF:8EF4A0ECFFE5FF88FBFCF90E107DF91A" box="[1030,1317,1678,1704]" country="China" county="Ningcheng County" municipality="Lower Cretaceous Yixian Formation" name="Dashuangmiao Township" pageId="1" pageNumber="241" stateProvince="Nei Mongol">Dashuangmiao Township</location>
,
<collectingCounty id="62F58EBBFFE5FF88FACAF90E1626F97F" pageId="1" pageNumber="241">Ningcheng County</collectingCounty>
,
<collectingRegion id="49EF38D5FFE5FF88FC7DF9331160F97E" box="[903,1080,1714,1741]" country="China" name="Nei Mongol" pageId="1" pageNumber="241">Inner Mongolia</collectingRegion>
,
<collectingCountry id="F33CB6A7FFE5FF88FBBBF93211DFF97E" box="[1089,1159,1714,1740]" name="China" pageId="1" pageNumber="241">China</collectingCountry>
;
<collectingMunicipality id="6BF06C4DFFE5FF88FB6BF93316C7F943" pageId="1" pageNumber="241">Lower Cretaceous Yixian Formation</collectingMunicipality>
(Barremian to Aptian)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C331A5BCFFE5FF8BFCA5F97B1153FCEF" lastPageId="2" lastPageNumber="242" pageId="1" pageNumber="241" type="description">
<paragraph id="8B94F637FFE5FF8BFCA5F97B1153FCEF" blockId="1.[810,1452,1129,2070]" lastBlockId="2.[810,1451,763,861]" lastPageId="2" lastPageNumber="242" pageId="1" pageNumber="241">
<emphasis id="B95F2A25FFE5FF88FCA5F97B16A9F8A7" bold="true" box="[863,1009,1787,1813]" pageId="1" pageNumber="241">Description.</emphasis>
A complete forewing, hyaline and very well-preserved; pterostigma dark brown; wing length
<quantity id="4CD35BD2FFE5FF88FA80F8A01600F8ED" metricMagnitude="-2" metricUnit="m" metricValue="4.67" pageId="1" pageNumber="241" unit="mm" value="46.7">46.7 mm</quantity>
,
<quantity id="4CD35BD2FFE5FF88FC9CF8C51695F8ED" box="[870,973,1861,1887]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="1" pageNumber="241" unit="mm" value="12.2">12.2 mm</quantity>
wide; distance between base and arculus 6.0 mm, between arculus and nodus
<quantity id="4CD35BD2FFE5FF88FB2DF8E91019F836" box="[1239,1345,1897,1924]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="1" pageNumber="241" unit="mm" value="17.2">17.2 mm</quantity>
; distance from nodus to pterostigma
<quantity id="4CD35BD2FFE5FF88FB91F80E118DF81A" box="[1131,1237,1934,1961]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="1" pageNumber="241" unit="mm" value="13.5">13.5 mm</quantity>
, from pterostigma to wing apex
<quantity id="4CD35BD2FFE5FF88FC32F832117BF87F" box="[968,1059,1970,1997]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="241" unit="mm" value="6.5">6.5 mm</quantity>
; Ax1 is well preserved but Ax2 is weakened and very similar to the secondary antenodal crossveins; distance between Ax1 and wing base 5.0 mm; 20 antenodal crossveins; arculus angular; pterostigma elongate and narrow,
<quantity id="4CD35BD2FFE6FF8BFE70FCA414BDFC8D" box="[394,485,804,831]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="242" unit="mm" value="4.5">4.5 mm</quantity>
long and 1.0 mm wide, covering four cells, not basally recessed; pterostigmal brace strong, well aligned with basal side of pterostigma, moderately oblique; 11 postnodal crossveins not aligned with 11 postsubnodal crossveins; median space free; submedian space only traversed by CuP-crossing and one distal cubito-anal crossvein above PsA; PsA distinctly sigmoidal; hypertriangle with two oblique and weak crossveins; discoidal triangle elongate, four-celled; MAb straight,
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long; a well-defined three-celled subdiscoidal triangle; base of IR2
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basal of nodus; base of RP3/4
<quantity id="4CD35BD2FFE6FF8BFEC3FB3114CBFB79" box="[313,403,1201,1228]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="242" unit="mm" value="6.5">6.5 mm</quantity>
basal of nodus; four crossveins between RP and IR2 basal of first oblique vein O; first oblique vein
<geoCoordinate id="EE1F90F0FFE6FF8BFED1FB791423FAA6" box="[299,379,1273,1300]" degrees="0.5" direction="west" orientation="longitude" pageId="2" pageNumber="242" precision="5555" value="-0.5">
O
<quantity id="4CD35BD2FFE6FF8BFEA2FB791423FAA6" box="[344,379,1273,1300]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="242" unit="mm" value="0.5">0.5</quantity>
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mm distad subnodus and second oblique vein three cells distally; pseudo-IR1 very short, originating from RP1 below distal side of pterostigma; area between RP1 and RP2 rather narrow with two-three rows of cells between them; base of RP2 well aligned with subnodus, RP2 smoothly undulated in its mid part and with a strong fork below distal end of pterostigma; IR2 only slightly undulated, area between it and RP2 strongly widened at their undulated parts, with three rows of cells in its broadest part; IR2 and RP2 basally parallel, with only one row of cells; a well-developed and strongly curved Rspl with three rows of cells between it and IR2; two short convex oblique and undulating secondary veins anastomosing between IR2 and RP3/4 directly basal of origin of Rspl; RP3/4 and MAa parallel and smoothly undulated, with one row of cells between them basally, two rows of cells more distally, and four rows along posterior margin of wing; MAa and MP more or less parallel, postdiscoidal area weakly widened with five rows of cells at level of nodus; no Mspl; MA and CuA basally parallel with one row of rectangular cells between them, distally strongly divergent with at least
<specimenCount id="9D2D3DBEFFE6FF8BFF72F7921466F79E" box="[136,318,2066,2092]" pageId="2" pageNumber="242" type="intercalary">three intercalary</specimenCount>
veins originating from MP; CuA with six posterior branches, cubito-anal area with seven rows of cells between CuA and posterior wing margin; two rows of cells in anal area.
</paragraph>
</subSubSection>
</treatment>
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